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<document id="3EEFD318E50A02484D68BBE40EBDA364" ID-DOI="10.5281/zenodo.185971" ID-GBIF-Dataset="1db39382-b4f0-46fa-ae08-dd5e57e0e26b" ID-ISSN="1175-5326" ID-Zenodo-Dep="185971" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460034098030" checkinUser="plazi" docAuthor="Pimvichai, Piyatida, Enghoff, Henrik &amp; Panha, Somsak" docDate="2009" docId="1644D538F440FFC1FF49FA279C16FD1E" docLanguage="en" docName="zt02016p050.pdf" docOrigin="Zootaxa 2016" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Thyropygus allevatus" docType="treatment" docVersion="11" lastPageNumber="20" masterDocId="EA7DAD40F442FFC2FFDEFFED9F66FFCD" masterDocTitle="A revision of the Thyropygus allevatus group. Part 1: the T. opinatus subgroup (Diplopoda: Spirostreptida: Harpagophoridae)" masterLastPageNumber="50" masterPageNumber="17" pageNumber="19" updateTime="1698234296060" updateUser="plazi">
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<mods:title id="C6585237C0EE66E484A0686093042DC1">A revision of the Thyropygus allevatus group. Part 1: the T. opinatus subgroup (Diplopoda: Spirostreptida: Harpagophoridae)</mods:title>
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<treatment id="1644D538F440FFC1FF49FA279C16FD1E" ID-DOI="http://doi.org/10.5281/zenodo.6218612" ID-GBIF-Taxon="119382580" ID-Zenodo-Dep="6218612" LSID="urn:lsid:plazi:treatment:1644D538F440FFC1FF49FA279C16FD1E" httpUri="http://treatment.plazi.org/id/1644D538F440FFC1FF49FA279C16FD1E" lastPageId="3" lastPageNumber="20" pageId="2" pageNumber="19">
<subSubSection id="D6F737A5F440FFC0FF49FA279E4DF9B6" pageId="2" pageNumber="19" type="nomenclature">
<paragraph id="9E52642EF440FFC0FF49FA279D44FA29" blockId="2.[151,546,1482,1508]" box="[151,546,1482,1508]" pageId="2" pageNumber="19">
<heading id="C51AD342F440FFC0FF49FA279D44FA29" bold="true" box="[151,546,1482,1508]" fontSize="11" level="1" pageId="2" pageNumber="19" reason="1">
<emphasis id="AC99B83CF440FFC0FF49FA279D44FA29" bold="true" box="[151,546,1482,1508]" pageId="2" pageNumber="19">
The
<taxonomicName id="59ED1FADF440FFC0FF11FA279EB7FA29" ID-CoL="56SYD" box="[207,465,1482,1508]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="2" pageNumber="19" phylum="Arthropoda" rank="species" species="allevatus">
<emphasis id="AC99B83CF440FFC0FF11FA279EB7FA29" bold="true" box="[207,465,1482,1508]" italics="true" pageId="2" pageNumber="19">Thyropygus allevatus</emphasis>
</taxonomicName>
group
</emphasis>
</heading>
</paragraph>
<paragraph id="9E52642EF440FFC0FF49F9FE9E4DF9B6" blockId="2.[151,1437,1555,1813]" pageId="2" pageNumber="19">
As defined by
<bibRefCitation id="FA7C19DFF440FFC0FE92F9FE9D70F9E0" author="Hoffman" box="[332,534,1555,1581]" pageId="2" pageNumber="50" refString="Hoffman, R. L. (1975) Studies on spirostreptoid millipeds. XI. A review of some Indonesian genera of the family Harpagophoridae. Journal of Natural History, 9, 121 - 152." type="journal article" year="1975">Hoffman (1975)</bibRefCitation>
, this group contains
<taxonomicName id="59ED1FADF440FFC0FCC9F9F99CC0F9E0" box="[791,934,1556,1581]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="2" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="AC99B83CF440FFC0FCC9F9F99CC0F9E0" box="[791,934,1556,1581]" italics="true" pageId="2" pageNumber="19">Thyropygus</emphasis>
</taxonomicName>
species in which both tibial and femoral spines are present on the gonopods, the tibial spine being very long and recurved proximad towards the femoral one.
