315 lines
53 KiB
XML
315 lines
53 KiB
XML
<document id="06FD471CC96E0CBCFE378B31A208F800" ID="10.11646/zootaxa.4200.3.9" ID-DOI="10.11646/zootaxa.4200.3.9" ID-GBIF-Dataset="8bbe4b55-50d2-4612-9145-78fb5d105ced" ID-ISSN="1175-5326" ID-Zenodo-Dep="187215" ID-ZooBank="66416B99-751B-4B33-BF76-45AF4A26650B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1480493788134" checkinUser="plazi" docAuthor="Maddison, Wayne P." docDate="2016" docId="1D4987B6FFABFF91FF0DFA0DF08EF186" docLanguage="en" docName="zootaxa.4200.3.9.pdf" docOrigin="Zootaxa 4200 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Papuaneon tualapa Maddison, sp. nov." docType="treatment" docVersion="6" lastPageNumber="441" masterDocId="E170FFCEFFA8FF95FF9AFFDBF25CF63E" masterDocTitle="Papuaneon, a new genus of jumping spiders from Papua New Guinea (Araneae: Salticidae: Neonini)" masterLastPageNumber="443" masterPageNumber="437" pageNumber="440" updateTime="1698395201330" updateUser="plazi">
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<mods:title id="9D9F8684B18507C217D6DA674BBF4DF7">Papuaneon, a new genus of jumping spiders from Papua New Guinea (Araneae: Salticidae: Neonini)</mods:title>
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</mods:titleInfo>
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<mods:name id="BE5C303971197408F9056E54E42C11DC" type="personal">
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<mods:roleTerm id="F179316D4C46A6EFE40E1BF750C8BA8F">Author</mods:roleTerm>
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<mods:namePart id="793069D2596D286912DF3471BE417031">Maddison, Wayne P.</mods:namePart>
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<mods:typeOfResource id="8F93927B72C9E2CF027243D578C89829">text</mods:typeOfResource>
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<mods:title id="BB929C2D9709D04533785F75A0AAD083">Zootaxa</mods:title>
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<mods:part id="28095C0ED29326D16AB74AC3DB151FFB">
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<mods:date id="2755EF401F64B20C7F30EE6231CC985F">2016</mods:date>
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<mods:detail id="07BB201ECDC4DD8DA7363967FBD45262" type="volume">
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<mods:number id="99AB752DA67345E086912E5DA9895118">4200</mods:number>
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</mods:detail>
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<mods:detail id="3FDBA3DCA83DA81ADFEE62FE5265B9D9" type="issue">
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<mods:number id="B1F6D9574C77FA0F60A01C1764E11BC1">3</mods:number>
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<mods:classification id="9F6B23A37649615A8DA8AB4068C3A304">journal article</mods:classification>
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<mods:identifier id="25DE577A53EFAD39436EE972F7FBDF99" type="DOI">10.11646/zootaxa.4200.3.9</mods:identifier>
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<mods:identifier id="21E0662A8DD97A315C58636D83ADD9D2" type="GBIF-Dataset">8bbe4b55-50d2-4612-9145-78fb5d105ced</mods:identifier>
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<mods:identifier id="B3825E322FE4CC99D09F52432ACFF25F" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="519CDD5B191786028DF9778A8ECBF47F" type="Zenodo-Dep">187215</mods:identifier>
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<treatment id="1D4987B6FFABFF91FF0DFA0DF08EF186" ID-DOI="http://doi.org/10.5281/zenodo.6075363" ID-GBIF-Taxon="124873361" ID-Zenodo-Dep="6075363" LSID="urn:lsid:plazi:treatment:1D4987B6FFABFF91FF0DFA0DF08EF186" httpUri="http://treatment.plazi.org/id/1D4987B6FFABFF91FF0DFA0DF08EF186" lastPageId="4" lastPageNumber="441" pageId="3" pageNumber="440">
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<subSubSection id="DDFA652BFFABFF96FF0DFA0DF35AF02C" pageId="3" pageNumber="440" type="nomenclature">
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<paragraph id="955F36A0FFABFF96FF0DFA0DF029F3CF" blockId="3.[151,629,1494,1554]" box="[151,629,1494,1521]" pageId="3" pageNumber="440">
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<heading id="CE1781CCFFABFF96FF0DFA0DF029F3CF" bold="true" box="[151,629,1494,1521]" fontSize="11" level="1" pageId="3" pageNumber="440" reason="1">
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<emphasis id="A794EAB2FFABFF96FF0DFA0DF029F3CF" bold="true" box="[151,629,1494,1521]" pageId="3" pageNumber="440">
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<taxonomicName id="52E04D23FFABFF96FF0DFA0DF057F3CF" ID-CoL="75NPB" authority="Maddison" authorityName="Maddison" box="[151,523,1494,1521]" class="Arachnida" family="Salticidae" genus="Papuaneon" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="440" phylum="Arthropoda" rank="species" species="tualapa" status="sp. nov.">
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<emphasis id="A794EAB2FFABFF96FF0DFA0DF3DFF3CE" bold="true" box="[151,387,1494,1520]" italics="true" pageId="3" pageNumber="440">Papuaneon tualapa</emphasis>
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Maddison
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</taxonomicName>
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,
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<taxonomicNameLabel id="BCA757C9FFABFF96FD8DFA0CF029F3CF" box="[535,629,1495,1521]" pageId="3" pageNumber="440" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="955F36A0FFABFF96FF0DFA22F35AF02C" blockId="3.[151,629,1494,1554]" box="[151,262,1529,1554]" pageId="3" pageNumber="440">
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<figureCitation id="0DDB2A25FFABFF96FF0DFA22F35AF02C" box="[151,262,1529,1554]" captionStart-0="FIGURES 1 – 10" captionStart-1="FIGURES 11 – 18" captionStartId-0="2.[151,264,1787,1810]" captionStartId-1="3.[151,264,1274,1297]" captionTargetBox-0="[151,1436,419,1763]" captionTargetBox-1="[151,1436,193,1253]" captionTargetId-0="figure@2.[151,1436,411,1763]" captionTargetId-1="figure@3.[151,1436,193,1253]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURES 1 – 10. Papuaneon tualapa sp. nov., male holotype (specimen 2008 PNG- 1342). 1 dorsal view of carapace; 2 dorsal view of abdomen; 3 face; 4 left palp cleared in clove oil, oblique view that is mostly prolateral, slightly ventral (e, embolus; be, base of embolus); 5 left palp, ventral view (scale bar 0.1 mm); 6 left palp, retrolateral view; 7 – 10 living holotype male. Figs 1 – 10 are copyright © 2016 W. P. Maddison, released under a Creative Commons Attribution (CC-BY) 3.0 license." captionText-1="FIGURES 11 – 18. Papuaneon tualapa sp. nov., female paratype (specimen 2008 PNG- 0845). 11 female paratype, dorsal view of carapace and abdomen; 12 lateral view; 13 oblique view of face and first leg; 14 retrolateral view of first leg femur (arrow highlights row of macrosetae); 15 – 16 living female; 17 ventral view of epigynum (scale bar 0.1 mm); 18 dorsal view of epigynum, cleared with clove oil. Figs 11 – 18 are copyright © 2016 W. P. Maddison, released under a Creative Commons Attribution (CC-BY) 3.0 license." httpUri-0="https://zenodo.org/record/187216/files/figure.png" httpUri-1="https://zenodo.org/record/187217/files/figure.png" pageId="3" pageNumber="440">Figs 1‒18</figureCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="DDFA652BFFABFF96FF0DF99AF6E1F0F8" pageId="3" pageNumber="440" type="materials_examined">
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<paragraph id="955F36A0FFABFF96FF0DF99AF6E1F0F8" blockId="3.[151,1437,1601,2022]" pageId="3" pageNumber="440">
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<materialsCitation id="25883CFDFFABFF96FF0DF99AF1B7F09C" ID-GBIF-Occurrence="1412767717" collectingDate="2008-07-11" collectingDateMax="2008-07-22" collectingDateMin="2008-07-11" collectionCode="DNA" collectorName="W. Maddison & Specimen" country="Papua New Guinea" elevation="1050" latitude="-5.283" location="Wanakipa" longLatPrecision="78" longitude="142.498" pageId="3" pageNumber="440" specimenCount="2" stateProvince="Southern Highlands Province" typeStatus="holotype">
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<emphasis id="A794EAB2FFABFF96FF0DF99AF39FF064" bold="true" box="[151,451,1601,1626]" pageId="3" pageNumber="440">
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<typeStatus id="4A5B8802FFABFF96FF0DF99AF28EF064" box="[151,210,1601,1626]" pageId="3" pageNumber="440">Type</typeStatus>
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material.
