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<document ID-DOI="http://dx.doi.org/10.3897/asp.81.e81961" ID-Pensoft-Pub="1864-8312-81-79" ID-Pensoft-UUID="CA16EA0C5A33560FBC955981DFC9D502" ID-ZooBank="5C1FDC828F9F4869ADDD83FA96E507ED" ModsDocID="1864-8312-81-79" checkinTime="1674309819629" checkinUser="pensoft" docAuthor="Domahovski, Alexandre Cruz &amp; Cavichioli, Rodney Ramiro" docDate="2023" docId="34519A1CC5195B04A203A3B3F09E66E5" docLanguage="en" docName="ArthropodSystPhyl 81: 79-164" docOrigin="Arthropod Systematics &amp; amp; Phylogeny 81" docPubDate="2023-01-20" docSource="http://dx.doi.org/10.3897/asp.81.e81961" docTitle="Coarctana nigromedia Domahovski &amp; Cavichioli 2023, sp. nov." docType="treatment" docUuid="DA9856A5-FD28-4FDE-AED4-EBCA9D85E623" docUuidSource="ZooBank" docVersion="1" id="CA16EA0C5A33560FBC955981DFC9D502" lastPageNumber="79" masterDocId="CA16EA0C5A33560FBC955981DFC9D502" masterDocTitle="Phylogenetic analysis and revision of the leafhopper genus Acuera DeLong &amp; Freytag (Hemiptera: Cicadellidae: Gyponini) based on morphological data" masterLastPageNumber="164" masterPageNumber="79" pageNumber="79" updateTime="1674309819629" updateUser="pensoft">
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<mods:title>Phylogenetic analysis and revision of the leafhopper genus Acuera DeLong &amp; Freytag (Hemiptera: Cicadellidae: Gyponini) based on morphological data</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Domahovski, Alexandre Cruz</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-4588-4236</mods:nameIdentifier>
<mods:affiliation>Departamento de Zoologia, Setor de Ciencias Biologicas, Universidade Federal do Parana, Av. Cel. Francisco H. dos Santos, 100, Caixa Postal 19020, 81531 - 980, Curitiba, PR, Brasil</mods:affiliation>
<mods:nameIdentifier type="email">domahovskiac@yahoo.com.br</mods:nameIdentifier>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Cavichioli, Rodney Ramiro</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-5996-0629</mods:nameIdentifier>
<mods:affiliation>Departamento de Zoologia, Setor de Ciencias Biologicas, Universidade Federal do Parana, Av. Cel. Francisco H. dos Santos, 100, Caixa Postal 19020, 81531 - 980, Curitiba, PR, Brasil</mods:affiliation>
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<mods:title>Arthropod Systematics &amp; amp; Phylogeny</mods:title>
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<mods:date>2023</mods:date>
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<mods:number>2023-01-20</mods:number>
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<mods:number>81</mods:number>
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<mods:start>79</mods:start>
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<mods:url>http://dx.doi.org/10.3897/asp.81.e81961</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/asp.81.e81961</mods:identifier>
<mods:identifier type="Pensoft-Pub">1864-8312-81-79</mods:identifier>
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<treatment LSID="urn:lsid:zoobank.org:act:DA9856A5-FD28-4FDE-AED4-EBCA9D85E623" httpUri="http://treatment.plazi.org/id/34519A1CC5195B04A203A3B3F09E66E5" lastPageNumber="79" pageId="0" pageNumber="79">
<subSubSection pageId="0" pageNumber="79" type="nomenclature">
<paragraph pageId="0" pageNumber="79">
<taxonomicName LSID="https://zoobank.org/DA9856A5-FD28-4FDE-AED4-EBCA9D85E623" authority="Domahovski &amp; Cavichioli, 2023" authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" family="Cicadellidae" genus="Coarctana" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Coarctana nigromedia" order="Hemiptera" pageId="0" pageNumber="79" rank="species" species="nigromedia" status="sp. nov.">Coarctana nigromedia</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="79">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="description">
<paragraph pageId="0" pageNumber="79">
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Coarctana nigromedia sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure20" httpUri="https://binary.pensoft.net/fig/798173" pageId="0" pageNumber="79">Figs 20</figureCitation>
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Coarctana nigromedia sp. nov., paratype female. A distal portion of abdomen, ventral view. B distal portion of abdomen, lateral view. C first valvifer and first valvula, lateral view. D apical portion of first valvula. E second valvula, lateral view. F apical portion of second valvula. G second valvifer and gonoplac, lateral view. H apical portion of gonoplac. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure21" httpUri="https://binary.pensoft.net/fig/798174" pageId="0" pageNumber="79">, 21</figureCitation>
<figureCitation captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Species of Coarctana gen. nov., dorsal (left) and lateral (right) views. A, B C. apena comb. nov., holotype male, AMNH. C, D C. asymmetrica sp. nov., holotype male, DZUP. E, F C. glabra sp. nov., holotype male, DZRJ. G, H C. nigromedia sp. nov., holotype male, DZUP. I, J C. occultata sp. nov., holotype male, DZUP. K, L C. oricula sp. nov., holotype male, DZUP. M, N C. parvula sp. nov., holotype male, DZUP." figureDoi="10.3897/asp.81.e81961.figure31" httpUri="https://binary.pensoft.net/fig/798184" pageId="0" pageNumber="79">, 31G, H</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="diagnosis">
<paragraph pageId="0" pageNumber="79">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="79">
Head (Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Coarctana nigromedia sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure20" httpUri="https://binary.pensoft.net/fig/798173" pageId="0" pageNumber="79">20A</figureCitation>
) with wide longitudinal dark-brown band between ocelli. Pygofer (Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Coarctana nigromedia sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure20" httpUri="https://binary.pensoft.net/fig/798173" pageId="0" pageNumber="79">20G</figureCitation>
) with apex excavated. Subgenital plate (Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Coarctana nigromedia sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure20" httpUri="https://binary.pensoft.net/fig/798173" pageId="0" pageNumber="79">20H</figureCitation>
) with long filiform setae. Style (Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Coarctana nigromedia sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure20" httpUri="https://binary.pensoft.net/fig/798173" pageId="0" pageNumber="79">20H</figureCitation>
) apex truncate, with dorsal process short and acute. Aedeagus (Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Coarctana nigromedia sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure20" httpUri="https://binary.pensoft.net/fig/798173" pageId="0" pageNumber="79">20L, M</figureCitation>
) with ventral margin of apodemal process forming rounded lobe produced ventrally; shaft with pair of apical processes directed ventrally and slightly curved laterally, with 1\4 length of shaft.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="description">
<paragraph pageId="0" pageNumber="79">Measurements.</paragraph>
<paragraph pageId="0" pageNumber="79">Total length: holotype male 7.3 mm; paratypes, males (n = 15) 6.8-7.5 mm, females (n = 6) 7.5-7.7 mm.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="description">
<paragraph pageId="0" pageNumber="79">Description.</paragraph>
<paragraph pageId="0" pageNumber="79">
Head, in frontal view (Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Coarctana nigromedia sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure20" httpUri="https://binary.pensoft.net/fig/798173" pageId="0" pageNumber="79">20B</figureCitation>
), frons 1.2
<normalizedToken originalValue="×">x</normalizedToken>
wider than long; frontogenal suture distant from eye margin by approximately half maximum width of clypeus; clypeus 1.2
<normalizedToken originalValue="×">x</normalizedToken>
wider than long; lateral margins straight, slightly convergent apically; apex straight. Head, in lateral view (Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Coarctana nigromedia sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure20" httpUri="https://binary.pensoft.net/fig/798173" pageId="0" pageNumber="79">20C</figureCitation>
), crown-face transition thin, with 3 transverse carinae; clypeus not inflated. Forewing (Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Coarctana nigromedia sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure20" httpUri="https://binary.pensoft.net/fig/798173" pageId="0" pageNumber="79">20D</figureCitation>
) with apex of anal veins bifid. Profemur AV row with 5-6 setae; PV row with 1 seta near apex. Protibia PD row with 3 setae and undifferentiated intercalary setae; PV row with 5 setae and undifferentiated intercalary setae. Metatibia PD, AD and AV rows with 22-25, 12 and 13-14 macrosetae respectively. Metatarsomere I 4
<normalizedToken originalValue="×">x</normalizedToken>
longer than apical width; inner row of the ventral surface with 5 setae. Other characteristics as in generic description.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="coloration">
<paragraph pageId="0" pageNumber="79">Coloration.</paragraph>
<paragraph pageId="0" pageNumber="79">
Head and thorax (Figs
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Coarctana nigromedia sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure20" httpUri="https://binary.pensoft.net/fig/798173" pageId="0" pageNumber="79">20A-D</figureCitation>
,
<figureCitation captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Species of Coarctana gen. nov., dorsal (left) and lateral (right) views. A, B C. apena comb. nov., holotype male, AMNH. C, D C. asymmetrica sp. nov., holotype male, DZUP. E, F C. glabra sp. nov., holotype male, DZRJ. G, H C. nigromedia sp. nov., holotype male, DZUP. I, J C. occultata sp. nov., holotype male, DZUP. K, L C. oricula sp. nov., holotype male, DZUP. M, N C. parvula sp. nov., holotype male, DZUP." figureDoi="10.3897/asp.81.e81961.figure31" httpUri="https://binary.pensoft.net/fig/798184" pageId="0" pageNumber="79">31G, H</figureCitation>
) yellowish-brown. Head (Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Coarctana nigromedia sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure20" httpUri="https://binary.pensoft.net/fig/798173" pageId="0" pageNumber="79">20A</figureCitation>
) with pair of small circular black spots, near posterior margin, behind ocelli; wide longitudinal dark-brown band between ocelli. Face (Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Coarctana nigromedia sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure20" httpUri="https://binary.pensoft.net/fig/798173" pageId="0" pageNumber="79">20B</figureCitation>
) yellowish; antennal pit dark brown. Pronotum (Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Coarctana nigromedia sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure20" httpUri="https://binary.pensoft.net/fig/798173" pageId="0" pageNumber="79">20A</figureCitation>
) with black punctures; small black maculae near anterior margin, behind eye and ocellus; median portion with wide longitudinal dark-brown band, continuous to the band of head and evanescent toward posterior margin of pronotum; proepimeron (Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Coarctana nigromedia sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure20" httpUri="https://binary.pensoft.net/fig/798173" pageId="0" pageNumber="79">20C</figureCitation>
) almost entirely black. Mesonotum (Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Coarctana nigromedia sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure20" httpUri="https://binary.pensoft.net/fig/798173" pageId="0" pageNumber="79">20A</figureCitation>
) with brown lateral angles, pair of small median brown spots near scutoscutellar suture. Forewing (Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Coarctana nigromedia sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure20" httpUri="https://binary.pensoft.net/fig/798173" pageId="0" pageNumber="79">20D</figureCitation>
) with brown veins, larger dark-brown maculae on apex of anal veins, outer discal cell, costal margin, cross veins of discal and apical cells, transverse brown stripe over subapical cells. Metatibia (Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Coarctana nigromedia sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure20" httpUri="https://binary.pensoft.net/fig/798173" pageId="0" pageNumber="79">20C</figureCitation>
) with cucullate bases of setae blacks.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="male terminalia">
<paragraph pageId="0" pageNumber="79">Male terminalia.</paragraph>
<paragraph pageId="0" pageNumber="79">
Sternite VIII (Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Coarctana nigromedia sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure20" httpUri="https://binary.pensoft.net/fig/798173" pageId="0" pageNumber="79">20E</figureCitation>
) 1.3
<normalizedToken originalValue="×">x</normalizedToken>
wider than long; lateral margins weakly convergent posterad; posterior margin straight. Valve (Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Coarctana nigromedia sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure20" httpUri="https://binary.pensoft.net/fig/798173" pageId="0" pageNumber="79">20F</figureCitation>
) 1.8
<normalizedToken originalValue="×">x</normalizedToken>
wider than long; posterior margin rounded. Pygofer, in lateral view (Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Coarctana nigromedia sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure20" httpUri="https://binary.pensoft.net/fig/798173" pageId="0" pageNumber="79">20G</figureCitation>
), 1.7
<normalizedToken originalValue="×">x</normalizedToken>
longer than maximum height; basodorsal process thin and short, extending ventrally for 1/5 height of lateral lobe; anteroventral margin rounded, posterodorsal and posteroventral margins straight and weakly convergent posterad; apex broad and excavated, forming two rounded tips. Subgenital plate, in lateral view (Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Coarctana nigromedia sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure20" httpUri="https://binary.pensoft.net/fig/798173" pageId="0" pageNumber="79">20G</figureCitation>
), not reaching pygofer apex; in ventral view (Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Coarctana nigromedia sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure20" httpUri="https://binary.pensoft.net/fig/798173" pageId="0" pageNumber="79">20H</figureCitation>
), elongated, 4
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide, maximum width near median third; long filiform setae on ventral surface near outer margin and dorsal surface near base; inner margin slightly rounded; outer margin straight; apex tapered, subacute. Connective (Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Coarctana nigromedia sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure20" httpUri="https://binary.pensoft.net/fig/798173" pageId="0" pageNumber="79">20I</figureCitation>
) about as wide as long; anterior margin excavated; dorsal keel reduced; stem short. Style, in dorsal view (Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Coarctana nigromedia sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure20" httpUri="https://binary.pensoft.net/fig/798173" pageId="0" pageNumber="79">20J</figureCitation>
), with outer lobe developed and rounded; in lateral view (Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Coarctana nigromedia sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure20" httpUri="https://binary.pensoft.net/fig/798173" pageId="0" pageNumber="79">20K</figureCitation>
), blade with approximately same width along entire length; apical third slightly curved dorsally, with ventral margin serrated; apex truncate, with dorsal process short and acute. Aedeagus (Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Coarctana nigromedia sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure20" httpUri="https://binary.pensoft.net/fig/798173" pageId="0" pageNumber="79">20L, M</figureCitation>
) preatrium weakly developed; dorsal apodeme with dorsal margin straight, lateral margins slightly produced laterally; apodemal processes narrow basally, curved in right angle dorsally in the basal third; ventral margin forming rounded lobe produced ventrally; distal portion straight, apex rounded and curved over the apex of shaft; shaft symmetrical and approximately cylindrical, strongly curved dorsally near base, with small basolateral expansion that surrounds the previous portion of shaft; apex with pair of processes directed ventrally and slightly curved laterally, with 1/4 length of shaft. Other characteristics as in generic description.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="female terminalia">
<paragraph pageId="0" pageNumber="79">Female terminalia.</paragraph>
<paragraph pageId="0" pageNumber="79">
Sternite VII (Fig.
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Coarctana nigromedia sp. nov., paratype female. A distal portion of abdomen, ventral view. B distal portion of abdomen, lateral view. C first valvifer and first valvula, lateral view. D apical portion of first valvula. E second valvula, lateral view. F apical portion of second valvula. G second valvifer and gonoplac, lateral view. H apical portion of gonoplac. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure21" httpUri="https://binary.pensoft.net/fig/798174" pageId="0" pageNumber="79">21A</figureCitation>
) 1.8
<normalizedToken originalValue="×">x</normalizedToken>
wider than long; posterolateral angles rounded; posterior margin deeply excavated each side of median lobe which occupies the median third and is V-shape notched medially. Internal sternite VIII membranous. Pygofer (Fig.
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Coarctana nigromedia sp. nov., paratype female. A distal portion of abdomen, ventral view. B distal portion of abdomen, lateral view. C first valvifer and first valvula, lateral view. D apical portion of first valvula. E second valvula, lateral view. F apical portion of second valvula. G second valvifer and gonoplac, lateral view. H apical portion of gonoplac. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure21" httpUri="https://binary.pensoft.net/fig/798174" pageId="0" pageNumber="79">21A, B</figureCitation>
) 1.8
<normalizedToken originalValue="×">x</normalizedToken>
longer than maximum height; ventral margin slightly rounded; dorsal margin straight; apex rounded. First valvifer (Fig.
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Coarctana nigromedia sp. nov., paratype female. A distal portion of abdomen, ventral view. B distal portion of abdomen, lateral view. C first valvifer and first valvula, lateral view. D apical portion of first valvula. E second valvula, lateral view. F apical portion of second valvula. G second valvifer and gonoplac, lateral view. H apical portion of gonoplac. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure21" httpUri="https://binary.pensoft.net/fig/798174" pageId="0" pageNumber="79">21C</figureCitation>
) triangular, 1.5
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide; anterior and dorsal margins straight; posterior margin slightly rounded. First valvula (Fig.
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Coarctana nigromedia sp. nov., paratype female. A distal portion of abdomen, ventral view. B distal portion of abdomen, lateral view. C first valvifer and first valvula, lateral view. D apical portion of first valvula. E second valvula, lateral view. F apical portion of second valvula. G second valvifer and gonoplac, lateral view. H apical portion of gonoplac. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure21" httpUri="https://binary.pensoft.net/fig/798174" pageId="0" pageNumber="79">21C</figureCitation>
) 6.5
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide; apex (Fig.
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Coarctana nigromedia sp. nov., paratype female. A distal portion of abdomen, ventral view. B distal portion of abdomen, lateral view. C first valvifer and first valvula, lateral view. D apical portion of first valvula. E second valvula, lateral view. F apical portion of second valvula. G second valvifer and gonoplac, lateral view. H apical portion of gonoplac. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure21" httpUri="https://binary.pensoft.net/fig/798174" pageId="0" pageNumber="79">21D</figureCitation>
) tapered and acute. Second valvula (Fig.
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Coarctana nigromedia sp. nov., paratype female. A distal portion of abdomen, ventral view. B distal portion of abdomen, lateral view. C first valvifer and first valvula, lateral view. D apical portion of first valvula. E second valvula, lateral view. F apical portion of second valvula. G second valvifer and gonoplac, lateral view. H apical portion of gonoplac. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure21" httpUri="https://binary.pensoft.net/fig/798174" pageId="0" pageNumber="79">21E</figureCitation>
) 4.5
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide; wider after mid-length; dorsal protuberance developed and rounded; dorsal margin (Fig.
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Coarctana nigromedia sp. nov., paratype female. A distal portion of abdomen, ventral view. B distal portion of abdomen, lateral view. C first valvifer and first valvula, lateral view. D apical portion of first valvula. E second valvula, lateral view. F apical portion of second valvula. G second valvifer and gonoplac, lateral view. H apical portion of gonoplac. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure21" httpUri="https://binary.pensoft.net/fig/798174" pageId="0" pageNumber="79">21F</figureCitation>
) with few minute teeth; apex abruptly tapered, subacute. Second valvifer (Fig.
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Coarctana nigromedia sp. nov., paratype female. A distal portion of abdomen, ventral view. B distal portion of abdomen, lateral view. C first valvifer and first valvula, lateral view. D apical portion of first valvula. E second valvula, lateral view. F apical portion of second valvula. G second valvifer and gonoplac, lateral view. H apical portion of gonoplac. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure21" httpUri="https://binary.pensoft.net/fig/798174" pageId="0" pageNumber="79">21G</figureCitation>
) 2.5
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide. Gonoplac (Fig.
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Coarctana nigromedia sp. nov., paratype female. A distal portion of abdomen, ventral view. B distal portion of abdomen, lateral view. C first valvifer and first valvula, lateral view. D apical portion of first valvula. E second valvula, lateral view. F apical portion of second valvula. G second valvifer and gonoplac, lateral view. H apical portion of gonoplac. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure21" httpUri="https://binary.pensoft.net/fig/798174" pageId="0" pageNumber="79">21G</figureCitation>
) 3.3
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide; apex (Fig.
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Coarctana nigromedia sp. nov., paratype female. A distal portion of abdomen, ventral view. B distal portion of abdomen, lateral view. C first valvifer and first valvula, lateral view. D apical portion of first valvula. E second valvula, lateral view. F apical portion of second valvula. G second valvifer and gonoplac, lateral view. H apical portion of gonoplac. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure21" httpUri="https://binary.pensoft.net/fig/798174" pageId="0" pageNumber="79">21H</figureCitation>
) rounded. Other characteristics as in generic description.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="materials_examined">
<paragraph pageId="0" pageNumber="79">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="79">
<materialsCitation collectingDate="1986-10-13" country="Brazil" county="Jundiai do Sul" location="Brasil" municipality="Fazenda Monte Verde" specimenCount="1" stateProvince="Parana" typeStatus="Holotype">
<emphasis bold="true" pageId="0" pageNumber="79">
<typeStatus>Holotype</typeStatus>
male:
<collectingCountry name="Brazil">Brazil</collectingCountry>
:
<normalizedToken originalValue="Paraná">
<collectingRegion country="Brazil" name="Parana">Parana</collectingRegion>
</normalizedToken>
</emphasis>
:
<normalizedToken originalValue="“Jundiaí">
&quot;
<collectingCounty>Jundiai do Sul</collectingCounty>
</normalizedToken>
-
<collectingRegion country="Brazil" name="Para">PR</collectingRegion>
\
<collectingMunicipality>Fazenda Monte Verde</collectingMunicipality>
\
<location LSID="urn:lsid:plazi:treatment:34519A1CC5195B04A203A3B3F09E66E5:6720AA8A4EFBC3AE6268CEB0BEB59467" country="Brazil" county="Jundiai do Sul" municipality="Fazenda Monte Verde" name="Brasil" stateProvince="Parana">Brasil</location>
<collectingDate value="1986-10-13">13.X.1986</collectingDate>
\ Lev. Ent. Profaupar\ Malaise&quot; (DZUP)
</materialsCitation>
.
<materialsCitation collectingDate="1986-10-20" country="Brazil" county="Jundiai do Sul" location="Brasil" municipality="Fazenda Monte Verde" specimenCount="1" specimenCount-female="1" stateProvince="Parana" typeStatus="Paratypes">
<emphasis bold="true" pageId="0" pageNumber="79">
<typeStatus>Paratypes</typeStatus>
</emphasis>
:
<specimenCount type="female">1♀</specimenCount>
, same data as holotype except
<collectingDate value="1986-10-20">20.X.1986</collectingDate>
(DZUP)
</materialsCitation>
;
<materialsCitation collectingDate="1987-03-02" country="Brazil" county="Jundiai do Sul" location="Brasil" municipality="Fazenda Monte Verde" specimenCount="1" specimenCount-male="1" stateProvince="Parana" typeStatus="paratype">
<specimenCount type="male">1♂</specimenCount>
, same data as holotype except
<collectingDate value="1987-03-02">02.III.1987</collectingDate>
(DZUP)
</materialsCitation>
;
<materialsCitation collectingDate="1987-10-19" country="Brazil" county="Jundiai do Sul" location="Brasil" municipality="Fazenda Monte Verde" specimenCount="1" specimenCount-male="1" stateProvince="Parana" typeStatus="paratype">
<specimenCount type="male">1♂</specimenCount>
, same data as holotype except
<collectingDate value="1987-10-19">19.X.1987</collectingDate>
(DZUP)
</materialsCitation>
;
<materialsCitation collectingDate="1988-04-11" country="Brazil" county="Jundiai do Sul" location="Brasil" municipality="Fazenda Monte Verde" specimenCount="1" specimenCount-male="1" stateProvince="Parana" typeStatus="paratype">
<specimenCount type="male">1♂</specimenCount>
, same data as holotype except
<collectingDate value="1988-04-11">11.IV.1988</collectingDate>
(DZUP)
</materialsCitation>
;
<materialsCitation collectingDate="2016-09-29" collectorName="M. Atlantica, Gomez" country="Brazil" elevation="545" latitude="-23.189693" location="Londrina" longLatPrecision="1" longitude="-51.171864" municipality="Brasil" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="PR" typeStatus="paratype">
<specimenCount type="male">1♂</specimenCount>
,
<specimenCount type="female">2♀</specimenCount>
, &quot;
<collectingMunicipality>Brasil</collectingMunicipality>
,
<collectingRegion country="Brazil" name="Para">PR</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:34519A1CC5195B04A203A3B3F09E66E5:E4D884CD747D679D0C27C8946161AC7B" country="Brazil" latitude="-23.189693" longLatPrecision="1" longitude="-51.171864" municipality="Brasil" name="Londrina" stateProvince="PR">Londrina</location>
,\
<collectorName>
M.
<normalizedToken originalValue="Atlântica">Atlantica</normalizedToken>
</collectorName>
,
<geoCoordinate degrees="23" direction="south" minutes="11" orientation="latitude" precision="1" seconds="22.9" value="-23.189693">23°1122.9″S</geoCoordinate>
,
<geoCoordinate degrees="51" direction="west" minutes="10" orientation="longitude" precision="1" seconds="18.7" value="-51.171864">51°1018.7″W</geoCoordinate>
,
<quantity metricMagnitude="2" metricUnit="m" metricValue="5.45" unit="m" value="545.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="5.45" unit="m" value="545.0">545m</elevation>
</quantity>
,\
<collectingDate value="2016-09-29">29.IX.2016</collectingDate>
, D. R. Sosa-\
<collectorName>Gomez</collectorName>
leg.&quot; (DZUP)
</materialsCitation>
;
<materialsCitation collectingDate="2015-09-17" collectorName="M. Atlantica &amp; Gomez" country="Brazil" elevation="545" latitude="-23.189693" location="Londrina" longLatPrecision="1" longitude="-51.171864" municipality="Brasil" specimenCount="1" specimenCount-male="1" stateProvince="Para" typeStatus="paratype">
<specimenCount type="male">1♂</specimenCount>
, same data as preceding except
<collectingDate value="2015-09-17">17.IX.2015</collectingDate>
(MZSP)
</materialsCitation>
;
<materialsCitation collectingDate="2016-09-30" collectorName="M. Atlantica &amp; Gomez" country="Brazil" elevation="545" latitude="-23.189693" location="Londrina" longLatPrecision="1" longitude="-51.171864" municipality="Brasil" specimenCount="1" specimenCount-male="1" stateProvince="Para" typeStatus="paratype">
<specimenCount type="male">1♂</specimenCount>
, same data as preceding except
<collectingDate value="2016-09-30">30.IX.2016</collectingDate>
(MZSP)
</materialsCitation>
;
<materialsCitation collectingDate="2017-02-22" collectorName="M. Atlantica &amp; Gomez" country="Brazil" elevation="545" latitude="-23.189693" location="Distrito Federal" longLatPrecision="1" longitude="-51.171864" municipality="Brasil" specimenCount="1" specimenCount-female="1" stateProvince="Distrito Federal" typeStatus="paratype">
<specimenCount type="female">1♀</specimenCount>
, same data as preceding except
<collectingDate value="2017-02-22">22.II.2017</collectingDate>
. -
<emphasis bold="true" pageId="0" pageNumber="79">
<collectingRegion country="Brazil" name="Distrito Federal">Distrito Federal</collectingRegion>
</emphasis>
</materialsCitation>
:
<materialsCitation collectingDate="2017-11-23" collectorName="M. Galeria, J. R. P. Luz" country="Brazil" county="Brasil" location="Fazenda Agua Limpa" municipality="Brasilia" specimenCount="16" specimenCount-female="4" specimenCount-male="12" stateProvince="Distrito Federal" typeStatus="paratype">
<specimenCount type="male">6♂</specimenCount>
,
<specimenCount type="female">2♀</specimenCount>
, &quot;
<collectingCounty>Brasil</collectingCounty>
,
<normalizedToken originalValue="Brasília">
<collectingMunicipality>Brasilia</collectingMunicipality>
</normalizedToken>
- DF,\
<location LSID="urn:lsid:plazi:treatment:34519A1CC5195B04A203A3B3F09E66E5:DDFACD5F5E3235F26B3D5E246E51FEC9" country="Brazil" county="Brasil" municipality="Brasilia" name="Fazenda Agua Limpa" stateProvince="Distrito Federal">
Fazenda
<normalizedToken originalValue="Água">Agua</normalizedToken>
Limpa
</location>
,\
<collectorName>M. Galeria</collectorName>
, malaise, 09-\
<collectingDate value="2017-11-23">23.XI.2017</collectingDate>
,
<collectorName>J.R.P. Luz</collectorName>
&quot; (
<specimenCount type="female">1♀</specimenCount>
DZRJ,
<specimenCount type="male">2♂</specimenCount>
MNRJ,
<specimenCount type="male">4♂</specimenCount>
,
<specimenCount type="female">1♀</specimenCount>
DZUP)
</materialsCitation>
;
<materialsCitation collectingDate="2017-04-27" collectorName="M. Galeria &amp; J. R. P. Luz" country="Brazil" county="Brasil" location="Minas Gerais" municipality="Brasilia" specimenCount="1" specimenCount-male="1" stateProvince="Minas Gerais" typeStatus="paratype">
<specimenCount type="male">1♂</specimenCount>
, same data as preceding except
<collectingDate value="2017-04-27">27.IV.2017</collectingDate>
(DZRJ). -
<emphasis bold="true" pageId="0" pageNumber="79">
<collectingRegion country="Brazil" name="Minas Gerais">Minas Gerais</collectingRegion>
</emphasis>
</materialsCitation>
:
<materialsCitation collectingDate="2017-01-30" collectorName="Moreira" country="Brazil" county="Brasil" elevation="784" latitude="-19.66252" location="Cerrado" longLatPrecision="1" longitude="-47.127544" municipality="Uberaba" specimenCount="1" specimenCount-male="1" stateProvince="Minas Gerais" typeStatus="paratype">
<specimenCount type="male">1♂</specimenCount>
, &quot;
<collectingCounty>Brasil</collectingCounty>
, MG,
<collectingMunicipality>Uberaba</collectingMunicipality>
,\
<location LSID="urn:lsid:plazi:treatment:34519A1CC5195B04A203A3B3F09E66E5:C5881102E7FE7ADC91634C2FF9E56C6A" country="Brazil" county="Brasil" latitude="-19.66252" longLatPrecision="1" longitude="-47.127544" municipality="Uberaba" name="Cerrado" stateProvince="Minas Gerais">Cerrado</location>
,
<geoCoordinate degrees="19" direction="south" minutes="39" orientation="latitude" precision="1" seconds="45.07" value="-19.66252">19°3945.07″S</geoCoordinate>
<geoCoordinate degrees="47" direction="west" minutes="7" orientation="longitude" precision="1" seconds="39.16" value="-47.127544">47°739.16″W</geoCoordinate>
,
<quantity metricMagnitude="2" metricUnit="m" metricValue="7.84" unit="m" value="784.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="7.84" unit="m" value="784.0">784m</elevation>
</quantity>
,\
<collectingDate value="2017-01-30">30.I.2017</collectingDate>
, B.M. \
<collectorName>Moreira</collectorName>
leg.&quot; (DZUP)
</materialsCitation>
;
<materialsCitation collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" country="Brazil" location="Sao Paulo" specimenCount="1" specimenCount-male="1" stateProvince="Sao Paulo" typeStatus="paratype">
<specimenCount type="male">1♂</specimenCount>
, ″
<collectingRegion country="Brazil" name="Sao Paulo">
<normalizedToken originalValue="São">Sao</normalizedToken>
Paulo
</collectingRegion>
</materialsCitation>
\
<materialsCitation collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" collectorName="Pres. Epitacio, Pte. Albano, J. Lane" country="Brazil" location="Brasil" specimenCount="1" specimenCount-male="1" stateProvince="Tocantins" typeStatus="paratype">
<collectorName>
Pres.
<normalizedToken originalValue="Epitácio">Epitacio</normalizedToken>
</collectorName>
\
<collectorName>Pte. Albano</collectorName>
\ X-54\
<collectorName>J. Lane</collectorName>
col.&quot; (MZSP). -
<emphasis bold="true" pageId="0" pageNumber="79">
<collectingRegion country="Brazil" name="Tocantins">Tocantins</collectingRegion>
</emphasis>
:
<specimenCount type="male">1♂</specimenCount>
, &quot;
<location LSID="urn:lsid:plazi:treatment:34519A1CC5195B04A203A3B3F09E66E5:EF2A850017F707C28EB54A30176BA046" country="Brazil" name="Brasil" stateProvince="Tocantins">Brasil</location>
,
<collectingRegion country="Brazil" name="Tocantins">Tocantins</collectingRegion>
</materialsCitation>
,
<materialsCitation collectingDate="2012-06-22" collectingDateMax="2012-06-29" collectingDateMin="2012-06-22" collectorName="Malaise, Krolow, Lima, H. I. L." country="Brazil" county="Palmas" latitude="-10.250639" location="Fazenda" longLatPrecision="1" longitude="-48.126" municipality="Taquarucu" specimenCount="1" stateProvince="Tocantins" typeStatus="paratype">
<collectingCounty>Palmas</collectingCounty>
\
<normalizedToken originalValue="Taquaruçu">
<collectingMunicipality>Taquarucu</collectingMunicipality>
</normalizedToken>
,
<location LSID="urn:lsid:plazi:treatment:34519A1CC5195B04A203A3B3F09E66E5:BEAD2E012840B20FB7EC5C9F3BF0D338" country="Brazil" county="Palmas" latitude="-10.250639" longLatPrecision="1" longitude="-48.126" municipality="Taquarucu" name="Fazenda" stateProvince="Tocantins">Fazenda</location>
\
<location LSID="urn:lsid:plazi:treatment:34519A1CC5195B04A203A3B3F09E66E5:C6E4A34D17C3D7802853621D9DFC2B3B" country="Brazil" county="Palmas" latitude="-10.250639" longLatPrecision="1" longitude="-48.126" municipality="Taquarucu" name="Encantada" stateProvince="Tocantins">Encantada</location>
<geoCoordinate degrees="10" direction="south" minutes="15" orientation="latitude" precision="1" seconds="2.3" value="-10.250639">10°152.3″S</geoCoordinate>
<geoCoordinate degrees="48" direction="west" minutes="07" orientation="longitude" precision="1" seconds="33.6" value="-48.126">48°0733.6″W</geoCoordinate>
<collectorName>Malaise</collectorName>
\
<collectingDate value="2012-06-22" valueMax="2012-06-29" valueMin="2012-06-22">22-29.vi.2012</collectingDate>
<collectorName>Krolow</collectorName>
,\ T.K. &amp;
<collectorName>Lima, H.I.L.</collectorName>
leg.&quot; (DZUP)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="etymology">
<paragraph pageId="0" pageNumber="79">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="79">The new species name refers to the wide dark-brown band between the ocelli.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="remarks">
<paragraph pageId="0" pageNumber="79">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="79">
<taxonomicName authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" family="Cicadellidae" genus="Coarctana" kingdom="Animalia" lsidName="Coarctana nigromedia" order="Hemiptera" pageId="0" pageNumber="79" rank="species" species="nigromedia">
<emphasis italics="true" pageId="0" pageNumber="79">Coarctana nigromedia</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="79">sp. nov.</emphasis>
can be distinguished from
<taxonomicName authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" family="Cicadellidae" kingdom="Animalia" lsidName="C. asymmetrica" order="Hemiptera" pageId="0" pageNumber="79" rank="species" species="asymmetrica">
<emphasis italics="true" pageId="0" pageNumber="79">C. asymmetrica</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="79">sp. nov.</emphasis>
by the style shape (Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Coarctana nigromedia sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure20" httpUri="https://binary.pensoft.net/fig/798173" pageId="0" pageNumber="79">20K</figureCitation>
) with apex lacking process on ventral margin and the aedeagal shaft (Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Coarctana nigromedia sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure20" httpUri="https://binary.pensoft.net/fig/798173" pageId="0" pageNumber="79">20L, M</figureCitation>
) with pair of apical processes symmetric in length.
</paragraph>
</subSubSection>
</treatment>
</document>