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<document id="D94587E25D1D8BF3486B6614D29ACFB6" ID-DOI="10.5281/zenodo.213112" ID-GBIF-Dataset="40e7a57f-5730-4375-83bf-eaa765ac079f" ID-ISSN="1175-5326" ID-Zenodo-Dep="213112" ID-ZooBank="A7CD3E37-26D3-417F-BEE2-62973E98893E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460527685773" checkinUser="plazi" docAuthor="Bird, Graham J." docDate="2012" docId="19228321FFCFFFC3CEF1FC1E1C3F04E3" docLanguage="en" docName="zt03481p026.pdf" docOrigin="Zootaxa 3481" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Heterotanoides Sieg 1977" docType="treatment" docVersion="10" lastPageNumber="3" masterDocId="E51BFB59FFCEFFC1CE66FFBB1F7A0644" masterDocTitle="A new leptochelioid family, Heterotanoididae (Crustacea: Peracarida: Tanaidacea), and a new species of Heterotanoides from New Zealand" masterLastPageNumber="26" masterPageNumber="1" pageNumber="2" updateTime="1698315313888" updateUser="plazi">
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<mods:title id="ACFCC923CAE21B9ECC6E92F112093901">A new leptochelioid family, Heterotanoididae (Crustacea: Peracarida: Tanaidacea), and a new species of Heterotanoides from New Zealand</mods:title>
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<mods:namePart id="93EB96B15B52CB94A88052E15AE1C90B">Bird, Graham J.</mods:namePart>
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<mods:date id="899DBA77C35A5A8A2C23452B88A1B12C">2012</mods:date>
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<mods:classification id="D0F147549E1F41A8F092E99539442BA1">journal article</mods:classification>
<mods:identifier id="56DD275CF2AD72C1DB97D7848C29CAD7" type="DOI">10.5281/zenodo.213112</mods:identifier>
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<treatment id="19228321FFCFFFC3CEF1FC1E1C3F04E3" ID-DOI="http://doi.org/10.5281/zenodo.5674341" ID-GBIF-Taxon="119526463" ID-Zenodo-Dep="5674341" LSID="urn:lsid:plazi:treatment:19228321FFCFFFC3CEF1FC1E1C3F04E3" httpUri="http://treatment.plazi.org/id/19228321FFCFFFC3CEF1FC1E1C3F04E3" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2">
<subSubSection id="D99161BCFFCFFFC0CEF1FC1E1CA7027B" pageId="1" pageNumber="2" type="nomenclature">
<paragraph id="91343237FFCFFFC0CEF1FC1E1D5005FB" blockId="1.[151,554,933,959]" box="[151,554,933,959]" pageId="1" pageNumber="2">
<heading id="CA7C855BFFCFFFC0CEF1FC1E1D5005FB" bold="true" box="[151,554,933,959]" fontSize="11" level="1" pageId="1" pageNumber="2" reason="1">
<emphasis id="A3FFEE25FFCFFFC0CEF1FC1E1D5005FB" bold="true" box="[151,554,933,959]" pageId="1" pageNumber="2">
Genus
<taxonomicName id="568B49B4FFCFFFC0CE8BFC1E1D5005FB" ID-CoL="84P4B" authority="Sieg, 1977" authorityName="Sieg" authorityYear="1977" box="[237,554,933,959]" class="Malacostraca" family="Heterotanoididae" genus="Heterotanoides" kingdom="Animalia" order="Tanaidacea" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="A3FFEE25FFCFFFC0CE8BFC1E1EDF05FB" bold="true" box="[237,421,933,959]" italics="true" pageId="1" pageNumber="2">Heterotanoides</emphasis>
Sieg, 1977
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="91343237FFCFFFC0CEF1FC501CA7027B" blockId="1.[151,1436,1003,1088]" pageId="1" pageNumber="2">
<bibRefCitation id="F51A4FC6FFCFFFC0CEF1FC501E670246" author="Sieg" box="[151,285,1003,1026]" pageId="1" pageNumber="2" refString="Sieg, J. (1977 a) Eine neue Gattung (Heterotanoides gen. n.) aus der Familie Paratanaidae Lang (Tanaidacea). Crustaceana, 33 (2), 203 - 209." type="journal article" year="1977" yearSuffix="a">Sieg (1977a)</bibRefCitation>
: 203 [diagnosis]; Sieg (1983): 442–443 [bibliography];