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<typeStatus id="4A5B8802FFABFF96FED7F99AF3E2F064" box="[333,446,1601,1626]" pageId="3" pageNumber="440" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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male from
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<taxonomicName id="52E04D23FFABFF96FDD2F99AF0F8F064" box="[584,676,1601,1626]" class="Arachnida" family="Salticidae" genus="Papuaneon" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="441" phylum="Arthropoda" rank="species" species="tualapa">Tualapa</taxonomicName>
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, near
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<location id="903F607BFFABFF96FD73F99AF107F064" LSID="urn:lsid:plazi:treatment:1D4987B6FFABFF91FF0DFA0DF08EF186:903F607BFFABFF96FD73F99AF107F064" box="[745,859,1601,1626]" country="Papua New Guinea" latitude="-5.283" longLatPrecision="78" longitude="142.498" name="Wanakipa" pageId="3" pageNumber="440" stateProvince="Southern Highlands Province">Wanakipa</location>
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,
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<geoCoordinate id="F0D45067FFABFF96FCFDF999F1E2F064" box="[871,958,1601,1626]" direction="south" orientation="latitude" pageId="3" pageNumber="440" precision="55" value="-5.283">S 5.283</geoCoordinate>
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<geoCoordinate id="F0D45067FFABFF96FC5CF999F661F064" box="[966,1085,1602,1626]" direction="east" orientation="longitude" pageId="3" pageNumber="440" precision="55" value="142.498">E 142.498</geoCoordinate>
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,
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<collectingRegion id="5724F842FFABFF96FBD3F99AF7C4F064" box="[1097,1432,1601,1626]" country="Papua New Guinea" name="Southern Highlands" pageId="3" pageNumber="440">
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Southern
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<collectingRegion id="5724F842FFABFF96FB2CF99AF7C4F064" box="[1206,1432,1601,1626]" inferredBySuffix="true" pageId="3" pageNumber="440">Highlands Province</collectingRegion>
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</collectingRegion>
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,
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<collectingCountry id="EDF77630FFABFF96FF0DF9BCF328F040" box="[151,372,1637,1662]" name="Papua New Guinea" pageId="3" pageNumber="440">Papua New Guinea</collectingCountry>
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, elev.
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<elevation id="1ECDD193FFABFF96FE21F9BDF0CFF040" box="[443,659,1637,1662]" metricMagnitude="3" metricUnit="m" metricValue="1.05" metricValueMax="1.1" metricValueMin="1.0" pageId="3" pageNumber="440" unit="m" value="1050.0" valueMax="1100.0" valueMin="1000.0">
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<quantity id="52189B45FFABFF96FE21F9BDF00AF040" box="[443,598,1638,1662]" metricMagnitude="3" metricUnit="m" metricValue="1.05" metricValueMax="1.1" metricValueMin="1.0" pageId="3" pageNumber="440" unit="m" value="1050.0" valueMax="1100.0" valueMin="1000.0">1000‒1100 m</quantity>
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a.s.l.
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</elevation>
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,
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<date id="E15E1060FFABFF96FD3AF9BDF102F040" box="[672,862,1637,1662]" pageId="3" pageNumber="440" value="2008-07-11" valueMax="2008-07-22" valueMin="2008-07-11">
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<collectingDate id="F11AE988FFABFF96FD3AF9BDF102F040" box="[672,862,1637,1662]" pageId="3" pageNumber="440" value="2008-07-11" valueMax="2008-07-22" valueMin="2008-07-11">11‒22 July 2008</collectingDate>
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</date>
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, forest interior and river side,
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<collectorName id="38155376FFABFF96FB2FF9BCF717F040" box="[1205,1355,1637,1662]" pageId="3" pageNumber="440">W. Maddison</collectorName>
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& Luc Fimo Tuki, WPM#08-008,
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<collectorName id="38155376FFABFF96FE52F952F06AF09C" box="[456,566,1673,1698]" pageId="3" pageNumber="440">Specimen</collectorName>
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#2008PNG-1342;
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||
<collectionCode id="F3F1AE65FFABFF96FC91F951F114F09C" box="[779,840,1674,1698]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="3" pageNumber="440">DNA</collectionCode>
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voucher d302
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</materialsCitation>
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||
.
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||
<materialsCitation id="25883CFDFFABFF96FC6EF952F6E1F0F8" ID-GBIF-Occurrence="1412767716" collectingDate="2008-07-11" collectingDateMax="2008-07-22" collectingDateMin="2008-07-11" collectionCode="DNA" collectorName="W. Maddison & Specimen" country="Papua New Guinea" elevation="1050" latitude="-5.283" location="Wanakipa" longLatPrecision="78" longitude="142.498" pageId="3" pageNumber="440" specimenCount="1" stateProvince="Southern Highlands Province" typeStatus="paratype">
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||
<emphasis id="A794EAB2FFABFF96FC6EF952F637F09C" bold="true" box="[1012,1131,1673,1698]" pageId="3" pageNumber="440">
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||
<typeStatus id="4A5B8802FFABFF96FC6EF952F639F09C" box="[1012,1125,1673,1698]" pageId="3" pageNumber="440" type="paratype">Paratype</typeStatus>
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||
:
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</emphasis>
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female, same data as male, Specimen #2008PNG-0845;
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<collectionCode id="F3F1AE65FFABFF96FE40F975F04BF0F8" box="[474,535,1710,1734]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="3" pageNumber="440">DNA</collectionCode>
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voucher JXZ267. These are the only two specimens known.
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</materialsCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="DDFA652BFFABFF96FF5DF90AF65BF0D4" box="[199,1031,1745,1770]" pageId="3" pageNumber="440" type="etymology">
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<paragraph id="955F36A0FFABFF96FF5DF90AF65BF0D4" blockId="3.[151,1437,1601,2022]" box="[199,1031,1745,1770]" pageId="3" pageNumber="440">
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<emphasis id="A794EAB2FFABFF96FF5DF90AF312F0D4" bold="true" box="[199,334,1745,1770]" pageId="3" pageNumber="440">Etymology.</emphasis>
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The name of the
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<typeStatus id="4A5B8802FFABFF96FD88F909F01EF0D4" box="[530,578,1746,1770]" pageId="3" pageNumber="440">type</typeStatus>
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locality, treated as a noun in apposition.
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</paragraph>
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</subSubSection>
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<subSubSection id="DDFA652BFFABFF96FF5DF92EF289F1A0" pageId="3" pageNumber="440" type="diagnosis">
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<paragraph id="955F36A0FFABFF96FF5DF92EF289F1A0" blockId="3.[151,1437,1601,2022]" pageId="3" pageNumber="440">
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<emphasis id="A794EAB2FFABFF96FF5DF92EF31DF130" bold="true" box="[199,321,1781,1806]" pageId="3" pageNumber="440">Diagnosis.</emphasis>
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The small broad hirsute body with long macrosetae under the first tibia (almost as long as the tibia), along with the cream clypeus of males and white palps of females, are distinctive among known New
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<collectingCountry id="EDF77630FFABFF96FAD1F8C2F7C0F10C" box="[1355,1436,1817,1842]" name="Guinea-Bissau" pageId="3" pageNumber="440">Guinea</collectingCountry>
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salticids. The embolus resembles that of euophryines in the
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<emphasis id="A794EAB2FFABFF96FCC4F8E5F600F168" box="[862,1116,1854,1878]" italics="true" pageId="3" pageNumber="440">
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||
<taxonomicName id="52E04D23FFABFF96FCC4F8E5F1ECF168" box="[862,944,1854,1878]" class="Arachnida" family="Salticidae" genus="Bulolia" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="440" phylum="Arthropoda" rank="genus">Bulolia</taxonomicName>
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-
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<taxonomicName id="52E04D23FFABFF96FC20F8E5F600F168" box="[954,1116,1854,1878]" class="Arachnida" family="Salticidae" genus="Coccorchestes" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="440" phylum="Arthropoda" rank="genus">Coccorchestes</taxonomicName>
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</emphasis>
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clade (
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||
<bibRefCitation id="F1714B51FFABFF96FB2FF8E6F28FF144" author="Zhang" pageId="3" pageNumber="440" refString="Zhang, J. X. & Maddison, W. P. (2015) Genera of euophryine jumping spiders (Araneae: Salticidae), with a combined molecular-morphological phylogeny. Zootaxa, 3938 (1), 1 - 147." type="journal article" year="2015">Zhang & Maddison 2015</bibRefCitation>
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), with a small spiral whose axis is parallel to the axis of the palp, but none of those species have this body form.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="DDFA652BFFABFF91FF5DF872F08EF186" lastPageId="4" lastPageNumber="441" pageId="3" pageNumber="440" type="description">
|
||
<paragraph id="955F36A0FFABFF91FF5DF872F712F4F3" blockId="3.[151,1437,1601,2022]" lastBlockId="4.[151,1437,151,1977]" lastPageId="4" lastPageNumber="441" pageId="3" pageNumber="440">
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<emphasis id="A794EAB2FFABFF96FF5DF872F3FEF1FC" bold="true" box="[199,418,1961,1986]" pageId="3" pageNumber="440">Description. Male</emphasis>
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||
(
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<typeStatus id="4A5B8802FFABFF96FE2DF872F043F1FC" box="[439,543,1961,1986]" pageId="3" pageNumber="440" type="holotype">holotype</typeStatus>
|
||
),
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||
<figureCitation id="0DDB2A25FFABFF96FDA2F872F0EDF1FC" box="[568,689,1961,1986]" captionStart="FIGURES 1 – 10" captionStartId="2.[151,264,1787,1810]" captionTargetBox="[151,1436,419,1763]" captionTargetId="figure@2.[151,1436,411,1763]" captionTargetPageId="2" captionText="FIGURES 1 – 10. Papuaneon tualapa sp. nov., male holotype (specimen 2008 PNG- 1342). 1 dorsal view of carapace; 2 dorsal view of abdomen; 3 face; 4 left palp cleared in clove oil, oblique view that is mostly prolateral, slightly ventral (e, embolus; be, base of embolus); 5 left palp, ventral view (scale bar 0.1 mm); 6 left palp, retrolateral view; 7 – 10 living holotype male. Figs 1 – 10 are copyright © 2016 W. P. Maddison, released under a Creative Commons Attribution (CC-BY) 3.0 license." httpUri="https://zenodo.org/record/187216/files/figure.png" pageId="3" pageNumber="440">Figs 1‒10</figureCitation>
|
||
. Carapace length 1.5; abdomen length 1.5. Chelicera (
|
||
<figureCitation id="0DDB2A25FFABFF96FAD8F872F7D1F1FC" box="[1346,1421,1961,1986]" captionStart="FIGURES 1 – 10" captionStartId="2.[151,264,1787,1810]" captionTargetBox="[151,1436,419,1763]" captionTargetId="figure@2.[151,1436,411,1763]" captionTargetPageId="2" captionText="FIGURES 1 – 10. Papuaneon tualapa sp. nov., male holotype (specimen 2008 PNG- 1342). 1 dorsal view of carapace; 2 dorsal view of abdomen; 3 face; 4 left palp cleared in clove oil, oblique view that is mostly prolateral, slightly ventral (e, embolus; be, base of embolus); 5 left palp, ventral view (scale bar 0.1 mm); 6 left palp, retrolateral view; 7 – 10 living holotype male. Figs 1 – 10 are copyright © 2016 W. P. Maddison, released under a Creative Commons Attribution (CC-BY) 3.0 license." httpUri="https://zenodo.org/record/187216/files/figure.png" pageId="3" pageNumber="440">Fig. 3</figureCitation>
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||
): vertical, slightly swollen distally. Promargin with 2 teeth; retromargin with 1 simple tooth. Palp (
|
||
<figureCitation id="0DDB2A25FFABFF96FA99F816F732F1D8" box="[1283,1390,1997,2022]" captionStart="FIGURES 1 – 10" captionStartId="2.[151,264,1787,1810]" captionTargetBox="[151,1436,419,1763]" captionTargetId="figure@2.[151,1436,411,1763]" captionTargetPageId="2" captionText="FIGURES 1 – 10. Papuaneon tualapa sp. nov., male holotype (specimen 2008 PNG- 1342). 1 dorsal view of carapace; 2 dorsal view of abdomen; 3 face; 4 left palp cleared in clove oil, oblique view that is mostly prolateral, slightly ventral (e, embolus; be, base of embolus); 5 left palp, ventral view (scale bar 0.1 mm); 6 left palp, retrolateral view; 7 – 10 living holotype male. Figs 1 – 10 are copyright © 2016 W. P. Maddison, released under a Creative Commons Attribution (CC-BY) 3.0 license." httpUri="https://zenodo.org/record/187216/files/figure.png" pageId="3" pageNumber="440">Figs 4‒6</figureCitation>
|
||
): of standard oval form with the embolus appearing to arise distally at 11‒12:00 in left palp (as if on a clock face). Terminal part of embolus a gentle spiral, though the larger base of the embolus extends fairly deep into the bulb (
|
||
<figureCitation id="0DDB2A25FFACFF91FF05FF04F2B9F6C6" box="[159,229,223,248]" captionStart="FIGURES 1 – 10" captionStartId="2.[151,264,1787,1810]" captionTargetBox="[151,1436,419,1763]" captionTargetId="figure@2.[151,1436,411,1763]" captionTargetPageId="2" captionText="FIGURES 1 – 10. Papuaneon tualapa sp. nov., male holotype (specimen 2008 PNG- 1342). 1 dorsal view of carapace; 2 dorsal view of abdomen; 3 face; 4 left palp cleared in clove oil, oblique view that is mostly prolateral, slightly ventral (e, embolus; be, base of embolus); 5 left palp, ventral view (scale bar 0.1 mm); 6 left palp, retrolateral view; 7 – 10 living holotype male. Figs 1 – 10 are copyright © 2016 W. P. Maddison, released under a Creative Commons Attribution (CC-BY) 3.0 license." httpUri="https://zenodo.org/record/187216/files/figure.png" pageId="4" pageNumber="441">Fig. 4</figureCitation>
|
||
). Tegular ledge cuts diagonally in front of shoulder of the tegulum (
|
||
<figureCitation id="0DDB2A25FFACFF91FC47FF04F61FF6C6" box="[989,1091,223,248]" captionStart="FIGURES 1 – 10" captionStartId="2.[151,264,1787,1810]" captionTargetBox="[151,1436,419,1763]" captionTargetId="figure@2.[151,1436,411,1763]" captionTargetPageId="2" captionText="FIGURES 1 – 10. Papuaneon tualapa sp. nov., male holotype (specimen 2008 PNG- 1342). 1 dorsal view of carapace; 2 dorsal view of abdomen; 3 face; 4 left palp cleared in clove oil, oblique view that is mostly prolateral, slightly ventral (e, embolus; be, base of embolus); 5 left palp, ventral view (scale bar 0.1 mm); 6 left palp, retrolateral view; 7 – 10 living holotype male. Figs 1 – 10 are copyright © 2016 W. P. Maddison, released under a Creative Commons Attribution (CC-BY) 3.0 license." httpUri="https://zenodo.org/record/187216/files/figure.png" pageId="4" pageNumber="441">Figs 5‒6</figureCitation>
|
||
). Even in the uncleared palp, a loop of the narrow part of the spermophore is visible through the prolateral proximal face of the tegulum (
|
||
<figureCitation id="0DDB2A25FFACFF91FAD2FEDFF7D1F723" box="[1352,1421,260,285]" captionStart="FIGURES 1 – 10" captionStartId="2.[151,264,1787,1810]" captionTargetBox="[151,1436,419,1763]" captionTargetId="figure@2.[151,1436,411,1763]" captionTargetPageId="2" captionText="FIGURES 1 – 10. Papuaneon tualapa sp. nov., male holotype (specimen 2008 PNG- 1342). 1 dorsal view of carapace; 2 dorsal view of abdomen; 3 face; 4 left palp cleared in clove oil, oblique view that is mostly prolateral, slightly ventral (e, embolus; be, base of embolus); 5 left palp, ventral view (scale bar 0.1 mm); 6 left palp, retrolateral view; 7 – 10 living holotype male. Figs 1 – 10 are copyright © 2016 W. P. Maddison, released under a Creative Commons Attribution (CC-BY) 3.0 license." httpUri="https://zenodo.org/record/187216/files/figure.png" pageId="4" pageNumber="441">Fig. 5</figureCitation>
|
||
), similar to that seen in
|
||
<taxonomicName id="52E04D23FFACFF91FE0BFEF2F391F77E" box="[401,461,297,320]" class="Arachnida" family="Salticidae" genus="Neon" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="441" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A794EAB2FFACFF91FE0BFEF2F391F77E" box="[401,461,297,320]" italics="true" pageId="4" pageNumber="441">Neon</emphasis>
|
||
</taxonomicName>
|
||
(e.g.,
|
||
<bibRefCitation id="F1714B51FFACFF91FD8EFEFCF084F77E" author="Lohmander" box="[532,728,295,320]" pageId="4" pageNumber="441" refString="Lohmander, H. (1945) Arachnologische Fragmente 3. Die Salticiden-Gattung Neon Simon in Sudschweden. Goteborgs Kungliga Uetenskaps och Uitterhets Samhalles Handlingar (6), 3 B (9), 31 - 75." type="journal article" year="1945">Lohmander 1945</bibRefCitation>
|
||
: fig. 43;
|
||
<bibRefCitation id="F1714B51FFACFF91FCA6FEFCF19AF77E" author="Zabka" box="[828,966,295,320]" pageId="4" pageNumber="441" refString="Zabka, M. (1997) Salticidae: Pajaki skaczace (Arachnida: Araneae). Fauna Polski, 19, 1 - 188." type="journal article" year="1997">Żabka 1997</bibRefCitation>
|
||
: figs 228, 237;
|
||
<bibRefCitation id="F1714B51FFACFF91FBEEFEFCF76BF77E" author="Richardson" box="[1140,1335,295,320]" pageId="4" pageNumber="441" refString="Richardson, B. J. (2013) New unidentate jumping spider genera (Araneae: Salticidae) from Australia. Zootaxa, 3716 (3), 460 - 474. http: // dx. doi. org / 10.11646 / zootaxa. 3716.3.8" type="journal article" year="2013">Richardson 2013</bibRefCitation>
|
||
: fig. 36). Legs: Relative lengths 1>4>3>2. As in
|
||
<taxonomicName id="52E04D23FFACFF91FDC8FE96F0D1F75A" box="[594,653,333,356]" class="Arachnida" family="Salticidae" genus="Neon" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="441" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A794EAB2FFACFF91FDC8FE96F0D1F75A" box="[594,653,333,356]" italics="true" pageId="4" pageNumber="441">Neon</emphasis>
|
||
</taxonomicName>
|
||
, first leg tibia with three pair of long ventral macrosetae, all of which terminate approximately at the end of the tibia. Although the femur apparently lacks the distinct row of macrosetae that the female has, there are one or two ventral prolateral and retrolateral macrosetae. Carapace (
|
||
<figureCitation id="0DDB2A25FFACFF91FB7DFE4FF711F792" box="[1255,1357,404,429]" captionStart="FIGURES 1 – 10" captionStartId="2.[151,264,1787,1810]" captionTargetBox="[151,1436,419,1763]" captionTargetId="figure@2.[151,1436,411,1763]" captionTargetPageId="2" captionText="FIGURES 1 – 10. Papuaneon tualapa sp. nov., male holotype (specimen 2008 PNG- 1342). 1 dorsal view of carapace; 2 dorsal view of abdomen; 3 face; 4 left palp cleared in clove oil, oblique view that is mostly prolateral, slightly ventral (e, embolus; be, base of embolus); 5 left palp, ventral view (scale bar 0.1 mm); 6 left palp, retrolateral view; 7 – 10 living holotype male. Figs 1 – 10 are copyright © 2016 W. P. Maddison, released under a Creative Commons Attribution (CC-BY) 3.0 license." httpUri="https://zenodo.org/record/187216/files/figure.png" pageId="4" pageNumber="441">Figs 1, 3</figureCitation>
|
||
): High and short, with anterior eye row wider than posterior. Abdomen (
|
||
<figureCitation id="0DDB2A25FFACFF91FCF0FE6CF1F1F7EE" box="[874,941,439,464]" captionStart="FIGURES 1 – 10" captionStartId="2.[151,264,1787,1810]" captionTargetBox="[151,1436,419,1763]" captionTargetId="figure@2.[151,1436,411,1763]" captionTargetPageId="2" captionText="FIGURES 1 – 10. Papuaneon tualapa sp. nov., male holotype (specimen 2008 PNG- 1342). 1 dorsal view of carapace; 2 dorsal view of abdomen; 3 face; 4 left palp cleared in clove oil, oblique view that is mostly prolateral, slightly ventral (e, embolus; be, base of embolus); 5 left palp, ventral view (scale bar 0.1 mm); 6 left palp, retrolateral view; 7 – 10 living holotype male. Figs 1 – 10 are copyright © 2016 W. P. Maddison, released under a Creative Commons Attribution (CC-BY) 3.0 license." httpUri="https://zenodo.org/record/187216/files/figure.png" pageId="4" pageNumber="441">Fig. 2</figureCitation>
|
||
) cordate, with long anterior-projecting setae along the anterior edge. Colour in alcohol (
|
||
<figureCitation id="0DDB2A25FFACFF91FDE4FE07F0B9F7CA" box="[638,741,476,501]" captionStart="FIGURES 1 – 10" captionStartId="2.[151,264,1787,1810]" captionTargetBox="[151,1436,419,1763]" captionTargetId="figure@2.[151,1436,411,1763]" captionTargetPageId="2" captionText="FIGURES 1 – 10. Papuaneon tualapa sp. nov., male holotype (specimen 2008 PNG- 1342). 1 dorsal view of carapace; 2 dorsal view of abdomen; 3 face; 4 left palp cleared in clove oil, oblique view that is mostly prolateral, slightly ventral (e, embolus; be, base of embolus); 5 left palp, ventral view (scale bar 0.1 mm); 6 left palp, retrolateral view; 7 – 10 living holotype male. Figs 1 – 10 are copyright © 2016 W. P. Maddison, released under a Creative Commons Attribution (CC-BY) 3.0 license." httpUri="https://zenodo.org/record/187216/files/figure.png" pageId="4" pageNumber="441">Figs 1‒3</figureCitation>
|
||
): Chelicera light brown. Palps black. All legs with femur and ventral surface of tibia dark brown to black, and dark annuli distally on patella, tibia and metatarsus. In addition, the first leg has a dark anteriorlateral surface and a ventral fringe of dark setae about as long as the tibia is deep. Clypeus covered with cream-coloured scales (
|
||
<figureCitation id="0DDB2A25FFACFF91FD06FD9CF0BDF45E" box="[668,737,583,608]" captionStart="FIGURES 1 – 10" captionStartId="2.[151,264,1787,1810]" captionTargetBox="[151,1436,419,1763]" captionTargetId="figure@2.[151,1436,411,1763]" captionTargetPageId="2" captionText="FIGURES 1 – 10. Papuaneon tualapa sp. nov., male holotype (specimen 2008 PNG- 1342). 1 dorsal view of carapace; 2 dorsal view of abdomen; 3 face; 4 left palp cleared in clove oil, oblique view that is mostly prolateral, slightly ventral (e, embolus; be, base of embolus); 5 left palp, ventral view (scale bar 0.1 mm); 6 left palp, retrolateral view; 7 – 10 living holotype male. Figs 1 – 10 are copyright © 2016 W. P. Maddison, released under a Creative Commons Attribution (CC-BY) 3.0 license." httpUri="https://zenodo.org/record/187216/files/figure.png" pageId="4" pageNumber="441">Fig. 3</figureCitation>
|
||
). Ocular and thoracic regions medium to dark brown, covered sparsely with some black, cream and brown scales. Abdomen grey-brown with indistinct wavy transverse bands. Colour in life (
|
||
<figureCitation id="0DDB2A25FFACFF91FED5FD54F397F496" box="[335,459,655,680]" captionStart="FIGURES 1 – 10" captionStartId="2.[151,264,1787,1810]" captionTargetBox="[151,1436,419,1763]" captionTargetId="figure@2.[151,1436,411,1763]" captionTargetPageId="2" captionText="FIGURES 1 – 10. Papuaneon tualapa sp. nov., male holotype (specimen 2008 PNG- 1342). 1 dorsal view of carapace; 2 dorsal view of abdomen; 3 face; 4 left palp cleared in clove oil, oblique view that is mostly prolateral, slightly ventral (e, embolus; be, base of embolus); 5 left palp, ventral view (scale bar 0.1 mm); 6 left palp, retrolateral view; 7 – 10 living holotype male. Figs 1 – 10 are copyright © 2016 W. P. Maddison, released under a Creative Commons Attribution (CC-BY) 3.0 license." httpUri="https://zenodo.org/record/187216/files/figure.png" pageId="4" pageNumber="441">Figs 7‒10</figureCitation>
|
||
): Except for the much brighter cream-coloured clypeus, the rest of the body and appendages are black and dark brown with scattered brown to tan scales, aligned in vague transverse bands.
|
||
</paragraph>
|
||
<paragraph id="955F36A0FFACFF91FF5DFD0CF0CEF243" blockId="4.[151,1437,151,1977]" pageId="4" pageNumber="441">
|
||
<emphasis id="A794EAB2FFACFF91FF5DFD0CF342F4CE" bold="true" box="[199,286,727,752]" pageId="4" pageNumber="441">Female</emphasis>
|
||
(
|
||
<typeStatus id="4A5B8802FFACFF91FEB4FD03F3C8F4CE" box="[302,404,728,752]" pageId="4" pageNumber="441" type="paratype">paratype</typeStatus>
|
||
),
|
||
<figureCitation id="0DDB2A25FFACFF91FE32FD0CF075F4CE" box="[424,553,727,752]" captionStart="FIGURES 11 – 18" captionStartId="3.[151,264,1274,1297]" captionTargetBox="[151,1436,193,1253]" captionTargetId="figure@3.[151,1436,193,1253]" captionTargetPageId="3" captionText="FIGURES 11 – 18. Papuaneon tualapa sp. nov., female paratype (specimen 2008 PNG- 0845). 11 female paratype, dorsal view of carapace and abdomen; 12 lateral view; 13 oblique view of face and first leg; 14 retrolateral view of first leg femur (arrow highlights row of macrosetae); 15 – 16 living female; 17 ventral view of epigynum (scale bar 0.1 mm); 18 dorsal view of epigynum, cleared with clove oil. Figs 11 – 18 are copyright © 2016 W. P. Maddison, released under a Creative Commons Attribution (CC-BY) 3.0 license." httpUri="https://zenodo.org/record/187217/files/figure.png" pageId="4" pageNumber="441">Figs 11‒18</figureCitation>
|
||
. Carapace length 1.3; abdomen length 1.4. Chelicera: Promargin with 2 teeth; retromargin with 1 simple tooth. Palp: tarsus swollen, as if a subadult male. Legs: Only a single leg remains on the specimen, the left first leg. Spination on first leg as in male (
|
||
<figureCitation id="0DDB2A25FFACFF91FCFBFCC4F1EAF506" box="[865,950,799,824]" captionStart="FIGURES 11 – 18" captionStartId="3.[151,264,1274,1297]" captionTargetBox="[151,1436,193,1253]" captionTargetId="figure@3.[151,1436,193,1253]" captionTargetPageId="3" captionText="FIGURES 11 – 18. Papuaneon tualapa sp. nov., female paratype (specimen 2008 PNG- 0845). 11 female paratype, dorsal view of carapace and abdomen; 12 lateral view; 13 oblique view of face and first leg; 14 retrolateral view of first leg femur (arrow highlights row of macrosetae); 15 – 16 living female; 17 ventral view of epigynum (scale bar 0.1 mm); 18 dorsal view of epigynum, cleared with clove oil. Figs 11 – 18 are copyright © 2016 W. P. Maddison, released under a Creative Commons Attribution (CC-BY) 3.0 license." httpUri="https://zenodo.org/record/187217/files/figure.png" pageId="4" pageNumber="441">Fig. 15</figureCitation>
|
||
), except that in addition there is a line of macrosetae on the posterior ventral edge of the first leg femur (
|
||
<figureCitation id="0DDB2A25FFACFF91FC14FC9FF1BAF562" box="[910,998,836,861]" captionStart="FIGURES 11 – 18" captionStartId="3.[151,264,1274,1297]" captionTargetBox="[151,1436,193,1253]" captionTargetId="figure@3.[151,1436,193,1253]" captionTargetPageId="3" captionText="FIGURES 11 – 18. Papuaneon tualapa sp. nov., female paratype (specimen 2008 PNG- 0845). 11 female paratype, dorsal view of carapace and abdomen; 12 lateral view; 13 oblique view of face and first leg; 14 retrolateral view of first leg femur (arrow highlights row of macrosetae); 15 – 16 living female; 17 ventral view of epigynum (scale bar 0.1 mm); 18 dorsal view of epigynum, cleared with clove oil. Figs 11 – 18 are copyright © 2016 W. P. Maddison, released under a Creative Commons Attribution (CC-BY) 3.0 license." httpUri="https://zenodo.org/record/187217/files/figure.png" pageId="4" pageNumber="441">Fig. 14</figureCitation>
|
||
). Carapace (
|
||
<figureCitation id="0DDB2A25FFACFF91FBE1FC9FF75EF562" box="[1147,1282,836,861]" captionStart="FIGURES 11 – 18" captionStartId="3.[151,264,1274,1297]" captionTargetBox="[151,1436,193,1253]" captionTargetId="figure@3.[151,1436,193,1253]" captionTargetPageId="3" captionText="FIGURES 11 – 18. Papuaneon tualapa sp. nov., female paratype (specimen 2008 PNG- 0845). 11 female paratype, dorsal view of carapace and abdomen; 12 lateral view; 13 oblique view of face and first leg; 14 retrolateral view of first leg femur (arrow highlights row of macrosetae); 15 – 16 living female; 17 ventral view of epigynum (scale bar 0.1 mm); 18 dorsal view of epigynum, cleared with clove oil. Figs 11 – 18 are copyright © 2016 W. P. Maddison, released under a Creative Commons Attribution (CC-BY) 3.0 license." httpUri="https://zenodo.org/record/187217/files/figure.png" pageId="4" pageNumber="441">Figs 11‒12</figureCitation>
|
||
): as in male. Abdomen: cordate, with long anterior setae as in male. Epigyne with circular openings and spermathecae closely resembling those of
|
||
<taxonomicName id="52E04D23FFACFF91FE18FC56F3E2F59A" box="[386,446,909,932]" class="Arachnida" family="Salticidae" genus="Neon" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="441" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A794EAB2FFACFF91FE18FC56F3E2F59A" box="[386,446,909,932]" italics="true" pageId="4" pageNumber="441">Neon</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="0DDB2A25FFACFF91FE54FC57F00EF59A" box="[462,594,908,933]" captionStart="FIGURES 11 – 18" captionStartId="3.[151,264,1274,1297]" captionTargetBox="[151,1436,193,1253]" captionTargetId="figure@3.[151,1436,193,1253]" captionTargetPageId="3" captionText="FIGURES 11 – 18. Papuaneon tualapa sp. nov., female paratype (specimen 2008 PNG- 0845). 11 female paratype, dorsal view of carapace and abdomen; 12 lateral view; 13 oblique view of face and first leg; 14 retrolateral view of first leg femur (arrow highlights row of macrosetae); 15 – 16 living female; 17 ventral view of epigynum (scale bar 0.1 mm); 18 dorsal view of epigynum, cleared with clove oil. Figs 11 – 18 are copyright © 2016 W. P. Maddison, released under a Creative Commons Attribution (CC-BY) 3.0 license." httpUri="https://zenodo.org/record/187217/files/figure.png" pageId="4" pageNumber="441">Figs 17‒18</figureCitation>
|
||
). As in
|
||
<taxonomicName id="52E04D23FFACFF91FD2AFC56F1ECF59A" authority="Logunov 1998" authorityName="Logunov" authorityYear="1998" box="[688,944,908,932]" class="Arachnida" family="Salticidae" genus="Neon" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="441" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A794EAB2FFACFF91FD2AFC56F0B0F59A" box="[688,748,909,932]" italics="true" pageId="4" pageNumber="441">Neon</emphasis>
|
||
(
|
||
<bibRefCitation id="F1714B51FFACFF91FD66FC57F1FBF59A" author="Logunov" box="[764,935,908,932]" pageId="4" pageNumber="441" refString="Logunov, D. V. (1998) The spider genus Neon Simon, 1876 (Araneae, Salticidae) in SE Asia, with notes on the genitalia and skin pore structures. Bulletin of the British Arachnological Society, 11, 15 - 22." type="journal article" year="1998">Logunov 1998</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, the spermatheca is divided into a primary receptacle bearing the fertilization ducts, and a simple-walled bulbous secondary receptacle (
|
||
<figureCitation id="0DDB2A25FFACFF91FB77FC74F71AF5F6" box="[1261,1350,943,968]" captionStart="FIGURES 11 – 18" captionStartId="3.[151,264,1274,1297]" captionTargetBox="[151,1436,193,1253]" captionTargetId="figure@3.[151,1436,193,1253]" captionTargetPageId="3" captionText="FIGURES 11 – 18. Papuaneon tualapa sp. nov., female paratype (specimen 2008 PNG- 0845). 11 female paratype, dorsal view of carapace and abdomen; 12 lateral view; 13 oblique view of face and first leg; 14 retrolateral view of first leg femur (arrow highlights row of macrosetae); 15 – 16 living female; 17 ventral view of epigynum (scale bar 0.1 mm); 18 dorsal view of epigynum, cleared with clove oil. Figs 11 – 18 are copyright © 2016 W. P. Maddison, released under a Creative Commons Attribution (CC-BY) 3.0 license." httpUri="https://zenodo.org/record/187217/files/figure.png" pageId="4" pageNumber="441">Fig. 18</figureCitation>
|
||
), as in Logunov’s (1998) figure 33. Colour in alcohol (
|
||
<figureCitation id="0DDB2A25FFACFF91FD2DFC0FF167F5D2" box="[695,827,980,1005]" captionStart="FIGURES 11 – 18" captionStartId="3.[151,264,1274,1297]" captionTargetBox="[151,1436,193,1253]" captionTargetId="figure@3.[151,1436,193,1253]" captionTargetPageId="3" captionText="FIGURES 11 – 18. Papuaneon tualapa sp. nov., female paratype (specimen 2008 PNG- 0845). 11 female paratype, dorsal view of carapace and abdomen; 12 lateral view; 13 oblique view of face and first leg; 14 retrolateral view of first leg femur (arrow highlights row of macrosetae); 15 – 16 living female; 17 ventral view of epigynum (scale bar 0.1 mm); 18 dorsal view of epigynum, cleared with clove oil. Figs 11 – 18 are copyright © 2016 W. P. Maddison, released under a Creative Commons Attribution (CC-BY) 3.0 license." httpUri="https://zenodo.org/record/187217/files/figure.png" pageId="4" pageNumber="441">Figs 11‒14</figureCitation>
|
||
): Chelicerae light brown. Palp with femur and patella dark, but tibia and tarsus bright white and with long white setae (
|
||
<figureCitation id="0DDB2A25FFACFF91FCE5FC2CF671F22E" box="[895,1069,1015,1040]" captionStart="FIGURES 11 – 18" captionStartId="3.[151,264,1274,1297]" captionTargetBox="[151,1436,193,1253]" captionTargetId="figure@3.[151,1436,193,1253]" captionTargetPageId="3" captionText="FIGURES 11 – 18. Papuaneon tualapa sp. nov., female paratype (specimen 2008 PNG- 0845). 11 female paratype, dorsal view of carapace and abdomen; 12 lateral view; 13 oblique view of face and first leg; 14 retrolateral view of first leg femur (arrow highlights row of macrosetae); 15 – 16 living female; 17 ventral view of epigynum (scale bar 0.1 mm); 18 dorsal view of epigynum, cleared with clove oil. Figs 11 – 18 are copyright © 2016 W. P. Maddison, released under a Creative Commons Attribution (CC-BY) 3.0 license." httpUri="https://zenodo.org/record/187217/files/figure.png" pageId="4" pageNumber="441">Figs 11, 13, 15</figureCitation>
|
||
). First leg as in male. Face dark brown with black setae. Carapace and abdomen as in male. Colour in life (
|
||
<figureCitation id="0DDB2A25FFACFF91FC42FBC7F607F20B" box="[984,1115,1052,1077]" captionStart="FIGURES 11 – 18" captionStartId="3.[151,264,1274,1297]" captionTargetBox="[151,1436,193,1253]" captionTargetId="figure@3.[151,1436,193,1253]" captionTargetPageId="3" captionText="FIGURES 11 – 18. Papuaneon tualapa sp. nov., female paratype (specimen 2008 PNG- 0845). 11 female paratype, dorsal view of carapace and abdomen; 12 lateral view; 13 oblique view of face and first leg; 14 retrolateral view of first leg femur (arrow highlights row of macrosetae); 15 – 16 living female; 17 ventral view of epigynum (scale bar 0.1 mm); 18 dorsal view of epigynum, cleared with clove oil. Figs 11 – 18 are copyright © 2016 W. P. Maddison, released under a Creative Commons Attribution (CC-BY) 3.0 license." httpUri="https://zenodo.org/record/187217/files/figure.png" pageId="4" pageNumber="441">Figs 15‒16</figureCitation>
|
||
): Except for the bright white tibia and tarsus of the palp, the rest of the body and appendages are black and dark brown with scattered brown to tan scales, aligned in vague transverse bands.
|
||
</paragraph>
|
||
<paragraph id="955F36A0FFACFF91FF5DFB5CF04BF30E" blockId="4.[151,1437,151,1977]" pageId="4" pageNumber="441">
|
||
<emphasis id="A794EAB2FFACFF91FF5DFB5CF315F29E" bold="true" box="[199,329,1159,1184]" pageId="4" pageNumber="441">Phylogeny.</emphasis>
|
||
Newly-obtained sequences are indicated in Table I. Alignment by MAFFT appeared reasonable and was not modified by hand. PartitionFinder determined the best partitioning scheme was to keep all partitions separate except to group the non-coding portion of 16SND1 with the first codon position of ND1, and to group first and second codon positions of Actin 5C. For all of these, GTR+I+G was determined as the best model, except for Actin 5C third position (GTR+G).
|
||
</paragraph>
|
||
<paragraph id="955F36A0FFACFF91FF5DFAE7F1F4F3FE" blockId="4.[151,1437,151,1977]" pageId="4" pageNumber="441">
|
||
The phylogenetic tree inferred is shown in
|
||
<figureCitation id="0DDB2A25FFACFF91FD2AFAE7F15FF36A" box="[688,771,1340,1365]" captionStart="FIGURE 19" captionStartId="5.[151,250,1026,1048]" captionTargetBox="[197,1402,202,985]" captionTargetId="figure@5.[181,1412,198,999]" captionTargetPageId="5" captionText="FIGURE 19. Maximum likelihood phylogeny based on analysis of 28 S, Actin 5 C and 16 SND 1 using RAxML. Numbers on branches are percentage of bootstrap replicates showing clade. Fig. 19 is copyright © 2016 W. P. Maddison, released under a Creative Commons Attribution (CC-BY) 3.0 license." httpUri="https://zenodo.org/record/187218/files/figure.png" pageId="4" pageNumber="441">Fig. 19</figureCitation>
|
||
.
|
||
<emphasis id="A794EAB2FFACFF91FC95FAE6F1D2F36A" box="[783,910,1341,1364]" italics="true" pageId="4" pageNumber="441">Papuaneon</emphasis>
|
||
is strongly supported as sister to
|
||
<taxonomicName id="52E04D23FFACFF91FA97FAE6F716F36A" box="[1293,1354,1341,1364]" class="Arachnida" family="Salticidae" genus="Neon" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="441" phylum="Arthropoda" rank="genus">Neon</taxonomicName>
|
||
(100% bootstrap support), though the branch separating them is long. As in previous analyses (
|
||
<bibRefCitation id="F1714B51FFACFF91FB00FA84F7C4F346" author="Maddison" box="[1178,1432,1375,1400]" pageId="4" pageNumber="441" refString="Maddison, W. P., Bodner, M. R., & Needham, K. (2008) Salticid spider phylogeny revisited, with the discovery of a large Australasian clade (Araneae: Salticidae). Zootaxa, 1893, 49 - 64." type="journal article" year="2008">
|
||
Maddison
|
||
<emphasis id="A794EAB2FFACFF91FA8DFABAF717F346" box="[1303,1355,1376,1400]" italics="true" pageId="4" pageNumber="441">et al</emphasis>
|
||
. 2008
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="F1714B51FFACFF91FF0DFA5FF39BF3A2" author="Bodner" box="[151,455,1412,1437]" pageId="4" pageNumber="441" refString="Bodner, M. R. & Maddison, W. P. (2012) The biogeography and age of salticid spider radiations (Araneae: Salticidae). Molecular Phylogenetics and Evolution, 65, 213 - 240. http: // dx. doi. org / 10.1016 / j. ympev. 2012.06.005" type="journal article" year="2012">Bodner & Maddison 2012</bibRefCitation>
|
||
), the five tribes of astioids (Mymarachninae,
|
||
<taxonomicName id="52E04D23FFACFF91FC4FFA5FF66EF3A3" box="[981,1074,1412,1437]" class="Arachnida" family="Salticidae" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="441" phylum="Arthropoda" rank="tribe" tribe="Neonini">Neonini</taxonomicName>
|
||
, Mopsini, Astiini, Viciriini) are well supported as distinct, but their interrelationships poorly resolved.
|
||
</paragraph>
|
||
<paragraph id="955F36A0FFACFF91FF5DFA17F08EF186" blockId="4.[151,1437,151,1977]" pageId="4" pageNumber="441">
|
||
The sample of
|
||
<taxonomicName id="52E04D23FFACFF91FEEBFA16F3F1F3DA" box="[369,429,1485,1508]" class="Arachnida" family="Salticidae" genus="Neon" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="441" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A794EAB2FFACFF91FEEBFA16F3F1F3DA" box="[369,429,1485,1508]" italics="true" pageId="4" pageNumber="441">Neon</emphasis>
|
||
</taxonomicName>
|
||
species is small within this analysis, which could lead to concerns that a broader sample might show
|
||
<emphasis id="A794EAB2FFACFF91FEB6FA2AF3F0F036" box="[300,428,1521,1544]" italics="true" pageId="4" pageNumber="441">Papuaneon</emphasis>
|
||
to be embedded within the full diversity of
|
||
<taxonomicName id="52E04D23FFACFF91FC52FA2AF65FF036" box="[968,1027,1521,1544]" class="Arachnida" family="Salticidae" genus="Neon" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="441" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A794EAB2FFACFF91FC52FA2AF65FF036" box="[968,1027,1521,1544]" italics="true" pageId="4" pageNumber="441">Neon</emphasis>
|
||
</taxonomicName>
|
||
species.
|
||
<taxonomicName id="52E04D23FFACFF91FBE9FA2AF6F2F036" box="[1139,1198,1521,1544]" class="Arachnida" family="Salticidae" genus="Neon" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="441" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A794EAB2FFACFF91FBE9FA2AF6F2F036" box="[1139,1198,1521,1544]" italics="true" pageId="4" pageNumber="441">Neon</emphasis>
|
||
</taxonomicName>
|
||
is divided into two subgenera (
|
||
<bibRefCitation id="F1714B51FFACFF91FE8CF9CFF387F012" author="Lohmander" box="[278,475,1556,1581]" pageId="4" pageNumber="441" refString="Lohmander, H. (1944) Vorlaufige Spinnennotizen. Arkiv for Zoologi, 35 (A, 16), 1 - 21." type="journal article" year="1944">Lohmander 1944</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="F1714B51FFACFF91FE7CF9CEF0D0F012" author="Logunov" box="[486,652,1556,1581]" pageId="4" pageNumber="441" refString="Logunov, D. V. (1998) The spider genus Neon Simon, 1876 (Araneae, Salticidae) in SE Asia, with notes on the genitalia and skin pore structures. Bulletin of the British Arachnological Society, 11, 15 - 22." type="journal article" year="1998">Logunov 1998</bibRefCitation>
|
||
), the nominate
|
||
<taxonomicName id="52E04D23FFACFF91FCADF9CEF12EF012" box="[823,882,1557,1580]" class="Arachnida" family="Salticidae" genus="Neon" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="441" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A794EAB2FFACFF91FCADF9CEF12EF012" box="[823,882,1557,1580]" italics="true" pageId="4" pageNumber="441">Neon</emphasis>
|
||
</taxonomicName>
|
||
as well as
|
||
<emphasis id="A794EAB2FFACFF91FC71F9CEF63EF012" box="[1003,1122,1557,1580]" italics="true" pageId="4" pageNumber="441">Dicroneon</emphasis>
|
||
<bibRefCitation id="F1714B51FFACFF91FBF2F9CFF76CF012" author="Lohmander" box="[1128,1328,1556,1581]" pageId="4" pageNumber="441" refString="Lohmander, H. (1944) Vorlaufige Spinnennotizen. Arkiv for Zoologi, 35 (A, 16), 1 - 21." type="journal article" year="1944">Lohmander, 1944</bibRefCitation>
|
||
, differing in the spicular lobe of the embolus (present in
|
||
<taxonomicName id="52E04D23FFACFF91FD07F9E2F084F06E" box="[669,728,1593,1616]" class="Arachnida" family="Salticidae" genus="Neon" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="441" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A794EAB2FFACFF91FD07F9E2F084F06E" box="[669,728,1593,1616]" italics="true" pageId="4" pageNumber="441">Neon</emphasis>
|
||
</taxonomicName>
|
||
), retrolateral tibial apophysis (more slender in
|
||
<emphasis id="A794EAB2FFACFF91FB7FF9E3F700F071" box="[1253,1372,1592,1615]" italics="true" pageId="4" pageNumber="441">Dicroneon</emphasis>
|
||
), and the secondary receptacle of the spermatheca (typically more elongate in
|
||
<emphasis id="A794EAB2FFACFF91FC5AF986F66BF04A" box="[960,1079,1629,1652]" italics="true" pageId="4" pageNumber="441">Dicroneon</emphasis>
|
||
) (see details in
|
||
<bibRefCitation id="F1714B51FFACFF91FB72F986F7CCF04A" author="Logunov" box="[1256,1424,1628,1653]" pageId="4" pageNumber="441" refString="Logunov, D. V. (1998) The spider genus Neon Simon, 1876 (Araneae, Salticidae) in SE Asia, with notes on the genitalia and skin pore structures. Bulletin of the British Arachnological Society, 11, 15 - 22." type="journal article" year="1998">Logunov 1998</bibRefCitation>
|
||
). However, all of the known species of
|
||
<taxonomicName id="52E04D23FFACFF91FDDFF95AF0DCF0A6" box="[581,640,1665,1688]" class="Arachnida" family="Salticidae" genus="Neon" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="441" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A794EAB2FFACFF91FDDFF95AF0DCF0A6" box="[581,640,1665,1688]" italics="true" pageId="4" pageNumber="441">Neon</emphasis>
|
||
</taxonomicName>
|
||
(including
|
||
<emphasis id="A794EAB2FFACFF91FC99F95BF126F0A9" box="[771,890,1664,1687]" italics="true" pageId="4" pageNumber="441">Dicroneon</emphasis>
|
||
) have the embolus arising on the prolateral side of the bulb (
|
||
<bibRefCitation id="F1714B51FFACFF91FEA5F97FF3A8F082" author="Logunov" box="[319,500,1700,1724]" pageId="4" pageNumber="441" refString="Logunov, D. V. (1998) The spider genus Neon Simon, 1876 (Araneae, Salticidae) in SE Asia, with notes on the genitalia and skin pore structures. Bulletin of the British Arachnological Society, 11, 15 - 22." type="journal article" year="1998">Logunov 1998</bibRefCitation>
|
||
). Among salticids with an embolus that is at least partially movable and counterclockwise-coiling (in the left palp), such as euophryines and marpissoids, the usual condition is for the embolus to arise distally on the bulb (e.g.,
|
||
<bibRefCitation id="F1714B51FFACFF91FDF5F937F17DF13B" author="Maddison" box="[623,801,1772,1797]" pageId="4" pageNumber="441" refString="Maddison, W. P. (1996) Pelegrina Franganillo and other jumping spiders formerly placed in the genus Metaphidippus (Araneae: Salticidae). Bulletin of the Museum of Comparative Zoology, 154, 215 - 368." type="journal article" year="1996">Maddison 1996</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="F1714B51FFACFF91FCB1F937F614F13B" author="Zhang" box="[811,1096,1772,1797]" pageId="4" pageNumber="441" refString="Zhang, J. X. & Maddison, W. P. (2015) Genera of euophryine jumping spiders (Araneae: Salticidae), with a combined molecular-morphological phylogeny. Zootaxa, 3938 (1), 1 - 147." type="journal article" year="2015">Zhang & Maddison 2015</bibRefCitation>
|
||
), and this is the condition seen in the relative of neonines, the mopsines. Thus, the prolateral embolus can be considered a synapomorphy delimiting the genus
|
||
<taxonomicName id="52E04D23FFACFF91FE1DF8EEF39FF172" box="[391,451,1845,1868]" class="Arachnida" family="Salticidae" genus="Neon" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="441" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A794EAB2FFACFF91FE1DF8EEF39FF172" box="[391,451,1845,1868]" italics="true" pageId="4" pageNumber="441">Neon</emphasis>
|
||
</taxonomicName>
|
||
and excluding
|
||
<emphasis id="A794EAB2FFACFF91FDEEF8EEF0AFF172" box="[628,755,1845,1868]" italics="true" pageId="4" pageNumber="441">Papuaneon</emphasis>
|
||
. In addition, even if some species currently placed in
|
||
<taxonomicName id="52E04D23FFACFF91FAFBF8EEF7C1F172" box="[1377,1437,1845,1868]" class="Arachnida" family="Salticidae" genus="Neon" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="441" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A794EAB2FFACFF91FAFBF8EEF7C1F172" box="[1377,1437,1845,1868]" italics="true" pageId="4" pageNumber="441">Neon</emphasis>
|
||
</taxonomicName>
|
||
were found to be as or more distantly related to the
|
||
<typeStatus id="4A5B8802FFACFF91FD43F882F155F14F" box="[729,777,1881,1905]" pageId="4" pageNumber="441">type</typeStatus>
|
||
species
|
||
<taxonomicName id="52E04D23FFACFF91FCF3F882F67EF14E" box="[873,1058,1880,1904]" class="Arachnida" family="Salticidae" genus="Neon" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="441" phylum="Arthropoda" rank="species" species="reticulatus">
|
||
<emphasis id="A794EAB2FFACFF91FCF3F882F67EF14E" box="[873,1058,1880,1904]" italics="true" pageId="4" pageNumber="441">Neon reticulatus</emphasis>
|
||
</taxonomicName>
|
||
than is
|
||
<taxonomicName id="52E04D23FFACFF91FBE0F882F709F14E" box="[1146,1365,1880,1904]" class="Arachnida" family="Salticidae" genus="Papuaneon" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="441" phylum="Arthropoda" rank="species" species="tualapa">
|
||
<emphasis id="A794EAB2FFACFF91FBE0F882F709F14E" box="[1146,1365,1880,1904]" italics="true" pageId="4" pageNumber="441">Papuaneon tualapa</emphasis>
|
||
</taxonomicName>
|
||
, there would be a strong argument for dividing
|
||
<taxonomicName id="52E04D23FFACFF91FD1AF8A6F0E0F1AA" box="[640,700,1917,1940]" class="Arachnida" family="Salticidae" genus="Neon" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="441" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A794EAB2FFACFF91FD1AF8A6F0E0F1AA" box="[640,700,1917,1940]" italics="true" pageId="4" pageNumber="441">Neon</emphasis>
|
||
</taxonomicName>
|
||
into multiple genera, given the depth of molecular divergence between
|
||
<emphasis id="A794EAB2FFACFF91FF61F87AF326F186" box="[251,378,1953,1976]" italics="true" pageId="4" pageNumber="441">Papuaneon</emphasis>
|
||
and the
|
||
<typeStatus id="4A5B8802FFACFF91FE40F87AF056F187" box="[474,522,1953,1977]" pageId="4" pageNumber="441">type</typeStatus>
|
||
species (
|
||
<figureCitation id="0DDB2A25FFACFF91FDE8F87BF099F186" box="[626,709,1952,1977]" captionStart="FIGURE 19" captionStartId="5.[151,250,1026,1048]" captionTargetBox="[197,1402,202,985]" captionTargetId="figure@5.[181,1412,198,999]" captionTargetPageId="5" captionText="FIGURE 19. Maximum likelihood phylogeny based on analysis of 28 S, Actin 5 C and 16 SND 1 using RAxML. Numbers on branches are percentage of bootstrap replicates showing clade. Fig. 19 is copyright © 2016 W. P. Maddison, released under a Creative Commons Attribution (CC-BY) 3.0 license." httpUri="https://zenodo.org/record/187218/files/figure.png" pageId="4" pageNumber="441">Fig. 19</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |