477 lines
50 KiB
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477 lines
50 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1097.82527" ID-GBIF-Dataset="4433ecc4-d495-4d4f-a8fb-baebcf8b7d4c" ID-PMC="PMC9046370" ID-Pensoft-Pub="1313-2970-1097-65" ID-Pensoft-UUID="8FCFCC1D8266501EA79AAE732D44FF0B" ID-PubMed="35837584" ID-ZooBank="6FC8EE30904F4E8D9EF9996F81F30693" ModsDocID="1313-2970-1097-65" checkinTime="1650499969150" checkinUser="pensoft" docAuthor="Hou, Yanmeng, Gao, Zhizhong & Zhang, Feng" docDate="2022" docId="385C611D235E5B0D9B1E4E54164EA1B9" docLanguage="en" docName="ZooKeys 1097: 65-83" docOrigin="ZooKeys 1097" docPubDate="2022-04-20" docSource="http://dx.doi.org/10.3897/zookeys.1097.82527" docTitle="Tyrannochthonius pandus Hou & Gao & Zhang 2022, sp. nov." docType="treatment" docUuid="D9B22241-9699-41ED-936C-5169456BD61A" docUuidSource="ZooBank" docVersion="4" id="8FCFCC1D8266501EA79AAE732D44FF0B" lastPageNumber="65" masterDocId="8FCFCC1D8266501EA79AAE732D44FF0B" masterDocTitle="Two new species of cave-adapted pseudoscorpions (Pseudoscorpiones, Chthoniidae) from Yunnan, China" masterLastPageNumber="83" masterPageNumber="65" pageNumber="65" updateTime="1668151692745" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Two new species of cave-adapted pseudoscorpions (Pseudoscorpiones, Chthoniidae) from Yunnan, China</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Hou, Yanmeng</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-0059-3419</mods:nameIdentifier>
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<mods:affiliation>The Key Laboratory of Zoological Systematics and Application, Institute of Life Science and Green Development, Hebei University, Baoding, Hebei 071002, China</mods:affiliation>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Gao, Zhizhong</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6666-8746</mods:nameIdentifier>
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<mods:affiliation>Department of Biology, Xinzhou Teachers University, Xinzhou 034000, Shanxi Province, China</mods:affiliation>
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<mods:nameIdentifier type="email">gaozhizhong1987@126.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Zhang, Feng</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-3347-1031</mods:nameIdentifier>
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<mods:affiliation>The Key Laboratory of Zoological Systematics and Application, Institute of Life Science and Green Development, Hebei University, Baoding, Hebei 071002, China</mods:affiliation>
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<mods:nameIdentifier type="email">dudu06042001@163.com</mods:nameIdentifier>
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<mods:title>ZooKeys</mods:title>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-04-20</mods:number>
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<mods:detail type="volume">
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<mods:number>1097</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>65</mods:start>
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<mods:end>83</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1097.82527</mods:url>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1097.82527</mods:identifier>
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<treatment ID-GBIF-Taxon="194685365" LSID="urn:lsid:zoobank.org:act:D9B22241-9699-41ED-936C-5169456BD61A" httpUri="http://treatment.plazi.org/id/385C611D235E5B0D9B1E4E54164EA1B9" lastPageNumber="65" pageId="0" pageNumber="65">
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<subSubSection pageId="0" pageNumber="65" type="nomenclature">
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<paragraph pageId="0" pageNumber="65">
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<taxonomicName LSID="http://zoobank.org/D9B22241-9699-41ED-936C-5169456BD61A" authority="Hou & Gao & Zhang, 2022" authorityName="Hou & Gao & Zhang" authorityYear="2022" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tyrannochthonius pandus" order="Pseudoscorpiones" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="pandus" status="sp. nov.">Tyrannochthonius pandus</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="65">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="65" type="description">
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<paragraph pageId="0" pageNumber="65">
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Tyrannochthonius pandus sp. nov., male habitus. Photographed when it crawled on stony natural habitat." figureDoi="10.3897/zookeys.1097.82527.figure2" httpUri="https://binary.pensoft.net/fig/674768" pageId="0" pageNumber="65">
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Chinese name.
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<normalizedToken originalValue="弯指暴伪蝎">弯指暴伪蝎</normalizedToken>
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Figs 2
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</figureCitation>
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Tyrannochthonius pandus sp. nov. A holotype male, dorsal view B paratype female, dorsal view. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1097.82527.figure3" httpUri="https://binary.pensoft.net/fig/674769" pageId="0" pageNumber="65">, 3</figureCitation>
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Tyrannochthonius pandus sp. nov., holotype male (A-D), paratype female (E) A chelal fingers (lateral view) B carapace (dorsal view) C left chelicera (dorsal view) D male genital area (ventral view) E female genital area (ventral view). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1097.82527.figure4" httpUri="https://binary.pensoft.net/fig/674770" pageId="0" pageNumber="65">, 4</figureCitation>
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tyrannochthonius pandus sp. nov., holotype male A carapace (dorsal view) with a detail of anterior margin B left chelicera (dorsal view) with details of teeth C coxal spines on coxae II (ventral view) D rallum E left pedipalp (minus chela, dorsal view) F left chela (lateral view) with details of teeth and with trichobothrial pattern (abbreviations explained in Material and methods) G finger tips of chela (lateral view) H left chela (dorsal view) I leg I (lateral view) J leg IV (lateral view). Scale bars: 0.25 mm (A-B, E-F, H-J); 0.10 mm (C-D, G)." figureDoi="10.3897/zookeys.1097.82527.figure5" httpUri="https://binary.pensoft.net/fig/674771" pageId="0" pageNumber="65">, 5</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="65" type="type material">
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<paragraph pageId="0" pageNumber="65">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="65">
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(Figs
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Study area, general cave locations, and type locality for each species, Yunnan Province, China A Biyu Cave, Tyrannochthonius pandus sp. nov. B Laoxueyan Cave, Lagynochthonius laoxueyanensis sp. nov." figureDoi="10.3897/zookeys.1097.82527.figure1" httpUri="https://binary.pensoft.net/fig/674767" pageId="0" pageNumber="65">1A</figureCitation>
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,
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Biyu Cave, type locality of Tyrannochthonius pandus sp. nov. A entrance B access to an undeveloped area C area where T. pandus specimens were collected. D, E beautiful cave landscapes inside the cave F exit." figureDoi="10.3897/zookeys.1097.82527.figure6" httpUri="https://binary.pensoft.net/fig/674772" pageId="0" pageNumber="65">6</figureCitation>
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)
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<emphasis bold="true" italics="true" pageId="0" pageNumber="65">Holotype</emphasis>
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: China • ♂; Yunnan Province, Luxi County, Luyuandong Village, the Ancient Alu Cave National Park of China, Biyu Cave;
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<geoCoordinate degrees="24" direction="north" minutes="34.01" orientation="latitude" precision="9" value="24.566833">24°34.01'N</geoCoordinate>
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,
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<geoCoordinate degrees="103" direction="east" minutes="45.16" orientation="longitude" precision="9" value="103.75267">103°45.16'E</geoCoordinate>
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; 1722 m a.s.l.; 13 Oct. 2021; Zegang Feng, Yanmeng Hou, Lu Zhang and Liu Fu leg.; dark zone; Ps.-
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<emphasis bold="true" pageId="0" pageNumber="65">MHBU</emphasis>
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-HBUARA#2021-438-01.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="65">Paratype</emphasis>
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: • 1♀; the same data as the holotype; Ps.-
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<emphasis bold="true" pageId="0" pageNumber="65">MSWU</emphasis>
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-HBUARA#2021-438-02.