<bibRefCitation id="F51A4FC6FFCFFFC0CD35FC501CA30245" author="Sieg" box="[851,985,1003,1025]" pageId="1" pageNumber="2" refString="Sieg, J. (1986 a) Crustacea Tanaidacea of the Antarctic and the Subantarctic. I. On material collected at Tierra del Fuego, Isla de los Estados, and the West Coast of the Antarctic Peninsula. Antarctic Research Series Vol. 45, Biology of the Antarctic Seas XVIII, 180 pp." type="book" year="1986" yearSuffix="a">Sieg (1986a)</bibRefCitation>
: 47 [new diagnosis, key to species];
<bibRefCitation id="F51A4FC6FFCFFFC0CB2AFC501E1B0265" author="Bamber" pageId="1" pageNumber="2" refString="Bamber, R. N. &amp; Bird, G. J. (1997) Peracarid crustaceans from Cape D'Aguilar and Hong Kong, III. Tanaidacea: Tanaidomorpha. In: Morton, B. [ed.], The marine Flora and Fauna of Hong Kong and Southern China IV. Proceedings of the Eighth International Marine Biological Workshop: The marine Flora and Fauna of Hong Kong and Southern China, Hong Kong, 2 - 20 April 1995. Hong Kong, Hong Kong University Press, pp. 103 - 142." type="journal article" year="1997">Bamber &amp; Bird (1997)</bibRefCitation>
: 117, 120 [comparison of species];
<bibRefCitation id="F51A4FC6FFCFFFC0CCBCFBB11C010265" author="Bamber" box="[730,891,1034,1057]" pageId="1" pageNumber="2" refString="Bamber, R. N. (1998) Zoogeographic trends in some Hong Kong arthropods. In: Morton, B. [ed.], The Marine Biology of the South China Sea. Proceedings of the Third International Conference on the Marine Biology of the South China Sea, Hong Kong, 26 October - 1 November 1996. Hong Kong, Hong Kong University Press, pp. 91 - 112." type="journal article" year="1998">Bamber (1998)</bibRefCitation>
: 101, fig. 8 [distribution map of species];
<bibRefCitation id="F51A4FC6FFCFFFC0CB5CFBB01E260204" author="Larsen" pageId="1" pageNumber="2" refString="Larsen, K. &amp; Wilson, G. D. F. (2002) Tanaidacean phylogeny, the first step: the Superfamily Paratanaidoidea. Journal of Zoological Systematics and Evolutionary Research, 40, 205 - 222." type="journal article" year="2002">Larsen &amp; Wilson (2002)</bibRefCitation>
: 13 [classification];
<bibRefCitation id="F51A4FC6FFCFFFC0CC4CFB921C730204" author="Bird" box="[554,777,1065,1088]" pageId="1" pageNumber="2" refString="Bird, G. J. &amp; Larsen, K. (2009) Tanaidacea phylogeny - the second step; the basal paratanaoidean families (Crustacea: Malacostraca). Arthropod Systematics &amp; Phylogeny, 67 (2), 137 - 158." type="journal article" year="2009">Bird &amp; Larsen (2009)</bibRefCitation>
: 156 [classification].
</paragraph>
</subSubSection>
<subSubSection id="D99161BCFFCFFFC0CEF1FBD01D8A0038" pageId="1" pageNumber="2" type="diagnosis">
<paragraph id="91343237FFCFFFC0CEF1FBD01D8A0038" blockId="1.[151,1437,1131,2020]" pageId="1" pageNumber="2">
<emphasis id="A3FFEE25FFCFFFC0CEF1FBD01E7002C0" bold="true" box="[151,266,1131,1156]" pageId="1" pageNumber="2">Diagnosis</emphasis>
. (Modified from
<bibRefCitation id="F51A4FC6FFCFFFC0CFB9FBD71D1C02C0" author="Sieg" box="[479,614,1132,1156]" pageId="1" pageNumber="2" refString="Sieg, J. (1986 a) Crustacea Tanaidacea of the Antarctic and the Subantarctic. I. On material collected at Tierra del Fuego, Isla de los Estados, and the West Coast of the Antarctic Peninsula. Antarctic Research Series Vol. 45, Biology of the Antarctic Seas XVIII, 180 pp." type="book" year="1986" yearSuffix="a">Sieg 1986a</bibRefCitation>
). Leptochelioid, with
<emphasis id="A3FFEE25FFCFFFC0CD0AFBD71B7202C0" box="[876,1032,1132,1156]" italics="true" pageId="1" pageNumber="2">pleonites 1–4</emphasis>
with or without delicate plumose epimeral setae.
<emphasis id="A3FFEE25FFCFFFC0CF2BFB2B1EC502EC" box="[333,447,1168,1192]" italics="true" pageId="1" pageNumber="2">Antennule</emphasis>
three-articled, with or without small cap-like terminal segment [‘article-4’]; article-3 with one or more groups of subterminal aesthetascs.
<emphasis id="A3FFEE25FFCFFFC0CC9DFB0F1C220288" box="[763,856,1204,1228]" italics="true" pageId="1" pageNumber="2">Antenna</emphasis>
seven-articled; article-2 larger than article-3, both with stiff superodistal seta;
<emphasis id="A3FFEE25FFCFFFC0CFB7FB631D6002B4" box="[465,538,1240,1264]" italics="true" pageId="1" pageNumber="2">article</emphasis>
-4 with long setae on mesial border.
<emphasis id="A3FFEE25FFCFFFC0CDDDFB631B6B02B4" box="[955,1041,1240,1264]" italics="true" pageId="1" pageNumber="2">Labium</emphasis>
with two pairs of lobes, outer pair dominant.
<emphasis id="A3FFEE25FFCFFFC0CF68FB471E000350" box="[270,378,1276,1300]" italics="true" pageId="1" pageNumber="2">Mandible</emphasis>
molar nodulose, without ridges.
<emphasis id="A3FFEE25FFCFFFC0CC8FFB471C2E0350" box="[745,852,1276,1300]" italics="true" pageId="1" pageNumber="2">Maxillule</emphasis>
endite with ten or eleven short terminal spines, palp with two long setae.
<emphasis id="A3FFEE25FFCFFFC0CFF6FA9B1D73037C" box="[400,521,1312,1336]" italics="true" pageId="1" pageNumber="2">Maxilliped</emphasis>
bases not fused; with one long distal seta; endite with 2–4 spines (acute and digitiform); palp article-2 without lateral seta.
<emphasis id="A3FFEE25FFCFFFC0CCC9FAFF1C6E0318" box="[687,788,1348,1372]" italics="true" pageId="1" pageNumber="2">Cheliped</emphasis>
sclerite triangular, dorsally inserted into basis, posterior margin confluent with rear of carapace; basis posterior lobe reaching pereonite-1; carpus with up to four superior setae; propodus weakly or distinctly flared; palm and fixed finger with three or more inferior setae; fixed finger incisive margin with or without small distal teeth.
<emphasis id="A3FFEE25FFCFFFC0CCBCFA0B1CF9038C" box="[730,899,1456,1480]" italics="true" pageId="1" pageNumber="2">Pereopods 1–3</emphasis>
merus, carpus and propodus with simple setae only.
<emphasis id="A3FFEE25FFCFFFC0CEBFFA6F1EFF03A8" box="[217,389,1492,1516]" italics="true" pageId="1" pageNumber="2">Pereopods 4–6</emphasis>
carpus with two or three setal loci [typically as two spines and superior seta]; propodus superodistal spines similar to inferodistal.
<emphasis id="A3FFEE25FFCFFFC0CC1BFA431DA00054" box="[637,730,1528,1552]" italics="true" pageId="1" pageNumber="2">Pleopod</emphasis>
peduncle with seta; endopod inner margin with seta.
<emphasis id="A3FFEE25FFCFFFC0CB20FA431AE60054" box="[1350,1436,1528,1552]" italics="true" pageId="1" pageNumber="2">Uropod</emphasis>
peduncle with seta; endopod three-segmented; exopod two-segmented, segment-1 much shorter than segment-2.
<emphasis id="A3FFEE25FFCFFFC0CEF1F9FB1FAB001C" box="[151,209,1600,1624]" italics="true" pageId="1" pageNumber="2">Male</emphasis>
similar to female but
<emphasis id="A3FFEE25FFCFFFC0CFAEF9FB1D4C001C" box="[456,566,1600,1624]" italics="true" pageId="1" pageNumber="2">antennule</emphasis>
more elongate, with or without additional aesthetasc bundles;
<emphasis id="A3FFEE25FFCFFFC0CA93F9FB1A2E001C" box="[1269,1364,1600,1624]" italics="true" pageId="1" pageNumber="2">cheliped</emphasis>
larger and more robust;
<emphasis id="A3FFEE25FFCFFFC0CF3AF9DF1EA60038" box="[348,476,1636,1660]" italics="true" pageId="1" pageNumber="2">mouthparts</emphasis>
functional, as in female.
</paragraph>
</subSubSection>
<subSubSection id="D99161BCFFCFFFC0CEA1F9331E2F0174" pageId="1" pageNumber="2" type="materials_examined">
<paragraph id="91343237FFCFFFC0CEA1F9331C2D00E4" blockId="1.[151,1437,1131,2020]" box="[199,855,1671,1696]" pageId="1" pageNumber="2">
<emphasis id="A3FFEE25FFCFFFC0CEA1F9331E1800E4" bold="true" box="[199,354,1671,1696]" pageId="1" pageNumber="2">
<typeStatus id="4E308C95FFCFFFC0CEA1F9331E7800E4" box="[199,258,1672,1696]" pageId="1" pageNumber="2">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="568B49B4FFCFFFC0CF0FF9331C2900E4" authority="Vanhoffen, 1914" authorityName="Vanhoffen" authorityYear="1914" box="[361,851,1672,1696]" class="Malacostraca" family="Heterotanoididae" genus="Heterotanoides" kingdom="Animalia" order="Tanaidacea" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="capensis">
<emphasis id="A3FFEE25FFCFFFC0CF0FF9331D0100E4" box="[361,635,1672,1696]" italics="true" pageId="1" pageNumber="2">Heterotanoides capensis</emphasis>
(
<bibRefCitation id="F51A4FC6FFCFFFC0CCECF9331C3100E4" author="Vanhoffen" box="[650,843,1672,1696]" pageId="1" pageNumber="2" refString="Vanhoffen, E. (1914) Die Isopoden der Deutschen Sudpolar-Expedition 1901 - 1903. Deutschen Sudpolar-Expedition, 15, 449 - 598." type="journal article" year="1914">Vanhöffen, 1914</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="91343237FFCFFFC0CEA1F9101E2F0174" blockId="1.[151,1437,1131,2020]" pageId="1" pageNumber="2">
<emphasis id="A3FFEE25FFCFFFC0CEA1F9101E250080" bold="true" box="[199,351,1707,1732]" pageId="1" pageNumber="2">Composition</emphasis>
.
<taxonomicName id="568B49B4FFCFFFC0CF08F9171C640081" authority="Simons Bay" authorityName="Simons Bay" box="[366,798,1708,1733]" class="Malacostraca" family="Heterotanoididae" genus="Heterotanoides" kingdom="Animalia" order="Tanaidacea" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="capensis">
<emphasis id="A3FFEE25FFCFFFC0CF08F9171DF10080" box="[366,651,1708,1732]" italics="true" pageId="1" pageNumber="2">Heterotanoides capensis:</emphasis>
Simons Bay
</taxonomicName>
,
<collectingCountry id="E99C72A7FFCFFFC0CD4DF9171CB80080" box="[811,962,1708,1732]" name="South Africa" pageId="1" pageNumber="2">South Africa</collectingCountry>
,?m, littoral?;
<taxonomicName id="568B49B4FFCFFFC0CA3DF9161FA900AC" authority="Bamber &amp; Bird 1997" authorityName="Bamber &amp; Bird" authorityYear="1997" class="Malacostraca" family="Heterotanoididae" genus="Heterotanoides" kingdom="Animalia" order="Tanaidacea" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="cognatus">
<emphasis id="A3FFEE25FFCFFFC0CA3DF9161B990080" box="[1115,1251,1709,1732]" italics="true" pageId="1" pageNumber="2">H. cognatus</emphasis>
<bibRefCitation id="F51A4FC6FFCFFFC0CA8AF9171FA900AC" author="Bamber" pageId="1" pageNumber="2" refString="Bamber, R. N. &amp; Bird, G. J. (1997) Peracarid crustaceans from Cape D'Aguilar and Hong Kong, III. Tanaidacea: Tanaidomorpha. In: Morton, B. [ed.], The marine Flora and Fauna of Hong Kong and Southern China IV. Proceedings of the Eighth International Marine Biological Workshop: The marine Flora and Fauna of Hong Kong and Southern China, Hong Kong, 2 - 20 April 1995. Hong Kong, Hong Kong University Press, pp. 103 - 142." type="journal article" year="1997">Bamber &amp; Bird 1997</bibRefCitation>
</taxonomicName>
:
<collectingCountry id="E99C72A7FFCFFFC0CEB9F96A1E1D00AD" box="[223,359,1745,1769]" name="Hong Kong" pageId="1" pageNumber="2">Hong Kong</collectingCountry>
,
<quantity id="56739FD2FFCFFFC0CF12F96B1E9900AC" box="[372,483,1744,1768]" metricMagnitude="0" metricUnit="m" metricValue="9.25" metricValueMax="14.5" metricValueMin="4.0" pageId="1" pageNumber="2" unit="m" value="9.25" valueMax="14.5" valueMin="4.0">4–14.5 m</quantity>
;
<emphasis id="A3FFEE25FFCFFFC0CF96F96B1DD000AC" bold="true" box="[496,682,1743,1768]" pageId="1" pageNumber="2">
<taxonomicName id="568B49B4FFCFFFC0CF96F96B1D0A00AC" box="[496,624,1743,1768]" class="Malacostraca" family="Heterotanoididae" genus="Heterotanoides" kingdom="Animalia" order="Tanaidacea" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="muimui">
<emphasis id="A3FFEE25FFCFFFC0CF96F96B1D0A00AC" bold="true" box="[496,624,1743,1768]" italics="true" pageId="1" pageNumber="2">H. muimui</emphasis>
</taxonomicName>
n.sp
</emphasis>
;
<taxonomicName id="568B49B4FFCFFFC0CCDCF96A1C8500AC" authority="Sieg, 1986" authorityName="Sieg" authorityYear="1986" box="[698,1023,1744,1768]" class="Malacostraca" family="Heterotanoididae" genus="Heterotanoides" kingdom="Animalia" order="Tanaidacea" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="meridionalis">
<emphasis id="A3FFEE25FFCFFFC0CCDCF96A1C1600AC" box="[698,876,1744,1768]" italics="true" pageId="1" pageNumber="2">H. meridionalis</emphasis>
<bibRefCitation id="F51A4FC6FFCFFFC0CD12F96B1C8500AC" author="Sieg" box="[884,1023,1744,1768]" pageId="1" pageNumber="2" refString="Sieg, J. (1986 a) Crustacea Tanaidacea of the Antarctic and the Subantarctic. I. On material collected at Tierra del Fuego, Isla de los Estados, and the West Coast of the Antarctic Peninsula. Antarctic Research Series Vol. 45, Biology of the Antarctic Seas XVIII, 180 pp." type="book" year="1986" yearSuffix="a">Sieg, 1986a</bibRefCitation>
</taxonomicName>
:
<collectingCountry id="E99C72A7FFCFFFC0CA6DF96B1B4E00AC" box="[1035,1076,1744,1768]" name="Iceland" pageId="1" pageNumber="2">Isla</collectingCountry>
de los Estados (
<collectingCountry id="E99C72A7FFCFFFC0CA96F96B1A1900AC" box="[1264,1379,1744,1768]" name="Argentina" pageId="1" pageNumber="2">Argentina</collectingCountry>
) and Tierra del Fuego,
<quantity id="56739FD2FFCFFFC0CF08F94F1EA30148" box="[366,473,1780,1804]" metricMagnitude="1" metricUnit="m" metricValue="2.85" metricValueMax="4.4" metricValueMin="1.3" pageId="1" pageNumber="2" unit="m" value="28.5" valueMax="44.0" valueMin="13.0">13–44 m</quantity>
;
<taxonomicName id="568B49B4FFCFFFC0CF8CF94E1CDD0148" authority="Kudinova-Pasternak, 1976" authorityName="Kudinova-Pasternak" authorityYear="1976" box="[490,935,1780,1804]" class="Malacostraca" family="Heterotanoididae" genus="Heterotanoides" kingdom="Animalia" order="Tanaidacea" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="ornatus">
<emphasis id="A3FFEE25FFCFFFC0CF8CF94E1D1D0148" box="[490,615,1781,1804]" italics="true" pageId="1" pageNumber="2">H. ornatus</emphasis>
<bibRefCitation id="F51A4FC6FFCFFFC0CC15F94F1CDD0148" author="Kudinova-Pasternak" box="[627,935,1780,1804]" pageId="1" pageNumber="2" refString="Kudinova-Pasternak, R. K. (1976) Additional notes on the fauna of Tanaidacea (Crustacea, Malacostraca) of the deep-water trenches. Trudy Instituta Okeanologii, Akademiya Nauk SSSR, 99, 115 - 135." type="journal article" year="1976">Kudinova-Pasternak, 1976</bibRefCitation>
</taxonomicName>
:
<collectingCountry id="E99C72A7FFCFFFC0CDD0F94E1C8C0148" box="[950,1014,1781,1804]" name="Japan" pageId="1" pageNumber="2">Japan</collectingCountry>
Trench,&gt;
<quantity id="56739FD2FFCFFFC0CA15F94F1BB40148" box="[1139,1230,1780,1804]" metricMagnitude="3" metricUnit="m" metricValue="7.2" pageId="1" pageNumber="2" unit="m" value="7200.0">7200 m</quantity>
(possibly sample contamination?).
</paragraph>
</subSubSection>
<subSubSection id="D99161BCFFCFFFC3CEA1F8871C3F04E3" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2" type="discussion">
<paragraph id="91343237FFCFFFC3CEA1F8871C7A0758" blockId="1.[151,1437,1131,2020]" lastBlockId="2.[151,1436,151,679]" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2">
<emphasis id="A3FFEE25FFCFFFC0CEA1F8871E410110" bold="true" box="[199,315,1852,1876]" pageId="1" pageNumber="2">Remarks.</emphasis>
The principal characters that distinguish
<taxonomicName id="568B49B4FFCFFFC0CD57F8871CA70110" box="[817,989,1852,1876]" class="Malacostraca" family="Heterotanoididae" genus="Heterotanoides" kingdom="Animalia" order="Tanaidacea" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="A3FFEE25FFCFFFC0CD57F8871CA70110" box="[817,989,1852,1876]" italics="true" pageId="1" pageNumber="2">Heterotanoides</emphasis>
</taxonomicName>
from all other leptochelioids are the presence of both subdistal antennular aesthetascs and a seven-articled antenna. Also, compared to genera such as
<taxonomicName id="568B49B4FFCFFFC0CEF1F83F1E0A01D8" authority="Dana" authorityName="Dana" box="[151,368,1924,1948]" class="Malacostraca" family="Leptocheliidae" genus="Leptochelia" kingdom="Animalia" order="Tanaidacea" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="A3FFEE25FFCFFFC0CEF1F83F1E6101D8" box="[151,283,1924,1948]" italics="true" pageId="1" pageNumber="2">Leptochelia</emphasis>
Dana
</taxonomicName>
,
<taxonomicName id="568B49B4FFCFFFC0CFEAF83F1DD401D9" authority="Lang" authorityName="Lang" box="[396,686,1924,1949]" class="Malacostraca" family="Leptocheliidae" genus="Pseudoleptochelia" kingdom="Animalia" order="Tanaidacea" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="A3FFEE25FFCFFFC0CFEAF83F1D2101D8" box="[396,603,1924,1948]" italics="true" pageId="1" pageNumber="2">Pseudoleptochelia</emphasis>
Lang
</taxonomicName>
, and
<taxonomicName id="568B49B4FFCFFFC0CD6CF83E1B5601D8" authority="G.O. Sars" authorityName="G.O. Sars" box="[778,1068,1924,1948]" class="Malacostraca" family="Leptocheliidae" genus="Heterotanais" kingdom="Animalia" order="Tanaidacea" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="A3FFEE25FFCFFFC0CD6CF83E1CE101D8" box="[778,923,1925,1948]" italics="true" pageId="1" pageNumber="2">Heterotanais</emphasis>
G.O. Sars
</taxonomicName>
(the last in the subfamily
<taxonomicName id="568B49B4FFCFFFC0CEF1F8131E210184" box="[151,347,1960,1984]" class="Malacostraca" family="Leptocheliidae" kingdom="Animalia" order="Tanaidacea" pageId="1" pageNumber="2" phylum="Arthropoda" rank="subFamily" subFamily="Heterotanaidinae">Heterotanaidinae</taxonomicName>
), sexual dimorphism is also only moderate in
<taxonomicName id="568B49B4FFCFFFC0CDE9F8131B410184" box="[911,1083,1960,1984]" class="Malacostraca" family="Heterotanoididae" genus="Heterotanoides" kingdom="Animalia" order="Tanaidacea" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="A3FFEE25FFCFFFC0CDE9F8131B410184" box="[911,1083,1960,1984]" italics="true" pageId="1" pageNumber="2">Heterotanoides</emphasis>
</taxonomicName>
, restricted to relatively small differences in cephalothorax, pereon and cheliped proportions, with antennular aesthetascs being only slightly more numerous in males. Other characters could help define the genus, but are variable in the literature, but it is unclear if this is due to oversights or is real; examples include the setation of the maxilliped palp articles 2–4. The propodal spines of pereopods 4–6 are also similar in both superior distal and inferior distal groups, a feature possibly useful for diagnoses and phylogenetic studies.
</paragraph>
<paragraph id="91343237FFCCFFC3CEA1FE931BD007B0" blockId="2.[151,1436,151,679]" pageId="2" pageNumber="3">
In his redescription of
<taxonomicName id="568B49B4FFCCFFC3CF85FE921C6E077B" authority="Sieg" authorityName="Sieg" authorityYear="1977" box="[483,788,295,320]" class="Malacostraca" family="Heterotanoididae" genus="Heterotanoides" kingdom="Animalia" order="Tanaidacea" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="capensis">
<emphasis id="A3FFEE25FFCCFFC3CF85FE921D170704" box="[483,621,297,320]" italics="true" pageId="2" pageNumber="3">H. capensis</emphasis>
<bibRefCitation id="F51A4FC6FFCCFFC3CC1AFE9C1C6E077B" author="Sieg" box="[636,788,295,319]" pageId="2" pageNumber="3" refString="Sieg, J. (1977 a) Eine neue Gattung (Heterotanoides gen. n.) aus der Familie Paratanaidae Lang (Tanaidacea). Crustaceana, 33 (2), 203 - 209." type="journal article" year="1977" yearSuffix="a">Sieg (1977a)</bibRefCitation>
</taxonomicName>
used a ‘female without oostegites’ but the pereonite proportions do not match those shown by
<bibRefCitation id="F51A4FC6FFCCFFC3CC11FEF71C440720" author="Vanhoffen" box="[631,830,332,356]" pageId="2" pageNumber="3" refString="Vanhoffen, E. (1914) Die Isopoden der Deutschen Sudpolar-Expedition 1901 - 1903. Deutschen Sudpolar-Expedition, 15, 449 - 598." type="journal article" year="1914">Vanhöffen (1914)</bibRefCitation>
for an ovigerous female; they more closely resemble the short pereonites of male
<taxonomicName id="568B49B4FFCCFFC3CF83FECA1D2507CC" box="[485,607,369,392]" class="Malacostraca" family="Heterotanoididae" genus="Heterotanoides" kingdom="Animalia" order="Tanaidacea" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="muimui" status="sp. nov.">
<emphasis id="A3FFEE25FFCCFFC3CF83FECA1D2507CC" box="[485,607,369,392]" italics="true" pageId="2" pageNumber="3">H. muimui</emphasis>
</taxonomicName>
<emphasis id="A3FFEE25FFCCFFC3CC0FFECB1DD007CC" bold="true" box="[617,682,368,392]" pageId="2" pageNumber="3">
<taxonomicNameLabel id="B8CC535EFFCCFFC3CC0FFECB1DD007CC" box="[617,682,368,392]" pageId="2" pageNumber="3" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
[see below]. In addition, the aesthetasc pattern (
<figureCitation id="09B02EB2FFCCFFC3CA85FED41A5207C3" box="[1251,1320,367,391]" captionStart="FIGURE 5" captionStartId="8.[151,249,1900,1923]" captionTargetBox="[172,1414,193,1863]" captionTargetId="figure@8.[172,1416,193,1879]" captionTargetPageId="8" captionText="FIGURE 5. Antennular aesthetascs. A, C Nototanoides; B, D Paranesotanais; E, J Heterotanoides meridionalis; F H. ornatus; G H. capensis; H, O H. muimui n. sp.; K, M H. cognatus; L, N Pseudozeuxo. All redrawn from original sources, rescaled, and setae omitted for clarity. Frame encloses nototanaid genera, with separate homology. Antennules E – O are arranged in approximate order of complexity." httpUri="https://zenodo.org/record/213117/files/figure.png" pageId="2" pageNumber="3">Fig. 5</figureCitation>
G) of two subterminal bundles corresponds to that of male
<taxonomicName id="568B49B4FFCCFFC3CCA7FE2E1C4307E8" box="[705,825,405,428]" class="Malacostraca" family="Heterotanoididae" genus="Heterotanoides" kingdom="Animalia" order="Tanaidacea" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="muimui">
<emphasis id="A3FFEE25FFCCFFC3CCA7FE2E1C4307E8" box="[705,825,405,428]" italics="true" pageId="2" pageNumber="3">H. muimui</emphasis>
</taxonomicName>
; these two observations suggest that Sieg’s figures of
<taxonomicName id="568B49B4FFCCFFC3CEF1FE021E670794" box="[151,285,441,464]" class="Malacostraca" family="Heterotanoididae" genus="Heterotanoides" kingdom="Animalia" order="Tanaidacea" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="capensis">
<emphasis id="A3FFEE25FFCCFFC3CEF1FE021E670794" box="[151,285,441,464]" italics="true" pageId="2" pageNumber="3">H. capensis</emphasis>
</taxonomicName>
are of a male. For similar reasons, and based on the relative stout cheliped, it is possible that the description of
<taxonomicName id="568B49B4FFCCFFC3CF5EFE661ECA07B0" box="[312,432,477,500]" class="Malacostraca" family="Heterotanoididae" genus="Heterotanoides" kingdom="Animalia" order="Tanaidacea" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="ornatus">
<emphasis id="A3FFEE25FFCCFFC3CF5EFE661ECA07B0" box="[312,432,477,500]" italics="true" pageId="2" pageNumber="3">H. ornatus</emphasis>
</taxonomicName>
by
<bibRefCitation id="F51A4FC6FFCCFFC3CFBCFE671C6A07B0" author="Kudinova-Pasternak" box="[474,784,476,500]" pageId="2" pageNumber="3" refString="Kudinova-Pasternak, R. K. (1976) Additional notes on the fauna of Tanaidacea (Crustacea, Malacostraca) of the deep-water trenches. Trudy Instituta Okeanologii, Akademiya Nauk SSSR, 99, 115 - 135." type="journal article" year="1976">Kudinova-Pasternak (1976)</bibRefCitation>
was also based on a male specimen.
</paragraph>
<paragraph id="91343237FFCCFFC3CEA1FE441C3F04E3" blockId="2.[151,1436,151,679]" pageId="2" pageNumber="3">
If the extent and details of sexual dimorphism in
<taxonomicName id="568B49B4FFCCFFC3CC8AFDBB1CE2045C" box="[748,920,512,536]" class="Malacostraca" family="Heterotanoididae" genus="Heterotanoides" kingdom="Animalia" order="Tanaidacea" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="A3FFEE25FFCCFFC3CC8AFDBB1CE2045C" box="[748,920,512,536]" italics="true" pageId="2" pageNumber="3">Heterotanoides</emphasis>
</taxonomicName>
are confused, it is also possible that the genus is polyphyletic. Of the five species,
<taxonomicName id="568B49B4FFCCFFC3CC41FD9E1DAC0478" box="[551,726,548,572]" class="Malacostraca" family="Heterotanoididae" genus="Heterotanoides" kingdom="Animalia" order="Tanaidacea" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="meridionalis">
<emphasis id="A3FFEE25FFCCFFC3CC41FD9E1DAC0478" box="[551,726,548,572]" italics="true" pageId="2" pageNumber="3">H. meridionalis</emphasis>
</taxonomicName>
appears as an outlier because of the setation of the propodus on pereopods 4–6 and the aesthetasc distribution (
<figureCitation id="09B02EB2FFCCFFC3CCCBFDFC1D8F041B" box="[685,757,583,607]" captionStart="FIGURE 5" captionStartId="8.[151,249,1900,1923]" captionTargetBox="[172,1414,193,1863]" captionTargetId="figure@8.[172,1416,193,1879]" captionTargetPageId="8" captionText="FIGURE 5. Antennular aesthetascs. A, C Nototanoides; B, D Paranesotanais; E, J Heterotanoides meridionalis; F H. ornatus; G H. capensis; H, O H. muimui n. sp.; K, M H. cognatus; L, N Pseudozeuxo. All redrawn from original sources, rescaled, and setae omitted for clarity. Frame encloses nototanaid genera, with separate homology. Antennules E – O are arranged in approximate order of complexity." httpUri="https://zenodo.org/record/213117/files/figure.png" pageId="2" pageNumber="3">Figs 5</figureCitation>
E, J). If the inferences about sexual dimorphism are correct (see above), the female antennule is exceptional in having two bundles of subterminal aesthetascs: one is proximal on article-3, and the more distal bundle has three aesthetascs.
</paragraph>
</subSubSection>
</treatment>
</document>
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