</paragraph>
</subSubSection>
<subSubSection id="D6F737A5F440FFC0FF1BF9659C1BF8D8" pageId="2" pageNumber="19" type="description">
<paragraph id="9E52642EF440FFC0FF1BF9659C1BF8D8" blockId="2.[151,1437,1555,1813]" pageId="2" pageNumber="19">
<bibRefCitation id="FA7C19DFF440FFC0FF1BF9659EF6F96F" author="Hoffman" box="[197,400,1672,1698]" pageId="2" pageNumber="50" refString="Hoffman, R. L. (1975) Studies on spirostreptoid millipeds. XI. A review of some Indonesian genera of the family Harpagophoridae. Journal of Natural History, 9, 121 - 152." type="journal article" year="1975">Hoffman (1975)</bibRefCitation>
referred 11 taxa to this group and suggested that two further species, described in
<taxonomicName id="59ED1FADF440FFC0FF49F9439E4CF90A" box="[151,298,1710,1735]" class="Diplopoda" family="Harpagophoridae" genus="Cornugonus" kingdom="Animalia" order="Spirostreptida" pageId="2" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="AC99B83CF440FFC0FF49F9439E4CF90A" box="[151,298,1710,1735]" italics="true" pageId="2" pageNumber="19">Cornugonus</emphasis>
</taxonomicName>
, might also belong to this group. Considering new species and taxonomic changes since 1975, the following species currently belong to the
<taxonomicName id="59ED1FADF440FFC0FD1AF9389C34F923" box="[708,850,1749,1774]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="2" pageNumber="19" phylum="Arthropoda" rank="species" species="allevatus">
<emphasis id="AC99B83CF440FFC0FD1AF9389C34F923" box="[708,850,1749,1774]" italics="true" pageId="2" pageNumber="19">T. allevatus</emphasis>
</taxonomicName>
group (names given here are as in
<bibRefCitation id="FA7C19DFF440FFC0FADDF9389AFEF922" author="Jeekel" box="[1283,1432,1749,1775]" pageId="2" pageNumber="50" refString="Jeekel, C. A. W. (2006) A bibliographic catalogue of the Oriental Harpagophoridae (Diplopoda, Spirostreptida). Myriapod Memoranda, 9, 5 - 58." type="journal article" year="2006">Jeekel 2006</bibRefCitation>
, except that
<taxonomicName id="59ED1FADF440FFC0FEC0F9119E30F8D8" box="[286,342,1788,1813]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="2" pageNumber="19" phylum="Arthropoda" rank="species" species="carli">
<emphasis id="AC99B83CF440FFC0FEC0F9119E30F8D8" box="[286,342,1788,1813]" italics="true" pageId="2" pageNumber="19">carli</emphasis>
</taxonomicName>
is given full species status, cf.
<bibRefCitation id="FA7C19DFF440FFC0FD1CF9169C17F8D8" author="Enghoff" box="[706,881,1787,1813]" pageId="2" pageNumber="50" refString="Enghoff, H. (2005) The millipedes of Thailand (Diplopoda). Steenstrupia, 29 (1), 87 - 103." type="journal article" year="2005">Enghoff, 2005</bibRefCitation>
):
</paragraph>
</subSubSection>
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<paragraph id="9E52642EF440FFC0FF49F8A49D65F847" blockId="2.[151,584,1865,2007]" pageId="2" pageNumber="19">
<taxonomicName id="59ED1FADF440FFC0FF49F8A49D2EF8AE" ID-CoL="56SYD" authority="Karsch, 1881" authorityName="Karsch" authorityYear="1881" box="[151,584,1865,1891]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="2" pageNumber="19" phylum="Arthropoda" rank="species" species="allevatus">
<emphasis id="AC99B83CF440FFC0FF49F8A49EF7F8AF" box="[151,401,1865,1890]" italics="true" pageId="2" pageNumber="19">Thyropygus allevatus</emphasis>
(
<bibRefCitation id="FA7C19DFF440FFC0FE7EF8A49D59F8AE" author="Karsch" box="[416,575,1865,1891]" pageId="2" pageNumber="50" refString="Karsch, F. (1881) Neue Juliden des Berliner Museums, als Prodromus einer Juliden-Monographie. Zeitschrift fur die Gesamten Naturwissenschaften, Dritte Folge, 6, 3 - 79." type="journal article" year="1881">Karsch, 1881</bibRefCitation>
)
</taxonomicName>
<taxonomicName id="59ED1FADF440FFC0FF49F89D9D65F847" ID-CoL="56SZ6" authority="Attems, 1938" authorityName="Attems" authorityYear="1938" box="[151,515,1904,1930]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="2" pageNumber="19" phylum="Arthropoda" rank="species" species="carli">
<emphasis id="AC99B83CF440FFC0FF49F89D9E39F844" box="[151,351,1904,1929]" italics="true" pageId="2" pageNumber="19">Thyropygus carli</emphasis>
<bibRefCitation id="FA7C19DFF440FFC0FEB8F89D9D65F847" author="Attems" box="[358,515,1904,1930]" pageId="2" pageNumber="50" refString="Attems, C. (1938) Die von Dr. C. Dawydoff in Franzosisch Indochina gesammelten Myriopoden. Memoires du Museum National d'Histoire Naturelle, N. S. 6, 187 - 321." type="book chapter" year="1938">Attems, 1938</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="9E52642EF440FFC0FF49F87B9D7FF81A" blockId="2.[151,584,1865,2007]" pageId="2" pageNumber="19">
<taxonomicName id="59ED1FADF440FFC0FF49F87B9D55F87D" ID-CoL="7BZC6" authority="Attems, 1938" authorityName="Attems" authorityYear="1938" box="[151,563,1942,1968]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="2" pageNumber="19" phylum="Arthropoda" rank="species" species="confusus">
<emphasis id="AC99B83CF440FFC0FF49F87B9EE8F862" box="[151,398,1942,1967]" italics="true" pageId="2" pageNumber="19">Thyropygus confusus</emphasis>
<bibRefCitation id="FA7C19DFF440FFC0FE4BF87B9D55F87D" author="Attems" box="[405,563,1942,1968]" pageId="2" pageNumber="50" refString="Attems, C. (1938) Die von Dr. C. Dawydoff in Franzosisch Indochina gesammelten Myriopoden. Memoires du Museum National d'Histoire Naturelle, N. S. 6, 187 - 321." type="book chapter" year="1938">Attems, 1938</bibRefCitation>
</taxonomicName>
<taxonomicName id="59ED1FADF440FFC0FF49F8509D7FF81A" ID-CoL="56SZL" authority="Carl, 1917" authorityName="Carl" authorityYear="1917" box="[151,537,1981,2007]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="2" pageNumber="19" phylum="Arthropoda" rank="species" species="cuisinieri">
<emphasis id="AC99B83CF440FFC0FF49F8509EF0F81B" box="[151,406,1981,2006]" italics="true" pageId="2" pageNumber="19">Thyropygus cuisinieri</emphasis>
Carl, 1917
</taxonomicName>
</paragraph>
<paragraph id="9E52642EF441FFC1FF49FF759D2AFF32" blockId="3.[151,617,152,487]" pageId="3" pageNumber="20">
<taxonomicName id="59ED1FADF441FFC1FF49FF759D06FF7F" ID-CoL="56T2B" authority="Demange, 1989" authorityName="Demange" authorityYear="1989" box="[151,608,152,178]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="3" pageNumber="20" phylum="Arthropoda" rank="species" species="globulus">
<emphasis id="AC99B83CF441FFC1FF49FF759EEAFF7C" box="[151,396,152,177]" italics="true" pageId="3" pageNumber="20">Thyropygus globulus</emphasis>
(
<bibRefCitation id="FA7C19DFF441FFC1FE42FF759D3EFF7F" author="Demange" box="[412,600,152,178]" pageId="3" pageNumber="50" refString="Demange, J. M. (1989) Sur quelques Harpagophoridae du Sud-Est asiatique et de l'Inde (Myriapoda, Diplopoda, Spirostreptoidea). Bulletin du Museum National d'Histoire Naturelle, Paris, 4 e ser., 11 sect. A, 773 - 781." type="journal article" year="1989">Demange, 1989</bibRefCitation>
)
</taxonomicName>
<taxonomicName id="59ED1FADF441FFC1FF49FF529D02FF15" ID-CoL="7BZBS" authority="Demange, 1961" authorityName="Demange" authorityYear="1961" box="[151,612,190,216]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="3" pageNumber="20" phylum="Arthropoda" rank="species" species="hoffmani">
<emphasis id="AC99B83CF441FFC1FF49FF529EE9FF15" box="[151,399,191,216]" italics="true" pageId="3" pageNumber="20">Thyropygus hoffmani</emphasis>
(
<bibRefCitation id="FA7C19DFF441FFC1FE41FF539D3DFF15" author="Demange" box="[415,603,190,216]" pageId="3" pageNumber="50" refString="Demange, J. M. (1961) Materiaux pour servir a une revision des Harpagophoridae. Memoires du Museum National d'Histoire Naturelle, Ser. A (Zool.), 24, 1 - 274." type="journal article" year="1961">Demange, 1961</bibRefCitation>
)
</taxonomicName>
<taxonomicName id="59ED1FADF441FFC1FF49FF089D2AFF32" ID-CoL="56T2X" authority="Demange, 1961" authorityName="Demange" authorityYear="1961" box="[151,588,229,255]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="3" pageNumber="20" phylum="Arthropoda" rank="species" species="ligulus">
<emphasis id="AC99B83CF441FFC1FF49FF089E11FF33" box="[151,375,229,254]" italics="true" pageId="3" pageNumber="20">Thyropygus ligulus</emphasis>
(
<bibRefCitation id="FA7C19DFF441FFC1FE58FF089D25FF32" author="Demange" box="[390,579,229,255]" pageId="3" pageNumber="50" refString="Demange, J. M. (1961) Materiaux pour servir a une revision des Harpagophoridae. Memoires du Museum National d'Histoire Naturelle, Ser. A (Zool.), 24, 1 - 274." type="journal article" year="1961">Demange, 1961</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="9E52642EF441FFC1FF49FEE19D24FEBE" blockId="3.[151,617,152,487]" pageId="3" pageNumber="20">
<taxonomicName id="59ED1FADF441FFC1FF49FEE19D04FEEB" ID-CoL="YGD8" authority="Demange, 1986" authorityName="Demange" authorityYear="1986" box="[151,610,268,294]" class="Diplopoda" family="Harpagophoridae" genus="Cornugonus" kingdom="Animalia" order="Spirostreptida" pageId="3" pageNumber="20" phylum="Arthropoda" rank="species" species="bifurcus">
<emphasis id="AC99B83CF441FFC1FF49FEE19EE8FEE8" box="[151,398,268,293]" italics="true" pageId="3" pageNumber="20">Cornugonus bifurcus</emphasis>
(
<bibRefCitation id="FA7C19DFF441FFC1FE43FEE19D3CFEEB" author="Demange" box="[413,602,268,294]" pageId="3" pageNumber="50" refString="Demange, J. M. (1986) Harpagophoridae de Thailande et de Malaisie (Myriapoda, Spirostreptida). Bulletin du Museum National d'Histoire Naturelle, Paris, 4 e ser., 8 sect. A, 851 - 865." type="journal article" year="1986">Demange, 1986</bibRefCitation>
)
</taxonomicName>
<taxonomicName id="59ED1FADF441FFC1FF49FEDF9D05FE81" ID-CoL="YGD9" authority="Demange, 1989" authorityName="Demange" authorityYear="1989" box="[151,611,306,332]" class="Diplopoda" family="Harpagophoridae" genus="Cornugonus" kingdom="Animalia" order="Spirostreptida" pageId="3" pageNumber="20" phylum="Arthropoda" rank="species" species="enghoffi">
<emphasis id="AC99B83CF441FFC1FF49FEDF9EE9FE86" box="[151,399,306,331]" italics="true" pageId="3" pageNumber="20">Cornugonus enghoffi</emphasis>
(
<bibRefCitation id="FA7C19DFF441FFC1FE40FEDF9D3DFE81" author="Demange" box="[414,603,306,332]" pageId="3" pageNumber="50" refString="Demange, J. M. (1989) Sur quelques Harpagophoridae du Sud-Est asiatique et de l'Inde (Myriapoda, Diplopoda, Spirostreptoidea). Bulletin du Museum National d'Histoire Naturelle, Paris, 4 e ser., 11 sect. A, 773 - 781." type="journal article" year="1989">Demange, 1989</bibRefCitation>
)
</taxonomicName>
<taxonomicName id="59ED1FADF441FFC1FF49FEB49D24FEBE" ID-CoL="6B4JQ" authority="Demange, 1961" authorityName="Demange" authorityYear="1961" box="[151,578,345,371]" class="Diplopoda" family="Harpagophoridae" genus="Cornugonus" kingdom="Animalia" order="Spirostreptida" pageId="3" pageNumber="20" phylum="Arthropoda" rank="species" species="floweri">
<emphasis id="AC99B83CF441FFC1FF49FEB49EE4FEBF" box="[151,386,345,370]" italics="true" pageId="3" pageNumber="20">Cornugonus floweri</emphasis>
<bibRefCitation id="FA7C19DFF441FFC1FE57FEB49D24FEBE" author="Demange" box="[393,578,345,371]" pageId="3" pageNumber="50" refString="Demange, J. M. (1961) Materiaux pour servir a une revision des Harpagophoridae. Memoires du Museum National d'Histoire Naturelle, Ser. A (Zool.), 24, 1 - 274." type="journal article" year="1961">Demange, 1961</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="9E52642EF441FFC1FF49FE6D9D2DFE0D" blockId="3.[151,617,152,487]" pageId="3" pageNumber="20">
<taxonomicName id="59ED1FADF441FFC1FF49FE6D9D0FFE57" ID-CoL="YGDB" authority="Demange, 1961" authorityName="Demange" authorityYear="1961" box="[151,617,384,410]" class="Diplopoda" family="Harpagophoridae" genus="Cornugonus" kingdom="Animalia" order="Spirostreptida" pageId="3" pageNumber="20" phylum="Arthropoda" rank="species" species="implicatus">
<emphasis id="AC99B83CF441FFC1FF49FE6D9ECFFE54" box="[151,425,384,409]" italics="true" pageId="3" pageNumber="20">Cornugonus implicatus</emphasis>
<bibRefCitation id="FA7C19DFF441FFC1FE6EFE6D9D0FFE57" author="Demange" box="[432,617,384,410]" pageId="3" pageNumber="50" refString="Demange, J. M. (1961) Materiaux pour servir a une revision des Harpagophoridae. Memoires du Museum National d'Histoire Naturelle, Ser. A (Zool.), 24, 1 - 274." type="journal article" year="1961">Demange, 1961</bibRefCitation>
</taxonomicName>
<taxonomicName id="59ED1FADF441FFC1FF49FE4A9D2DFE0D" ID-CoL="YGDC" authority="Demange, 1989" authorityName="Demange" authorityYear="1989" box="[151,587,422,448]" class="Diplopoda" family="Harpagophoridae" genus="Cornugonus" kingdom="Animalia" order="Spirostreptida" pageId="3" pageNumber="20" phylum="Arthropoda" rank="species" species="inflexus">
<emphasis id="AC99B83CF441FFC1FF49FE4A9EECFE0D" box="[151,394,423,448]" italics="true" pageId="3" pageNumber="20">Cornugonus inflexus</emphasis>
<bibRefCitation id="FA7C19DFF441FFC1FE4CFE4B9D2DFE0D" author="Demange" box="[402,587,422,448]" pageId="3" pageNumber="50" refString="Demange, J. M. (1989) Sur quelques Harpagophoridae du Sud-Est asiatique et de l'Inde (Myriapoda, Diplopoda, Spirostreptoidea). Bulletin du Museum National d'Histoire Naturelle, Paris, 4 e ser., 11 sect. A, 773 - 781." type="journal article" year="1989">Demange, 1989</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="9E52642EF441FFC1FF49FE209D2DFE2A" blockId="3.[151,617,152,487]" box="[151,587,461,487]" pageId="3" pageNumber="20">
<taxonomicName id="59ED1FADF441FFC1FF49FE209D2DFE2A" ID-CoL="YGDD" authority="Karsch, 1881" authorityName="Karsch" authorityYear="1881" box="[151,587,461,487]" class="Diplopoda" family="Harpagophoridae" genus="Cornugonus" kingdom="Animalia" order="Spirostreptida" pageId="3" pageNumber="20" phylum="Arthropoda" rank="species" species="opinatus">
<emphasis id="AC99B83CF441FFC1FF49FE209EF2FE2B" box="[151,404,461,486]" italics="true" pageId="3" pageNumber="20">Cornugonus opinatus</emphasis>
(
<bibRefCitation id="FA7C19DFF441FFC1FE7DFE209D25FE2A" author="Karsch" box="[419,579,461,487]" pageId="3" pageNumber="50" refString="Karsch, F. (1881) Neue Juliden des Berliner Museums, als Prodromus einer Juliden-Monographie. Zeitschrift fur die Gesamten Naturwissenschaften, Dritte Folge, 6, 3 - 79." type="journal article" year="1881">Karsch, 1881</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="9E52642EF441FFC1FF18FDF29C16FD1E" blockId="3.[151,1436,542,723]" pageId="3" pageNumber="20">
<taxonomicName id="59ED1FADF441FFC1FF18FDF29DE7FDF5" ID-CoL="56SYD" authority="Karsch, 1881" authorityName="Karsch" authorityYear="1881" box="[198,641,542,568]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="3" pageNumber="20" phylum="Arthropoda" rank="species" species="allevatus">
<emphasis id="AC99B83CF441FFC1FF18FDF29EA5FDF5" box="[198,451,543,568]" italics="true" pageId="3" pageNumber="20">Thyropygus allevatus</emphasis>
(
<bibRefCitation id="FA7C19DFF441FFC1FE0BFDF39D1EFDF5" author="Karsch" box="[469,632,542,568]" pageId="3" pageNumber="50" refString="Karsch, F. (1881) Neue Juliden des Berliner Museums, als Prodromus einer Juliden-Monographie. Zeitschrift fur die Gesamten Naturwissenschaften, Dritte Folge, 6, 3 - 79." type="journal article" year="1881">Karsch, 1881</bibRefCitation>
)
</taxonomicName>
, the
<typeStatus id="4156DA8CF441FFC1FD1CFDF39D9FFDF5" box="[706,761,542,568]" pageId="3" pageNumber="20">type</typeStatus>
species of the
<taxonomicName id="59ED1FADF441FFC1FC65FDF29B24FDF5" ID-CoL="56SYD" box="[955,1090,543,568]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="3" pageNumber="20" phylum="Arthropoda" rank="species" species="allevatus">
<emphasis id="AC99B83CF441FFC1FC65FDF29B24FDF5" box="[955,1090,543,568]" italics="true" pageId="3" pageNumber="20">T. allevatus</emphasis>
</taxonomicName>
group, is a common species recorded from all over
<collectingCountry id="E6FA24BEF441FFC1FE79FDA89D69FD92" box="[423,527,581,607]" name="Thailand" pageId="3" pageNumber="20">Thailand</collectingCountry>
and also from
<collectingCountry id="E6FA24BEF441FFC1FD1FFDA89C4EFD92" box="[705,808,581,607]" name="Vietnam" pageId="3" pageNumber="20">Vietnam</collectingCountry>
,
<collectingCountry id="E6FA24BEF441FFC1FCEAFDA89C16FD92" box="[820,880,581,607]" name="Laos" pageId="3" pageNumber="20">Laos</collectingCountry>
,
<collectingCountry id="E6FA24BEF441FFC1FCA2FDA89C92FD92" box="[892,1012,581,607]" name="Cambodia" pageId="3" pageNumber="20">Cambodia</collectingCountry>
and continental
<collectingCountry id="E6FA24BEF441FFC1FB66FDA89A45FD92" box="[1208,1315,581,607]" name="Malaysia" pageId="3" pageNumber="20">Malaysia</collectingCountry>
(
<bibRefCitation id="FA7C19DFF441FFC1FAEDFDA89FBEFD48" author="Enghoff" pageId="3" pageNumber="50" refString="Enghoff, H. (2005) The millipedes of Thailand (Diplopoda). Steenstrupia, 29 (1), 87 - 103." type="journal article" year="2005">Enghoff, 2005</bibRefCitation>
). However, it still presents many taxonomic problems. There are many synonyms of
<taxonomicName id="59ED1FADF441FFC1FB07FD819A05FD48" ID-CoL="56SYD" box="[1241,1379,620,645]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="3" pageNumber="20" phylum="Arthropoda" rank="species" species="allevatus">
<emphasis id="AC99B83CF441FFC1FB07FD819A05FD48" box="[1241,1379,620,645]" italics="true" pageId="3" pageNumber="20">T. allevatus</emphasis>
</taxonomicName>
(see
<bibRefCitation id="FA7C19DFF441FFC1FF49FD7F9E58FD61" author="Enghoff" box="[151,318,658,684]" pageId="3" pageNumber="50" refString="Enghoff, H. (2005) The millipedes of Thailand (Diplopoda). Steenstrupia, 29 (1), 87 - 103." type="journal article" year="2005">Enghoff 2005</bibRefCitation>
), and several subspecies have been recognized, but the considerable variation in both somatic characters and gonopods still needs a comprehensive analysis.
</paragraph>
</subSubSection>
</treatment>
</document>