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<mods:title id="A2CAE432FB645CC4848FE8EE1A769A14">New data on the millipede genus Shearia Mikhaljova, 2000 from the Russian and Mongolian Altais (Diplopoda: Chordeumatida: Diplomaragnidae)</mods:title>
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<taxonomicName id="50B24D5B327EFFEEFEBBEC2CE4DDFDA0" ID-CoL="6YM88" authority="Mikhaljova, 2012" authorityName="Mikhaljova" authorityYear="2012" box="[306,657,517,541]" class="Diplopoda" family="Diplomaragnidae" genus="Shearia" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="223" phylum="Arthropoda" rank="species" species="longa">
<emphasis id="A5C6EACA327EFFEEFEBBEC2CE786FDA0" box="[306,458,517,541]" italics="true" pageId="4" pageNumber="223">Shearia longa</emphasis>
<bibRefCitation id="F3234B29327EFFEEFE5BEC2CE4DDFDA0" author="Mikhaljova E. V." box="[466,657,517,541]" pageId="4" pageNumber="223" pagination="41 - 50" refId="ref3476" refString="Mikhaljova E. V. 2012. The class Diplopoda in Mongolia, with description of a new species // Zootaxa. Vol. 3418. P. 41 - 50." type="journal article" year="2012">Mikhaljova, 2012</bibRefCitation>
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<figureCitation id="0F892A5D327EFFEEFE31EC0BE444FD87" box="[440,520,546,570]" captionStart="Fig" captionStartId="4.[226,257,1969,1988]" captionTargetBox="[189,1399,1100,1955]" captionTargetId="figure-429@4.[189,1399,1100,1955]" captionTargetPageId="4" captionText="Fig. 14. Distribution of Shearia khakassica (circle) and S. longa (square). Previously known localities marked in black, new records given in white. Рис. 14. Распространение Shearia khakassica (круг) и S. longa (квадрат). Чёрным отмечены ранее иЗвестные места находок, новые находки отмечены белым." figureDoi="http://doi.org/10.5281/zenodo.13163598" httpUri="https://zenodo.org/record/13163598/files/figure.png" pageId="4" pageNumber="223">Fig. 14</figureCitation>
.
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<taxonomicName id="50B24D5B327EFFEEFF6BEC6EE46AFDE7" ID-CoL="6YM88" authority="Mikhaljova, 2012: 42" authorityName="Mikhaljova" authorityPageNumber="42" authorityYear="2012" box="[226,550,583,602]" class="Diplopoda" family="Diplomaragnidae" genus="Shearia" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="223" phylum="Arthropoda" rank="species" species="longa">
<emphasis id="A5C6EACA327EFFEEFF6BEC6EE712FDE7" box="[226,350,583,602]" italics="true" pageId="4" pageNumber="223">Shearia longa</emphasis>
<bibRefCitation id="F3234B29327EFFEEFEECEC6EE46AFDE7" author="Mikhaljova E. V." box="[357,550,583,602]" pageId="4" pageNumber="223" pagination="41 - 50" refId="ref3476" refString="Mikhaljova E. V. 2012. The class Diplopoda in Mongolia, with description of a new species // Zootaxa. Vol. 3418. P. 41 - 50." type="journal article" year="2012">Mikhaljova, 2012: 42</bibRefCitation>
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, 4346: figs.
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<materialsCitation id="27DA3C85327EFFEEFF6BEC76E4D6FD07" collectingDate="2015-07-11" collectionCode="MATERIAL, EXAMINED" collectorName="A. A. Fomichev." country="Mongolia" elevation="2700" latitude="48.15" location="Tsengel Hayrhan Nuruu Mt. Range" longLatPrecision="1112" longitude="89.23333" municipality="Altai District" pageId="4" pageNumber="223" specimenCount="9" specimenCount-female="2" specimenCount-male="7" stateProvince="Bayan-Olgiy">
<collectionCode id="F1A3AE1D327EFFEEFF6BEC76E784FDCF" box="[226,456,607,626]" pageId="4" pageNumber="223">MATERIAL EXAMINED</collectionCode>
.
<specimenCount id="81B4FD51327EFFEEFE47EC49E448FDCE" box="[462,516,605,627]" count="7" pageId="4" pageNumber="223" type="male">7 ♂♂</specimenCount>
,
<specimenCount id="81B4FD51327EFFEEFD87EC76E478FDCE" box="[526,564,605,627]" count="2" pageId="4" pageNumber="223" type="female">2 ♀♀</specimenCount>
,
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<collectingCountry id="EFA57648327EFFEEFDB6EC76E4D8FDCF" box="[575,660,607,626]" name="Mongolia" pageId="4" pageNumber="223">Mongolia</collectingCountry>
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,
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(Aimag),
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(Somon),
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,
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W of Mt Elt Uul
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,
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;
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, screes, under stones,
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a.s.l.
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,
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<collectingDate id="F348E9F0327EFFEEFEFEEC8EE7ACFD07" box="[375,480,679,698]" pageId="4" pageNumber="223" value="2015-07-11">11.VII.2015</collectingDate>
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, leg.
<collectorName id="3A47530E327EFFEEFD86EC8EE4D6FD07" box="[527,666,679,698]" pageId="4" pageNumber="223">A.A. Fomichev.</collectorName>
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</paragraph>
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<paragraph id="970D36D8327EFFEEFF6BEC96E4C5FD52" blockId="4.[189,775,517,859]" pageId="4" pageNumber="223">
DISTRIBUTION. This species is known only from the Mongolian Altais, western
<collectingCountry id="EFA57648327EFFEEFE4EECF3E469FD4D" box="[455,549,730,752]" name="Mongolia" pageId="4" pageNumber="223">Mongolia</collectingCountry>
(
<figureCitation id="0F892A5D327EFFEEFDBDECF3E431FD52" box="[564,637,730,752]" captionStart="Fig" captionStartId="4.[226,257,1969,1988]" captionTargetBox="[189,1399,1100,1955]" captionTargetId="figure-429@4.[189,1399,1100,1955]" captionTargetPageId="4" captionText="Fig. 14. Distribution of Shearia khakassica (circle) and S. longa (square). Previously known localities marked in black, new records given in white. Рис. 14. Распространение Shearia khakassica (круг) и S. longa (квадрат). Чёрным отмечены ранее иЗвестные места находок, новые находки отмечены белым." figureDoi="http://doi.org/10.5281/zenodo.13163598" httpUri="https://zenodo.org/record/13163598/files/figure.png" pageId="4" pageNumber="223">Fig. 14</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="DFA86553327EFFEEFF6BECDCE33FFC56" pageId="4" pageNumber="223" type="discussion">
<paragraph id="970D36D8327EFFEEFF6BECDCE6A3FCE6" blockId="4.[189,775,517,859]" pageId="4" pageNumber="223">
REMARKS.Originally described from the Tsengel Somon of the
<collectingRegion id="5576F83A327EFFEEFF75ED25E7F1FC98" box="[252,445,780,805]" country="Mongolia" name="Bayan-Olgiy" pageId="4" pageNumber="223">Bayan-Ölgii Aimag</collectingRegion>
[see
<bibRefCitation id="F3234B29327EFFEEFE7BED26E4E8FC99" author="Mikhaljova E. V." box="[498,676,783,805]" pageId="4" pageNumber="223" pagination="41 - 50" refId="ref3476" refString="Mikhaljova E. V. 2012. The class Diplopoda in Mongolia, with description of a new species // Zootaxa. Vol. 3418. P. 41 - 50." type="journal article" year="2012">Mikhaljova, 2012</bibRefCitation>
], this species is now recorded from outside its
<emphasis id="A5C6EACA327EFFEEFDADED02E4D9FCFD" box="[548,661,811,832]" italics="true" pageId="4" pageNumber="223">terra typica</emphasis>
for the first time.
</paragraph>
<paragraph id="970D36D8327EFFEEFF6BED53E4B0FC2D" blockId="4.[189,775,890,992]" box="[226,764,890,912]" pageId="4" pageNumber="223">
The following key can be proposed to
<taxonomicName id="50B24D5B327EFFEEFDD4ED53E4B0FC2D" box="[605,764,890,912]" class="Diplopoda" family="Diplomaragnidae" genus="Shearia" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="223" phylum="Arthropoda" rank="species" species="undefined-1">
<emphasis id="A5C6EACA327EFFEEFDD4ED53E4E4FC32" box="[605,680,890,911]" italics="true" pageId="4" pageNumber="223">Shearia</emphasis>
species.
</taxonomicName>
</paragraph>
<paragraph id="970D36D8327EFFEEFF34EDBCE54FFC79" blockId="4.[189,775,890,992]" pageId="4" pageNumber="223">1(4) Anterior angiocoxal processes of posterior gonopods with very large front branches and small outer ones ............ 2</paragraph>
<paragraph id="970D36D8327EFFEEFF34EDE3E33FFED6" blockId="4.[189,775,890,992]" lastBlockId="4.[812,1399,315,1004]" pageId="4" pageNumber="223">
2(3) Colpocoxites of posterior gonopods strongly curved caudad, with external lateral lobes ................................. ....................................
<emphasis id="A5C6EACA327EFFEEFBB0EF7FE2F2FED6" box="[1081,1214,342,363]" italics="true" pageId="4" pageNumber="223">shushenskaya</emphasis>
<bibRefCitation id="F3234B29327EFFEEFB4FEF7FE33FFED6" author="Mikhaljova E. V." box="[1222,1395,342,364]" pageId="4" pageNumber="223" pagination="153 - 181" refId="ref3405" refString="Mikhaljova E. V. 2000. Review of the millipede family Diplomaragnidae (Diplopoda: Chordeumatida) // Arthropoda Selecta (for 1999). Vol. 8. No. 3. P. 153 - 181." type="book chapter" year="2000">Mikhaljova, 2000</bibRefCitation>
</paragraph>
<paragraph id="970D36D8327EFFEEFCA5EF58E33FFE06" blockId="4.[812,1399,315,1004]" pageId="4" pageNumber="223">
3(2) Colpocoxites of posterior gonopods regularly curved caudad, without external lateral lobes ............................ ..........................................
<emphasis id="A5C6EACA327EFFEEFBD7EF8FE2F2FE06" box="[1118,1214,422,443]" italics="true" pageId="4" pageNumber="223">teletskaya</emphasis>
<bibRefCitation id="F3234B29327EFFEEFB4FEF8FE33FFE06" author="Mikhaljova E. V." box="[1222,1395,422,444]" pageId="4" pageNumber="223" pagination="153 - 181" refId="ref3405" refString="Mikhaljova E. V. 2000. Review of the millipede family Diplomaragnidae (Diplopoda: Chordeumatida) // Arthropoda Selecta (for 1999). Vol. 8. No. 3. P. 153 - 181." type="book chapter" year="2000">Mikhaljova, 2000</bibRefCitation>
</paragraph>
<paragraph id="970D36D8327EFFEEFCA5EFE8E33FFE4C" blockId="4.[812,1399,315,1004]" pageId="4" pageNumber="223">4(1) Anterior angiocoxal processes of posterior gonopods small, both branches equal in height ........................... 5</paragraph>
<paragraph id="970D36D8327EFFEEFCA5EFDFE33FFD9B" blockId="4.[812,1399,315,1004]" pageId="4" pageNumber="223">
5(6) Colpocoxite tips of posterior gonopods with subquadrate excavations .........................
<emphasis id="A5C6EACA327EFFEEFBE7EC38E2F2FD9B" box="[1134,1214,529,550]" italics="true" pageId="4" pageNumber="223">rybalovi</emphasis>
<bibRefCitation id="F3234B29327EFFEEFB4FEC38E33FFD9B" author="Mikhaljova E. V." box="[1222,1395,529,551]" pageId="4" pageNumber="223" pagination="153 - 181" refId="ref3405" refString="Mikhaljova E. V. 2000. Review of the millipede family Diplomaragnidae (Diplopoda: Chordeumatida) // Arthropoda Selecta (for 1999). Vol. 8. No. 3. P. 153 - 181." type="book chapter" year="2000">Mikhaljova, 2000</bibRefCitation>
</paragraph>
<paragraph id="970D36D8327EFFEEFCA5EC02E33FFDFC" blockId="4.[812,1399,315,1004]" box="[812,1395,555,577]" pageId="4" pageNumber="223">6(5) Colpocoxite tips of posterior gonopods different ..... 7</paragraph>
<paragraph id="970D36D8327EFFEEFCA5EC6FE33FFD2D" blockId="4.[812,1399,315,1004]" pageId="4" pageNumber="223">
7(8) Front branches of anterior angiocoxal processes of posterior gonopods distinctly triangular in shape, with point- ed tips (
<figureCitation id="0F892A5D327EFFEEFC28EC52E5A4FD2D" box="[929,1000,635,657]" captionStart="Figs 1520" captionStartId="5.[226,263,1966,1985]" captionTargetBox="[189,1399,563,1929]" captionTargetId="figure-220@5.[189,1399,563,1929]" captionTargetPageId="5" captionText="Figs 1520. Shearia khakassica, male (Republic of Altai, near Lak-Yyash Lakes): 1518 — gonopods; 19 — coxae 10; 20 — coxae 11; 15, 17, 1920 — oral view; 16, 18 — caudal view. Scale bars: 1 mm (1518) and 0.4 mm (1920). Designations explained in text. Рис. 1520. Shearia khakassica, самец: 1518 — гоноподы; 19 — коксы 10-й пары ног; 20 — коксы 11-й пары ног; 15, 17, 19 20 — спереди; 16, 18 — сЗади. МасШтаб: 1 мм (1518) и 0,4 мм (1920). ОбЪЯснение обоЗначений дано в тексте." figureDoi="http://doi.org/10.5281/zenodo.13163600" httpUri="https://zenodo.org/record/13163600/files/figure.png" pageId="4" pageNumber="223">Fig. 17</figureCitation>
) ............................................
<taxonomicName id="50B24D5B327EFFEEFA80EC52E33FFD2D" authorityName="Mikhaljova" authorityYear="2000" box="[1289,1395,635,656]" class="Diplopoda" family="Diplomaragnidae" genus="Shearia" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="223" phylum="Arthropoda" rank="species" species="khakassica">
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</taxonomicName>
</paragraph>
<paragraph id="970D36D8327EFFEEFCA5ECBFE33FFD7B" blockId="4.[812,1399,315,1004]" pageId="4" pageNumber="223">8(7) Front branches of anterior angiocoxal processes of posterior gonopods different, without pointed tips .......... 9</paragraph>
<paragraph id="970D36D8327EFFEEFCA5ECE2E33FFD46" blockId="4.[812,1399,315,1004]" pageId="4" pageNumber="223">9(12) Colpocoxites of posterior gonopods tapering distally, straight ........................................................................ 10</paragraph>
<paragraph id="970D36D8327EFFEEFCA5ED28E33FFC8D" blockId="4.[812,1399,315,1004]" pageId="4" pageNumber="223">
10(11) Anterior gonopod telopodites very long and twisted ...............................................................................
<taxonomicName id="50B24D5B327EFFEEFAB5ED32E33FFC8D" authorityName="Mikhaljova" authorityYear="2012" box="[1340,1395,795,816]" class="Diplopoda" family="Diplomaragnidae" genus="Shearia" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="223" phylum="Arthropoda" rank="species" species="longa">
<emphasis id="A5C6EACA327EFFEEFAB5ED32E33FFC8D" box="[1340,1395,795,816]" italics="true" pageId="4" pageNumber="223">longa</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="970D36D8327EFFEEFCA5ED1FE33FFCDB" blockId="4.[812,1399,315,1004]" pageId="4" pageNumber="223">
11(10) Anterior gonopod telopodites shorter, not twisted .. ..........................................................................
<taxonomicName id="50B24D5B327EFFEEFAA8ED78E33FFCDB" authorityName="Mikhaljova" authorityYear="2000" box="[1313,1395,849,870]" class="Diplopoda" family="Diplomaragnidae" genus="Shearia" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="223" phylum="Arthropoda" rank="species" species="calycina">
<emphasis id="A5C6EACA327EFFEEFAA8ED78E33FFCDB" box="[1313,1395,849,870]" italics="true" pageId="4" pageNumber="223">calycina</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="970D36D8327EFFEEFCA5ED42E33FFC26" blockId="4.[812,1399,315,1004]" pageId="4" pageNumber="223">12(13) Colpocoxites of posterior gonopods broad, concave .................................................................................... 13</paragraph>
<paragraph id="970D36D8327EFFEEFCA5ED88E33FFC0B" blockId="4.[812,1399,315,1004]" box="[812,1395,929,951]" pageId="4" pageNumber="223">
13(14) Mesal sheath plate small .........................
<taxonomicName id="50B24D5B327EFFEEFA86ED88E33FFC0B" baseAuthorityName="Gulicka" baseAuthorityYear="1972" box="[1295,1395,929,950]" class="Diplopoda" family="Diplomaragnidae" genus="Shearia" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="223" phylum="Arthropoda" rank="species" species="densecava">
<emphasis id="A5C6EACA327EFFEEFA86ED88E33FFC0B" box="[1295,1395,929,950]" italics="true" pageId="4" pageNumber="223">densecava</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="970D36D8327EFFEEFCA5ED92E33FFC56" blockId="4.[812,1399,315,1004]" pageId="4" pageNumber="223">
14(13) Mesal sheath plate very large ................................... ..............................................
<taxonomicName id="50B24D5B327EFFEEFBFCEDFFE33FFC56" authority="Mikhaljova, 2000" authorityName="Mikhaljova" authorityYear="2000" box="[1141,1395,982,1004]" class="Diplopoda" family="Diplomaragnidae" genus="Shearia" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="223" phylum="Arthropoda" rank="species" species="oiskaya">
<emphasis id="A5C6EACA327EFFEEFBFCEDFFE2F3FC56" box="[1141,1215,982,1003]" italics="true" pageId="4" pageNumber="223">oiskaya</emphasis>
<bibRefCitation id="F3234B29327EFFEEFB4FEDFFE33FFC56" author="Mikhaljova E. V." box="[1222,1395,982,1004]" pageId="4" pageNumber="223" pagination="153 - 181" refId="ref3405" refString="Mikhaljova E. V. 2000. Review of the millipede family Diplomaragnidae (Diplopoda: Chordeumatida) // Arthropoda Selecta (for 1999). Vol. 8. No. 3. P. 153 - 181." type="book chapter" year="2000">Mikhaljova, 2000</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>