treatments-xml/data/33/57/87/335787BBE01AB160C82DFEC1D3040E08.xml

<document id="0B74ACEDF53FF16D9FF4AEFF4620FF6B" ID-CLB-Dataset="51437" ID-DOI="10.5281/zenodo.155768" ID-GBIF-Dataset="ba446901-2a3b-43c5-826a-8f2f75a7495b" ID-ISSN="1175­5326" ID-Zenodo-Dep="155768" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459435950089" checkinUser="plazi" docAuthor="Gusarov, Vladimir I. &amp; Koval, Alexander G." docDate="2002" docId="335787BBE01AB160C82DFEC1D3040E08" docLanguage="en" docName="zt00069.pdf" docOrigin="Zootaxa 69" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Heinzia caucasica Gusarov &amp; Koval, sp. nov." docType="treatment" docVersion="15" lastPageNumber="11" masterDocId="CF6EFFC3E01EB16AC925FFDAD7230A0E" masterDocTitle="A revision of the genus Heinzia Korge, 1971 (Coleoptera: Staphylinidae: Quediina), with description of a new species and its probable larva" masterLastPageNumber="19" masterPageNumber="1" pageNumber="5" updateTime="1698192890222" updateUser="ExternalLinkService">
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<mods:title id="E45FA04E2EEFE4E226AD9AA249D78797">A revision of the genus Heinzia Korge, 1971 (Coleoptera: Staphylinidae: Quediina), with description of a new species and its probable larva</mods:title>
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<mods:namePart id="CB48B1135616B2DC4C07CBE53971D6A2">Gusarov, Vladimir I.</mods:namePart>
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<mods:namePart id="97C09129D4CAA1BF1EDA7D6260C2BDD7">Koval, Alexander G.</mods:namePart>
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<mods:date id="8D68F39313A84D77019FC30AA4207F34">2002</mods:date>
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<treatment id="335787BBE01AB160C82DFEC1D3040E08" ID-DOI="http://doi.org/10.5281/zenodo.6277686" ID-GBIF-Taxon="119329877" ID-Zenodo-Dep="6277686" LSID="urn:lsid:plazi:treatment:335787BBE01AB160C82DFEC1D3040E08" httpUri="http://treatment.plazi.org/id/335787BBE01AB160C82DFEC1D3040E08" lastPageId="10" lastPageNumber="11" pageId="4" pageNumber="5">
<subSubSection id="F3E46526E01AB16EC82DFEC1D6F90B53" pageId="4" pageNumber="5" type="nomenclature">
<paragraph id="BB4136ADE01AB16EC82DFEC1D6F90B53" blockId="4.[264,1323,283,349]" pageId="4" pageNumber="5">
<heading id="E00981C1E01AB16EC82DFEC1D6F90B53" bold="true" fontSize="11" level="1" pageId="4" pageNumber="5" reason="1">
<emphasis id="898AEABFE01AB16EC82DFEC1D6F90B53" bold="true" pageId="4" pageNumber="5">
1. 
<taxonomicName id="7CFE4D2EE01AB16EC803FEC6D5CB0B3B" ID-CoL="3K3VC" authority="Gusarov &amp; Koval" authorityName="Gusarov &amp; Koval" box="[294,744,283,310]" class="Insecta" family="Staphylinidae" genus="Heinzia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="caucasica" status="sp. nov.">
<emphasis id="898AEABFE01AB16EC803FEC6D5210B38" bold="true" box="[294,514,284,310]" italics="true" pageId="4" pageNumber="5">Heinzia caucasica</emphasis>
Gusarov &amp; Koval
</taxonomicName>
, 
<taxonomicNameLabel id="92B957C4E01AB16ECBD1FEC1D4710B3B" box="[756,850,283,309]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="23C52A28E01AB16ECA44FEC1D3800B3B" box="[865,1187,283,309]" captionStart-0="FIGURES 1 ­ 7" captionStart-1="FIGURES 8 ­ 13" captionStart-2="FIGURES 14 ­ 19" captionStartId-0="3.[264,382,1783,1807]" captionStartId-1="5.[264,383,1583,1607]" captionStartId-2="6.[264,383,1576,1600]" captionTargetBox-0="[289,1293,450,1752]" captionTargetBox-1="[287,1301,335,1549]" captionTargetBox-2="[286,1299,499,1546]" captionTargetId-0="figure@3.[283,1304,440,1756]" captionTargetId-1="figure@5.[278,1308,329,1556]" captionTargetId-2="figure@6.[282,1306,489,1549]" captionTargetPageId-0="3" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="FIGURES 1 ­ 7. Mouthparts of Heinzia caucasica Gusarov &amp; Koval, sp. nov. (paratype from Baribana cave). 1 – epipharynx; 2 – labrum; 3 – left mandible, dorsal view; 4 – right maxilla, ventral view; 5 – prementum; 6 – hypopharynx, left portion not shown; 7 – mentum. Scale bar 0.4 mm (1 ­ 5), 0.2 mm (6 ­ 7)." captionText-1="FIGURES 8 ­ 13. Details of Heinzia caucasica Gusarov &amp; Koval, sp. nov. (8, 10, holotype; 12 ­ 13, paratype from Baribana cave) and H. variabilis Korge (9, Kalkanl Da {lar; 11, holotype). 8 ­ 9 – head; 10 ­ 11 – pronotum; 12 – left antenna; 13 – right metatarsus. Scale bar 1 mm (8 ­ 12), 0.4 mm (13)." captionText-2="FIGURES 14 ­ 19. Details of Heinzia caucasica Gusarov &amp; Koval, sp. nov. (14, 16, 18, paratype from Dolgaya cave) and H. variabilis Korge (15, 17, 19, Kalkanl Da {lar). 14 ­ 15 – male sternum 8; 16 ­ 17 – apex of mail sternum 8; 18 ­ 19 – male sternum 9. Scale bar 1 mm (14 ­ 15), 0.5 mm (16 ­ 19)." httpUri-0="https://zenodo.org/record/155769/files/figure.png" httpUri-1="https://zenodo.org/record/155770/files/figure.png" httpUri-2="https://zenodo.org/record/155772/files/figure.png" pageId="4" pageNumber="5">Figs. 1­8, 10, 12­14, 16, 18</figureCitation>
, 
<figureCitation id="23C52A28E01AB16ECD94FEC1D3D50B3B" box="[1201,1270,283,309]" captionStart="FIGURES 20 ­ 23" captionStartId="7.[264,383,1764,1788]" captionTargetBox="[300,1282,579,1733]" captionTargetId="figure@7.[295,1292,569,1737]" captionTargetPageId="7" captionText="FIGURES 20 ­ 23. Aedeagus of Heinzia caucasica Gusarov &amp; Koval, sp. nov. (20 ­ 21, paratype from Mezmay) and H. variabilis Korge (22 ­ 23, Kalkanl Da {lar). 20, 22 – ventral view; 21, 23, lateral view. Scale bar 0.4 mm." httpUri="https://zenodo.org/record/155773/files/figure.png" pageId="4" pageNumber="5">20­21</figureCitation>
, 
<figureCitation id="23C52A28E01AB16ECC21FEC1D6040B53" captionStart="FIGURES 24 ­ 27" captionStartId="8.[264,383,1791,1815]" captionTargetBox="[281,1306,495,1759]" captionTargetId="figure@8.[272,1315,489,1765]" captionTargetPageId="8" captionText="FIGURES 24 ­ 27. Details of aedeagus of Heinzia caucasica Gusarov &amp; Koval, sp. nov. (24 ­ 25, holotype) and H. variabilis Korge (26 ­ 27, holotype). 24, 26 – median lobe, ventral view; 25, 27, paramere. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/155774/files/figure.png" pageId="4" pageNumber="5">24­ 25</figureCitation>
, 
<figureCitation id="23C52A28E01AB16EC811FE99D65A0B53" box="[308,377,323,349]" captionStart="FIGURES 28 ­ 32" captionStartId="9.[264,383,1416,1440]" captionTargetBox="[324,1261,853,1386]" captionTargetId="figure@9.[318,1269,849,1389]" captionTargetPageId="9" captionText="FIGURES 28 ­ 32. Apex of median lobe of aedeagus (lateral view) of Heinzia caucasica Gusarov &amp; Koval, sp. nov. (28, paratypes from Mezmay; 29 ­ 30, Baribana cave; 31, Akhty) and H. variabilis Korge (32, Kalkanl Da {lar). Scale bar 0.2 mm." httpUri="https://zenodo.org/record/155775/files/figure.png" pageId="4" pageNumber="5">28­31</figureCitation>
, 
<figureCitation id="23C52A28E01AB16EC8A2FE99D6F20B53" box="[391,465,323,349]" captionStart="FIGURES 33 ­ 35" captionStartId="11.[264,383,1747,1771]" captionTargetBox="[315,1269,719,1731]" captionTargetId="figure@11.[310,1278,713,1736]" captionTargetPageId="11" captionText="FIGURES 33 ­ 35. Details of female tergum 10 of Heinzia caucasica Gusarov &amp; Koval, sp. nov. (33, 35, Baribana cave) and H. variabilis Korge (34, paratype). 33 ­ 34 – dorsal view; 35 – lateral view; LL – lateral lobe of female tergum 10; MP – medial process of female tergum 10. Scale bar 0.3 mm." httpUri="https://zenodo.org/record/155776/files/figure.png" pageId="4" pageNumber="5">33, 35</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="F3E46526E01AB16EC82DFE4ED4940BA5" box="[264,951,404,427]" pageId="4" pageNumber="5" type="reference_group">
<paragraph id="BB4136ADE01AB16EC82DFE4ED4940BA5" blockId="4.[264,951,404,427]" box="[264,951,404,427]" pageId="4" pageNumber="5">
<treatmentCitationGroup id="9BEE1183E01AB16EC82DFE4ED4940BA5" box="[264,951,404,427]" pageId="4" pageNumber="5">
<taxonomicName id="7CFE4D2EE01AB16EC82DFE4ED5C00BA5" ID-CoL="3K3VD" authority="Solodovnikov, 1998: 16" authorityName="Solodovnikov" authorityPageNumber="16" authorityYear="1998" box="[264,739,404,427]" class="Insecta" family="Staphylinidae" genus="Heinzia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis id="898AEABFE01AB16EC82DFE4ED6E40BA5" box="[264,455,404,427]" italics="true" pageId="4" pageNumber="5">Heinzia variabilis</emphasis>
: 
<treatmentCitation id="3A5F10BCE01AB16EC8F8FE4ED5C00BA5" author="Solodovnikov" box="[477,739,404,427]" page="16" pageId="4" pageNumber="5" year="1998">
<bibRefCitation id="DF6F4B5CE01AB16EC8F8FE4ED5940BA5" author="Solodovnikov" box="[477,695,404,427]" pageId="4" pageNumber="19" refString="Solodovnikov, A. (1998) On diagnostics, distribution and ecology of the little-known staphylinid beetles (Coleoptera, Staphylinidae) of the Caucasus. [In Russian] Vestnik Sankt-Peterburgskogo Universiteta, Seria 3, Biologia, Vypusk 3 (No. 17), 13 - 20." type="journal article" year="1998">Solodovnikov, 1998</bibRefCitation>
: 16
</treatmentCitation>
</taxonomicName>
(misidentification).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="F3E46526E01AB16EC82DFE04D3000EC7" pageId="4" pageNumber="5" type="materials_examined">
<paragraph id="BB4136ADE01AB16EC82DFE04D4B4082F" blockId="4.[264,1325,478,1864]" pageId="4" pageNumber="5">
<emphasis id="898AEABFE01AB16EC82DFE04D69F0BF6" bold="true" box="[264,444,478,504]" pageId="4" pageNumber="5">
<typeStatus id="6445880FE01AB16EC82DFE04D6600BF6" box="[264,323,478,504]" pageId="4" pageNumber="5">Type</typeStatus>
material.
</emphasis>
<typeStatus id="6445880FE01AB16EC8E8FE05D51C0BF7" box="[461,575,479,505]" pageId="4" pageNumber="5" type="holotype">Holotype</typeStatus>
: 
<collectingCountry id="C3E9763DE01AB16ECB74FE05D5870BF7" box="[593,676,479,505]" name="Russia" pageId="4" pageNumber="5">Russia</collectingCountry>
: Krasnodar Terr.:, Sochi, Alek Mountain Ridge, Baribana Cave, 
<quantity id="7C069B48E01AB16EC886FDDDD6CD082F" box="[419,494,519,545]" metricMagnitude="2" metricUnit="m" metricValue="7.4" pageId="4" pageNumber="5" unit="m" value="740.0">740 m</quantity>
(A.G.Koval) 
<date id="CF40106DE01AB16ECBB5FDDDD43D082F" box="[656,798,519,545]" pageId="4" pageNumber="5" value="1996-08-22">22.viii.1996</date>
(
<collectionCode id="DDEFAE68E01AB16ECA0BFDDDD4A8082F" box="[814,907,519,545]" pageId="4" pageNumber="5">ZINAS</collectionCode>
).
</paragraph>
<paragraph id="BB4136ADE01AB16EC81DFDF5D3000EC7" blockId="4.[264,1325,478,1864]" pageId="4" pageNumber="5">
<typeStatus id="6445880FE01AB16EC81DFDF5D68C0847" box="[312,431,559,585]" pageId="4" pageNumber="5" type="paratype">Paratypes</typeStatus>
: 
<emphasis id="898AEABFE01AB16EC89AFDF4D53F0846" bold="true" box="[447,540,558,584]" pageId="4" pageNumber="5">
<collectingCountry id="C3E9763DE01AB16EC89AFDF4D5340846" box="[447,535,558,584]" name="Russia" pageId="4" pageNumber="5">Russia</collectingCountry>
:
</emphasis>
Krasnodar Terr.: 3, 5, 
<specimenCount id="ADF8FD24E01AB16ECA58FDF5D3220847" box="[893,1025,559,585]" pageId="4" pageNumber="5" type="generic">1 specimen</specimenCount>
(with missing abdomen), Sochi, Alek Mountain Ridge, Baribana Cave, traps, 
<quantity id="7C069B48E01AB16ECA83FD8DD4DA087F" box="[934,1017,599,625]" metricMagnitude="2" metricUnit="m" metricValue="7.4" pageId="4" pageNumber="5" unit="m" value="740.0">740 m</quantity>
(A.G.Koval), 
<date id="CF40106DE01AB16ECD91FD8DD207087F" box="[1204,1316,599,625]" pageId="4" pageNumber="5" value="1995-06-04">4.vi.1995</date>
­ 
<date id="CF40106DE01AB16EC82DFDA5D64E0897" box="[264,365,639,665]" pageId="4" pageNumber="5" value="1996-05-07">7.v.1996</date>
(
<collectionCode id="DDEFAE68E01AB16EC85AFDA5D6F20897" box="[383,465,639,665]" pageId="4" pageNumber="5">AKCS</collectionCode>
, 
<collectionCode id="DDEFAE68E01AB16EC8C5FDA5D5130897" box="[480,560,639,665]" httpUri="http://grbio.org/cool/nwhf-jc3k" name="Adams State College Collection" pageId="4" pageNumber="5">ASCC</collectionCode>
, 
<collectionCode id="DDEFAE68E01AB16ECB1BFDA5D5AA0897" box="[574,649,639,665]" pageId="4" pageNumber="5">SPSU</collectionCode>
); 2, 5, ditto but 
<date id="CF40106DE01AB16ECABBFDA5D30E0897" box="[926,1069,639,665]" pageId="4" pageNumber="5" value="1997-08-19">19.viii.1997</date>
­ 
<date id="CF40106DE01AB16ECD10FDA5D3B90897" box="[1077,1178,639,665]" pageId="4" pageNumber="5" value="1998-05-09">9.v.1998</date>
(
<collectionCode id="DDEFAE68E01AB16ECD88FDA5D3DA0897" box="[1197,1273,639,665]" pageId="4" pageNumber="5">SPSU</collectionCode>
);,, ditto but 
<date id="CF40106DE01AB16EC880FD7DD51708CF" box="[421,564,679,705]" pageId="4" pageNumber="5" value="1998-08-19">19.viii.1998</date>
­ 
<date id="CF40106DE01AB16ECB19FD7DD5E908CF" box="[572,714,679,705]" pageId="4" pageNumber="5" value="1999-08-18">18.viii.1999</date>
(
<collectionCode id="DDEFAE68E01AB16ECBC5FD7DD40F08CF" box="[736,812,679,705]" pageId="4" pageNumber="5">SPSU</collectionCode>
); 3, ditto but 
<date id="CF40106DE01AB16ECD23FD7DD3B508CF" box="[1030,1174,679,705]" pageId="4" pageNumber="5" value="2000-08-21">21.viii.2000</date>
­ 
<date id="CF40106DE01AB16ECDBBFD7DD20808CF" box="[1182,1323,679,705]" pageId="4" pageNumber="5" value="2001-08-16">16.viii.2001</date>
(
<collectionCode id="DDEFAE68E01AB16EC835FD15D64108E7" box="[272,354,719,745]" pageId="4" pageNumber="5">AKCS</collectionCode>
, 
<collectionCode id="DDEFAE68E01AB16EC854FD15D69F08E7" box="[369,444,719,745]" pageId="4" pageNumber="5">SPSU</collectionCode>
);, ditto but (I.A.Solodovnikov), 
<date id="CF40106DE01AB16ECA7EFD15D4DC08E7" box="[859,1023,719,745]" pageId="4" pageNumber="5" value="1999-06-16" valueMax="1999-06-27" valueMin="1999-06-16">16­27.vi.1999</date>
(
<collectionCode id="DDEFAE68E01AB16ECD34FD15D34708E7" box="[1041,1124,719,745]" httpUri="http://grbio.org/cool/nwhf-jc3k" name="Adams State College Collection" pageId="4" pageNumber="5">ASCC</collectionCode>
);, Sochi, Alek Mountain Ridge, Sokolova (=Atsinskaya) Cave, traps, 
<quantity id="7C069B48E01AB16ECAF0FD2DD30F091F" box="[981,1068,759,785]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="5" unit="m" value="300.0">300 m</quantity>
(A.G.Koval), 
<date id="CF40106DE01AB16ECDCBFD2DD6B50937" pageId="4" pageNumber="5" value="1998-05-11" valueMax="1998-08-21" valueMin="1998-05-11">11.v­ 21.viii.1998</date>
(
<collectionCode id="DDEFAE68E01AB16EC883FCC5D6D10937" box="[422,498,799,825]" pageId="4" pageNumber="5">SPSU</collectionCode>
); 2,, ditto but 
<date id="CF40106DE01AB16ECBF9FCC5D4490937" box="[732,874,799,825]" pageId="4" pageNumber="5" value="1998-08-21">21.viii.1998</date>
(
<collectionCode id="DDEFAE68E01AB16ECA5EFCC5D4E40937" box="[891,967,799,825]" pageId="4" pageNumber="5">SPSU</collectionCode>
);,, 
<specimenCount id="ADF8FD24E01AB16ECD06FCC5D38B0937" box="[1059,1192,799,825]" pageId="4" pageNumber="5" type="generic">1 specimen</specimenCount>
(with missing apex of abdomen), Sochi, Dolgaya Cave, traps, 
<quantity id="7C069B48E01AB16ECA5DFC9DD4E0096F" box="[888,963,839,865]" metricMagnitude="2" metricUnit="m" metricValue="7.2" pageId="4" pageNumber="5" unit="m" value="720.0">720 m</quantity>
(A.G.Koval), 
<date id="CF40106DE01AB16ECD4BFC9DD208096F" box="[1134,1323,839,865]" pageId="4" pageNumber="5" value="1998-05-03" valueMax="1998-08-19" valueMin="1998-05-03">3.v­19.viii.1998</date>
(
<collectionCode id="DDEFAE68E01AB16EC834FCB5D67E0987" box="[273,349,879,905]" pageId="4" pageNumber="5">SPSU</collectionCode>
); 11, 40, ditto but 
<date id="CF40106DE01AB16ECBB4FCB5D4020987" box="[657,801,879,905]" pageId="4" pageNumber="5" value="1998-08-19">19.viii.1998</date>
­ 
<date id="CF40106DE01AB16ECA0FFCB5D4860987" box="[810,933,879,905]" pageId="4" pageNumber="5" value="1999-06-11">11.vi.1999</date>
(
<collectionCode id="DDEFAE68E01AB16ECA9DFCB5D3280987" box="[952,1035,879,905]" pageId="4" pageNumber="5">AKCS</collectionCode>
, 
<collectionCode id="DDEFAE68E01AB16ECD3FFCB5D3480987" box="[1050,1131,879,905]" pageId="4" pageNumber="5">ASCO</collectionCode>
, 
<collectionCode id="DDEFAE68E01AB16ECD5FFCB5D3ED0987" box="[1146,1230,879,905]" pageId="4" pageNumber="5">KSEM</collectionCode>
, 
<collectionCode id="DDEFAE68E01AB16ECDFBFCB5D2040987" box="[1246,1319,879,905]" pageId="4" pageNumber="5">SPSU</collectionCode>
, 
<collectionCode id="DDEFAE68E01AB16EC82DFC4DD64009BF" box="[264,355,919,945]" pageId="4" pageNumber="5">ZINAS</collectionCode>
);, ditto but 
<date id="CF40106DE01AB16ECB22FC4DD5FB09BF" box="[519,728,919,945]" pageId="4" pageNumber="5" value="1999-06-11" valueMax="1999-08-16" valueMin="1999-06-11">11.vi­16.viii.1999</date>
(
<collectionCode id="DDEFAE68E01AB16ECBCCFC4DD41509BF" box="[745,822,919,945]" pageId="4" pageNumber="5">SPSU</collectionCode>
); 3, Sochi, Vorontsovskaya Cave Complex, Labirintovaya Cave, traps, 
<quantity id="7C069B48E01AB16ECBB7FC65D5C309D7" box="[658,736,959,985]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="4" pageNumber="5" unit="m" value="550.0">550 m</quantity>
(A.G.Koval), 
<date id="CF40106DE01AB16ECAAAFC65D33E09D7" box="[911,1053,959,985]" pageId="4" pageNumber="5" value="1994-08-13">13.viii.1994</date>
­ 
<date id="CF40106DE01AB16ECD00FC65D3B909D7" box="[1061,1178,959,985]" pageId="4" pageNumber="5" value="1995-05-20">20.v.1995</date>
(
<collectionCode id="DDEFAE68E01AB16ECD89FC65D3DB09D7" box="[1196,1272,959,985]" pageId="4" pageNumber="5">SPSU</collectionCode>
);, 5, 
<specimenCount id="ADF8FD24E01AB16EC87DFC3DD6CA0E0F" box="[344,489,999,1025]" pageId="4" pageNumber="5" type="generic">1 specimen</specimenCount>
(without mesometathorax and abdomen), ditto but 
<date id="CF40106DE01AB16ECDB1FC3DD2000E0F" box="[1172,1315,999,1025]" pageId="4" pageNumber="5" value="1996-08-20">20.viii.1996</date>
­ 
<date id="CF40106DE01AB16EC82DFBD5D6B50E27" box="[264,406,1039,1065]" pageId="4" pageNumber="5" value="1997-08-17">17.viii.1997</date>
(
<collectionCode id="DDEFAE68E01AB16EC893FBD5D5200E27" box="[438,515,1039,1065]" pageId="4" pageNumber="5">SPSU</collectionCode>
); 4, ditto but 
<date id="CF40106DE01AB16ECA20FBD5D4B70E27" box="[773,916,1039,1065]" pageId="4" pageNumber="5" value="1997-08-17">17.viii.1997</date>
­ 
<date id="CF40106DE01AB16ECAB9FBD5D3210E27" box="[924,1026,1039,1065]" pageId="4" pageNumber="5" value="1998-05-02">2.v.1998</date>
(
<collectionCode id="DDEFAE68E01AB16ECD07FBD5D34D0E27" box="[1058,1134,1039,1065]" pageId="4" pageNumber="5">SPSU</collectionCode>
);, Mezmay (E.A.Khachikov), 
<date id="CF40106DE01AB16EC8C1FBEDD5660E5F" box="[484,581,1079,1105]" pageId="4" pageNumber="5" value="1991-07">vii.1991</date>
(
<collectionCode id="DDEFAE68E01AB16ECB7DFBEDD5870E5F" box="[600,676,1079,1105]" pageId="4" pageNumber="5">SPSU</collectionCode>
);, SE Krasnaya Polyana, Western portion of A9bga Mountain Ridge, 
<quantity id="7C069B48E01AB16EC8FAFB85D5A10E77" box="[479,642,1119,1145]" metricMagnitude="3" metricUnit="m" metricValue="1.6" metricValueMax="1.8" metricValueMin="1.4" pageId="4" pageNumber="5" unit="m" value="1600.0" valueMax="1800.0" valueMin="1400.0">1400­1800 m</quantity>
(V.Savitsky) 
<date id="CF40106DE01AB16ECA0FFB85D49B0E77" box="[810,952,1119,1145]" pageId="4" pageNumber="5" value="1995-08-23">23.viii.1995</date>
(
<collectionCode id="DDEFAE68E01AB16ECAE8FB85D3030E77" box="[973,1056,1119,1145]" httpUri="http://grbio.org/cool/nwhf-jc3k" name="Adams State College Collection" pageId="4" pageNumber="5">ASCC</collectionCode>
); Daghestan:, SSW Akhty, W of Shalbuzdaghm (V. &amp; M.Savitsky), 
<date id="CF40106DE01AB16ECA5FFB5DD32A0EAF" box="[890,1033,1159,1185]" pageId="4" pageNumber="5" value="1994-07-03" valueMax="1994-07-04" valueMin="1994-07-03">3­4.vii.1994</date>
(
<collectionCode id="DDEFAE68E01AB16ECD04FB5DD3570EAF" box="[1057,1140,1159,1185]" httpUri="http://grbio.org/cool/nwhf-jc3k" name="Adams State College Collection" pageId="4" pageNumber="5">ASCC</collectionCode>
); 
<emphasis id="898AEABFE01AB16ECDB4FB5CD3DC0EAE" bold="true" box="[1169,1279,1158,1184]" pageId="4" pageNumber="5">
<collectingCountry id="C3E9763DE01AB16ECDB4FB5CD3D90EAE" box="[1169,1274,1158,1184]" name="Georgia" pageId="4" pageNumber="5">Georgia</collectingCountry>
:
</emphasis>
, Lagodekhi Nature Reserve, 
<quantity id="7C069B48E01AB16ECB74FB75D5B80EC7" box="[593,667,1199,1225]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="4" pageNumber="5" unit="m" value="800.0">800 m</quantity>
(M.Kozlov), 
<date id="CF40106DE01AB16ECA19FB75D4980EC7" box="[828,955,1199,1225]" pageId="4" pageNumber="5" value="1989-08-01">1.viii.1989</date>
(
<collectionCode id="DDEFAE68E01AB16ECAEEFB75D3340EC7" box="[971,1047,1199,1225]" pageId="4" pageNumber="5">SPSU</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="F3E46526E01AB16EC81DFB0CD31D0FB7" pageId="4" pageNumber="5" type="diagnosis">
<paragraph id="BB4136ADE01AB16EC81DFB0CD31D0FB7" blockId="4.[264,1325,478,1864]" pageId="4" pageNumber="5">
<emphasis id="898AEABFE01AB16EC81DFB0CD6940EFE" bold="true" box="[312,439,1238,1264]" pageId="4" pageNumber="5">Diagnosis.</emphasis>
In comparison with 
<taxonomicName id="7CFE4D2EE01AB16ECB8BFB0DD4620EFE" box="[686,833,1239,1264]" class="Insecta" family="Staphylinidae" genus="Heinzia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis id="898AEABFE01AB16ECB8BFB0DD4620EFE" box="[686,833,1239,1264]" italics="true" pageId="4" pageNumber="5">H. variabilis</emphasis>
</taxonomicName>
, in 
<taxonomicName id="7CFE4D2EE01AB16ECA48FB0DD3260EFE" box="[877,1029,1239,1264]" class="Insecta" family="Staphylinidae" genus="Heinzia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="caucasica">
<emphasis id="898AEABFE01AB16ECA48FB0DD3260EFE" box="[877,1029,1239,1264]" italics="true" pageId="4" pageNumber="5">H. caucasica</emphasis>
</taxonomicName>
the disc of pronotum has denser punctation and no extensive impunctate areas (
<figureCitation id="23C52A28E01AB16ECAB1FB25D3040F17" box="[916,1063,1279,1305]" captionStart="FIGURES 8 ­ 13" captionStartId="5.[264,383,1583,1607]" captionTargetBox="[287,1301,335,1549]" captionTargetId="figure@5.[278,1308,329,1556]" captionTargetPageId="5" captionText="FIGURES 8 ­ 13. Details of Heinzia caucasica Gusarov &amp; Koval, sp. nov. (8, 10, holotype; 12 ­ 13, paratype from Baribana cave) and H. variabilis Korge (9, Kalkanl Da {lar; 11, holotype). 8 ­ 9 – head; 10 ­ 11 – pronotum; 12 – left antenna; 13 – right metatarsus. Scale bar 1 mm (8 ­ 12), 0.4 mm (13)." httpUri="https://zenodo.org/record/155770/files/figure.png" pageId="4" pageNumber="5">Figs. 10, 11</figureCitation>
); on head impunctate area is restricted to anterior margin of the disc (
<figureCitation id="23C52A28E01AB16ECA1AFAFDD4A50F4F" box="[831,902,1319,1345]" captionStart="FIGURES 8 ­ 13" captionStartId="5.[264,383,1583,1607]" captionTargetBox="[287,1301,335,1549]" captionTargetId="figure@5.[278,1308,329,1556]" captionTargetPageId="5" captionText="FIGURES 8 ­ 13. Details of Heinzia caucasica Gusarov &amp; Koval, sp. nov. (8, 10, holotype; 12 ­ 13, paratype from Baribana cave) and H. variabilis Korge (9, Kalkanl Da {lar; 11, holotype). 8 ­ 9 – head; 10 ­ 11 – pronotum; 12 – left antenna; 13 – right metatarsus. Scale bar 1 mm (8 ­ 12), 0.4 mm (13)." httpUri="https://zenodo.org/record/155770/files/figure.png" pageId="4" pageNumber="5">Fig. 8</figureCitation>
) and does not extend posteriorly to the center of the disc as in 
<taxonomicName id="7CFE4D2EE01AB16ECB6CFA95D5FD0F66" box="[585,734,1359,1384]" class="Insecta" family="Staphylinidae" genus="Heinzia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis id="898AEABFE01AB16ECB6CFA95D5FD0F66" box="[585,734,1359,1384]" italics="true" pageId="4" pageNumber="5">H. variabilis</emphasis>
</taxonomicName>
(
<figureCitation id="23C52A28E01AB16ECBD5FA95D41A0F67" box="[752,825,1359,1385]" captionStart="FIGURES 8 ­ 13" captionStartId="5.[264,383,1583,1607]" captionTargetBox="[287,1301,335,1549]" captionTargetId="figure@5.[278,1308,329,1556]" captionTargetPageId="5" captionText="FIGURES 8 ­ 13. Details of Heinzia caucasica Gusarov &amp; Koval, sp. nov. (8, 10, holotype; 12 ­ 13, paratype from Baribana cave) and H. variabilis Korge (9, Kalkanl Da {lar; 11, holotype). 8 ­ 9 – head; 10 ­ 11 – pronotum; 12 – left antenna; 13 – right metatarsus. Scale bar 1 mm (8 ­ 12), 0.4 mm (13)." httpUri="https://zenodo.org/record/155770/files/figure.png" pageId="4" pageNumber="5">Fig. 9</figureCitation>
). Additionally, 
<taxonomicName id="7CFE4D2EE01AB16ECADEFA95D3B50F66" box="[1019,1174,1359,1384]" class="Insecta" family="Staphylinidae" genus="Heinzia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="caucasica">
<emphasis id="898AEABFE01AB16ECADEFA95D3B50F66" box="[1019,1174,1359,1384]" italics="true" pageId="4" pageNumber="5">H. caucasica</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="7CFE4D2EE01AB16EC82DFAADD6BF0F9E" box="[264,412,1399,1424]" class="Insecta" family="Staphylinidae" genus="Heinzia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis id="898AEABFE01AB16EC82DFAADD6BF0F9E" box="[264,412,1399,1424]" italics="true" pageId="4" pageNumber="5">H. variabilis</emphasis>
</taxonomicName>
in having more obtuse subapical tooth of median lobe of aedeagus (
<figureCitation id="23C52A28E01AB16ECDE4FAADD6700FB7" captionStart="FIGURES 28 ­ 32" captionStartId="9.[264,383,1416,1440]" captionTargetBox="[324,1261,853,1386]" captionTargetId="figure@9.[318,1269,849,1389]" captionTargetPageId="9" captionText="FIGURES 28 ­ 32. Apex of median lobe of aedeagus (lateral view) of Heinzia caucasica Gusarov &amp; Koval, sp. nov. (28, paratypes from Mezmay; 29 ­ 30, Baribana cave; 31, Akhty) and H. variabilis Korge (32, Kalkanl Da {lar). Scale bar 0.2 mm." httpUri="https://zenodo.org/record/155775/files/figure.png" pageId="4" pageNumber="5">Figs. 28­ 31, 32</figureCitation>
) and shorter medial process of female tergum 10 (
<figureCitation id="23C52A28E01AB16ECA84FA45D3120FB7" box="[929,1073,1439,1465]" captionStart="FIGURES 33 ­ 35" captionStartId="11.[264,383,1747,1771]" captionTargetBox="[315,1269,719,1731]" captionTargetId="figure@11.[310,1278,713,1736]" captionTargetPageId="11" captionText="FIGURES 33 ­ 35. Details of female tergum 10 of Heinzia caucasica Gusarov &amp; Koval, sp. nov. (33, 35, Baribana cave) and H. variabilis Korge (34, paratype). 33 ­ 34 – dorsal view; 35 – lateral view; LL – lateral lobe of female tergum 10; MP – medial process of female tergum 10. Scale bar 0.3 mm." httpUri="https://zenodo.org/record/155776/files/figure.png" pageId="4" pageNumber="5">Figs. 33, 34</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="F3E46526E01AB162C81DFA1CD6F40B88" lastPageId="8" lastPageNumber="9" pageId="4" pageNumber="5" type="description">
<paragraph id="BB4136ADE01AB16EC81DFA1CD54B0C3F" blockId="4.[264,1325,478,1864]" pageId="4" pageNumber="5">
<emphasis id="898AEABFE01AB16EC81DFA1CD6F30FEE" bold="true" box="[312,464,1478,1504]" pageId="4" pageNumber="5">Description.</emphasis>
Length 
<quantity id="7C069B48E01AB16ECB1BFA1DD5F90FEF" box="[574,730,1479,1505]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.15" metricValueMin="0.85" pageId="4" pageNumber="5" unit="mm" value="10.0" valueMax="11.5" valueMin="8.5">8.5­11.5 mm</quantity>
. Body from brownish black to black, legs brown with lighter tarsi and black inner surface of tibia, mouthparts and bases of antennal segments brown to brownish red.
</paragraph>
<paragraph id="BB4136ADE01AB16EC81DF9E4D3A00D46" blockId="4.[264,1325,478,1864]" pageId="4" pageNumber="5">
Head as long as wide; on disc with dense punctation except impunctate area near anterior margin (
<figureCitation id="23C52A28E01AB16EC8BAF9BCD6CB0C8E" box="[415,488,1638,1664]" captionStart="FIGURES 8 ­ 13" captionStartId="5.[264,383,1583,1607]" captionTargetBox="[287,1301,335,1549]" captionTargetId="figure@5.[278,1308,329,1556]" captionTargetPageId="5" captionText="FIGURES 8 ­ 13. Details of Heinzia caucasica Gusarov &amp; Koval, sp. nov. (8, 10, holotype; 12 ­ 13, paratype from Baribana cave) and H. variabilis Korge (9, Kalkanl Da {lar; 11, holotype). 8 ­ 9 – head; 10 ­ 11 – pronotum; 12 – left antenna; 13 – right metatarsus. Scale bar 1 mm (8 ­ 12), 0.4 mm (13)." httpUri="https://zenodo.org/record/155770/files/figure.png" pageId="4" pageNumber="5">Fig. 8</figureCitation>
); with microsculpture consisting of strongly transverse meshes, medially meshes weakly transverse or isodiametric. Posterior frontal puncture closer to posterior margin of eye than to posterior margin of head. Temples 1.1­1.2 times longer than eyes (in dorsal view). All antennal articles longer than wide, first article shorter than second and third combined, second twice as long as wide, third 2.8 times, 4th­6th 1.9­2.0, 7th 1.7, 8th­9th 1.4, 10th 1.3, last article 1.8 times as long as wide (
<figureCitation id="23C52A28E01AB16ECD3BF8F4D3550D46" box="[1054,1142,1838,1864]" captionStart="FIGURES 8 ­ 13" captionStartId="5.[264,383,1583,1607]" captionTargetBox="[287,1301,335,1549]" captionTargetId="figure@5.[278,1308,329,1556]" captionTargetPageId="5" captionText="FIGURES 8 ­ 13. Details of Heinzia caucasica Gusarov &amp; Koval, sp. nov. (8, 10, holotype; 12 ­ 13, paratype from Baribana cave) and H. variabilis Korge (9, Kalkanl Da {lar; 11, holotype). 8 ­ 9 – head; 10 ­ 11 – pronotum; 12 – left antenna; 13 – right metatarsus. Scale bar 1 mm (8 ­ 12), 0.4 mm (13)." httpUri="https://zenodo.org/record/155770/files/figure.png" pageId="4" pageNumber="5">Fig. 12</figureCitation>
).
</paragraph>
<caption id="EF816625E01BB16FC82DF9F5D6180CA6" httpUri="https://zenodo.org/record/155770/files/figure.png" pageId="5" pageNumber="6" targetBox="[287,1301,335,1549]" targetPageId="5">
<paragraph id="BB4136ADE01BB16FC82DF9F5D6180CA6" blockId="5.[264,1323,1583,1704]" pageId="5" pageNumber="6">
<emphasis id="898AEABFE01BB16FC82DF9F5D6950C49" bold="true" box="[264,438,1583,1607]" pageId="5" pageNumber="6">FIGURES 8­13</emphasis>
. Details of 
<taxonomicName id="7CFE4D2EE01BB16FCB1EF9F5D4E40C48" authority="Gusarov &amp; Koval" authorityName="Gusarov &amp; Koval" box="[571,967,1583,1606]" class="Insecta" family="Staphylinidae" genus="Heinzia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="caucasica" status="sp. nov.">
<emphasis id="898AEABFE01BB16FCB1EF9F5D4230C48" box="[571,768,1583,1606]" italics="true" pageId="5" pageNumber="6">Heinzia caucasica</emphasis>
Gusarov &amp; Koval
</taxonomicName>
, 
<taxonomicNameLabel id="92B957C4E01BB16FCAF7F9F5D3040C49" box="[978,1063,1583,1607]" pageId="5" pageNumber="6" rank="species">
<emphasis id="898AEABFE01BB16FCAF7F9F5D3040C49" bold="true" box="[978,1063,1583,1607]" pageId="5" pageNumber="6">sp. nov.</emphasis>
</taxonomicNameLabel>
(8, 10, holotype; 12­13, paratype from Baribana cave) and 
<taxonomicName id="7CFE4D2EE01BB16FCBA6F98BD4750C66" authority="Korge" authorityName="Korge" box="[643,854,1617,1640]" class="Insecta" family="Staphylinidae" genus="Heinzia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis id="898AEABFE01BB16FCBA6F98BD4280C66" box="[643,779,1617,1640]" italics="true" pageId="5" pageNumber="6">H. variabilis</emphasis>
Korge
</taxonomicName>
(9, Kalkanl Da{lar; 11, holotype). 8­9 – head; 10­11 – pronotum; 12 – left antenna; 13 – right metatarsus. Scale bar 1 mm (8­12), 0.4 mm (13).
</paragraph>
</caption>
<paragraph id="BB4136ADE01BB16CC81DF8D8D4600BA0" blockId="5.[264,1324,1794,1900]" lastBlockId="6.[264,1324,284,430]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
Pronotum 1.1 times as wide as long; on disc with dense scattered punctation without extensive impunctate areas (cf. 
<figureCitation id="23C52A28E01BB16FCB5EF8F0D4110D4A" box="[635,818,1834,1860]" captionStart="FIGURES 8 ­ 13" captionStartId="5.[264,383,1583,1607]" captionTargetBox="[287,1301,335,1549]" captionTargetId="figure@5.[278,1308,329,1556]" captionTargetPageId="5" captionText="FIGURES 8 ­ 13. Details of Heinzia caucasica Gusarov &amp; Koval, sp. nov. (8, 10, holotype; 12 ­ 13, paratype from Baribana cave) and H. variabilis Korge (9, Kalkanl Da {lar; 11, holotype). 8 ­ 9 – head; 10 ­ 11 – pronotum; 12 – left antenna; 13 – right metatarsus. Scale bar 1 mm (8 ­ 12), 0.4 mm (13)." httpUri="https://zenodo.org/record/155770/files/figure.png" pageId="5" pageNumber="6">Figs. 10 and 11</figureCitation>
); dorsal rows with 3 (in some specimens 2 or 4) punctures; punctures of sublateral rows indistinguishable from scattered punctures; microsculpture consists of isodiametric or slightly transverse meshes. Elytra (measured from humeral angle) 1.4 times as long as pronotum, 1.1 times as long as wide; punctation as on pronotum, distance between punctures equal to their diameter; without visible microsculpture (at 
<date id="CF40106DE018B16CC8C0FE4ED5350BA0" box="[485,534,404,430]" pageId="6" pageNumber="7" value="1970-10">70x</date>
). Wings fully developed.
</paragraph>
<caption id="EF816625E018B16CC82DF9F2D6110CAC" httpUri="https://zenodo.org/record/155772/files/figure.png" pageId="6" pageNumber="7" targetBox="[286,1299,499,1546]" targetPageId="6">
<paragraph id="BB4136ADE018B16CC82DF9F2D6110CAC" blockId="6.[264,1323,1576,1698]" pageId="6" pageNumber="7">
<emphasis id="898AEABFE018B16CC82DF9F2D6E60C4E" bold="true" box="[264,453,1576,1600]" pageId="6" pageNumber="7">FIGURES 14­19</emphasis>
. Details of 
<taxonomicName id="7CFE4D2EE018B16CCB6BF9F2D4FC0C31" authority="Gusarov &amp; Koval" authorityName="Gusarov &amp; Koval" box="[590,991,1576,1599]" class="Insecta" family="Staphylinidae" genus="Heinzia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="caucasica" status="sp. nov.">
<emphasis id="898AEABFE018B16CCB6BF9F2D4370C31" box="[590,788,1576,1599]" italics="true" pageId="6" pageNumber="7">Heinzia caucasica</emphasis>
Gusarov &amp; Koval
</taxonomicName>
, 
<emphasis id="898AEABFE018B16CCACEF9F2D3610C4E" bold="true" box="[1003,1090,1576,1600]" pageId="6" pageNumber="7">
<taxonomicNameLabel id="92B957C4E018B16CCACEF9F2D3610C4E" box="[1003,1090,1576,1600]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(14, 16, 18, paratype from Dolgaya cave) and 
<taxonomicName id="7CFE4D2EE018B16CCB31F990D5C00C6E" authority="Korge" authorityName="Korge" box="[532,739,1609,1633]" class="Insecta" family="Staphylinidae" genus="Heinzia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis id="898AEABFE018B16CCB31F990D5B80C6F" box="[532,667,1610,1633]" italics="true" pageId="6" pageNumber="7">H. variabilis</emphasis>
Korge
</taxonomicName>
(15, 17, 19, Kalkanl Da{lar). 14­15 – male sternum 8; 16­17 – apex of mail sternum 8; 18­19 – male sternum 9. Scale bar 1 mm (14­15), 0.5 mm (16­ 19).
</paragraph>
</caption>
<paragraph id="BB4136ADE018B16DC81DF921D6C50BD8" blockId="6.[264,1324,1787,1893]" lastBlockId="7.[264,1324,284,470]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
Abdominal terga covered with black semierect microsetae; with very fine microsculpture consisting of transverse waves; punctation finer than on elytra, distance between punctures equals 1­3 times their diameter. Abdominal tergum 7 with wide white palisade fringe. Posterior margin of male sternum 7 concave. Male sternum 8 with medial emargination (
<figureCitation id="23C52A28E019B16DC85CFE9ED5290B50" box="[377,522,324,350]" captionStart="FIGURES 14 ­ 19" captionStartId="6.[264,383,1576,1600]" captionTargetBox="[286,1299,499,1546]" captionTargetId="figure@6.[282,1306,489,1549]" captionTargetPageId="6" captionText="FIGURES 14 ­ 19. Details of Heinzia caucasica Gusarov &amp; Koval, sp. nov. (14, 16, 18, paratype from Dolgaya cave) and H. variabilis Korge (15, 17, 19, Kalkanl Da {lar). 14 ­ 15 – male sternum 8; 16 ­ 17 – apex of mail sternum 8; 18 ­ 19 – male sternum 9. Scale bar 1 mm (14 ­ 15), 0.5 mm (16 ­ 19)." httpUri="https://zenodo.org/record/155772/files/figure.png" pageId="7" pageNumber="8">Figs. 14, 16</figureCitation>
). Female tergum 10 split into two broad lateral lobes (
<figureCitation id="23C52A28E019B16DCDBDFE9ED3D30B50" box="[1176,1264,324,350]" captionStart="FIGURES 33 ­ 35" captionStartId="11.[264,383,1747,1771]" captionTargetBox="[315,1269,719,1731]" captionTargetId="figure@11.[310,1278,713,1736]" captionTargetPageId="11" captionText="FIGURES 33 ­ 35. Details of female tergum 10 of Heinzia caucasica Gusarov &amp; Koval, sp. nov. (33, 35, Baribana cave) and H. variabilis Korge (34, paratype). 33 ­ 34 – dorsal view; 35 – lateral view; LL – lateral lobe of female tergum 10; MP – medial process of female tergum 10. Scale bar 0.3 mm." httpUri="https://zenodo.org/record/155776/files/figure.png" pageId="7" pageNumber="8">Fig. 33</figureCitation>
), the lobes poorly sclerotized and devoid of setae. Long, sclerotized and setose medial process attached at basis of tergum where the lobes meet, and extends posteriad beyond margin of the lobes (
<figureCitation id="23C52A28E019B16DC8A4FE66D6FA0BD8" box="[385,473,444,470]" captionStart="FIGURES 33 ­ 35" captionStartId="11.[264,383,1747,1771]" captionTargetBox="[315,1269,719,1731]" captionTargetId="figure@11.[310,1278,713,1736]" captionTargetPageId="11" captionText="FIGURES 33 ­ 35. Details of female tergum 10 of Heinzia caucasica Gusarov &amp; Koval, sp. nov. (33, 35, Baribana cave) and H. variabilis Korge (34, paratype). 33 ­ 34 – dorsal view; 35 – lateral view; LL – lateral lobe of female tergum 10; MP – medial process of female tergum 10. Scale bar 0.3 mm." httpUri="https://zenodo.org/record/155776/files/figure.png" pageId="7" pageNumber="8">Fig. 33</figureCitation>
).
</paragraph>
<caption id="EF816625E019B16DC82DF93ED57B0D33" httpUri="https://zenodo.org/record/155773/files/figure.png" pageId="7" pageNumber="8" targetBox="[300,1282,579,1733]" targetPageId="7">
<paragraph id="BB4136ADE019B16DC82DF93ED57B0D33" blockId="7.[264,1323,1764,1853]" pageId="7" pageNumber="8">
<emphasis id="898AEABFE019B16DC82DF93ED6E40CF2" bold="true" box="[264,455,1764,1788]" pageId="7" pageNumber="8">FIGURES 20­23</emphasis>
. Aedeagus of 
<taxonomicName id="7CFE4D2EE019B16DCB57F93ED32A0CF5" authority="Gusarov &amp; Koval" authorityName="Gusarov &amp; Koval" box="[626,1033,1764,1787]" class="Insecta" family="Staphylinidae" genus="Heinzia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="caucasica" status="sp. nov.">
<emphasis id="898AEABFE019B16DCB57F93ED4190CF5" box="[626,826,1764,1787]" italics="true" pageId="7" pageNumber="8">Heinzia caucasica</emphasis>
Gusarov &amp; Koval
</taxonomicName>
, 
<taxonomicNameLabel id="92B957C4E019B16DCD33F93ED34D0CF2" box="[1046,1134,1764,1788]" pageId="7" pageNumber="8" rank="species">
<emphasis id="898AEABFE019B16DCD33F93ED34D0CF2" bold="true" box="[1046,1134,1764,1788]" pageId="7" pageNumber="8">sp. nov.</emphasis>
</taxonomicNameLabel>
(20­21, paratype from Mezmay) and 
<taxonomicName id="7CFE4D2EE019B16DC8C7F8DCD5970D12" authority="Korge" authorityName="Korge" box="[482,692,1797,1821]" class="Insecta" family="Staphylinidae" genus="Heinzia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis id="898AEABFE019B16DC8C7F8DCD5490D13" box="[482,618,1798,1821]" italics="true" pageId="7" pageNumber="8">H. variabilis</emphasis>
Korge
</taxonomicName>
(22­23, Kalkanl Da{lar). 20, 22 – ventral view; 21, 23, lateral view. Scale bar 0.4 mm.
</paragraph>
</caption>
<paragraph id="BB4136ADE016B162C81DFEC6D6F40B88" blockId="8.[264,1324,284,390]" pageId="8" pageNumber="9">
Median lobe of aedeagus with single subapical tooth on ventral surface (
<figureCitation id="23C52A28E016B162CDB3FEC6D3DA0B38" box="[1174,1273,284,310]" captionStart="FIGURES 20 ­ 23" captionStartId="7.[264,383,1764,1788]" captionTargetBox="[300,1282,579,1733]" captionTargetId="figure@7.[295,1292,569,1737]" captionTargetPageId="7" captionText="FIGURES 20 ­ 23. Aedeagus of Heinzia caucasica Gusarov &amp; Koval, sp. nov. (20 ­ 21, paratype from Mezmay) and H. variabilis Korge (22 ­ 23, Kalkanl Da {lar). 20, 22 – ventral view; 21, 23, lateral view. Scale bar 0.4 mm." httpUri="https://zenodo.org/record/155773/files/figure.png" pageId="8" pageNumber="9">Figs. 21</figureCitation>
, 
<figureCitation id="23C52A28E016B162CC22FEC6D2040B38" box="[1287,1319,284,310]" captionStart="FIGURES 24 ­ 27" captionStartId="8.[264,383,1791,1815]" captionTargetBox="[281,1306,495,1759]" captionTargetId="figure@8.[272,1315,489,1765]" captionTargetPageId="8" captionText="FIGURES 24 ­ 27. Details of aedeagus of Heinzia caucasica Gusarov &amp; Koval, sp. nov. (24 ­ 25, holotype) and H. variabilis Korge (26 ­ 27, holotype). 24, 26 – median lobe, ventral view; 25, 27, paramere. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/155774/files/figure.png" pageId="8" pageNumber="9">24</figureCitation>
, 
<figureCitation id="23C52A28E016B162C82DFE9ED6720B50" box="[264,337,324,350]" captionStart="FIGURES 28 ­ 32" captionStartId="9.[264,383,1416,1440]" captionTargetBox="[324,1261,853,1386]" captionTargetId="figure@9.[318,1269,849,1389]" captionTargetPageId="9" captionText="FIGURES 28 ­ 32. Apex of median lobe of aedeagus (lateral view) of Heinzia caucasica Gusarov &amp; Koval, sp. nov. (28, paratypes from Mezmay; 29 ­ 30, Baribana cave; 31, Akhty) and H. variabilis Korge (32, Kalkanl Da {lar). Scale bar 0.2 mm." httpUri="https://zenodo.org/record/155775/files/figure.png" pageId="8" pageNumber="9">28­31</figureCitation>
). Paramere with 15­20 peg­like setae (
<figureCitation id="23C52A28E016B162CA3DFE9ED4530B50" box="[792,880,324,350]" captionStart="FIGURES 24 ­ 27" captionStartId="8.[264,383,1791,1815]" captionTargetBox="[281,1306,495,1759]" captionTargetId="figure@8.[272,1315,489,1765]" captionTargetPageId="8" captionText="FIGURES 24 ­ 27. Details of aedeagus of Heinzia caucasica Gusarov &amp; Koval, sp. nov. (24 ­ 25, holotype) and H. variabilis Korge (26 ­ 27, holotype). 24, 26 – median lobe, ventral view; 25, 27, paramere. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/155774/files/figure.png" pageId="8" pageNumber="9">Fig. 25</figureCitation>
). Internal sac with single basal diverticulum (
<figureCitation id="23C52A28E016B162C854FEB6D6EB0B88" box="[369,456,364,390]" captionStart="FIGURES 20 ­ 23" captionStartId="7.[264,383,1764,1788]" captionTargetBox="[300,1282,579,1733]" captionTargetId="figure@7.[295,1292,569,1737]" captionTargetPageId="7" captionText="FIGURES 20 ­ 23. Aedeagus of Heinzia caucasica Gusarov &amp; Koval, sp. nov. (20 ­ 21, paratype from Mezmay) and H. variabilis Korge (22 ­ 23, Kalkanl Da {lar). 20, 22 – ventral view; 21, 23, lateral view. Scale bar 0.4 mm." httpUri="https://zenodo.org/record/155773/files/figure.png" pageId="8" pageNumber="9">Fig. 21</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="EF816625E016B162C82DF925D5620D52" httpUri="https://zenodo.org/record/155774/files/figure.png" pageId="8" pageNumber="9" targetBox="[281,1306,495,1759]" targetPageId="8">
<paragraph id="BB4136ADE016B162C82DF925D5620D52" blockId="8.[264,1322,1791,1884]" pageId="8" pageNumber="9">
<emphasis id="898AEABFE016B162C82DF925D6E60D19" bold="true" box="[264,453,1791,1815]" pageId="8" pageNumber="9">FIGURES 24­27</emphasis>
. Details of aedeagus of 
<taxonomicName id="7CFE4D2EE016B162CBF9F8DAD3530D19" authority="Gusarov &amp; Koval" authorityName="Gusarov &amp; Koval" box="[732,1136,1792,1815]" class="Insecta" family="Staphylinidae" genus="Heinzia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="caucasica" status="sp. nov.">
<emphasis id="898AEABFE016B162CBF9F8DAD4800D19" box="[732,931,1792,1815]" italics="true" pageId="8" pageNumber="9">Heinzia caucasica</emphasis>
Gusarov &amp; Koval
</taxonomicName>
, 
<taxonomicNameLabel id="92B957C4E016B162CD58F925D3F70D19" box="[1149,1236,1791,1815]" pageId="8" pageNumber="9" rank="species">
<emphasis id="898AEABFE016B162CD58F925D3F70D19" bold="true" box="[1149,1236,1791,1815]" pageId="8" pageNumber="9">sp. nov.</emphasis>
</taxonomicNameLabel>
(24­25, holotype) and 
<taxonomicName id="7CFE4D2EE016B162C883F8F8D55B0D37" authority="Korge" authorityName="Korge" box="[422,632,1826,1849]" class="Insecta" family="Staphylinidae" genus="Heinzia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis id="898AEABFE016B162C883F8F8D50D0D37" box="[422,558,1826,1849]" italics="true" pageId="8" pageNumber="9">H. variabilis</emphasis>
Korge
</taxonomicName>
(26­27, holotype). 24, 26 – median lobe, ventral view; 25, 27, paramere. Scale bar 0.2 mm.
</paragraph>
</caption>
<subSubSection id="F3E46526E017B163C81DFEC1D5F008E0" pageId="9" pageNumber="10" type="discussion">
<paragraph id="BB4136ADE017B163C81DFEC1D5F008E0" blockId="9.[264,1324,283,750]" pageId="9" pageNumber="10">
<emphasis id="898AEABFE017B163C81DFEC1D6E00B3B" bold="true" box="[312,451,283,309]" pageId="9" pageNumber="10">Discussion.</emphasis>
In most groups of staphylinids the characters of genitalia are more useful for distinguishing close species than external characters. Our examination of available series of 
<taxonomicName id="7CFE4D2EE017B163C851FEB6D52D0B8B" box="[372,526,364,389]" class="Insecta" family="Staphylinidae" genus="Heinzia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="caucasica">
<emphasis id="898AEABFE017B163C851FEB6D52D0B8B" box="[372,526,364,389]" italics="true" pageId="9" pageNumber="10">H. caucasica</emphasis>
</taxonomicName>
demonstrated that in this species the aedeagus is subject to significant variation. The shape of paramere, the arrangement of peg­like setae and the distance between the apex of median lobe and the subapical tooth (see 
<figureCitation id="23C52A28E017B163CADBFE66D3A40BD8" box="[1022,1159,444,470]" captionStart="FIGURES 28 ­ 32" captionStartId="9.[264,383,1416,1440]" captionTargetBox="[324,1261,853,1386]" captionTargetId="figure@9.[318,1269,849,1389]" captionTargetPageId="9" captionText="FIGURES 28 ­ 32. Apex of median lobe of aedeagus (lateral view) of Heinzia caucasica Gusarov &amp; Koval, sp. nov. (28, paratypes from Mezmay; 29 ­ 30, Baribana cave; 31, Akhty) and H. variabilis Korge (32, Kalkanl Da {lar). Scale bar 0.2 mm." httpUri="https://zenodo.org/record/155775/files/figure.png" pageId="9" pageNumber="10">Figs. 28­31</figureCitation>
) vary, like in some species of the genus 
<taxonomicName id="7CFE4D2EE017B163CB66FE3ED4400BF0" authority="Stephens, 1829" authorityName="Stephens" authorityYear="1829" box="[579,867,484,510]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="898AEABFE017B163CB66FE3ED5860BF3" box="[579,677,484,509]" italics="true" pageId="9" pageNumber="10">Quedius</emphasis>
Stephens, 1829
</taxonomicName>
. The only difference in male genitalia between the two species of 
<taxonomicName id="7CFE4D2EE017B163CB71FDD6D58C082B" box="[596,687,524,549]" class="Insecta" family="Staphylinidae" genus="Heinzia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="898AEABFE017B163CB71FDD6D58C082B" box="[596,687,524,549]" italics="true" pageId="9" pageNumber="10">Heinzia</emphasis>
</taxonomicName>
, that we were able to find, is somewhat sharper subapical tooth of median lobe of aedeagus in 
<taxonomicName id="7CFE4D2EE017B163CA25FDEED4B70843" box="[768,916,564,589]" class="Insecta" family="Staphylinidae" genus="Heinzia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis id="898AEABFE017B163CA25FDEED4B70843" box="[768,916,564,589]" italics="true" pageId="9" pageNumber="10">H. variabilis</emphasis>
</taxonomicName>
. On the other hand, the density of punctation of the head and pronotum displays relatively little variation and allows to distinguish between 
<taxonomicName id="7CFE4D2EE017B163C8F2FD5ED5520893" box="[471,625,644,669]" class="Insecta" family="Staphylinidae" genus="Heinzia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="caucasica">
<emphasis id="898AEABFE017B163C8F2FD5ED5520893" box="[471,625,644,669]" italics="true" pageId="9" pageNumber="10">H. caucasica</emphasis>
</taxonomicName>
and 
<taxonomicName id="7CFE4D2EE017B163CB8FFD5ED41D0893" box="[682,830,644,669]" class="Insecta" family="Staphylinidae" genus="Heinzia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis id="898AEABFE017B163CB8FFD5ED41D0893" box="[682,830,644,669]" italics="true" pageId="9" pageNumber="10">H. variabilis</emphasis>
</taxonomicName>
. The presence of the clear gap in punctation between the beetles from the Caucasus and 
<collectingCountry id="C3E9763DE017B163CA56FD76D4E408C8" box="[883,967,684,710]" name="Turkey" pageId="9" pageNumber="10">Turkey</collectingCountry>
implies that these allopatric populations represent different species.
</paragraph>
</subSubSection>
<caption id="EF816625E017B163C82DFA52D4300FE8" httpUri="https://zenodo.org/record/155775/files/figure.png" pageId="9" pageNumber="10" targetBox="[324,1261,853,1386]" targetPageId="9">
<paragraph id="BB4136ADE017B163C82DFA52D4300FE8" blockId="9.[264,1322,1416,1511]" pageId="9" pageNumber="10">
<emphasis id="898AEABFE017B163C82DFA52D6E60FAE" bold="true" box="[264,453,1416,1440]" pageId="9" pageNumber="10">FIGURES 28­32</emphasis>
. Apex of median lobe of aedeagus (lateral view) of 
<taxonomicName id="7CFE4D2EE017B163CD26FA53D6440FCC" authority="Gusarov &amp; Koval" authorityName="Gusarov &amp; Koval" class="Insecta" family="Staphylinidae" genus="Heinzia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="caucasica" status="sp. nov.">
<emphasis id="898AEABFE017B163CD26FA53D3EA0FAE" box="[1027,1225,1417,1440]" italics="true" pageId="9" pageNumber="10">Heinzia caucasica</emphasis>
Gusarov &amp; Koval
</taxonomicName>
, 
<taxonomicNameLabel id="92B957C4E017B163C854FA71D6E50FCD" box="[369,454,1451,1475]" pageId="9" pageNumber="10" rank="species">
<emphasis id="898AEABFE017B163C854FA71D6E50FCD" bold="true" box="[369,454,1451,1475]" pageId="9" pageNumber="10">sp. nov.</emphasis>
</taxonomicNameLabel>
(28, paratypes from Mezmay; 29­30, Baribana cave; 31, Akhty) and 
<taxonomicName id="7CFE4D2EE017B163CD81FA71D66A0FE8" authority="Korge" authorityName="Korge" class="Insecta" family="Staphylinidae" genus="Heinzia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis id="898AEABFE017B163CD81FA71D2090FCC" box="[1188,1322,1451,1474]" italics="true" pageId="9" pageNumber="10">H. variabilis</emphasis>
Korge
</taxonomicName>
(32, Kalkanl Da{lar). Scale bar 0.2 mm.
</paragraph>
</caption>
<subSubSection id="F3E46526E017B160C81DF998D3040E08" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10" type="distribution">
<paragraph id="BB4136ADE017B163C81DF998D5E10CA3" blockId="9.[264,1324,1602,1869]" pageId="9" pageNumber="10">
<emphasis id="898AEABFE017B163C81DF998D6FB0C52" bold="true" box="[312,472,1602,1628]" pageId="9" pageNumber="10">Distribution.</emphasis>
<taxonomicName id="7CFE4D2EE017B163C8C3F999D59D0C52" box="[486,702,1603,1628]" class="Insecta" family="Staphylinidae" genus="Heinzia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="caucasica">
<emphasis id="898AEABFE017B163C8C3F999D59D0C52" box="[486,702,1603,1628]" italics="true" pageId="9" pageNumber="10">Heinzia caucasica</emphasis>
</taxonomicName>
is known from the Caucasus (Main Caucasian Ridge (Glavnyy Kavkazskiy Ridge) from Krasnodar Territory in the West to Daghestan in the East; altitudes 
<quantity id="7C069B48E017B163C890F949D5670CA3" box="[437,580,1683,1709]" metricMagnitude="3" metricUnit="m" metricValue="1.05" metricValueMax="1.8" metricValueMin="0.3" pageId="9" pageNumber="10" unit="m" value="1050.0" valueMax="1800.0" valueMin="300.0">300­1800 m</quantity>
) (
<figureCitation id="23C52A28E017B163CB78F949D5960CA3" box="[605,693,1683,1709]" captionStart="FIGURE 48" captionStartId="16.[264,367,1711,1735]" captionTargetBox="[324,1266,798,1690]" captionTargetId="figure@16.[317,1270,793,1693]" captionTargetPageId="16" captionText="FIGURE 48. Geographical distribution of Heinzia Korge. Solid circles – H. caucasica Gusarov &amp; Koval, sp. nov.; open circles – H. variabilis Korge." httpUri="https://zenodo.org/record/155780/files/figure.png" pageId="9" pageNumber="10">Fig. 48</figureCitation>
).
</paragraph>
<paragraph id="BB4136ADE017B163C81DF960D41B0D43" blockId="9.[264,1324,1602,1869]" pageId="9" pageNumber="10">
<emphasis id="898AEABFE017B163C81DF960D5240CDA" bold="true" box="[312,519,1722,1748]" pageId="9" pageNumber="10">Natural History.</emphasis>
Big series of 
<taxonomicName id="7CFE4D2EE017B163CB9BF961D47A0CDA" box="[702,857,1723,1748]" class="Insecta" family="Staphylinidae" genus="Heinzia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="caucasica">
<emphasis id="898AEABFE017B163CB9BF961D47A0CDA" box="[702,857,1723,1748]" italics="true" pageId="9" pageNumber="10">H. caucasica</emphasis>
</taxonomicName>
were collected in caves by the second author who used pitfall traps, as described by 
<bibRefCitation id="DF6F4B5CE017B163CA04F939D4EA0CF3" author="Barber" box="[801,969,1763,1789]" pageId="9" pageNumber="18" refString="Barber, H. S. (1931) Traps for cave-inhabiting insects. Journal of the Elisha Mitchell Scientific Society, 46, 259 - 267." type="journal article" year="1931">Barber (1931)</bibRefCitation>
, but with some modifications. The traps were filled with 1:1 solution of ethylene­glycol and beer, and baited with old cheese and sausage suspended above the liquid.
</paragraph>
<paragraph id="BB4136ADE014B160C81DFEC6D4250BA0" blockId="10.[264,1324,284,1030]" pageId="10" pageNumber="11">
As a cave dweller, 
<taxonomicName id="7CFE4D2EE014B160CB0BFEC6D5EE0B3B" box="[558,717,284,309]" class="Insecta" family="Staphylinidae" genus="Heinzia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="caucasica">
<emphasis id="898AEABFE014B160CB0BFEC6D5EE0B3B" box="[558,717,284,309]" italics="true" pageId="10" pageNumber="11">H. caucasica</emphasis>
</taxonomicName>
apparently belongs to the group of troglophiles, defined by Racovitza (1907) as inhabitants of caves and different kinds of large and small subterranean caverns, including hollows under big boulders. Unlike troglobiontic species, the troglophiles occasionally occur outside.
</paragraph>
<paragraph id="BB4136ADE014B160C81DFE66D6E608E0" blockId="10.[264,1324,284,1030]" pageId="10" pageNumber="11">
The four caves where the specimens of 
<taxonomicName id="7CFE4D2EE014B160CA2CFE66D4810BDB" box="[777,930,444,469]" class="Insecta" family="Staphylinidae" genus="Heinzia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="caucasica">
<emphasis id="898AEABFE014B160CA2CFE66D4810BDB" box="[777,930,444,469]" italics="true" pageId="10" pageNumber="11">H. caucasica</emphasis>
</taxonomicName>
were collected are situated in the Sochi area of the Western Caucasus and have plenty of water. Three caves (Sokolova, Dolgaya and Labirintovaya Caves) are remarkable for having subterranean brooks or even a river (Sokolova Cave). Baribana Cave has no brooks but it is still very moist and has many pools as a result of water dripping from the ceiling and trickling down the walls and the floor. It is in the wettest parts of the caves that the specimens of 
<taxonomicName id="7CFE4D2EE014B160CD68FD5ED3CB0893" box="[1101,1256,644,669]" class="Insecta" family="Staphylinidae" genus="Heinzia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="caucasica">
<emphasis id="898AEABFE014B160CD68FD5ED3CB0893" box="[1101,1256,644,669]" italics="true" pageId="10" pageNumber="11">H. caucasica</emphasis>
</taxonomicName>
were collected. The air temperature in the four caves is similar and fluctuates annually between 8.0 and 11.5 °C.
</paragraph>
<paragraph id="BB4136ADE014B160C81DFD26D3BC0980" blockId="10.[264,1324,284,1030]" pageId="10" pageNumber="11">
Despite intensive sampling no specimens of 
<taxonomicName id="7CFE4D2EE014B160CA69FD26D4C4091B" box="[844,999,764,789]" class="Insecta" family="Staphylinidae" genus="Heinzia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="caucasica">
<emphasis id="898AEABFE014B160CA69FD26D4C4091B" box="[844,999,764,789]" italics="true" pageId="10" pageNumber="11">H. caucasica</emphasis>
</taxonomicName>
were collected in other visited caves of the Sochi area (Partizanskaya, Muzeynaya, Kolokolnaya and Beloskalskaya Caves). These caves are relatively dry, have no subterranean brooks or strong drippings, and apparently do not provide suitable habitats for hygrophilous 
<taxonomicName id="7CFE4D2EE014B160CADAFCAED3BB0983" box="[1023,1176,884,909]" class="Insecta" family="Staphylinidae" genus="Heinzia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="caucasica">
<emphasis id="898AEABFE014B160CADAFCAED3BB0983" box="[1023,1176,884,909]" italics="true" pageId="10" pageNumber="11">H. caucasica</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="BB4136ADE014B160C81DFC46D3040E08" blockId="10.[264,1324,284,1030]" pageId="10" pageNumber="11">
Outside caves the beetles were collected only occasionally (single specimens). It is possible that outside 
<taxonomicName id="7CFE4D2EE014B160CB24FC1ED5B909D3" box="[513,666,964,989]" class="Insecta" family="Staphylinidae" genus="Heinzia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="caucasica">
<emphasis id="898AEABFE014B160CB24FC1ED5B909D3" box="[513,666,964,989]" italics="true" pageId="10" pageNumber="11">H. caucasica</emphasis>
</taxonomicName>
inhabits the caverns between stones in deeper layers of talus­like creek banks, the typical habitat of 
<taxonomicName id="7CFE4D2EE014B160CA2AFC36D3000E08" authority="Smetana 1977" authorityName="Smetana" authorityYear="1977" box="[783,1059,1004,1030]" class="Insecta" family="Lymantriidae" genus="Beeria" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="898AEABFE014B160CA2AFC36D47E0E0B" box="[783,861,1004,1029]" italics="true" pageId="10" pageNumber="11">Beeria</emphasis>
(
<bibRefCitation id="DF6F4B5CE014B160CA48FC36D3390E08" author="Smetana" box="[877,1050,1004,1030]" pageId="10" pageNumber="19" refString="Smetana, A. (1977) The Nearctic genus Beeria Hatch. Taxonomy, distribution and ecology (Coleoptera: Staphylinidae). Entomologica Scandinavica, 8, 177 - 190." type="journal article" year="1977">Smetana 1977</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>