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</paragraph>
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<caption doi="10.3897/zookeys.1097.82527.figure1" httpUri="https://binary.pensoft.net/fig/674767" pageId="0" pageNumber="65" start="Figure 1" startId="F1">
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<paragraph pageId="0" pageNumber="65">
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<emphasis bold="true" pageId="0" pageNumber="65">Figure 1.</emphasis>
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Study area, general cave locations, and type locality for each species, Yunnan Province, China
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<emphasis bold="true" pageId="0" pageNumber="65">A</emphasis>
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Biyu Cave,
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<taxonomicName authorityName="Hou & Gao & Zhang" authorityYear="2022" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tyrannochthonius pandus" order="Pseudoscorpiones" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="pandus">
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<emphasis italics="true" pageId="0" pageNumber="65">Tyrannochthonius pandus</emphasis>
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</taxonomicName>
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sp. nov.
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<emphasis bold="true" pageId="0" pageNumber="65">B</emphasis>
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Laoxueyan Cave,
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<taxonomicName authorityName="Hou & Gao & Zhang" authorityYear="2022" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lagynochthonius laoxueyanensis" order="Pseudoscorpiones" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="laoxueyanensis">
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<emphasis italics="true" pageId="0" pageNumber="65">Lagynochthonius laoxueyanensis</emphasis>
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</taxonomicName>
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sp. nov.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="65" type="diagnosis">
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<paragraph pageId="0" pageNumber="65">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="65">Moderately sized troglomorphic species with elongate appendages; carapace without eyes or eyespots; anterior margin of carapace gently serrate, epistome small, pointed, triangular, with 2 setae flanking base; posterior margin of carapace with 2 setae; tergites I-III with 2 setae; lacking chemosensory setae on dorsum of chelal hand; chelal fingers distinctly curved in dorsal view, with numerous large, gently curved, well-spaced teeth.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="65" type="etymology">
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<paragraph pageId="0" pageNumber="65">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="65">
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The specific name is derived from the Latin word "
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<emphasis italics="true" pageId="0" pageNumber="65">pandus</emphasis>
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", meaning curved, refers to the curved chelal finger.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="65" type="description">
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<paragraph pageId="0" pageNumber="65">Description.</paragraph>
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<paragraph pageId="0" pageNumber="65">
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<emphasis bold="true" pageId="0" pageNumber="65">Adult male</emphasis>
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(Figs
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Tyrannochthonius pandus sp. nov., male habitus. Photographed when it crawled on stony natural habitat." figureDoi="10.3897/zookeys.1097.82527.figure2" httpUri="https://binary.pensoft.net/fig/674768" pageId="0" pageNumber="65">2</figureCitation>
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,
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Tyrannochthonius pandus sp. nov. A holotype male, dorsal view B paratype female, dorsal view. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1097.82527.figure3" httpUri="https://binary.pensoft.net/fig/674769" pageId="0" pageNumber="65">3A</figureCitation>
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,
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Tyrannochthonius pandus sp. nov., holotype male (A-D), paratype female (E) A chelal fingers (lateral view) B carapace (dorsal view) C left chelicera (dorsal view) D male genital area (ventral view) E female genital area (ventral view). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1097.82527.figure4" httpUri="https://binary.pensoft.net/fig/674770" pageId="0" pageNumber="65">4A-D</figureCitation>
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,
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tyrannochthonius pandus sp. nov., holotype male A carapace (dorsal view) with a detail of anterior margin B left chelicera (dorsal view) with details of teeth C coxal spines on coxae II (ventral view) D rallum E left pedipalp (minus chela, dorsal view) F left chela (lateral view) with details of teeth and with trichobothrial pattern (abbreviations explained in Material and methods) G finger tips of chela (lateral view) H left chela (dorsal view) I leg I (lateral view) J leg IV (lateral view). Scale bars: 0.25 mm (A-B, E-F, H-J); 0.10 mm (C-D, G)." figureDoi="10.3897/zookeys.1097.82527.figure5" httpUri="https://binary.pensoft.net/fig/674771" pageId="0" pageNumber="65">5</figureCitation>
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).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="65">Color</emphasis>
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: generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale (Figs
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Tyrannochthonius pandus sp. nov., male habitus. Photographed when it crawled on stony natural habitat." figureDoi="10.3897/zookeys.1097.82527.figure2" httpUri="https://binary.pensoft.net/fig/674768" pageId="0" pageNumber="65">2</figureCitation>
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,
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Tyrannochthonius pandus sp. nov. A holotype male, dorsal view B paratype female, dorsal view. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1097.82527.figure3" httpUri="https://binary.pensoft.net/fig/674769" pageId="0" pageNumber="65">3A</figureCitation>
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).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="65">Cephalothorax</emphasis>
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(Figs
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Tyrannochthonius pandus sp. nov., holotype male (A-D), paratype female (E) A chelal fingers (lateral view) B carapace (dorsal view) C left chelicera (dorsal view) D male genital area (ventral view) E female genital area (ventral view). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1097.82527.figure4" httpUri="https://binary.pensoft.net/fig/674770" pageId="0" pageNumber="65">4B</figureCitation>
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,
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tyrannochthonius pandus sp. nov., holotype male A carapace (dorsal view) with a detail of anterior margin B left chelicera (dorsal view) with details of teeth C coxal spines on coxae II (ventral view) D rallum E left pedipalp (minus chela, dorsal view) F left chela (lateral view) with details of teeth and with trichobothrial pattern (abbreviations explained in Material and methods) G finger tips of chela (lateral view) H left chela (dorsal view) I leg I (lateral view) J leg IV (lateral view). Scale bars: 0.25 mm (A-B, E-F, H-J); 0.10 mm (C-D, G)." figureDoi="10.3897/zookeys.1097.82527.figure5" httpUri="https://binary.pensoft.net/fig/674771" pageId="0" pageNumber="65">5A</figureCitation>
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): carapace 1.07 times longer than broad, gently narrowed posteriorly, surface smooth; anterior margin slightly serrate; without any traces of eyes; epistome very pointed and small, triangular, with 2 setae flanking base; with 18 setae arranged s4s: 4: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; without furrows but with 4 lyrifissures (Fig.
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tyrannochthonius pandus sp. nov., holotype male A carapace (dorsal view) with a detail of anterior margin B left chelicera (dorsal view) with details of teeth C coxal spines on coxae II (ventral view) D rallum E left pedipalp (minus chela, dorsal view) F left chela (lateral view) with details of teeth and with trichobothrial pattern (abbreviations explained in Material and methods) G finger tips of chela (lateral view) H left chela (dorsal view) I leg I (lateral view) J leg IV (lateral view). Scale bars: 0.25 mm (A-B, E-F, H-J); 0.10 mm (C-D, G)." figureDoi="10.3897/zookeys.1097.82527.figure5" httpUri="https://binary.pensoft.net/fig/674771" pageId="0" pageNumber="65">5A</figureCitation>
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). Chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta less than 1/2 length of medial seta; apex of coxa I with small, rounded anteromedial process; coxae II with 10 terminally indented coxal spines on each side, set as an oblique row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (Fig.
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tyrannochthonius pandus sp. nov., holotype male A carapace (dorsal view) with a detail of anterior margin B left chelicera (dorsal view) with details of teeth C coxal spines on coxae II (ventral view) D rallum E left pedipalp (minus chela, dorsal view) F left chela (lateral view) with details of teeth and with trichobothrial pattern (abbreviations explained in Material and methods) G finger tips of chela (lateral view) H left chela (dorsal view) I leg I (lateral view) J leg IV (lateral view). Scale bars: 0.25 mm (A-B, E-F, H-J); 0.10 mm (C-D, G)." figureDoi="10.3897/zookeys.1097.82527.figure5" httpUri="https://binary.pensoft.net/fig/674771" pageId="0" pageNumber="65">5C</figureCitation>
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); intercoxal tubercle absent; without sub-oral seta.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="65">Chelicera</emphasis>
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(Figs
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Tyrannochthonius pandus sp. nov., holotype male (A-D), paratype female (E) A chelal fingers (lateral view) B carapace (dorsal view) C left chelicera (dorsal view) D male genital area (ventral view) E female genital area (ventral view). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1097.82527.figure4" httpUri="https://binary.pensoft.net/fig/674770" pageId="0" pageNumber="65">4C</figureCitation>
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,
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tyrannochthonius pandus sp. nov., holotype male A carapace (dorsal view) with a detail of anterior margin B left chelicera (dorsal view) with details of teeth C coxal spines on coxae II (ventral view) D rallum E left pedipalp (minus chela, dorsal view) F left chela (lateral view) with details of teeth and with trichobothrial pattern (abbreviations explained in Material and methods) G finger tips of chela (lateral view) H left chela (dorsal view) I leg I (lateral view) J leg IV (lateral view). Scale bars: 0.25 mm (A-B, E-F, H-J); 0.10 mm (C-D, G)." figureDoi="10.3897/zookeys.1097.82527.figure5" httpUri="https://binary.pensoft.net/fig/674771" pageId="0" pageNumber="65">5B</figureCitation>
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): large, about as long as carapace, 2.44 times longer than broad; 5 setae on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation dorsally. Both fingers well provided with teeth, fixed finger with 11 teeth, distal one largest; movable finger with 8 retrorse continuous small teeth; galea completely vestigial (Fig.
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tyrannochthonius pandus sp. nov., holotype male A carapace (dorsal view) with a detail of anterior margin B left chelicera (dorsal view) with details of teeth C coxal spines on coxae II (ventral view) D rallum E left pedipalp (minus chela, dorsal view) F left chela (lateral view) with details of teeth and with trichobothrial pattern (abbreviations explained in Material and methods) G finger tips of chela (lateral view) H left chela (dorsal view) I leg I (lateral view) J leg IV (lateral view). Scale bars: 0.25 mm (A-B, E-F, H-J); 0.10 mm (C-D, G)." figureDoi="10.3897/zookeys.1097.82527.figure5" httpUri="https://binary.pensoft.net/fig/674771" pageId="0" pageNumber="65">5B</figureCitation>
|
||
). Rallum with 8 blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tyrannochthonius pandus sp. nov., holotype male A carapace (dorsal view) with a detail of anterior margin B left chelicera (dorsal view) with details of teeth C coxal spines on coxae II (ventral view) D rallum E left pedipalp (minus chela, dorsal view) F left chela (lateral view) with details of teeth and with trichobothrial pattern (abbreviations explained in Material and methods) G finger tips of chela (lateral view) H left chela (dorsal view) I leg I (lateral view) J leg IV (lateral view). Scale bars: 0.25 mm (A-B, E-F, H-J); 0.10 mm (C-D, G)." figureDoi="10.3897/zookeys.1097.82527.figure5" httpUri="https://binary.pensoft.net/fig/674771" pageId="0" pageNumber="65">5D</figureCitation>
|
||
).
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="65">Pedipalp</emphasis>
|
||
(Figs
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Tyrannochthonius pandus sp. nov., holotype male (A-D), paratype female (E) A chelal fingers (lateral view) B carapace (dorsal view) C left chelicera (dorsal view) D male genital area (ventral view) E female genital area (ventral view). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1097.82527.figure4" httpUri="https://binary.pensoft.net/fig/674770" pageId="0" pageNumber="65">4A</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tyrannochthonius pandus sp. nov., holotype male A carapace (dorsal view) with a detail of anterior margin B left chelicera (dorsal view) with details of teeth C coxal spines on coxae II (ventral view) D rallum E left pedipalp (minus chela, dorsal view) F left chela (lateral view) with details of teeth and with trichobothrial pattern (abbreviations explained in Material and methods) G finger tips of chela (lateral view) H left chela (dorsal view) I leg I (lateral view) J leg IV (lateral view). Scale bars: 0.25 mm (A-B, E-F, H-J); 0.10 mm (C-D, G)." figureDoi="10.3897/zookeys.1097.82527.figure5" httpUri="https://binary.pensoft.net/fig/674771" pageId="0" pageNumber="65">5E-H</figureCitation>
|
||
): surface of palpal segments smooth; setae generally long and acuminate; femur 7.36, patella 2.55, chela 7.47, hand 2.80 times longer than deep; movable finger 1.71 times longer than hand and 0.64 times longer than chela, without large basal apodeme, only slightly sclerotized section present. Femur and dorsal hand without tactile setae but with 1 distal lyrifissure on patella (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tyrannochthonius pandus sp. nov., holotype male A carapace (dorsal view) with a detail of anterior margin B left chelicera (dorsal view) with details of teeth C coxal spines on coxae II (ventral view) D rallum E left pedipalp (minus chela, dorsal view) F left chela (lateral view) with details of teeth and with trichobothrial pattern (abbreviations explained in Material and methods) G finger tips of chela (lateral view) H left chela (dorsal view) I leg I (lateral view) J leg IV (lateral view). Scale bars: 0.25 mm (A-B, E-F, H-J); 0.10 mm (C-D, G)." figureDoi="10.3897/zookeys.1097.82527.figure5" httpUri="https://binary.pensoft.net/fig/674771" pageId="0" pageNumber="65">5E</figureCitation>
|
||
). Fixed chelal finger and hand with 8 trichobothria, movable chelal finger with 4 trichobothria,
|
||
<emphasis italics="true" pageId="0" pageNumber="65">ib</emphasis>
|
||
and
|
||
<emphasis italics="true" pageId="0" pageNumber="65">isb</emphasis>
|
||
situated close together, submedially on dorsum of chelal hand;
|
||
<emphasis italics="true" pageId="0" pageNumber="65">eb</emphasis>
|
||
,
|
||
<emphasis italics="true" pageId="0" pageNumber="65">esb</emphasis>
|
||
and
|
||
<emphasis italics="true" pageId="0" pageNumber="65">ist</emphasis>
|
||
forming a straight oblique row at base of fixed chelal finger;
|
||
<emphasis italics="true" pageId="0" pageNumber="65">it</emphasis>
|
||
slightly distal to
|
||
<emphasis italics="true" pageId="0" pageNumber="65">est</emphasis>
|
||
, situated subdistally;
|
||
<emphasis italics="true" pageId="0" pageNumber="65">et</emphasis>
|
||
slightly near to tip of fixed finger, very close to chelal teeth;
|
||
<emphasis italics="true" pageId="0" pageNumber="65">dx</emphasis>
|
||
situated distal to
|
||
<emphasis italics="true" pageId="0" pageNumber="65">et</emphasis>
|
||
;
|
||
<emphasis italics="true" pageId="0" pageNumber="65">sb</emphasis>
|
||
situated closer to
|
||
<emphasis italics="true" pageId="0" pageNumber="65">st</emphasis>
|
||
than to
|
||
<emphasis italics="true" pageId="0" pageNumber="65">b</emphasis>
|
||
;
|
||
<emphasis italics="true" pageId="0" pageNumber="65">b</emphasis>
|
||
and
|
||
<emphasis italics="true" pageId="0" pageNumber="65">t</emphasis>
|
||
situated subdistally,
|
||
<emphasis italics="true" pageId="0" pageNumber="65">b</emphasis>
|
||
situated at same level as
|
||
<emphasis italics="true" pageId="0" pageNumber="65">est</emphasis>
|
||
;
|
||
<emphasis italics="true" pageId="0" pageNumber="65">t</emphasis>
|
||
situated distal to
|
||
<emphasis italics="true" pageId="0" pageNumber="65">est</emphasis>
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tyrannochthonius pandus sp. nov., holotype male A carapace (dorsal view) with a detail of anterior margin B left chelicera (dorsal view) with details of teeth C coxal spines on coxae II (ventral view) D rallum E left pedipalp (minus chela, dorsal view) F left chela (lateral view) with details of teeth and with trichobothrial pattern (abbreviations explained in Material and methods) G finger tips of chela (lateral view) H left chela (dorsal view) I leg I (lateral view) J leg IV (lateral view). Scale bars: 0.25 mm (A-B, E-F, H-J); 0.10 mm (C-D, G)." figureDoi="10.3897/zookeys.1097.82527.figure5" httpUri="https://binary.pensoft.net/fig/674771" pageId="0" pageNumber="65">5F</figureCitation>
|
||
). Microsetae (chemosensory setae) absent on hand and both chelal fingers. Sensilla absent. A tiny antiaxial lyrifissure present at base of fixed finger (slightly distal to
|
||
<emphasis italics="true" pageId="0" pageNumber="65">ist</emphasis>
|
||
). Both chelal fingers with a row of teeth, homodentate, spaced regularly along the margin, larger teeth present in the middle of the row, becoming smaller distally and proximally: fixed finger with 45 large, gently curved, well-spaced teeth, without intercalary teeth; movable finger with 44 small (slightly smaller than the teeth on fixed finger), retrorse, gently curved and well-spaced teeth (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tyrannochthonius pandus sp. nov., holotype male A carapace (dorsal view) with a detail of anterior margin B left chelicera (dorsal view) with details of teeth C coxal spines on coxae II (ventral view) D rallum E left pedipalp (minus chela, dorsal view) F left chela (lateral view) with details of teeth and with trichobothrial pattern (abbreviations explained in Material and methods) G finger tips of chela (lateral view) H left chela (dorsal view) I leg I (lateral view) J leg IV (lateral view). Scale bars: 0.25 mm (A-B, E-F, H-J); 0.10 mm (C-D, G)." figureDoi="10.3897/zookeys.1097.82527.figure5" httpUri="https://binary.pensoft.net/fig/674771" pageId="0" pageNumber="65">5F</figureCitation>
|
||
). Chelal fingers distinctly curved in dorsal view (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tyrannochthonius pandus sp. nov., holotype male A carapace (dorsal view) with a detail of anterior margin B left chelicera (dorsal view) with details of teeth C coxal spines on coxae II (ventral view) D rallum E left pedipalp (minus chela, dorsal view) F left chela (lateral view) with details of teeth and with trichobothrial pattern (abbreviations explained in Material and methods) G finger tips of chela (lateral view) H left chela (dorsal view) I leg I (lateral view) J leg IV (lateral view). Scale bars: 0.25 mm (A-B, E-F, H-J); 0.10 mm (C-D, G)." figureDoi="10.3897/zookeys.1097.82527.figure5" httpUri="https://binary.pensoft.net/fig/674771" pageId="0" pageNumber="65">5H</figureCitation>
|
||
).
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="65">Opisthosoma</emphasis>
|
||
: generally typical; pleural membrane finely granulate. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I-XII: 2: 2: 2: 3: 4: 4: 4: 5: 5: 4: T2T: 0; tergites VIII and IX each with an unpaired median seta. Sternal chaetotaxy IV-XII: 10: 10: 9: 9: 9: 9: 8: 0: 2; sternites VI-IX with unpaired median seta. Anterior genital operculum with 10 setae, genital opening slit-like, with 15 marginal setae on each side (Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Tyrannochthonius pandus sp. nov., holotype male (A-D), paratype female (E) A chelal fingers (lateral view) B carapace (dorsal view) C left chelicera (dorsal view) D male genital area (ventral view) E female genital area (ventral view). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1097.82527.figure4" httpUri="https://binary.pensoft.net/fig/674770" pageId="0" pageNumber="65">4D</figureCitation>
|
||
).
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="65">Legs</emphasis>
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tyrannochthonius pandus sp. nov., holotype male A carapace (dorsal view) with a detail of anterior margin B left chelicera (dorsal view) with details of teeth C coxal spines on coxae II (ventral view) D rallum E left pedipalp (minus chela, dorsal view) F left chela (lateral view) with details of teeth and with trichobothrial pattern (abbreviations explained in Material and methods) G finger tips of chela (lateral view) H left chela (dorsal view) I leg I (lateral view) J leg IV (lateral view). Scale bars: 0.25 mm (A-B, E-F, H-J); 0.10 mm (C-D, G)." figureDoi="10.3897/zookeys.1097.82527.figure5" httpUri="https://binary.pensoft.net/fig/674771" pageId="0" pageNumber="65">5I, J</figureCitation>
|
||
): generally typical, long and slender. Femur of leg I 1.92 times longer than patella and with 1 lyrifissure at the base of femur; tarsus 2.50 times longer than tibia. Femoropatella of leg IV 3.67 times longer than deep; tibia 6.29 times longer than deep; with basal tactile setae on both tarsal segments: basitarsus 4.00 times longer than deep (TS = 0.35), telotarsus 14.50 times longer than deep and 2.90 times longer than basitarsus (TS = 0.33). Setae of leg I (trochanter to tibia) 3: 13: 12: 9, setae of leg IV (trochanter to basitarsus) 1: 3: 6: 9: 8. Arolium slightly shorter than the claws, not divided; claws simple.
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="65">Dimensions of male holotype</emphasis>
|
||
(length/width or, in the case of the legs, length/depth in mm; ratios in parentheses). Body length 1.41. Pedipalps: trochanter 0.18/0.13 (1.38), femur 0.81/0.11 (7.36), patella 0.28/0.11 (2.55), chela 1.12/0.15 (7.47), hand length 0.42/0.15 (2.80), movable finger length 0.72. Chelicera 0.44/0.18 (2.44), movable finger length 0.24. Carapace 0.45/0.42 (1.07). Leg I: trochanter 0.13/0.11 (1.18), femur 0.46/0.06 (7.67), patella 0.24/0.05 (4.80), tibia 0.20/0.04 (5.00), tarsus 0.50/0.04 (12.50). Leg IV: trochanter 0.20/0.11 (1.82), femoropatella 0.66/0.18 (3.67), tibia 0.44/0.07 (6.29), basitarsus 0.20/0.05 (4.00), telotarsus 0.58/0.04 (14.50).
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.1097.82527.figure2" httpUri="https://binary.pensoft.net/fig/674768" pageId="0" pageNumber="65" start="Figure 2" startId="F2">
|
||
<paragraph pageId="0" pageNumber="65">
|
||
<emphasis bold="true" pageId="0" pageNumber="65">Figure 2.</emphasis>
|
||
<taxonomicName authorityName="Hou & Gao & Zhang" authorityYear="2022" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tyrannochthonius pandus" order="Pseudoscorpiones" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="pandus">
|
||
<emphasis italics="true" pageId="0" pageNumber="65">Tyrannochthonius pandus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., male habitus. Photographed when it crawled on stony natural habitat.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="65">
|
||
<emphasis bold="true" pageId="0" pageNumber="65">Adult female</emphasis>
|
||
(Figs
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Tyrannochthonius pandus sp. nov. A holotype male, dorsal view B paratype female, dorsal view. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1097.82527.figure3" httpUri="https://binary.pensoft.net/fig/674769" pageId="0" pageNumber="65">3B</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Tyrannochthonius pandus sp. nov., holotype male (A-D), paratype female (E) A chelal fingers (lateral view) B carapace (dorsal view) C left chelicera (dorsal view) D male genital area (ventral view) E female genital area (ventral view). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1097.82527.figure4" httpUri="https://binary.pensoft.net/fig/674770" pageId="0" pageNumber="65">4E</figureCitation>
|
||
). Mostly same as males. Anterior genital operculum with 10 setae plus 7 setae on posterior margin. Body length 1.67. Pedipalps: trochanter 0.24/0.13 (1.85), femur 0.88/0.13 (6.77), patella 0.28/0.11 (2.55), chela 1.20/0.17 (7.06), hand length 0.49/0.17 (2.88), movable finger length 0.76. Chelicera 0.45/0.21 (2.14), movable finger length 0.23. Carapace 0.46/0.46 (1.00). Leg I: trochanter 0.14/0.11 (1.27), femur 0.50/0.06 (8.33), patella 0.26/0.06 (4.33), tibia 0.21/0.05 (4.20), tarsus 0.53/0.04 (13.25). Leg IV: trochanter 0.21/0.12 (1.75), femoropatella 0.70/0.19 (3.68), tibia 0.46/0.07 (6.57), basitarsus 0.21/0.06 (3.50), telotarsus 0.62/0.04 (15.50).
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.1097.82527.figure3" httpUri="https://binary.pensoft.net/fig/674769" pageId="0" pageNumber="65" start="Figure 3" startId="F3">
|
||
<paragraph pageId="0" pageNumber="65">
|
||
<emphasis bold="true" pageId="0" pageNumber="65">Figure 3.</emphasis>
|
||
<taxonomicName authorityName="Hou & Gao & Zhang" authorityYear="2022" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tyrannochthonius pandus" order="Pseudoscorpiones" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="pandus">
|
||
<emphasis italics="true" pageId="0" pageNumber="65">Tyrannochthonius pandus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
<emphasis bold="true" pageId="0" pageNumber="65">A</emphasis>
|
||
holotype male, dorsal view
|
||
<emphasis bold="true" pageId="0" pageNumber="65">B</emphasis>
|
||
paratype female, dorsal view. Scale bars: 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/zookeys.1097.82527.figure4" httpUri="https://binary.pensoft.net/fig/674770" pageId="0" pageNumber="65" start="Figure 4" startId="F4">
|
||
<paragraph pageId="0" pageNumber="65">
|
||
<emphasis bold="true" pageId="0" pageNumber="65">Figure 4.</emphasis>
|
||
<taxonomicName authorityName="Hou & Gao & Zhang" authorityYear="2022" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tyrannochthonius pandus" order="Pseudoscorpiones" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="pandus">
|
||
<emphasis italics="true" pageId="0" pageNumber="65">Tyrannochthonius pandus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype male (
|
||
<emphasis bold="true" pageId="0" pageNumber="65">A-D</emphasis>
|
||
), paratype female (
|
||
<emphasis bold="true" pageId="0" pageNumber="65">E</emphasis>
|
||
)
|
||
<emphasis bold="true" pageId="0" pageNumber="65">A</emphasis>
|
||
chelal fingers (lateral view)
|
||
<emphasis bold="true" pageId="0" pageNumber="65">B</emphasis>
|
||
carapace (dorsal view)
|
||
<emphasis bold="true" pageId="0" pageNumber="65">C</emphasis>
|
||
left chelicera (dorsal view)
|
||
<emphasis bold="true" pageId="0" pageNumber="65">D</emphasis>
|
||
male genital area (ventral view)
|
||
<emphasis bold="true" pageId="0" pageNumber="65">E</emphasis>
|
||
female genital area (ventral view). Scale bars: 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="65" type="remarks">
|
||
<paragraph pageId="0" pageNumber="65">Remarks.</paragraph>
|
||
<paragraph pageId="0" pageNumber="65">
|
||
Compared with the other six cave-dwelling species of the genus in China,
|
||
<taxonomicName authorityName="Hou & Gao & Zhang" authorityYear="2022" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tyrannochthonius pandus" order="Pseudoscorpiones" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="pandus">
|
||
<emphasis italics="true" pageId="0" pageNumber="65">Tyrannochthonius pandus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. is most similar to
|
||
<taxonomicName lsidName="T. ganshuanensis" pageId="0" pageNumber="65" rank="species" species="ganshuanensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="65">T. ganshuanensis</emphasis>
|
||
</taxonomicName>
|
||
in having only 2 setae on tergites I-III, the same chaetotaxy of carapace and triangular, a small epistome, but differs in the shape of teeth on chelal fingers (large, gently curved, well-spaced teeth, without intercalary teeth in
|
||
<taxonomicName lsidName="T. pandus" pageId="0" pageNumber="65" rank="species" species="pandus">
|
||
<emphasis italics="true" pageId="0" pageNumber="65">T. pandus</emphasis>
|
||
</taxonomicName>
|
||
, but with pointed, well-spaced and intercalary teeth in
|
||
<taxonomicName lsidName="T. ganshuanensis" pageId="0" pageNumber="65" rank="species" species="ganshuanensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="65">T. ganshuanensis</emphasis>
|
||
</taxonomicName>
|
||
), the relative position of the trichobothria on the movable chelal finger (
|
||
<emphasis italics="true" pageId="0" pageNumber="65">sb</emphasis>
|
||
situated closer to
|
||
<emphasis italics="true" pageId="0" pageNumber="65">st</emphasis>
|
||
than to
|
||
<emphasis italics="true" pageId="0" pageNumber="65">b</emphasis>
|
||
in
|
||
<taxonomicName lsidName="T. pandus" pageId="0" pageNumber="65" rank="species" species="pandus">
|
||
<emphasis italics="true" pageId="0" pageNumber="65">T. pandus</emphasis>
|
||
</taxonomicName>
|
||
, but
|
||
<emphasis italics="true" pageId="0" pageNumber="65">sb</emphasis>
|
||
situated closer to
|
||
<emphasis italics="true" pageId="0" pageNumber="65">b</emphasis>
|
||
in
|
||
<taxonomicName lsidName="T. ganshuanensis" pageId="0" pageNumber="65" rank="species" species="ganshuanensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="65">T. ganshuanensis</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<taxonomicName authorityName="Hou & Gao & Zhang" authorityYear="2022" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tyrannochthonius pandus" order="Pseudoscorpiones" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="pandus">
|
||
<emphasis italics="true" pageId="0" pageNumber="65">Tyrannochthonius pandus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. can be easily separated from
|
||
<taxonomicName lsidName="T. akaleus" pageId="0" pageNumber="65" rank="species" species="akaleus">
|
||
<emphasis italics="true" pageId="0" pageNumber="65">T. akaleus</emphasis>
|
||
</taxonomicName>
|
||
by a smaller body size (1.67 vs. 2.10 mm in female), the teeth pattern on chelal fingers (intercalary teeth absent in
|
||
<taxonomicName lsidName="T. pandus" pageId="0" pageNumber="65" rank="species" species="pandus">
|
||
<emphasis italics="true" pageId="0" pageNumber="65">T. pandus</emphasis>
|
||
</taxonomicName>
|
||
, but present in
|
||
<taxonomicName lsidName="T. akaleus" pageId="0" pageNumber="65" rank="species" species="akaleus">
|
||
<emphasis italics="true" pageId="0" pageNumber="65">T. akaleus</emphasis>
|
||
</taxonomicName>
|
||
); from
|
||
<taxonomicName lsidName="T. harveyi" pageId="0" pageNumber="65" rank="species" species="harveyi">
|
||
<emphasis italics="true" pageId="0" pageNumber="65">T. harveyi</emphasis>
|
||
</taxonomicName>
|
||
by the different setae number on the anterior and posterior margins of the carapace (
|
||
<taxonomicName lsidName="T. pandus" pageId="0" pageNumber="65" rank="species" species="pandus">
|
||
<emphasis italics="true" pageId="0" pageNumber="65">T. pandus</emphasis>
|
||
</taxonomicName>
|
||
with 6 and 2 setae, respectively, but
|
||
<taxonomicName lsidName="T. harveyi" pageId="0" pageNumber="65" rank="species" species="harveyi">
|
||
<emphasis italics="true" pageId="0" pageNumber="65">T. harveyi</emphasis>
|
||
</taxonomicName>
|
||
with 4 and 4 setae, respectively), the shape of the epistome (long and pointed in
|
||
<taxonomicName lsidName="T. pandus" pageId="0" pageNumber="65" rank="species" species="pandus">
|
||
<emphasis italics="true" pageId="0" pageNumber="65">T. pandus</emphasis>
|
||
</taxonomicName>
|
||
, but rounded and inconspicuous in
|
||
<taxonomicName lsidName="T. harveyi" pageId="0" pageNumber="65" rank="species" species="harveyi">
|
||
<emphasis italics="true" pageId="0" pageNumber="65">T. harveyi</emphasis>
|
||
</taxonomicName>
|
||
), the number of rallar blades (8 in
|
||
<taxonomicName lsidName="T. pandus" pageId="0" pageNumber="65" rank="species" species="pandus">
|
||
<emphasis italics="true" pageId="0" pageNumber="65">T. pandus</emphasis>
|
||
</taxonomicName>
|
||
, but 6 in
|
||
<taxonomicName lsidName="T. harveyi" pageId="0" pageNumber="65" rank="species" species="harveyi">
|
||
<emphasis italics="true" pageId="0" pageNumber="65">T. harveyi</emphasis>
|
||
</taxonomicName>
|
||
); and from
|
||
<taxonomicName lsidName="T. zhai" pageId="0" pageNumber="65" rank="species" species="zhai">
|
||
<emphasis italics="true" pageId="0" pageNumber="65">T. zhai</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName lsidName="T. chixingi" pageId="0" pageNumber="65" rank="species" species="chixingi">
|
||
<emphasis italics="true" pageId="0" pageNumber="65">T. chixingi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName lsidName="T. antridraconis" pageId="0" pageNumber="65" rank="species" species="antridraconis">
|
||
<emphasis italics="true" pageId="0" pageNumber="65">T. antridraconis</emphasis>
|
||
</taxonomicName>
|
||
by the number of setae on the anterior tergites (tergites I-III with 2 setae in
|
||
<taxonomicName lsidName="T. pandus" pageId="0" pageNumber="65" rank="species" species="pandus">
|
||
<emphasis italics="true" pageId="0" pageNumber="65">T. pandus</emphasis>
|
||
</taxonomicName>
|
||
, but the other three with 4 setae). In addition, compared to the new species,
|
||
<taxonomicName lsidName="T. zhai" pageId="0" pageNumber="65" rank="species" species="zhai">
|
||
<emphasis italics="true" pageId="0" pageNumber="65">T. zhai</emphasis>
|
||
</taxonomicName>
|
||
differs by the shorter body length (1.40 vs. 1.67 mm in female) and lacking an epistome;
|
||
<taxonomicName lsidName="T. chixingi" pageId="0" pageNumber="65" rank="species" species="chixingi">
|
||
<emphasis italics="true" pageId="0" pageNumber="65">T. chixingi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName lsidName="T. antridraconis" pageId="0" pageNumber="65" rank="species" species="antridraconis">
|
||
<emphasis italics="true" pageId="0" pageNumber="65">T. antridraconis</emphasis>
|
||
</taxonomicName>
|
||
differs from the new species also by the presence of intercalary teeth on the fixed chelal finger (
|
||
<bibRefCitation DOI="https://doi.org/10.5962/bhl.part.79492" author="Mahnert, V" journalOrPublisher="Revue Suisse de Zoologie" pageId="0" pageNumber="65" pagination="185 - 201" refId="B17" refString="Mahnert, V, 2009. New species of pseudoscorpions (Arachnida, Pseudoscorpiones, Chthoniidae, Chernetidae) from caves in China. Revue Suisse de Zoologie 116 (2): 185 - 201, DOI: https://doi.org/10.5962/bhl.part.79492" title="New species of pseudoscorpions (Arachnida, Pseudoscorpiones, Chthoniidae, Chernetidae) from caves in China." url="https://doi.org/10.5962/bhl.part.79492" volume="116" year="2009">Mahnert 2009</bibRefCitation>
|
||
;
|
||
<bibRefCitation DOI="https://doi.org/10.1636/JoA-S-17-063.1" author="Gao, ZZ" journalOrPublisher="The Journal of Arachnology" pageId="0" pageNumber="65" pagination="345 - 354" refId="B7" refString="Gao, ZZ, Wynne, JJ, Zhang, F, 2018. Two new species of cave-adapted pseudoscorpions (Pseudoscorpiones: Neobisiidae, Chthoniidae) from Guangxi, China. The Journal of Arachnology 46 (2): 345 - 354, DOI: https://doi.org/10.1636/JoA-S-17-063.1" title="Two new species of cave-adapted pseudoscorpions (Pseudoscorpiones: Neobisiidae, Chthoniidae) from Guangxi, China." url="https://doi.org/10.1636/JoA-S-17-063.1" volume="46" year="2018">Gao et al. 2018</bibRefCitation>
|
||
;
|
||
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4853.4.6" author="Gao, ZZ" journalOrPublisher="Zootaxa" pageId="0" pageNumber="65" pagination="572 - 580" refId="B8" refString="Gao, ZZ, Zhang, F, Chen, HM, 2020. Two new cave-dwelling species of Tyrannochthonius Chamberlin 1929 (Pseudoscorpiones: Chthoniidae) from the Guizhou karst, China. Zootaxa 4853 (4): 572 - 580, DOI: https://doi.org/10.11646/zootaxa.4853.4.6" title="Two new cave-dwelling species of Tyrannochthonius Chamberlin 1929 (Pseudoscorpiones: Chthoniidae) from the Guizhou karst, China." url="https://doi.org/10.11646/zootaxa.4853.4.6" volume="4853" year="2020">Gao et al. 2020</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.1097.82527.figure5" httpUri="https://binary.pensoft.net/fig/674771" pageId="0" pageNumber="65" start="Figure 5" startId="F5">
|
||
<paragraph pageId="0" pageNumber="65">
|
||
<emphasis bold="true" pageId="0" pageNumber="65">Figure 5.</emphasis>
|
||
<taxonomicName authorityName="Hou & Gao & Zhang" authorityYear="2022" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tyrannochthonius pandus" order="Pseudoscorpiones" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="pandus">
|
||
<emphasis italics="true" pageId="0" pageNumber="65">Tyrannochthonius pandus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype male
|
||
<emphasis bold="true" pageId="0" pageNumber="65">A</emphasis>
|
||
carapace (dorsal view) with a detail of anterior margin
|
||
<emphasis bold="true" pageId="0" pageNumber="65">B</emphasis>
|
||
left chelicera (dorsal view) with details of teeth
|
||
<emphasis bold="true" pageId="0" pageNumber="65">C</emphasis>
|
||
coxal spines on coxae II (ventral view)
|
||
<emphasis bold="true" pageId="0" pageNumber="65">D</emphasis>
|
||
rallum
|
||
<emphasis bold="true" pageId="0" pageNumber="65">E</emphasis>
|
||
left pedipalp (minus chela, dorsal view)
|
||
<emphasis bold="true" pageId="0" pageNumber="65">F</emphasis>
|
||
left chela (lateral view) with details of teeth and with trichobothrial pattern (abbreviations explained in Material and methods)
|
||
<emphasis bold="true" pageId="0" pageNumber="65">G</emphasis>
|
||
finger tips of chela (lateral view)
|
||
<emphasis bold="true" pageId="0" pageNumber="65">H</emphasis>
|
||
left chela (dorsal view)
|
||
<emphasis bold="true" pageId="0" pageNumber="65">I</emphasis>
|
||
leg I (lateral view)
|
||
<emphasis bold="true" pageId="0" pageNumber="65">J</emphasis>
|
||
leg IV (lateral view). Scale bars: 0.25 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="65">A-B, E-F, H-J</emphasis>
|
||
); 0.10 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="65">C-D, G</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="65" type="distribution">
|
||
<paragraph pageId="0" pageNumber="65">Distribution.</paragraph>
|
||
<paragraph pageId="0" pageNumber="65">
|
||
This species is known only from the type locality, Biyu Cave (Figs
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Study area, general cave locations, and type locality for each species, Yunnan Province, China A Biyu Cave, Tyrannochthonius pandus sp. nov. B Laoxueyan Cave, Lagynochthonius laoxueyanensis sp. nov." figureDoi="10.3897/zookeys.1097.82527.figure1" httpUri="https://binary.pensoft.net/fig/674767" pageId="0" pageNumber="65">1A</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Biyu Cave, type locality of Tyrannochthonius pandus sp. nov. A entrance B access to an undeveloped area C area where T. pandus specimens were collected. D, E beautiful cave landscapes inside the cave F exit." figureDoi="10.3897/zookeys.1097.82527.figure6" httpUri="https://binary.pensoft.net/fig/674772" pageId="0" pageNumber="65">6</figureCitation>
|
||
). Biyu Cave is one of the tourist caves in the Ancient Alu Cave National Park of China, with the entrance located in the Jilong hillside. This beautiful cave is a valley type karst cave, with an internal height of 2 to 5 m and a width of 1 to 30 m. The cave has a latticed distribution. The stalactites in this cave are jasper colored, so it is called Biyu Cave (a quote from the
|
||
<normalizedToken originalValue="cave’s">cave's</normalizedToken>
|
||
interior slogan). The specimens of this new species were collected from under a stone and on a stone wall in an undeveloped area of the cave that is still in a natural condition. It is a small, dark, high humidity and low temperature space (temperature: 11 °C, humidity: 90%), which is suitable for the survival of the species.
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.1097.82527.figure6" httpUri="https://binary.pensoft.net/fig/674772" pageId="0" pageNumber="65" start="Figure 6" startId="F6">
|
||
<paragraph pageId="0" pageNumber="65">
|
||
<emphasis bold="true" pageId="0" pageNumber="65">Figure 6.</emphasis>
|
||
Biyu Cave, type locality of
|
||
<taxonomicName authorityName="Hou & Gao & Zhang" authorityYear="2022" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tyrannochthonius pandus" order="Pseudoscorpiones" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="pandus">
|
||
<emphasis italics="true" pageId="0" pageNumber="65">Tyrannochthonius pandus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
<emphasis bold="true" pageId="0" pageNumber="65">A</emphasis>
|
||
entrance
|
||
<emphasis bold="true" pageId="0" pageNumber="65">B</emphasis>
|
||
access to an undeveloped area
|
||
<emphasis bold="true" pageId="0" pageNumber="65">C</emphasis>
|
||
area where
|
||
<taxonomicName lsidName="T. pandus" pageId="0" pageNumber="65" rank="species" species="pandus">
|
||
<emphasis italics="true" pageId="0" pageNumber="65">T. pandus</emphasis>
|
||
</taxonomicName>
|
||
specimens were collected.
|
||
<emphasis bold="true" pageId="0" pageNumber="65">D, E</emphasis>
|
||
beautiful cave landscapes inside the cave
|
||
<emphasis bold="true" pageId="0" pageNumber="65">F</emphasis>
|
||
exit.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |