466 lines
50 KiB
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466 lines
50 KiB
XML
<document id="741C6731D1E60F54D1D754AAA3281667" ID-DOI="10.11646/phytotaxa.518.2.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="5453884" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630653214218" checkinUser="plazi" docAuthor="Martel, Carlos, Egoavil, Luis, Ocupa, Luis, Tello, Juan Andre, Patrón, Federico Rizo, Laura, César, León, Marco & Collantes, Benjamín" docDate="2021" docId="331487BDFFA66676FF2AFDC9FD4BF603" docLanguage="en" docName="phytotaxa.518.2.1.pdf" docOrigin="Phytotaxa 518 (2)" docSource="http://dx.doi.org/10.11646/phytotaxa.518.2.1" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.7:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="7" docTitle="Maxillaria pyhalae D. E. Benn. & Christenson" docType="treatment" docVersion="4" lastPageNumber="91" masterDocId="CF2DFFC5FFA46672FFA2FF9BFFF0F37E" masterDocTitle="An homage to Peru’s bicentenary: Maxillaria bicentenaria (Orchidaceae), a new species previously misidentified as M. pyhalae" masterLastPageNumber="99" masterPageNumber="87" pageNumber="89" updateTime="1699281583604" updateUser="plazi" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="DF09F3A84189ECF2CE9CE8985E482748">An homage to Peru’s bicentenary: Maxillaria bicentenaria (Orchidaceae), a new species previously misidentified as M. pyhalae</mods:title>
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<mods:namePart id="3930453F30613BC0492C3311F3FB950B">Martel, Carlos</mods:namePart>
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<mods:nameIdentifier id="38D9419D1C4D7D59AA5A94CE3AFF642E" type="ORCID">0000-0001-9892-1999</mods:nameIdentifier>
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<mods:affiliation id="503F1B3A99ED3AEA0564DB2AE5B6E6A2">Instituto de Ciencias Ómicas y Biotecnología Aplicada, Pontificia Universidad Católica del Perú, Av. Universitaria 1801, San Miguel, Lima 15088, Peru & carlos. martel @ pucp. pe; https: // orcid. org / 0000 - 0001 - 9892 - 1999</mods:affiliation>
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<mods:nameIdentifier id="71A4789FB5321C5FDEF509B7438431C1" type="email">carlos.martel@pucp.pe</mods:nameIdentifier>
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<mods:namePart id="F24C9C6D50B7A739A7C49356C6A56396">Egoavil, Luis</mods:namePart>
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<mods:nameIdentifier id="F88D586361FF95AA24FDAB7A43E7CA14" type="ORCID">0000-0001-9690-3455</mods:nameIdentifier>
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<mods:affiliation id="FF0785A0C83B12CFD4F5AA7D0224360F">Facultad de Ciencias, Forestales y del Ambiente, Universidad Nacional del Centro del Perú, Huancayo 12006, Peru & lenriq 225 @ gmail. com; https: // orcid. org / 0000 - 0001 - 9690 - 3455</mods:affiliation>
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<mods:nameIdentifier id="DD18B7919730DEA8D3387609C832396A" type="email">lenriq225@gmail.com</mods:nameIdentifier>
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<mods:roleTerm id="ECA8943B40C7DE690082B1D808EFD846">Author</mods:roleTerm>
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<mods:namePart id="6B9036ADB55975301936852E6D6511C0">Ocupa, Luis</mods:namePart>
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<mods:nameIdentifier id="E8D9B9027AFB5C2F7634BE133ABE45F0" type="ORCID">0000-0002-4904-1060</mods:nameIdentifier>
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<mods:affiliation id="7BC6292B69267D7310FCA0838D516336">Unidad de Postgrado, Facultad de Ciencias Biológicas, Universidad Nacional Mayor de San Marcos, Av. Germán Amézaga 375, Cercado de Lima, Lima 15081, Peru & luisocupa. horna @ gmail. com; https: // orcid. org / 0000 - 0002 - 4904 - 1060</mods:affiliation>
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<mods:nameIdentifier id="7B2616E861DFA8B6EF920B2FFF6FABD2" type="email">luisocupa.horna@gmail.com</mods:nameIdentifier>
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<mods:namePart id="42D64ECBEDCD22DDD443B6C1DC2914A7">Tello, Juan Andre</mods:namePart>
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<mods:nameIdentifier id="9C108C96037A5E9D0A176268DD053CE3" type="ORCID">0000-0002-2674-9170</mods:nameIdentifier>
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<mods:affiliation id="62118066CD15E8B69B61D08073A66A51">Unidad de Postgrado, Facultad de Ciencias Biológicas, Universidad Nacional Mayor de San Marcos, Av. Germán Amézaga 375, Cercado de Lima, Lima 15081, Peru & juandretf 22 @ gmail. com; https: // orcid. org / 0000 - 0002 - 2674 - 9170</mods:affiliation>
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<mods:nameIdentifier id="8DFC630B6E227E7EE6E09E3D3B64B58B" type="email">juandretf22@gmail.com</mods:nameIdentifier>
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<mods:namePart id="BB8D26202E60E135218B92FCF5397D98">Patrón, Federico Rizo</mods:namePart>
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<mods:nameIdentifier id="420121631CB0D27EE379DFE1FA5EE990" type="ORCID">0000-0002-2261-1078</mods:nameIdentifier>
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<mods:affiliation id="FCDA03812FD0BC38A4D3A157F0082D96">Centro Neotropical de Entrenamiento en Humedales - Perú, Carretera San Alberto km 5.5, Oxapampa 15302, Peru & feluse 30 @ gmail. com; https: // orcid. org / 0000 - 0002 - 2261 - 1078</mods:affiliation>
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<mods:nameIdentifier id="2008AE5136AA27DB212CA09246A367BB" type="email">feluse30@gmail.com</mods:nameIdentifier>
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<mods:roleTerm id="C6040995AF56954731A0C55211359B1A">Author</mods:roleTerm>
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<mods:namePart id="99C38B70F473AA027D4753A6D9EF4D0F">Laura, César</mods:namePart>
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<mods:nameIdentifier id="A4D2B5B1238BAF77249D349E0E039A90" type="ORCID">0000-0002-9918-3489</mods:nameIdentifier>
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<mods:affiliation id="B36FA270ABBD8DF0C1B0EE4479F2208F">Instituto del Bien Común, Jr. Frey 1122, Oxapampa 19230, Peru & crlaurac @ gmail. com; https: // orcid. org / 0000 - 0002 - 9918 - 3489</mods:affiliation>
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<mods:nameIdentifier id="6B24D645B36FD35DB9B5C1831076BCB6" type="email">crlaurac@gmail.com</mods:nameIdentifier>
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<mods:namePart id="A997FB8D4338119FDE0FE330EB3135E5">León, Marco</mods:namePart>
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<mods:nameIdentifier id="052792B2B2A987FAFC018A2473D77569" type="ORCID">0000-0003-3774-4743</mods:nameIdentifier>
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<mods:affiliation id="42E0127D0F3AC641C03F983E2338EFB5">Instituto de Investigación Biológica de las Cordilleras Orientales, Jr. José Olaya 510 dpto. 201, Tarapoto 22201, Peru & mleon @ inibico. org; https: // orcid. org / 0000 - 0003 - 3774 - 4743</mods:affiliation>
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<mods:nameIdentifier id="BB8C95285A7230E341BF4B104D399BB2" type="email">mleon@inibico.org</mods:nameIdentifier>
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<mods:namePart id="B67A6597ED0D3A1AECF59F7351B1FBB7">Collantes, Benjamín</mods:namePart>
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<mods:nameIdentifier id="58B8A5F2894817F0BD41DB96D2CBE7F7" type="ORCID">0000-0003-2346-9576</mods:nameIdentifier>
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<mods:affiliation id="3EBB1AA74BB1D5C5BC0352C038F664DD">Inka-Terra Association, Andalucía 174, Miraflores, Lima 15074, Peru & bcollantes @ hotmail. com; https: // orcid. org / 0000 - 0003 - 2346 - 9576</mods:affiliation>
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<mods:nameIdentifier id="79491FB1932EE8F92FD4650D8FE54AFA" type="email">bcollantes@hotmail.com</mods:nameIdentifier>
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<mods:title id="8BEF55E5BDBFC7CDF58E1BD10A368706">Phytotaxa</mods:title>
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<mods:part id="CCC8693919602419F06204AFC2A08975">
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<mods:date id="3A48936E263FEA6CF44553F0AF5C0868">2021</mods:date>
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<mods:number id="59BC90DF6A111F6AF8822F768641D9AB">2021-09-03</mods:number>
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<mods:number id="15E17BF4E2DD4120806FE21601AB121E">518</mods:number>
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<mods:start id="9B7AAB7C436F099AFD1CC251B617F697">87</mods:start>
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<mods:url id="D4B00367A88CB77947F65D6E23373439">http://dx.doi.org/10.11646/phytotaxa.518.2.1</mods:url>
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<mods:classification id="8BB0E920BB953AC526F611B04699E56E">journal article</mods:classification>
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<mods:identifier id="56F3D234BD40E52A81EB2FA197DF22AA" type="DOI">10.11646/phytotaxa.518.2.1</mods:identifier>
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<treatment id="331487BDFFA66676FF2AFDC9FD4BF603" ID-DOI="http://doi.org/10.5281/zenodo.5495287" ID-Zenodo-Dep="5495287" LSID="urn:lsid:plazi:treatment:331487BDFFA66676FF2AFDC9FD4BF603" httpUri="http://treatment.plazi.org/id/331487BDFFA66676FF2AFDC9FD4BF603" lastPageId="4" lastPageNumber="91" pageId="2" pageNumber="89">
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<subSubSection id="F3A76520FFA66670FF2AFDC9FCFCF113" box="[136,780,594,622]" pageId="2" pageNumber="89" type="nomenclature">
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<paragraph id="BB0236ABFFA66670FF2AFDC9FCFCF113" blockId="2.[136,780,594,622]" box="[136,780,594,622]" pageId="2" pageNumber="89">
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<heading id="E04A81C7FFA66670FF2AFDC9FCFCF113" box="[136,780,594,622]" fontSize="11" level="2" pageId="2" pageNumber="89" reason="3">
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<taxonomicName id="7CBD4D28FFA66670FF2AFDC9FD7AF113" ID-CoL="3YGTJ" authority="D. E. Benn. & Christenson" authorityName="D. E. Benn. & Christenson" box="[136,650,594,622]" class="Liliopsida" family="Orchidaceae" genus="Maxillaria" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="89" phylum="Tracheophyta" rank="species" species="pyhalae">
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||
<emphasis id="89C9EAB9FFA66670FF2AFDC9FE94F113" bold="true" box="[136,356,594,621]" italics="true" pageId="2" pageNumber="89">Maxillaria pyhalae</emphasis>
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||
D.E.Benn. & Christenson
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||
</taxonomicName>
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||
(
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||
<figureCitation id="23862A2EFFA66670FD3BFDC8FCF3F113" box="[665,771,595,621]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="3.[136,229,1889,1911]" captionStartId-1="4.[136,229,1062,1084]" captionTargetBox-0="[238,1343,266,1849]" captionTargetBox-1="[161,1426,200,1028]" captionTargetId-0="figure-50@3.[215,1371,241,1861]" captionTargetId-1="figure-27@4.[151,1436,190,1038]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURE 1. Maxillaria pyhalae. A. Habit. B. Flower, frontal view. C. Dissected flower with extended perianth and details of the dorsal sepal apex. D. Labellum and column, lateral view. E. Labellum, frontal view with detail of the abaxial side of the extended apex. F. Protuberance at the apex centre of the labellum, coronal view. G. Column, frontal and lateral views with details of the margins.G. Column, lateral view with details of the surface. H. Anther and pollinarium, ventral and dorsal views. Note the flower is non-resupinate and it is shown as resupinate in C for graphical purposes only. Drawn from M. León & B. Collantes 2975 (USM) by B. Collantes (personal plate nº 21)." captionText-1="FIGURE 2. Maxillaria pyhalae. A. Habit. B. Flower, frontal view. Photographs by L. Egoavil (A) and B. Collantes (B)." figureDoi-0="http://doi.org/10.5281/zenodo.5453886" figureDoi-1="http://doi.org/10.5281/zenodo.5453888" httpUri-0="https://zenodo.org/record/5453886/files/figure.png" httpUri-1="https://zenodo.org/record/5453888/files/figure.png" pageId="2" pageNumber="89">Figs. 1–2</figureCitation>
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||
)
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="F3A76520FFA66670FF2AFD05FC1CF5E8" pageId="2" pageNumber="89" type="materials_examined">
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<paragraph id="BB0236ABFFA66670FF2AFD05FE6EF1A6" blockId="2.[136,1451,670,728]" pageId="2" pageNumber="89">
|
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<emphasis id="89C9EAB9FFA66670FF2AFD05FF4AF1CD" bold="true" box="[136,186,670,691]" pageId="2" pageNumber="89">Type</emphasis>
|
||
:—
|
||
<materialsCitation id="0BD53CF6FFA66670FF74FD04FE6AF1A6" collectingDate="1999-02-14" collectionCode="MOL" collectorName="Leon & Collantes" country="Peru" elevation="2600" location="Llactapata" municipality="Prov. of Tayacaja" pageId="2" pageNumber="89" specimenCount="1" stateProvince="Huancavelica" typeStatus="holotype">
|
||
<collectingCountry id="C3AA763BFFA66670FF74FD04FEE6F1CA" box="[214,278,671,692]" name="Peru" pageId="2" pageNumber="89">PERU</collectingCountry>
|
||
.
|
||
<collectingRegion id="7979F849FFA66670FE80FD05FE58F1CA" box="[290,424,670,692]" country="Peru" name="Huancavelica" pageId="2" pageNumber="89">Huancavelica</collectingRegion>
|
||
:
|
||
<collectingMunicipality id="5B66ACD1FFA66670FE13FD04FDAAF1CA" box="[433,602,670,692]" pageId="2" pageNumber="89">Prov. of Tayacaja</collectingMunicipality>
|
||
,
|
||
<location id="BE626070FFA66670FDC1FD05FD39F1CA" LSID="urn:lsid:plazi:treatment:331487BDFFA66676FF2AFDC9FD4BF603:BE626070FFA66670FDC1FD05FD39F1CA" box="[611,713,670,692]" country="Peru" municipality="Prov. of Tayacaja" name="Llactapata" pageId="2" pageNumber="89" stateProvince="Huancavelica">Llactapata</location>
|
||
,
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||
<quantity id="7C459B4EFFA66670FD71FD05FCEEF1CA" box="[723,798,670,692]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="2" pageNumber="89" unit="m" value="2600.0">
|
||
<elevation id="3090D198FFA66670FD71FD05FCEEF1CA" box="[723,798,670,692]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="2" pageNumber="89" unit="m" value="2600.0">2600 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="CF03106BFFA66670FC85FD05FC23F1CD" box="[807,979,670,692]" pageId="2" pageNumber="89" value="1999-02-14">
|
||
<collectingDate id="DF47E983FFA66670FC85FD05FC23F1CD" box="[807,979,670,692]" pageId="2" pageNumber="89" value="1999-02-14">14 February 1999</collectingDate>
|
||
</date>
|
||
,
|
||
<emphasis id="89C9EAB9FFA66670FC7FFD05FB4DF1CD" box="[989,1213,670,691]" italics="true" pageId="2" pageNumber="89">
|
||
<collectorName id="1648537DFFA66670FC7FFD05FBFDF1CD" box="[989,1037,670,691]" pageId="2" pageNumber="89">León</collectorName>
|
||
&
|
||
<collectorName id="1648537DFFA66670FB89FD05FB77F1CD" box="[1067,1159,670,691]" pageId="2" pageNumber="89">Collantes</collectorName>
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||
2975
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||
</emphasis>
|
||
(
|
||
<typeStatus id="64068809FFA66670FB6BFD05FAEFF1CA" box="[1225,1311,670,692]" pageId="2" pageNumber="89" type="holotype">holotype</typeStatus>
|
||
<collectionCode id="DDACAE6EFFA66670FA87FD05FAADF1CD" box="[1317,1373,670,691]" country="Peru" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15530" lsid="urn:lsid:biocol.org:col:15530" name="Universidad Nacional Agraria La Molina" pageId="2" pageNumber="89" type="Herbarium">MOL</collectionCode>
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||
, photo!; isotype
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||
<collectionCode id="DDACAE6EFFA66670FEABFD59FEB1F1A9" box="[265,321,706,727]" country="Peru" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15530" lsid="urn:lsid:biocol.org:col:15530" name="Universidad Nacional Agraria La Molina" pageId="2" pageNumber="89" type="Herbarium">MOL</collectionCode>
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||
, phtoto!)
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||
</materialsCitation>
|
||
.
|
||
</paragraph>
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||
<paragraph id="BB0236ABFFA66670FF2AFC9EFCFCF50C" blockId="2.[136,1452,773,2077]" pageId="2" pageNumber="89">
|
||
<emphasis id="89C9EAB9FFA66670FF2AFC9EFF35F061" box="[136,197,773,799]" italics="true" pageId="2" pageNumber="89">Plant</emphasis>
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||
up to
|
||
<quantity id="7C459B4EFFA66670FEABFC9EFEBDF05E" box="[265,333,773,800]" metricMagnitude="-1" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="89" unit="cm" value="60.0">60 cm</quantity>
|
||
tall (including the inflorescence), terrestrial and occasionally epiphytic.
|
||
<emphasis id="89C9EAB9FFA66670FBD3FC9DFB41F061" box="[1137,1201,774,799]" italics="true" pageId="2" pageNumber="89">Roots</emphasis>
|
||
slender, born from the rhizome at the base of the pseudobulb.
|
||
<emphasis id="89C9EAB9FFA66670FD9FFCB2FD6DF03D" box="[573,669,809,835]" italics="true" pageId="2" pageNumber="89">Rhizome</emphasis>
|
||
stout, 8–10 ×
|
||
<quantity id="7C459B4EFFA66670FC9FFCB2FC7CF03D" box="[829,908,809,835]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="89" unit="cm" value="0.8">0.8 cm</quantity>
|
||
, covered by scarious bracts.
|
||
<emphasis id="89C9EAB9FFA66670FB6EFCB2FAA0F03D" box="[1228,1360,809,835]" italics="true" pageId="2" pageNumber="89">Pseudobulb</emphasis>
|
||
1.5–2.1 ×
|
||
<quantity id="7C459B4EFFA66670FF02FCD7FED1F019" box="[160,289,844,871]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="2" pageNumber="89" unit="cm" value="0.65" valueMax="0.8" valueMin="0.5">0.5–0.8 cm</quantity>
|
||
, covered by two scarious bracts, 1–2 articulated foliaceous bracts of
|
||
<quantity id="7C459B4EFFA66670FB92FCD7FB50F019" box="[1072,1184,844,871]" metricMagnitude="-1" metricUnit="m" metricValue="2.7" metricValueMax="2.9" metricValueMin="2.5" pageId="2" pageNumber="89" unit="cm" value="27.0" valueMax="29.0" valueMin="25.0">25–29 cm</quantity>
|
||
long.
|
||
<emphasis id="89C9EAB9FFA66670FB48FCD7FAECF018" box="[1258,1308,844,870]" italics="true" pageId="2" pageNumber="89">Leaf</emphasis>
|
||
20.5–39.5 ×
|
||
<quantity id="7C459B4EFFA66670FF2AFCEBFEF6F0F4" box="[136,262,880,906]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" metricValueMax="3.8" metricValueMin="3.2" pageId="2" pageNumber="89" unit="cm" value="3.5" valueMax="3.8" valueMin="3.2">3.2-3.8 cm</quantity>
|
||
, unifoliate; petiole
|
||
<quantity id="7C459B4EFFA66670FE47FCEBFDADF0F4" box="[485,605,880,907]" metricMagnitude="-2" metricUnit="m" metricValue="7.25" metricValueMax="10.0" metricValueMin="4.5" pageId="2" pageNumber="89" unit="cm" value="7.25" valueMax="10.0" valueMin="4.5">4.5–10 cm</quantity>
|
||
long; blade loriform, acute to acuminate apex.
|
||
<emphasis id="89C9EAB9FFA66670FB23FCEBFAE7F0F4" box="[1153,1303,880,906]" italics="true" pageId="2" pageNumber="89">Inflorescence</emphasis>
|
||
up to
|
||
<quantity id="7C459B4EFFA66670FAC6FCEBFA5BF0F4" box="[1380,1451,880,906]" metricMagnitude="-1" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="89" unit="cm" value="60.0">60 cm</quantity>
|
||
long, basal, slender scape covered by 7–8 bracts;
|
||
<emphasis id="89C9EAB9FFA66670FD19FC08FCF1F0D3" box="[699,769,915,941]" italics="true" pageId="2" pageNumber="89">bracts</emphasis>
|
||
4.0–
|
||
<quantity id="7C459B4EFFA66670FC99FC08FC77F0D0" box="[827,903,915,942]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" pageId="2" pageNumber="89" unit="cm" value="6.5">6.5 cm</quantity>
|
||
long;
|
||
<emphasis id="89C9EAB9FFA66670FC73FC08FBA3F0D3" box="[977,1107,915,941]" italics="true" pageId="2" pageNumber="89">floral bract</emphasis>
|
||
5.0 ×
|
||
<quantity id="7C459B4EFFA66670FB3CFC08FB1EF0D0" box="[1182,1262,915,942]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" pageId="2" pageNumber="89" unit="cm" value="2.4">2.4 cm</quantity>
|
||
, ovate, cucullate and carinate on the back, acuminate, basally tubular and unguiculate along
|
||
<quantity id="7C459B4EFFA66670FC43FC2CFBCCF0AF" box="[993,1084,951,977]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="89" unit="mm" value="0.8">0.8 mm</quantity>
|
||
.
|
||
<emphasis id="89C9EAB9FFA66670FBEAFC2CFB69F0AF" box="[1096,1177,951,977]" italics="true" pageId="2" pageNumber="89">Flower</emphasis>
|
||
non-resupinate, red and white,
|
||
<quantity id="7C459B4EFFA66670FF77FC41FED8F08B" box="[213,296,986,1013]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="2" pageNumber="89" unit="cm" value="4.5" valueMax="5.0" valueMin="4.0">4–5 cm</quantity>
|
||
long, facing downwards.
|
||
<emphasis id="89C9EAB9FFA66670FDEEFC40FD62F08B" box="[588,658,987,1013]" italics="true" pageId="2" pageNumber="89">Ovary</emphasis>
|
||
dark purple, strongly recurved.
|
||
<emphasis id="89C9EAB9FFA66670FC5EFC41FBB5F08A" box="[1020,1093,986,1012]" italics="true" pageId="2" pageNumber="89">Sepals</emphasis>
|
||
with a longitudinal white stripe in the middle, flanked by two red stripes on the sides;
|
||
<emphasis id="89C9EAB9FFA66670FD4AFC65FC82F766" box="[744,882,1022,1048]" italics="true" pageId="2" pageNumber="89">dorsal sepal</emphasis>
|
||
4.2–4.8 ×
|
||
<quantity id="7C459B4EFFA66670FC49FC65FB9BF766" box="[1003,1131,1022,1048]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="2" pageNumber="89" unit="cm" value="1.15" valueMax="1.2" valueMin="1.1">1.1–1.2 cm</quantity>
|
||
, triangular oblong, apiculate apex, margin irregularly entire, 17-nerved;
|
||
<emphasis id="89C9EAB9FFA66670FDCEFBBAFCF4F745" box="[620,772,1057,1083]" italics="true" pageId="2" pageNumber="89">lateral sepals</emphasis>
|
||
3.6–4.3 × 1.8–2.0 cm, triangular falcate, acute and apiculate apex, 15–17-nerved.
|
||
<emphasis id="89C9EAB9FFA66670FED0FBDFFE48F720" box="[370,440,1092,1118]" italics="true" pageId="2" pageNumber="89">Petals</emphasis>
|
||
3.4–4.1 ×
|
||
<quantity id="7C459B4EFFA66670FD8CFBDEFD5DF721" box="[558,685,1093,1119]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="2" pageNumber="89" unit="cm" value="1.25" valueMax="1.3" valueMin="1.2">1.2–1.3 cm</quantity>
|
||
, triangular sub-falcate, with a longitudinal white stripe in the middle, flanked by two red stripes on the sides, 11–13-nerved.
|
||
<emphasis id="89C9EAB9FFA66670FCAEFBF3FC86F7FC" box="[780,886,1128,1154]" italics="true" pageId="2" pageNumber="89">Labellum</emphasis>
|
||
2.0–2.1 ×
|
||
<quantity id="7C459B4EFFA66670FC5BFBF3FBBBF7FD" box="[1017,1099,1128,1155]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="89" unit="cm" value="1.5">1.5 cm</quantity>
|
||
, sub-pyriform, adaxial surface covered by abundant spherical and sub-spherical yellow pseudopollen granules forming plush clumps towards the disk base, a semi-circular callus at the middle on the adaxial surface, an arc of
|
||
<quantity id="7C459B4EFFA66670FC4DFB34FBC0F7B4" box="[1007,1072,1199,1226]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="89" unit="mm" value="6.0">6 mm</quantity>
|
||
long on the abaxial surface;
|
||
<emphasis id="89C9EAB9FFA66670FACDFB34FF44F792" italics="true" pageId="2" pageNumber="89">basal half</emphasis>
|
||
hemi-rounded with entire, dark red margins;
|
||
<emphasis id="89C9EAB9FFA66670FD60FB49FCC6F792" box="[706,822,1234,1260]" italics="true" pageId="2" pageNumber="89">distal half</emphasis>
|
||
transversally extended in two dolabriform expansions, margins irregularly denticulate, dark cream, with irregular and enlarged dark garnet spots;
|
||
<emphasis id="89C9EAB9FFA66670FBD2FB6CFB55F66E" box="[1136,1189,1271,1296]" italics="true" pageId="2" pageNumber="89">apex</emphasis>
|
||
dark mustard with dark brown spots or completely yellowish, margin revolute of
|
||
<quantity id="7C459B4EFFA66670FCB0FA81FC80F64A" box="[786,880,1306,1332]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="2" pageNumber="89" unit="mm" value="8.5" valueMax="9.0" valueMin="8.0">8–9 mm</quantity>
|
||
long;
|
||
<emphasis id="89C9EAB9FFA66670FC1BFA81FBCAF64D" box="[953,1082,1306,1331]" italics="true" pageId="2" pageNumber="89">apex centre</emphasis>
|
||
a semi-ellipsoidal to cupuliform protuberance of 2.5–3.0 mm long, covered by pseudopollen, revolute margins dorsally as closed or open lips.
|
||
<emphasis id="89C9EAB9FFA66670FAF1FAA6FA5CF629" box="[1363,1452,1341,1367]" italics="true" pageId="2" pageNumber="89">Column</emphasis>
|
||
12 ×
|
||
<quantity id="7C459B4EFFA66670FF65FAFBFEF3F605" box="[199,259,1376,1403]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="89" unit="cm" value="5.0">5 cm</quantity>
|
||
, arcuate, semi-terete, slightly concave ventral surface, surface with minute dactylar papillae, cream and ventrally pale yellow with small and irregular dark red dots;
|
||
<emphasis id="89C9EAB9FFA66670FCFAFA1FFC11F6E0" box="[856,993,1412,1438]" italics="true" pageId="2" pageNumber="89">clinandrium</emphasis>
|
||
with minuscule dactylar appendices of variable size;
|
||
<emphasis id="89C9EAB9FFA66670FE85FA3CFE5EF6BF" box="[295,430,1447,1473]" italics="true" pageId="2" pageNumber="89">column foot</emphasis>
|
||
16–18 ×
|
||
<quantity id="7C459B4EFFA66670FDBFFA3CFD92F6BC" box="[541,610,1447,1474]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="89" unit="mm" value="5.0">5 mm</quantity>
|
||
, arcuate, dorsoventrally flattened and slightly concave with lateral ridges.
|
||
<emphasis id="89C9EAB9FFA66670FF2AFA50FF24F69B" box="[136,212,1483,1509]" italics="true" pageId="2" pageNumber="89">Anther</emphasis>
|
||
5.8–6.4 ×
|
||
<quantity id="7C459B4EFFA66670FEE9FA50FE4EF69B" box="[331,446,1483,1510]" metricMagnitude="-3" metricUnit="m" metricValue="5.25" metricValueMax="5.5" metricValueMin="5.0" pageId="2" pageNumber="89" unit="mm" value="5.25" valueMax="5.5" valueMin="5.0">5–5.5 mm</quantity>
|
||
, widely obovoid, slightly complanate dorsoventrally, thick dark cream with reddish brown spots at the centre or entirely cream, dorsal surface slightly concave, a thick carina extending rearward with a rounded superior margin from the middle to the apex, a conspicuous V-shaped promontory at the end of the carina.
|
||
<emphasis id="89C9EAB9FFA66670FA86F989FA5BF552" box="[1316,1451,1554,1580]" italics="true" pageId="2" pageNumber="89">Pollinarium</emphasis>
|
||
5 ×
|
||
<quantity id="7C459B4EFFA66670FF17F9AEFF08F52E" box="[181,248,1589,1616]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="89" unit="mm" value="3.0">3 mm</quantity>
|
||
, with four pollinia;
|
||
<emphasis id="89C9EAB9FFA66670FE7AF9AEFDB3F531" box="[472,579,1589,1615]" italics="true" pageId="2" pageNumber="89">viscidium</emphasis>
|
||
saddle shaped;
|
||
<emphasis id="89C9EAB9FFA66670FD54F9AEFC91F531" box="[758,865,1589,1615]" italics="true" pageId="2" pageNumber="89">pollinium</emphasis>
|
||
claviform, complanate, convex-concave, larger pair 5 ×
|
||
<quantity id="7C459B4EFFA66670FF16F9C2FF07F50D" box="[180,247,1625,1651]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="89" unit="mm" value="2.0">2 mm</quantity>
|
||
, smaller pair 3.4 ×
|
||
<quantity id="7C459B4EFFA66670FE6DF9C3FDD8F50D" box="[463,552,1624,1651]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="89" unit="mm" value="1.5">1.5 mm</quantity>
|
||
. Fruit, not recorded.
|
||
</paragraph>
|
||
<paragraph id="BB0236ABFFA66670FF1FF9E7FC1CF5E8" blockId="2.[136,1452,773,2077]" box="[189,1004,1660,1686]" pageId="2" pageNumber="89">
|
||
<emphasis id="89C9EAB9FFA66670FF1FF9E7FEB3F5E8" bold="true" box="[189,323,1660,1686]" pageId="2" pageNumber="89">Phenology:</emphasis>
|
||
—Flowering between February and March (
|
||
<bibRefCitation id="DF2C4B5AFFA66670FC93F9E7FC10F5E8" author="Collantes, B." box="[817,992,1660,1686]" pageId="2" pageNumber="89" refId="ref5956" refString="Collantes, B. (2014) Huancavelica: Amaru, paraiso de orquideas. Impresso Grafica S. A., Lima, 340 pp." type="book" year="2014">Collantes 2014</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F3A76520FFA66671FF1FF904FAB5F3A2" lastPageId="3" lastPageNumber="90" pageId="2" pageNumber="89" type="distribution">
|
||
<paragraph id="BB0236ABFFA66670FF1FF904FAAEF45A" blockId="2.[136,1452,773,2077]" pageId="2" pageNumber="89">
|
||
<emphasis id="89C9EAB9FFA66670FF1FF904FD94F5C4" bold="true" box="[189,612,1695,1722]" pageId="2" pageNumber="89">Distribution, habitat, and ecology:</emphasis>
|
||
—
|
||
<taxonomicName id="7CBD4D28FFA66670FD22F904FCACF5C7" box="[640,860,1695,1721]" class="Liliopsida" family="Orchidaceae" genus="Maxillaria" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="89" phylum="Tracheophyta" rank="species" species="pyhalae">
|
||
<emphasis id="89C9EAB9FFA66670FD22F904FCACF5C7" box="[640,860,1695,1721]" italics="true" pageId="2" pageNumber="89">Maxillaria pyhalae</emphasis>
|
||
</taxonomicName>
|
||
is only known from three localities (
|
||
<emphasis id="89C9EAB9FFA66670FABBF93BFACBF5C7" box="[1305,1339,1696,1721]" italics="true" pageId="2" pageNumber="89">i.e.</emphasis>
|
||
Regional Conservation Area Amaru-Chihuana, Velapaccha, and Roble) in the district of Huachocolpa, department of
|
||
<collectingRegion id="7979F849FFA66670FF2AF97DFED3F47E" box="[136,291,1766,1792]" country="Peru" name="Huancavelica" pageId="2" pageNumber="89">Huancavelica</collectingRegion>
|
||
,
|
||
<collectingCountry id="C3AA763BFFA66670FE8DF97CFE92F47F" box="[303,354,1767,1793]" name="Peru" pageId="2" pageNumber="89">Peru</collectingCountry>
|
||
(
|
||
<figureCitation id="23862A2EFFA66670FED1F97DFE48F47F" box="[371,440,1766,1793]" captionStart="FIGURE 3" captionStartId="5.[136,229,2001,2023]" captionTargetBox="[163,1427,833,1965]" captionTargetId="figure-320@5.[151,1436,821,1977]" captionTargetPageId="5" captionText="FIGURE 3. Geographical locations of Maxillaria bicentenaria (squares) and M. pyhalae (triangles) in Peru. Names in capital letters indicate political departments and names associated to red dots indicate provincial capitals. Colour scale indicates elevation." figureDoi="http://doi.org/10.5281/zenodo.5453892" httpUri="https://zenodo.org/record/5453892/files/figure.png" pageId="2" pageNumber="89">Fig. 3</figureCitation>
|
||
;
|
||
<bibRefCitation id="DF2C4B5AFFA66670FE67F97DFD84F47F" author="Collantes, B." box="[453,628,1766,1793]" pageId="2" pageNumber="89" refId="ref5956" refString="Collantes, B. (2014) Huancavelica: Amaru, paraiso de orquideas. Impresso Grafica S. A., Lima, 340 pp." type="book" year="2014">Collantes 2014</bibRefCitation>
|
||
). The species is locally abundant and can be found at elevations between 2300 and
|
||
<quantity id="7C459B4EFFA66670FF54F891FEBEF45A" box="[246,334,1802,1828]" metricMagnitude="3" metricUnit="m" metricValue="2.9" pageId="2" pageNumber="89" unit="m" value="2900.0">2900 m</quantity>
|
||
. They grow in humid and shaded undergrowth on rocks, but may also be found as epiphytes.
|
||
</paragraph>
|
||
<paragraph id="BB0236ABFFA66670FF1FF8B6FB22F4F0" blockId="2.[136,1452,773,2077]" pageId="2" pageNumber="89">
|
||
<emphasis id="89C9EAB9FFA66670FF1FF8B6FDB3F439" bold="true" box="[189,579,1837,1863]" pageId="2" pageNumber="89">Additional specimens examined:</emphasis>
|
||
—
|
||
<materialsCitation id="0BD53CF6FFA66670FDC2F8B5FC24F415" collectingDate="1999-02-14" collectionCode="MOL, USM" collectorName="Leon & Collantes" country="Peru" county="Prov. of Tayacaja" elevation="2600" location="Llactapata" municipality="Dist. of Huachocolpa" pageId="2" pageNumber="89" specimenCount="1" stateProvince="Huancavelica">
|
||
<collectingCountry id="C3AA763BFFA66670FDC2F8B5FD5AF436" box="[608,682,1838,1864]" name="Peru" pageId="2" pageNumber="89">PERU</collectingCountry>
|
||
.
|
||
<collectingRegion id="7979F849FFA66670FD17F8B6FCA2F439" box="[693,850,1837,1863]" country="Peru" name="Huancavelica" pageId="2" pageNumber="89">Huancavelica</collectingRegion>
|
||
:
|
||
<collectingCounty id="52634E27FFA66670FCFFF8B5FBD4F439" box="[861,1060,1837,1864]" pageId="2" pageNumber="89">Prov. of Tayacaja</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="5B66ACD1FFA66670FB92F8B6FAD4F439" box="[1072,1316,1837,1863]" pageId="2" pageNumber="89">Dist. of Huachocolpa</collectingMunicipality>
|
||
,
|
||
<location id="BE626070FFA66670FA92F8B6FA58F439" LSID="urn:lsid:plazi:treatment:331487BDFFA66676FF2AFDC9FD4BF603:BE626070FFA66670FA92F8B6FA58F439" box="[1328,1448,1837,1863]" country="Peru" county="Prov. of Tayacaja" municipality="Dist. of Huachocolpa" name="Llactapata" pageId="2" pageNumber="89" stateProvince="Huancavelica">Llactapata</location>
|
||
,
|
||
<quantity id="7C459B4EFFA66670FF2AF8CAFF2EF415" box="[136,222,1873,1899]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="2" pageNumber="89" unit="m" value="2600.0">
|
||
<elevation id="3090D198FFA66670FF2AF8CAFF2EF415" box="[136,222,1873,1899]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="2" pageNumber="89" unit="m" value="2600.0">2600 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="CF03106BFFA66670FF4BF8CAFE41F415" box="[233,433,1873,1899]" pageId="2" pageNumber="89" value="1999-02-14">
|
||
<collectingDate id="DF47E983FFA66670FF4BF8CAFE41F415" box="[233,433,1873,1899]" pageId="2" pageNumber="89" value="1999-02-14">14 February 1999</collectingDate>
|
||
</date>
|
||
,
|
||
<emphasis id="89C9EAB9FFA66670FE19F8CAFD12F415" box="[443,738,1873,1899]" italics="true" pageId="2" pageNumber="89">
|
||
<collectorName id="1648537DFFA66670FE19F8CAFE03F415" box="[443,499,1873,1899]" pageId="2" pageNumber="89">León</collectorName>
|
||
&
|
||
<collectorName id="1648537DFFA66670FDB7F8CAFD70F415" box="[533,640,1873,1899]" pageId="2" pageNumber="89">Collantes</collectorName>
|
||
2975[b]
|
||
</emphasis>
|
||
(
|
||
<collectionCode id="DDACAE6EFFA66670FD52F8CAFCC3F415" box="[752,819,1873,1899]" country="Peru" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15530" lsid="urn:lsid:biocol.org:col:15530" name="Universidad Nacional Agraria La Molina" pageId="2" pageNumber="89" type="Herbarium">MOL</collectionCode>
|
||
, photo;
|
||
<collectionCode id="DDACAE6EFFA66670FC28F8CAFC3BF415" box="[906,971,1873,1899]" country="Malaysia" httpUri="http://grbio.org/cool/11pi-mg53" name="Universiti Sains Malaysia" pageId="2" pageNumber="89">USM</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="0BD53CF6FFA66670FC7CF8CAFEEAF4F0" collectingDate="2021-03-22" collectionCode="USM" collectorName="Leon" country="Peru" elevation="2700" location="Inquilpata" pageId="2" pageNumber="89" specimenCount="1" stateProvince="Huancavelica">
|
||
<location id="BE626070FFA66670FC7CF8CAFBA2F415" LSID="urn:lsid:plazi:treatment:331487BDFFA66676FF2AFDC9FD4BF603:BE626070FFA66670FC7CF8CAFBA2F415" box="[990,1106,1873,1899]" country="Peru" name="Inquilpata" pageId="2" pageNumber="89" stateProvince="Huancavelica">Inquilpata</location>
|
||
,
|
||
<quantity id="7C459B4EFFA66670FBFEF8CAFB42F415" box="[1116,1202,1873,1899]" metricMagnitude="3" metricUnit="m" metricValue="2.7" pageId="2" pageNumber="89" unit="m" value="2700.0">
|
||
<elevation id="3090D198FFA66670FBFEF8CAFB42F415" box="[1116,1202,1873,1899]" metricMagnitude="3" metricUnit="m" metricValue="2.7" pageId="2" pageNumber="89" unit="m" value="2700.0">2700 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="CF03106BFFA66670FB1FF8CAFA99F415" box="[1213,1385,1873,1899]" pageId="2" pageNumber="89" value="2021-03-22">
|
||
<collectingDate id="DF47E983FFA66670FB1FF8CAFA99F415" box="[1213,1385,1873,1899]" pageId="2" pageNumber="89" value="2021-03-22">22 March 2021</collectingDate>
|
||
</date>
|
||
,
|
||
<emphasis id="89C9EAB9FFA66670FAD1F8CAFF30F4F0" italics="true" pageId="2" pageNumber="89">
|
||
<collectorName id="1648537DFFA66670FAD1F8CAFA5BF415" box="[1395,1451,1873,1899]" pageId="2" pageNumber="89">León</collectorName>
|
||
8651
|
||
</emphasis>
|
||
(
|
||
<collectionCode id="DDACAE6EFFA66670FF6DF8EFFEE1F4F0" box="[207,273,1908,1934]" country="Malaysia" httpUri="http://grbio.org/cool/11pi-mg53" name="Universiti Sains Malaysia" pageId="2" pageNumber="89">USM</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="0BD53CF6FFA66670FE87F8EFFB3EF4F0" box="[293,1230,1908,1935]" collectingDate="2014-02-17" collectionCode="USM" collectorName="de Robles & Egoavil" country="Peru" elevation="2630" location="Bosque de Robles" municipality="Dist. of Robles" pageId="2" pageNumber="89" specimenCount="1" stateProvince="Huancavelica">
|
||
<collectingMunicipality id="5B66ACD1FFA66670FE87F8EFFE21F4F0" box="[293,465,1908,1934]" pageId="2" pageNumber="89">Dist. of Robles</collectingMunicipality>
|
||
,
|
||
<location id="BE626070FFA66670FE7EF8EEFD59F4F0" LSID="urn:lsid:plazi:treatment:331487BDFFA66676FF2AFDC9FD4BF603:BE626070FFA66670FE7EF8EEFD59F4F0" box="[476,681,1908,1935]" country="Peru" municipality="Dist. of Robles" name="Bosque de Robles" pageId="2" pageNumber="89" stateProvince="Huancavelica">
|
||
Bosque
|
||
<collectorName id="1648537DFFA66670FD95F8EFFD59F4F0" box="[567,681,1908,1934]" pageId="2" pageNumber="89">de Robles</collectorName>
|
||
</location>
|
||
,
|
||
<quantity id="7C459B4EFFA66670FD16F8EFFCFBF4F1" box="[692,779,1908,1935]" metricMagnitude="3" metricUnit="m" metricValue="2.63" pageId="2" pageNumber="89" unit="m" value="2630.0">
|
||
<elevation id="3090D198FFA66670FD16F8EFFCFBF4F1" box="[692,779,1908,1935]" metricMagnitude="3" metricUnit="m" metricValue="2.63" pageId="2" pageNumber="89" unit="m" value="2630.0">2630 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="CF03106BFFA66670FCB5F8EFFC11F4F0" box="[791,993,1908,1934]" pageId="2" pageNumber="89" value="2014-02-17">
|
||
<collectingDate id="DF47E983FFA66670FCB5F8EFFC11F4F0" box="[791,993,1908,1934]" pageId="2" pageNumber="89" value="2014-02-17">17 February 2014</collectingDate>
|
||
</date>
|
||
,
|
||
<emphasis id="89C9EAB9FFA66670FC4EF8EFFB84F4F0" box="[1004,1140,1908,1934]" italics="true" pageId="2" pageNumber="89">
|
||
<collectorName id="1648537DFFA66670FC4EF8EFFBB3F4F0" box="[1004,1091,1908,1934]" pageId="2" pageNumber="89">Egoavil</collectorName>
|
||
152
|
||
</emphasis>
|
||
(
|
||
<collectionCode id="DDACAE6EFFA66670FB21F8EFFB36F4F0" box="[1155,1222,1908,1934]" country="Malaysia" httpUri="http://grbio.org/cool/11pi-mg53" name="Universiti Sains Malaysia" pageId="2" pageNumber="89">USM</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="BB0236ABFFA66671FF1FF80CFAB5F3A2" blockId="2.[136,1452,773,2077]" lastBlockId="3.[136,1452,158,220]" lastPageId="3" lastPageNumber="90" pageId="2" pageNumber="89">
|
||
<emphasis id="89C9EAB9FFA66670FF1FF80CFE48F4CC" bold="true" box="[189,440,1943,1970]" pageId="2" pageNumber="89">Conservation status:</emphasis>
|
||
—
|
||
<taxonomicName id="7CBD4D28FFA66670FE76F803FD5FF4CC" box="[468,687,1944,1970]" class="Liliopsida" family="Orchidaceae" genus="Maxillaria" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="89" phylum="Tracheophyta" rank="species" species="pyhalae">
|
||
<emphasis id="89C9EAB9FFA66670FE76F803FD5FF4CC" box="[468,687,1944,1970]" italics="true" pageId="2" pageNumber="89">Maxillaria pyhalae</emphasis>
|
||
</taxonomicName>
|
||
is known only from three localities where it is locally abundant. However, the observed population is calculated as less than 1000 individuals given that most growth points are the result of vegetative growth from the rhizome. Therefore, following the IUCN Red List (
|
||
<bibRefCitation id="DF2C4B5AFFA66670FB2EF844FAE7F487" author="IUCN" box="[1164,1303,2015,2041]" pageId="2" pageNumber="89" refId="ref6277" refString="IUCN. (2019) Guidelines for using the IUCN Red List categories and criteria. Version 14. Prepared by the Standards and Petitions Committee. Available from: http: // www. iucnredlist. org / documents / RedListGuidelines. pdf. (accessed 15 March 2021)" type="book" year="2019">IUCN 2019</bibRefCitation>
|
||
) criteria, the species would likely be categorized as Endangered (EN), criterion D, based on the number of mature individuals; the same categorization was previously assessed (
|
||
<bibRefCitation id="DF2C4B5AFFA76671FD35FF05FC77F3C7" author="Roque, J. E. & Leon, B." box="[663,903,158,185]" pageId="3" pageNumber="90" pagination="759 - 878" refId="ref6526" refString="Roque, J. E. & Leon, B. (2006) Orchidaceae endemicas del Peru. Revista Peruana de Biologia 13: 759 - 878. https: // doi. org / 10.15381 / rpb. v 13 i 2.1953" type="journal article" year="2006">Roque & León 2006</bibRefCitation>
|
||
) The species has been included in the Peruvian CITES list (Ministerio del Ambiente 2018) and the national plan for orchid conservation (SERNANP 2021).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="EFC26623FFA76671FF2AF8FAFF38FB55" ID-DOI="http://doi.org/10.5281/zenodo.5453886" ID-Zenodo-Dep="5453886" httpUri="https://zenodo.org/record/5453886/files/figure.png" pageId="3" pageNumber="90" startId="3.[136,229,1889,1911]" targetBox="[238,1343,266,1849]" targetPageId="3">
|
||
<paragraph id="BB0236ABFFA76671FF2AF8FAFF38FB55" blockId="3.[136,1452,1889,2091]" pageId="3" pageNumber="90">
|
||
<emphasis id="89C9EAB9FFA76671FF2AF8FAFF08F408" bold="true" box="[136,248,1889,1911]" pageId="3" pageNumber="90">FIGURE 1</emphasis>
|
||
.
|
||
<taxonomicName id="7CBD4D28FFA76671FEA7F8FAFE4EF408" box="[261,446,1889,1910]" class="Liliopsida" family="Orchidaceae" genus="Maxillaria" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="90" phylum="Tracheophyta" rank="species" species="pyhalae">
|
||
<emphasis id="89C9EAB9FFA76671FEA7F8FAFE4EF408" box="[261,446,1889,1910]" italics="true" pageId="3" pageNumber="90">Maxillaria pyhalae</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="89C9EAB9FFA76671FE69F8FAFE2CF408" bold="true" box="[459,476,1889,1910]" pageId="3" pageNumber="90">A</emphasis>
|
||
. Habit.
|
||
<emphasis id="89C9EAB9FFA76671FD89F8FAFDCBF408" bold="true" box="[555,571,1889,1910]" pageId="3" pageNumber="90">B</emphasis>
|
||
. Flower, frontal view.
|
||
<emphasis id="89C9EAB9FFA76671FCBAF8FAFCD9F409" bold="true" box="[792,809,1889,1911]" pageId="3" pageNumber="90">C</emphasis>
|
||
. Dissected flower with extended perianth and details of the dorsal sepal apex.
|
||
<emphasis id="89C9EAB9FFA76671FF5FF81EFEFEF4E4" bold="true" box="[253,270,1925,1946]" pageId="3" pageNumber="90">D</emphasis>
|
||
. Labellum and column, lateral view.
|
||
<emphasis id="89C9EAB9FFA76671FD21F81EFD63F4E4" bold="true" box="[643,659,1925,1946]" pageId="3" pageNumber="90">E</emphasis>
|
||
. Labellum, frontal view with detail of the abaxial side of the extended apex.
|
||
<emphasis id="89C9EAB9FFA76671FA35F81EFA56F4E4" bold="true" box="[1431,1446,1925,1946]" pageId="3" pageNumber="90">F</emphasis>
|
||
. Protuberance at the apex centre of the labellum, coronal view.
|
||
<emphasis id="89C9EAB9FFA76671FD77F832FD18F4C1" bold="true" box="[725,744,1961,1983]" pageId="3" pageNumber="90">G</emphasis>
|
||
. Column, frontal and lateral views with details of the margins.
|
||
<emphasis id="89C9EAB9FFA76671FA98F832FABDF4C1" bold="true" box="[1338,1357,1961,1983]" pageId="3" pageNumber="90">G</emphasis>
|
||
. Column, lateral view with details of the surface.
|
||
<emphasis id="89C9EAB9FFA76671FDABF856FDECF49C" bold="true" box="[521,540,1997,2018]" pageId="3" pageNumber="90">H</emphasis>
|
||
. Anther and pollinarium, ventral and dorsal views. Note the flower is non-resupinate and it is shown as resupinate in C for graphical purposes only. Drawn from
|
||
<emphasis id="89C9EAB9FFA76671FCB0F869FBC0FB78" box="[786,1072,2033,2055]" italics="true" pageId="3" pageNumber="90">M. León & B. Collantes 2975</emphasis>
|
||
(USM) by B. Collantes (personal plate nº 21).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="EFC26623FFA06676FF2AFBBDFAFBF743" ID-DOI="http://doi.org/10.5281/zenodo.5453888" ID-Zenodo-Dep="5453888" box="[136,1291,1062,1085]" httpUri="https://zenodo.org/record/5453888/files/figure.png" pageId="4" pageNumber="91" startId="4.[136,229,1062,1084]" targetBox="[161,1426,200,1028]" targetPageId="4">
|
||
<paragraph id="BB0236ABFFA06676FF2AFBBDFAFBF743" blockId="4.[136,1291,1062,1085]" box="[136,1291,1062,1085]" pageId="4" pageNumber="91">
|
||
<emphasis id="89C9EAB9FFA06676FF2AFBBDFF07F745" bold="true" box="[136,247,1062,1084]" pageId="4" pageNumber="91">FIGURE 2</emphasis>
|
||
.
|
||
<taxonomicName id="7CBD4D28FFA06676FEA1FBBDFE4CF745" box="[259,444,1062,1083]" class="Liliopsida" family="Orchidaceae" genus="Maxillaria" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="91" phylum="Tracheophyta" rank="species" species="pyhalae">
|
||
<emphasis id="89C9EAB9FFA06676FEA1FBBDFE4CF745" box="[259,444,1062,1083]" italics="true" pageId="4" pageNumber="91">Maxillaria pyhalae</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="89C9EAB9FFA06676FE6AFBBDFE29F745" bold="true" box="[456,473,1062,1083]" pageId="4" pageNumber="91">A</emphasis>
|
||
. Habit.
|
||
<emphasis id="89C9EAB9FFA06676FD85FBBCFDC7F742" bold="true" box="[551,567,1063,1084]" pageId="4" pageNumber="91">B</emphasis>
|
||
. Flower, frontal view. Photographs by L. Egoavil (A) and B. Collantes (B).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="F3A76520FFA06676FF1FFBF1FD4BF603" pageId="4" pageNumber="91" type="discussion">
|
||
<paragraph id="BB0236ABFFA06676FF1FFBF1FD4BF603" blockId="4.[136,1452,1130,1405]" pageId="4" pageNumber="91">
|
||
<emphasis id="89C9EAB9FFA06676FF1FFBF1FEB8F7FA" bold="true" box="[189,328,1130,1156]" pageId="4" pageNumber="91">Comments:</emphasis>
|
||
—Besides the new taxon proposed herein, we believe
|
||
<taxonomicName id="7CBD4D28FFA06676FC04FBF0FBD4F7FA" box="[934,1060,1130,1156]" class="Liliopsida" family="Orchidaceae" genus="Maxillaria" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="91" phylum="Tracheophyta" rank="species" species="pyhalae">
|
||
<emphasis id="89C9EAB9FFA06676FC04FBF0FBD4F7FA" box="[934,1060,1130,1156]" italics="true" pageId="4" pageNumber="91">M. pyhalae</emphasis>
|
||
</taxonomicName>
|
||
is morphologically most similar to
|
||
<taxonomicName id="7CBD4D28FFA06676FF2AFB15FD9FF7D7" authority="Reichenbach (1886: 616)" authorityName="Reichenbach" authorityPageNumber="616" authorityYear="1886" box="[136,623,1166,1193]" class="Liliopsida" family="Orchidaceae" genus="Maxillaria" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="91" phylum="Tracheophyta" rank="species" species="fucata">
|
||
<emphasis id="89C9EAB9FFA06676FF2AFB15FEBCF7D6" box="[136,332,1166,1192]" italics="true" pageId="4" pageNumber="91">Maxillaria fucata</emphasis>
|
||
<bibRefCitation id="DF2C4B5AFFA06676FEF1FB15FD9FF7D7" author="Reichenbach, H. G." box="[339,623,1166,1193]" pageId="4" pageNumber="91" pagination="616" refId="ref6570" refString="Reichenbach, H. G. (1886) New garden plants. The Gardeners' chronicle: a weekly illustrated journal of horticulture and allied subjects, new series 26: 616." type="book chapter" year="1886">Reichenbach (1886: 616)</bibRefCitation>
|
||
</taxonomicName>
|
||
, rather than
|
||
<taxonomicName id="7CBD4D28FFA06676FD5BFB15FAD2F7D7" authority="R. H. Pearson (1913: 258)" authorityName="R. H. Pearson" authorityPageNumber="258" authorityYear="1913" box="[761,1314,1166,1193]" class="Liliopsida" family="Orchidaceae" genus="Maxillaria" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="91" phylum="Tracheophyta" rank="species" species="fletcheriana">
|
||
<emphasis id="89C9EAB9FFA06676FD5BFB15FC0EF7D6" box="[761,1022,1166,1192]" italics="true" pageId="4" pageNumber="91">Maxillaria fletcheriana</emphasis>
|
||
R.H. Pearson (1913: 258)
|
||
</taxonomicName>
|
||
(a synonym of
|
||
<taxonomicName id="7CBD4D28FFA06676FF04FB29FE13F7B2" authority="Rolfe [1922: 25]" authorityName="Rolfe, Bull. Misc. Inform. Kew" authorityPageNumber="25" authorityYear="1922" box="[166,483,1201,1228]" class="Liliopsida" family="Orchidaceae" genus="Maxillaria" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="91" phylum="Tracheophyta" rank="species" species="insignis">
|
||
<emphasis id="89C9EAB9FFA06676FF04FB29FF4DF7B5" box="[166,189,1202,1227]" italics="true" pageId="4" pageNumber="91">M</emphasis>
|
||
.
|
||
<emphasis id="89C9EAB9FFA06676FF69FB29FED2F7B5" box="[203,290,1202,1227]" italics="true" pageId="4" pageNumber="91">insignis</emphasis>
|
||
<bibRefCitation id="DF2C4B5AFFA06676FE8BFB2AFE13F7B2" author="Rolfe, R. A." box="[297,483,1201,1228]" pageId="4" pageNumber="91" pagination="22 - 26" refId="ref6638" refString="Rolfe, R. A. (1922) New orchids: decas XLIX. Bulletin of miscellaneous information 1922: 22 - 26. https: // doi. org / 10.2307 / 4118619" type="journal article" year="1922">Rolfe [1922: 25]</bibRefCitation>
|
||
</taxonomicName>
|
||
) as suggested by Bennet & Christenson [2001] in the protologue. The line drawings of
|
||
<taxonomicName id="7CBD4D28FFA06676FF2AFB4EFEF5F791" box="[136,261,1237,1263]" class="Liliopsida" family="Orchidaceae" genus="Maxillaria" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="91" phylum="Tracheophyta" rank="species" species="pyhalae">
|
||
<emphasis id="89C9EAB9FFA06676FF2AFB4EFEF5F791" box="[136,261,1237,1263]" italics="true" pageId="4" pageNumber="91">M. pyhalae</emphasis>
|
||
</taxonomicName>
|
||
published in the IOP misrepresents important details of the plant specimens, especially the labellum, which makes the identification of this taxon challenging at times. The labellum shape is also not described in detail in the protologue. This has caused confusion with regard to the identity and correct interpretation of this taxon. Additionally, it has been determined that the
|
||
<typeStatus id="64068809FFA06676FE49FAA4FDBDF627" box="[491,589,1343,1369]" pageId="4" pageNumber="91" type="holotype">holotype</typeStatus>
|
||
and an isotype of
|
||
<taxonomicName id="7CBD4D28FFA06676FCBDFADBFC60F627" box="[799,912,1343,1370]" class="Liliopsida" family="Orchidaceae" genus="Maxillaria" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="91" phylum="Tracheophyta" rank="species" species="phylae">
|
||
<emphasis id="89C9EAB9FFA06676FCBDFADBFCC6F627" box="[799,822,1344,1369]" italics="true" pageId="4" pageNumber="91">M</emphasis>
|
||
.
|
||
<emphasis id="89C9EAB9FFA06676FCE7FAA4FC60F627" box="[837,912,1343,1369]" italics="true" pageId="4" pageNumber="91">phylae</emphasis>
|
||
</taxonomicName>
|
||
are gathered at MOL under the same collection number (i.e.
|
||
<emphasis id="89C9EAB9FFA06676FEB5FAF8FE3DF603" box="[279,461,1378,1405]" italics="true" pageId="4" pageNumber="91">León et al. 2975</emphasis>
|
||
, MO;
|
||
<bibRefCitation id="DF2C4B5AFFA06676FDB4FAF9FD5FF603" author="Trujillo, D." box="[534,687,1378,1405]" pageId="4" pageNumber="91" pagination="1 - 88" refId="ref6905" refString="Trujillo, D. (2014) Annotated list of orchidaceae types of the Bennett collection at the forestry herbarium MOL. Lankesteriana 14: 1 - 88. https: // doi. org / 10.15517 / lank. v 14 i 1.15584" type="journal article" year="2014">Trujillo 2014</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |