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<document id="A8C3A6C87537BEF7CF5F3483D12B0187" ID-DOI="10.15298/rusentj.28.2.16" ID-ISSN="0132-8069" ID-Zenodo-Dep="13163590" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722578434347" checkinUser="felipe" docAuthor="Nefediev, P. S." docDate="2019" docId="1F1B87CE327BFFEBFEAAEFEBE441FBAA" docLanguage="en" docName="RussEntomolJ.28.2.219-225.pdf" docOrigin="Russian Entomological Journal 28 (2)" docSource="http://dx.doi.org/10.15298/rusentj.28.2.16" docStyle="DocumentStyle:0D5BAC5AA0C130B3809AF578A601D837.2:RussEntomolJ.2019-.journal_article" docStyleId="0D5BAC5AA0C130B3809AF578A601D837" docStyleName="RussEntomolJ.2019-.journal_article" docStyleVersion="2" docTitle="Shearia calycina Mikhaljova 2000" docType="treatment" docVersion="2" lastPageNumber="220" masterDocId="E322FFB6327AFFEAFF89EE29E64CFFBD" masterDocTitle="New data on the millipede genus Shearia Mikhaljova, 2000 from the Russian and Mongolian Altais (Diplopoda: Chordeumatida: Diplomaragnidae)" masterLastPageNumber="225" masterPageNumber="219" pageNumber="220" updateTime="1722579031342" updateUser="ExternalLinkService">
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<mods:title id="4D1D826F6BEBBD03E33A0512E5BFE3D4">New data on the millipede genus Shearia Mikhaljova, 2000 from the Russian and Mongolian Altais (Diplopoda: Chordeumatida: Diplomaragnidae)</mods:title>
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<subSubSection id="DFA86553327BFFEBFEAAEFEBE4ECFE67" box="[291,672,450,474]" pageId="1" pageNumber="220" type="nomenclature">
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<taxonomicName id="50B24D5B327BFFEBFEAAEFEBE4ECFE67" ID-CoL="4X495" authority="Mikhaljova, 2000" authorityName="Mikhaljova" authorityYear="2000" box="[291,672,450,474]" class="Diplopoda" family="Diplomaragnidae" genus="Shearia" kingdom="Animalia" order="Chordeumatida" pageId="1" pageNumber="220" phylum="Arthropoda" rank="species" species="calycina">
<emphasis id="A5C6EACA327BFFEBFEAAEFEBE794FE67" box="[291,472,450,474]" italics="true" pageId="1" pageNumber="220">Shearia calycina</emphasis>
<bibRefCitation id="F3234B29327BFFEBFE68EFEBE4ECFE67" author="Mikhaljova E. V." box="[481,672,450,474]" pageId="1" pageNumber="220" pagination="153 - 181" refId="ref3405" refString="Mikhaljova E. V. 2000. Review of the millipede family Diplomaragnidae (Diplopoda: Chordeumatida) // Arthropoda Selecta (for 1999). Vol. 8. No. 3. P. 153 - 181." type="book chapter" year="2000">Mikhaljova, 2000</bibRefCitation>
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</paragraph>
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<paragraph id="970D36D8327BFFEBFE26EFF6E458FE45" blockId="1.[189,776,376,1048]" box="[431,532,479,504]" pageId="1" pageNumber="220">
<figureCitation id="0F892A5D327BFFEBFE26EFF6E45CFE45" box="[431,528,479,504]" captionStart-0="Fig" captionStart-1="Figs 2–7" captionStartId-0="1.[226,257,1969,1988]" captionStartId-1="2.[226,263,1947,1966]" captionTargetBox-0="[189,1399,1085,1940]" captionTargetBox-1="[189,1399,568,1933]" captionTargetId-0="figure-502@1.[189,1399,1085,1940]" captionTargetId-1="figure-126@2.[189,1399,568,1933]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="Fig. 1. Distribution of Shearia calycina (triangle) and S. densecava (circle). Previously known localities marked in black, new records given in white. Рис. 1. Распространение Shearia calycina (треугольник) и S. densecava (круг). Чёрным отмечены ранее иЗвестные места находок, новые находки отмечены белым." captionText-1="Figs 2–7. Shearia calycina, male (Republic of Altai, floodplain of Karagem River): 2–5 — gonopods; 6 — coxae 10; 7 — coxae 11; 2, 4, 6–7 — oral view; 3–5 — caudal view. Scale bars: 1 mm (2–3), 0.4 mm (4–5) and 0.3 mm (6–7). Designations explained in text. Рис. 2–7. Shearia calycina, самец (Республика Алтай, пойма р. Карагем): 2–5 — гоноподы; 6 — коксы 10-й пары ног; 7 — коксы 11-й пары ног; 2, 4, 6–7 — спереди; 3–5 — сЗади. МасШтаб: 1 мм (2–3), 0,4 мм (4–5) и 0,3 мм (6–7). ОбЪЯснение обоЗначений дано в тексте." figureDoi-0="http://doi.org/10.5281/zenodo.13163592" figureDoi-1="http://doi.org/10.5281/zenodo.13163594" httpUri-0="https://zenodo.org/record/13163592/files/figure.png" httpUri-1="https://zenodo.org/record/13163594/files/figure.png" pageId="1" pageNumber="220">Figs 1–7</figureCitation>
.
</paragraph>
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<treatmentCitationGroup id="B7A211F6327BFFEBFF6BEC2DE4E5FDAA" box="[226,681,516,536]" pageId="1" pageNumber="220">
<treatmentCitation id="161310C9327BFFEBFF6BEC2DE405FDAA" author="Mikhaljova E. V." box="[226,585,516,536]" page="166" pageId="1" pageNumber="220" year="2000">
<taxonomicName id="50B24D5B327BFFEBFF6BEC2DE405FDAA" ID-CoL="4X495" authority="Mikhaljova, 2000: 166" authorityName="Mikhaljova" authorityPageNumber="166" authorityYear="2000" box="[226,585,516,536]" class="Diplopoda" family="Diplomaragnidae" genus="Shearia" kingdom="Animalia" order="Chordeumatida" pageId="1" pageNumber="220" phylum="Arthropoda" rank="species" species="calycina">
<emphasis id="A5C6EACA327BFFEBFF6BEC2DE73AFDAA" box="[226,374,516,535]" italics="true" pageId="1" pageNumber="220">Shearia calycina</emphasis>
<bibRefCitation id="F3234B29327BFFEBFEF4EC2DE405FDAA" author="Mikhaljova E. V." box="[381,585,516,536]" pageId="1" pageNumber="220" pagination="153 - 181" refId="ref3405" refString="Mikhaljova E. V. 2000. Review of the millipede family Diplomaragnidae (Diplopoda: Chordeumatida) // Arthropoda Selecta (for 1999). Vol. 8. No. 3. P. 153 - 181." type="book chapter" year="2000">Mikhaljova, 2000: 166</bibRefCitation>
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, 167: figs.
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<treatmentCitationGroup id="B7A211F6327BFFEBFF6BEC35E495FDE2" pageId="1" pageNumber="220">
<taxonomicName id="50B24D5B327BFFEBFF6BEC35E73EFD92" ID-CoL="4X495" authorityName="Mikhaljova" authorityYear="2000" box="[226,370,540,559]" class="Diplopoda" family="Diplomaragnidae" genus="Shearia" kingdom="Animalia" order="Chordeumatida" pageId="1" pageNumber="220" phylum="Arthropoda" rank="species" species="calycina">
<emphasis id="A5C6EACA327BFFEBFF6BEC35E73EFD92" box="[226,370,540,559]" italics="true" pageId="1" pageNumber="220">Shearia calycina</emphasis>
</taxonomicName>
—
<treatmentCitation id="161310C9327BFFEBFE19EC35E4F5FD8D" author="Mikhaljova E. V. &amp; Golovatch S. I." box="[400,697,540,560]" page="110" pageId="1" pageNumber="220" year="2001">
<bibRefCitation id="F3234B29327BFFEBFE19EC35E4F5FD8D" author="Mikhaljova E. V. &amp; Golovatch S. I." box="[400,697,540,560]" pageId="1" pageNumber="220" pagination="103 - 118" refId="ref3620" refString="Mikhaljova E. V., Golovatch S. I. 2001. A review of the millipede fauna of Siberia (Diplopoda) // Arthropoda Selecta. Vol. 9 (for 2000). No. 2. P. 103 - 118." type="book chapter" year="2001">Mikhaljova, Golovatch, 2001: 110</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="161310C9327BFFEBFD48EC35E707FDFA" author="Mikhaljova E. V." page="163" pageId="1" pageNumber="220" year="2004">
<bibRefCitation id="F3234B29327BFFEBFD48EC35E707FDFA" author="Mikhaljova E. V." pageId="1" pageNumber="220" refId="ref3445" refString="Mikhaljova E. V. 2004. The millipedes (Diplopoda) of the Asian part of Russia. Sofia-Moscow: Pensoft Publishers. Series Faunistica 39. 292 pp." type="book" year="2004">Mikhaljova, 2004: 163</bibRefCitation>
</treatmentCitation>
, figs, 164: map; 2017: 200, 201: figs, 134: map;
<treatmentCitation id="161310C9327BFFEBFF34EC65E794FDE2" author="Nefediev P. S. &amp; Nefedieva J. S." box="[189,472,588,608]" page="162" pageId="1" pageNumber="220" year="2007">
<bibRefCitation id="F3234B29327BFFEBFF34EC65E794FDE2" author="Nefediev P. S. &amp; Nefedieva J. S." box="[189,472,588,608]" pageId="1" pageNumber="220" pagination="159 - 164" refId="ref3844" refString="Nefediev P. S., Nefedieva J. S. 2007. [Biogeographical characteristic of the millipede fauna in the southeast of Western Siberia] // G. P. Ostroverkhova (ed.). Bioraznoobrazie bespozvonochnykh zhivotnykh. Sbornik materialov II Vserossiiskoi shkoly-seminara s mezhdunarodnym uchastiem. 24 - 26 October 2007, Tomsk. Tomsk: Deltaplan Publ. P. 159 - 164 [in Russian]." type="book chapter" year="2007">Nefediev, Nefedieva, 2007: 162</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="161310C9327BFFEBFE6BEC65E499FDE2" ID-DOI="http://doi.org/10.5281/zenodo.6227656" ID-Zenodo-Dep="6227656" author="Mikhaljova E. V. &amp; Nefediev P. S. &amp; Nefedieva J. S." box="[482,725,588,608]" httpUri="http://treatment.plazi.org/id/AA1987BCFF82194BFF2B29E4FEE0C8BE" page="53" pageId="1" pageNumber="220" year="2008">
<bibRefCitation id="F3234B29327BFFEBFE6BEC65E499FDE2" author="Mikhaljova E. V. &amp; Nefediev P. S. &amp; Nefedieva J. S." box="[482,725,588,608]" pageId="1" pageNumber="220" pagination="49 - 56" refId="ref3712" refString="Mikhaljova E. V., Nefediev P. S., Nefedieva J. S. 2008. A new species and new records of millipedes of the family Diplomaragnidae (Diplopoda, Chordeumatida) from Altai // Zootaxa. Vol. 1931. P. 49 - 56." type="book chapter" year="2008">Mikhaljova et al., 2008: 53</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
</subSubSection>
<subSubSection id="DFA86553327BFFEBFF6BEC4CE4F0FD6A" pageId="1" pageNumber="220" type="materials_examined">
<paragraph id="970D36D8327BFFEBFF6BEC4CE4F0FD6A" blockId="1.[189,776,376,1048]" pageId="1" pageNumber="220">
<materialsCitation id="27DA3C85327BFFEBFF6BEC4CE4F0FD6A" collectingDate="2018-07-29" collectionCode="MATERIAL, EXAMINED" collectorName="P. S. Nefediev." country="Russia" county="Republic" elevation="1355" latitude="49.88593" location="Karagem River" longLatPrecision="1" longitude="87.19024" municipality="Kosh-Agach District" pageId="1" pageNumber="220" specimenCount="14" specimenCount-female="1" specimenCount-male="5" stateProvince="Altay">
<collectionCode id="F1A3AE1D327BFFEBFF6BEC4CE786FDC5" box="[226,458,613,632]" pageId="1" pageNumber="220">MATERIAL EXAMINED</collectionCode>
.
<specimenCount id="81B4FD51327BFFEBFE58EC4DE444FDC5" box="[465,520,610,632]" count="5" pageId="1" pageNumber="220" type="male">5 ♂♂</specimenCount>
,
<specimenCount id="81B4FD51327BFFEBFD9BEC4CE461FDC5" box="[530,557,610,632]" count="1" pageId="1" pageNumber="220" type="female">1 ♀</specimenCount>
,
<specimenCount id="81B4FD51327BFFEBFDBEEC4CE426FDCA" box="[567,618,612,632]" count="8" pageId="1" pageNumber="220" type="generic">8 juv.</specimenCount>
,
<collectingCountry id="EFA57648327BFFEBFDF8EC4DE4E1FDCA" box="[625,685,612,631]" name="Russia" pageId="1" pageNumber="220">Russia</collectingCountry>
,
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<collectingCounty id="7E6C4E54327BFFEBFD3DEC4DE54AFDCA" box="[692,774,612,631]" pageId="1" pageNumber="220">Republic</collectingCounty>
of
<collectingRegion id="5576F83A327BFFEBFF5EEC55E749FD32" box="[215,261,636,655]" country="Russia" name="Altay" pageId="1" pageNumber="220">Altai</collectingRegion>
</emphasis>
,
<collectingMunicipality id="7769ACA2327BFFEBFE9BEC55E786FD32" box="[274,458,636,655]" pageId="1" pageNumber="220">Kosh-Agach District</collectingMunicipality>
, 26 air-km
<locationDeviation id="86BF2FFA327BFFEBFDBCEC54E49AFD32" box="[565,726,636,656]" location="Belyashi" pageId="1" pageNumber="220">NNW of Belyashi</locationDeviation>
(Dzhazator), floodplain of
<location id="926D6003327BFFEBFE07ECBCE454FD1A" LSID="urn:lsid:plazi:treatment:1F1B87CE327BFFEBFEAAEFEBE441FBAA:926D6003327BFFEBFE07ECBCE454FD1A" box="[398,536,660,680]" country="Russia" county="Republic" latitude="49.88593" longLatPrecision="1" longitude="87.19024" municipality="Kosh-Agach District" name="Karagem River" pageId="1" pageNumber="220" stateProvince="Altay">Karagem River</location>
, on left bank near bridge,
<geoCoordinate id="F286501F327BFFEBFF34EC85E764FD02" box="[189,296,684,703]" degrees="49.88593" direction="north" orientation="latitude" pageId="1" pageNumber="220" precision="1" value="49.88593">49.88593°N</geoCoordinate>
,
<geoCoordinate id="F286501F327BFFEBFEB8EC84E7D5FD7D" box="[305,409,685,704]" degrees="87.19024" direction="east" orientation="longitude" pageId="1" pageNumber="220" precision="1" value="87.19024">87.19024°E</geoCoordinate>
, valley
<taxonomicName id="50B24D5B327BFFEBFE55EC85E46EFD02" box="[476,546,684,703]" class="Magnoliopsida" family="Salicaceae" genus="Populus" kingdom="Plantae" order="Malpighiales" pageId="1" pageNumber="220" phylum="Tracheophyta" rank="genus">
<emphasis id="A5C6EACA327BFFEBFE55EC85E46EFD02" box="[476,546,684,703]" italics="true" pageId="1" pageNumber="220">Populus</emphasis>
</taxonomicName>
forest with
<taxonomicName id="50B24D5B327BFFEBFD02EC84E54DFD02" box="[651,769,684,703]" class="Pinopsida" family="Pinaceae" genus="Larix" kingdom="Plantae" order="Pinales" pageId="1" pageNumber="220" phylum="Tracheophyta" rank="species" species="sibirica">
<emphasis id="A5C6EACA327BFFEBFD02EC84E54DFD02" box="[651,769,684,703]" italics="true" pageId="1" pageNumber="220">Larix sibirica</emphasis>
</taxonomicName>
, ca
<elevation id="1C9FD1EB327BFFEBFF5FECEDE705FD6A" box="[214,329,708,728]" metricMagnitude="3" metricUnit="m" metricValue="1.355" pageId="1" pageNumber="220" unit="m" value="1355.0">
<quantity id="504A9B3D327BFFEBFF5FECEDE755FD65" box="[214,281,708,728]" metricMagnitude="3" metricUnit="m" metricValue="1.355" pageId="1" pageNumber="220" unit="m" value="1355.0">1355 m</quantity>
a.s.l.
</elevation>
, in litter,
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<collectingDate id="F348E9F0327BFFEBFE29ECECE440FD65" box="[416,524,709,728]" pageId="1" pageNumber="220" value="2018-07-29">29.VII.2018</collectingDate>
</date>
, leg.
<collectorName id="3A47530E327BFFEBFDB5ECECE4F0FD6A" box="[572,700,708,728]" pageId="1" pageNumber="220">P.S. Nefediev.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="DFA86553327BFFEBFF6BECF4E45EFCB1" pageId="1" pageNumber="220" type="distribution">
<paragraph id="970D36D8327BFFEBFF6BECF4E45EFCB1" blockId="1.[189,776,376,1048]" pageId="1" pageNumber="220">
DISTRIBUTION. This species inhabits SE districts of the Republic of
<collectingRegion id="5576F83A327BFFEBFED3ECDEE7C6FCB0" box="[346,394,759,781]" country="Russia" name="Altay" pageId="1" pageNumber="220">Altai</collectingRegion>
only (
<figureCitation id="0F892A5D327BFFEBFE43ECDEE44FFCB1" box="[458,515,759,781]" captionStart="Fig" captionStartId="1.[226,257,1969,1988]" captionTargetBox="[189,1399,1085,1940]" captionTargetId="figure-502@1.[189,1399,1085,1940]" captionTargetPageId="1" captionText="Fig. 1. Distribution of Shearia calycina (triangle) and S. densecava (circle). Previously known localities marked in black, new records given in white. Рис. 1. Распространение Shearia calycina (треугольник) и S. densecava (круг). Чёрным отмечены ранее иЗвестные места находок, новые находки отмечены белым." figureDoi="http://doi.org/10.5281/zenodo.13163592" httpUri="https://zenodo.org/record/13163592/files/figure.png" pageId="1" pageNumber="220">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="DFA86553327BFFEBFF6BED3BE441FBAA" pageId="1" pageNumber="220" type="discussion">
<paragraph id="970D36D8327BFFEBFF6BED3BE441FBAA" blockId="1.[189,776,376,1048]" pageId="1" pageNumber="220">
REMARKS. Endemic to the Russian Altais. Neither the gonopods nor coxae 10, nor coxae 11 of this species have ever been shown using SEM. Therefore, they are given here (
<figureCitation id="0F892A5D327BFFEBFF4AED4BE750FCCA" box="[195,284,866,888]" captionStart="Figs 2–7" captionStartId="2.[226,263,1947,1966]" captionTargetBox="[189,1399,568,1933]" captionTargetId="figure-126@2.[189,1399,568,1933]" captionTargetPageId="2" captionText="Figs 2–7. Shearia calycina, male (Republic of Altai, floodplain of Karagem River): 2–5 — gonopods; 6 — coxae 10; 7 — coxae 11; 2, 4, 6–7 — oral view; 3–5 — caudal view. Scale bars: 1 mm (2–3), 0.4 mm (4–5) and 0.3 mm (6–7). Designations explained in text. Рис. 2–7. Shearia calycina, самец (Республика Алтай, пойма р. Карагем): 2–5 — гоноподы; 6 — коксы 10-й пары ног; 7 — коксы 11-й пары ног; 2, 4, 6–7 — спереди; 3–5 — сЗади. МасШтаб: 1 мм (2–3), 0,4 мм (4–5) и 0,3 мм (6–7). ОбЪЯснение обоЗначений дано в тексте." figureDoi="http://doi.org/10.5281/zenodo.13163594" httpUri="https://zenodo.org/record/13163594/files/figure.png" pageId="1" pageNumber="220">Figs 2–7</figureCitation>
). An examination of males of the species from the ASU collection has revealed a small, two-branched anterior process (
<emphasis id="A5C6EACA327BFFEBFE9DEDBEE761FC11" bold="true" box="[276,301,919,940]" pageId="1" pageNumber="220">ap</emphasis>
) on each angiocoxite of the posterior gonopods. This character is well visible in a SEM micrograph (
<figureCitation id="0F892A5D327BFFEBFD37ED9BE4B6FC7A" box="[702,762,946,968]" captionStart="Figs 2–7" captionStartId="2.[226,263,1947,1966]" captionTargetBox="[189,1399,568,1933]" captionTargetId="figure-126@2.[189,1399,568,1933]" captionTargetPageId="2" captionText="Figs 2–7. Shearia calycina, male (Republic of Altai, floodplain of Karagem River): 2–5 — gonopods; 6 — coxae 10; 7 — coxae 11; 2, 4, 6–7 — oral view; 3–5 — caudal view. Scale bars: 1 mm (2–3), 0.4 mm (4–5) and 0.3 mm (6–7). Designations explained in text. Рис. 2–7. Shearia calycina, самец (Республика Алтай, пойма р. Карагем): 2–5 — гоноподы; 6 — коксы 10-й пары ног; 7 — коксы 11-й пары ног; 2, 4, 6–7 — спереди; 3–5 — сЗади. МасШтаб: 1 мм (2–3), 0,4 мм (4–5) и 0,3 мм (6–7). ОбЪЯснение обоЗначений дано в тексте." figureDoi="http://doi.org/10.5281/zenodo.13163594" httpUri="https://zenodo.org/record/13163594/files/figure.png" pageId="1" pageNumber="220">Fig. 4</figureCitation>
), but it was not specified in the original description [see
<bibRefCitation id="F3234B29327BFFEBFF34EDCEE73DFC41" author="Mikhaljova E. V." box="[189,369,999,1021]" pageId="1" pageNumber="220" pagination="153 - 181" refId="ref3405" refString="Mikhaljova E. V. 2000. Review of the millipede family Diplomaragnidae (Diplopoda: Chordeumatida) // Arthropoda Selecta (for 1999). Vol. 8. No. 3. P. 153 - 181." type="book chapter" year="2000">Mikhaljova, 2000</bibRefCitation>
]. Hence, this structure must be included into the description of
<taxonomicName id="50B24D5B327BFFEBFE15EA2BE444FBAA" authorityName="Mikhaljova" authorityYear="2000" box="[412,520,1026,1047]" class="Diplopoda" family="Diplomaragnidae" genus="Shearia" kingdom="Animalia" order="Chordeumatida" pageId="1" pageNumber="220" phylum="Arthropoda" rank="species" species="calycina">
<emphasis id="A5C6EACA327BFFEBFE15EA2BE7E4FBAA" box="[412,424,1026,1047]" italics="true" pageId="1" pageNumber="220">S</emphasis>
.
<emphasis id="A5C6EACA327BFFEBFE3FEA2BE444FBAA" box="[438,520,1026,1047]" italics="true" pageId="1" pageNumber="220">calycina</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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