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<document id="9DBFF8A10250FF800E673F7E8D241637" ID-DOI="10.5281/zenodo.5404276" ID-ISSN="0374-1036" ID-Zenodo-Dep="5404276" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630132078781" checkinUser="felipe" docAuthor="Hromádka, Lubomír" docDate="2009" docId="142187F5FFD7FF9DF5E0FE27FC35FD54" docLanguage="en" docName="ActaEntMusNatPra.2009.49.1.161-190.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 49 (1)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Philonthus iridicollis M. Bernhauer 1932" docType="treatment" docVersion="4" lastPageNumber="179" masterDocId="E818FF8DFFC6FF8FF464FFBCFFA1FFC7" masterDocTitle="Revision of the Afrotropical species of the Philonthus caffer species group (Coleoptera: Staphylinidae: Philonthina)" masterLastPageNumber="190" masterPageNumber="161" pageNumber="178" updateTime="1699162834734" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="879BA45CD116D331B5AEC66FF392FECB">Revision of the Afrotropical species of the Philonthus caffer species group (Coleoptera: Staphylinidae: Philonthina)</mods:title>
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<mods:namePart id="D4E26D6375B044C08E5B6149BADAEE11">Hromádka, Lubomír</mods:namePart>
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<mods:title id="E3F76E8C1565592E0FCAFF823DA29C0A">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:part id="F1D5C7296566DC1AC1CF8B4ECE1536E2">
<mods:date id="0803475681D5DD15AE3FDE9D1517DC10">2009</mods:date>
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<mods:number id="CF72BF54319DE2549EDB083E158266DC">49</mods:number>
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<mods:identifier id="4BC36868A860070CE8BD024BBC6EFB7E" type="ISSN">0374-1036</mods:identifier>
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<treatment id="142187F5FFD7FF9DF5E0FE27FC35FD54" ID-DOI="http://doi.org/10.5281/zenodo.5337810" ID-Zenodo-Dep="5337810" LSID="urn:lsid:plazi:treatment:142187F5FFD7FF9DF5E0FE27FC35FD54" httpUri="http://treatment.plazi.org/id/142187F5FFD7FF9DF5E0FE27FC35FD54" lastPageId="18" lastPageNumber="179" pageId="17" pageNumber="178">
<subSubSection id="D4926568FFD7FF9EF5E0FE27FC8CFE74" box="[388,813,411,436]" pageId="17" pageNumber="178" type="nomenclature">
<paragraph id="9C3736E3FFD7FF9EF5E0FE27FC8CFE74" blockId="17.[388,813,411,436]" box="[388,813,411,436]" pageId="17" pageNumber="178">
<heading id="C77F818FFFD7FF9EF5E0FE27FC8CFE74" bold="true" box="[388,813,411,436]" centered="true" fontSize="8" level="2" pageId="17" pageNumber="178" reason="3">
<taxonomicName id="5B884D60FFD7FF9EF5E0FE27FC8CFE74" ID-CoL="4GB27" authority="Bernhauer, 1932" authorityName="M. Bernhauer" authorityYear="1932" box="[388,813,411,436]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="178" phylum="Arthropoda" rank="species" species="iridicollis">
<emphasis id="AEFCEAF1FFD7FF9EF5E0FE27FC8CFE74" bold="true" box="[388,813,411,436]" pageId="17" pageNumber="178">
<emphasis id="AEFCEAF1FFD7FF9EF5E0FE27FDCDFE73" bold="true" box="[388,620,411,436]" italics="true" pageId="17" pageNumber="178">Philonthus iridicollis</emphasis>
<bibRefCitation id="F8194B12FFD7FF9EF617FE20FC8CFE74" author="BERNHAUER M." box="[627,813,411,436]" pageId="17" pageNumber="178" pagination="140 - 174" refId="ref15986" refString="BERNHAUER M. 1932: Neue Kurzflugler aus dem belgischen Kongostaate. Revue de Zoologie et de Botanique Africaines 22: 140 - 174." type="journal article" year="1932">Bernhauer, 1932</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="D4926568FFD7FF9EF67BFE06FD32FE0A" box="[543,659,442,461]" pageId="17" pageNumber="178" type="description">
<paragraph id="9C3736E3FFD7FF9EF67BFE06FD32FE0A" blockId="17.[113,659,442,495]" box="[543,659,442,461]" pageId="17" pageNumber="178">
(
<figureCitation id="04B32A66FFD7FF9EF642FE06FD2DFE0A" box="[550,652,442,461]" captionStart="Figs" captionStartId="20.[151,191,1474,1493]" captionTargetBox="[202,1071,219,1393]" captionTargetId="figure-10@20.[202,1078,210,1393]" captionTargetPageId="20" captionText="Figs. 3853. 3844 Philonthus iridicollis Bernhauer, 1932; 4548 P. kenyanus Bernhauer, 1939; 4953 P. leptophilus Hromádka, 2006. 38, 45, 49 aedeagus, ventral view; 39, 46, 50 aedeagus, lateral view; 40, 51 paramere with sensory peg setae, ventral view; 41, 52 apical portion of male sternite VIII, ventral view; 42, 48, 53 male sternite IX, ventral view; 43 female tergite X, ventral view; 44 gonocoxite of female genital segment; 47 aedeagus without paramere, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5404284" httpUri="https://zenodo.org/record/5404284/files/figure.png" pageId="17" pageNumber="178">Figs. 3844</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="D4926568FFD7FF9EF415FE61FE4AFE28" box="[113,491,476,495]" pageId="17" pageNumber="178" type="reference_group">
<paragraph id="9C3736E3FFD7FF9EF415FE61FE4AFE28" blockId="17.[113,659,442,495]" box="[113,491,476,495]" pageId="17" pageNumber="178">
<treatmentCitationGroup id="BC9811CDFFD7FF9EF415FE61FE4AFE28" box="[113,491,476,495]" pageId="17" pageNumber="178">
<treatmentCitation id="1D2910F2FFD7FF9EF415FE61FE46FE28" author="BERNHAUER M." box="[113,487,476,495]" page="152" pageId="17" pageNumber="178" year="1932">
<taxonomicName id="5B884D60FFD7FF9EF415FE61FE46FE28" ID-CoL="4GB27" authority="Bernhauer, 1932: 152" authorityName="M. Bernhauer" authorityPageNumber="152" authorityYear="1932" box="[113,487,476,495]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="178" phylum="Arthropoda" rank="species" species="iridicollis">
<emphasis id="AEFCEAF1FFD7FF9EF415FE61FE87FE28" box="[113,294,477,495]" italics="true" pageId="17" pageNumber="178">Philonthus iridicollis</emphasis>
<bibRefCitation id="F8194B12FFD7FF9EF54FFE60FE46FE28" author="BERNHAUER M." box="[299,487,476,495]" pageId="17" pageNumber="178" pagination="140 - 174" refId="ref15986" refString="BERNHAUER M. 1932: Neue Kurzflugler aus dem belgischen Kongostaate. Revue de Zoologie et de Botanique Africaines 22: 140 - 174." type="journal article" year="1932">Bernhauer, 1932: 152</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="D4926568FFD7FF9EF415FDAEFC71FD6D" pageId="17" pageNumber="178" type="materials_examined">
<paragraph id="9C3736E3FFD7FF9EF415FDAEFD59FDE3" blockId="17.[113,1090,529,682]" box="[113,760,529,549]" pageId="17" pageNumber="178">
<emphasis id="AEFCEAF1FFD7FF9EF415FDAEFF4BFDE2" bold="true" box="[113,234,529,549]" pageId="17" pageNumber="178">
<typeStatus id="43338841FFD7FF9EF415FDAEFF3FFDE2" box="[113,158,530,549]" pageId="17" pageNumber="178">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="E49F7673FFD7FF9EF48BFDADFDBDFDE2" box="[239,540,529,549]" name="Democratic Republic of the Congo" pageId="17" pageNumber="178">Democratic Republic of the Congo</collectingCountry>
, Moreo, Kiambi, Niunzu.
</paragraph>
<paragraph id="9C3736E3FFD7FF9EF415FD90FCC6FD9D" blockId="17.[113,1090,529,682]" pageId="17" pageNumber="178">
<emphasis id="AEFCEAF1FFD7FF9EF415FD90FF53FDF8" bold="true" box="[113,242,555,575]" pageId="17" pageNumber="178">Type material.</emphasis>
<materialsCitation id="2CE03CBEFFD7FF9EF492FD91FCC6FD9D" collectingDate="1930-06" collectingDateMax="1930-07-31" collectingDateMin="1930-06" collectionCode="FMNH" collectorName="Moreo &amp; Kiambi &amp; Dr. P. Gerard &amp; Chicago &amp; M. Bernhauer Collection &amp; Type" pageId="17" pageNumber="178" specimenCount="1" typeStatus="holotype">
<typeStatus id="43338841FFD7FF9EF492FD91FEE9FDF9" box="[246,328,557,576]" pageId="17" pageNumber="178" type="holotype">HOLOTYPE</typeStatus>
: J,
<collectorName id="317D5335FFD7FF9EF515FD91FE0AFD87" box="[369,427,557,576]" pageId="17" pageNumber="178">Moreo</collectorName>
,
<collectorName id="317D5335FFD7FF9EF5D6FD90FE52FDF8" box="[434,499,556,575]" pageId="17" pageNumber="178">Kiambi</collectorName>
,
<date id="E8361023FFD7FF9EF59DFD90FDFCFDF8" box="[505,605,556,575]" pageId="17" pageNumber="178" value="1930-06" valueMax="1930-07-31" valueMin="1930-06-00">
<collectingDate id="F872E9CBFFD7FF9EF59DFD90FDFCFDF8" box="[505,605,556,575]" pageId="17" pageNumber="178" value="1930-06" valueMax="1930-07-31" valueMin="1930-06-00">vi.-vii.1930</collectingDate>
</date>
,
<collectorName id="317D5335FFD7FF9EF600FD91FD71FDF8" box="[612,720,556,576]" pageId="17" pageNumber="178">Dr. P.Gérard</collectorName>
,
<collectorName id="317D5335FFD7FF9EF6B3FD90FCBCFDF8" box="[727,797,556,575]" pageId="17" pageNumber="178">Chicago</collectorName>
NHMus.
<collectorName id="317D5335FFD7FF9EF709FD91FB9FFDF8" box="[877,1086,556,576]" pageId="17" pageNumber="178">M.Bernhauer Collection</collectorName>
, //
<collectorName id="317D5335FFD7FF9EF4E6FDFBFF0EFD9D" box="[130,175,583,602]" pageId="17" pageNumber="178">Type</collectorName>
,
<taxonomicName id="5B884D60FFD7FF9EF4D3FDF4FE49FD9D" authority="M. Bernhauer" authorityName="M. Bernhauer" authorityYear="1932" box="[183,488,583,602]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="178" phylum="Arthropoda" rank="species" species="iridicollis">
<emphasis id="AEFCEAF1FFD7FF9EF4D3FDF4FECDFD9D" box="[183,364,584,602]" italics="true" pageId="17" pageNumber="178">Philonthus iridicollis</emphasis>
M. Bernhauer
</taxonomicName>
[ochre oblong label, handwritten] (
<collectionCode id="FA99AE26FFD7FF9EF77FFDFBFCFFFD9D" box="[795,862,583,602]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="17" pageNumber="178" type="Museum">FMNH</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="9C3736E3FFD7FF9EF415FDDDFC71FD6D" blockId="17.[113,1090,529,682]" pageId="17" pageNumber="178">
<emphasis id="AEFCEAF1FFD7FF9EF415FDDDFE2BFDB2" bold="true" box="[113,394,609,629]" pageId="17" pageNumber="178">Additional material examined.</emphasis>
<materialsCitation id="2CE03CBEFFD7FF9EF5F4FDDDFE2BFD48" collectionCode="BMNH, LHPC" collectorName="Jimma Kaffa, Ft. &amp; R. O. S. Clark" country="Ethiopia" location="ETHIOPIA" pageId="17" pageNumber="178" specimenCount="5" specimenCount-female="5">
<emphasis id="AEFCEAF1FFD7FF9EF5F4FDDDFDA3FDB2" bold="true" box="[400,514,609,629]" pageId="17" pageNumber="178">
<collectingCountry id="E49F7673FFD7FF9EF5F4FDDDFE5FFDB2" box="[400,510,609,629]" name="Ethiopia" pageId="17" pageNumber="178">ETHIOPIA</collectingCountry>
:
</emphasis>
4 JJ
<specimenCount id="8A8EFD6AFFD7FF9EF65AFDDDFDCFFDB3" box="[574,622,603,628]" pageId="17" pageNumber="178" type="female">5 ♀♀</specimenCount>
,
<collectorName id="317D5335FFD7FF9EF619FDDDFCB4FDB2" box="[637,789,609,629]" pageId="17" pageNumber="178">Jimma Kaffa, Ft.</collectorName>
1971, grass cuttings,
<collectorName id="317D5335FFD7FF9EF7B5FDDEFBE1FDB3" box="[977,1088,609,629]" pageId="17" pageNumber="178">R.O.S. Clark</collectorName>
B.M. 1973-450 (
<collectionCode id="FA99AE26FFD7FF9EF564FDC0FEE2FD48" box="[256,323,636,655]" pageId="17" pageNumber="178">BMNH</collectionCode>
,
<collectionCode id="FA99AE26FFD7FF9EF52EFDC0FE25FD48" box="[330,388,636,655]" pageId="17" pageNumber="178">LHPC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="2CE03CBEFFD7FF9EF5F5FDC0FF63FD6D" collectingDate="1957-01-15" collectionCode="BMNH" collectorName="Bamenda, V. F &amp; C. E. Tottenham &amp; C. H. BOHEMAN" country="Ethiopia" pageId="17" pageNumber="178" specimenCount="1" specimenCount-female="1">
<specimenCount id="8A8EFD6AFFD7FF9EF5F5FDC0FE0FFD48" box="[401,430,630,655]" pageId="17" pageNumber="178" type="female">1♀</specimenCount>
,
<collectorName id="317D5335FFD7FF9EF5DFFDC0FD97FD57" box="[443,566,636,656]" pageId="17" pageNumber="178">Bamenda,V.F</collectorName>
.Easrop,
<date id="E8361023FFD7FF9EF6E4FDC0FD70FD48" box="[640,721,636,655]" pageId="17" pageNumber="178" value="1957-01-15">
<collectingDate id="F872E9CBFFD7FF9EF6E4FDC0FD70FD48" box="[640,721,636,655]" pageId="17" pageNumber="178" value="1957-01-15">15.i.1957</collectingDate>
</date>
,
<collectorName id="317D5335FFD7FF9EF6BCFDC0FCFAFD48" box="[728,859,636,655]" pageId="17" pageNumber="178">C.E.Tottenham</collectorName>
collection,
<collectorName id="317D5335FFD7FF9EF7DFFDC1FC49FD57" box="[955,1000,637,656]" name="C. H. BOHEMAN" pageId="17" pageNumber="178">B.M.</collectorName>
1974-587 (
<collectionCode id="FA99AE26FFD7FF9EF413FD2BFF1DFD6D" box="[119,188,663,682]" pageId="17" pageNumber="178">BMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="2CE03CBEFFD7FF9EF4AEFD2AFC6DFD6D" box="[202,972,657,682]" collectingDate="1997-04-18" collectionCode="LHPC" collectorName="Kutsaga &amp; W. Rossi" country="Zimbabwe" location="Harare" pageId="17" pageNumber="178" specimenCount="1" specimenCount-female="1" stateProvince="Harare">
<emphasis id="AEFCEAF1FFD7FF9EF4AEFD2AFEEDFD6D" bold="true" box="[202,332,662,682]" pageId="17" pageNumber="178">
<collectingCountry id="E49F7673FFD7FF9EF4AEFD2AFEE6FD6D" box="[202,327,662,682]" name="Zimbabwe" pageId="17" pageNumber="178">ZIMBABWE</collectingCountry>
:
</emphasis>
<specimenCount id="8A8EFD6AFFD7FF9EF535FD2BFED0FD6D" box="[337,369,657,682]" pageId="17" pageNumber="178" type="female">1 ♀</specimenCount>
, loc.,
<collectorName id="317D5335FFD7FF9EF5C8FD2BFE53FD6D" box="[428,498,663,682]" pageId="17" pageNumber="178">Kutsaga</collectorName>
near
<collectingRegion id="5E4CF801FFD7FF9EF646FD2BFDFDFD6D" box="[546,604,663,682]" country="Zimbabwe" name="Harare" pageId="17" pageNumber="178">Harare</collectingRegion>
airport,
<date id="E8361023FFD7FF9EF6C0FD2BFCA0FD6D" box="[676,769,663,682]" pageId="17" pageNumber="178" value="1997-04-18">
<collectingDate id="F872E9CBFFD7FF9EF6C0FD2BFCA0FD6D" box="[676,769,663,682]" pageId="17" pageNumber="178" value="1997-04-18">18.iv.1997</collectingDate>
</date>
,
<collectorName id="317D5335FFD7FF9EF76DFD2BFCF5FD6D" box="[777,852,663,682]" pageId="17" pageNumber="178">W. Rossi</collectorName>
leg. (
<collectionCode id="FA99AE26FFD7FF9EF7EEFD2BFC64FD6D" box="[906,965,663,682]" pageId="17" pageNumber="178">LHPC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="D4926568FFD7FF9DF415FD7EFF5DFDD3" lastPageId="18" lastPageNumber="179" pageId="17" pageNumber="178" type="description">
<paragraph id="9C3736E3FFD7FF9EF415FD7EFF79FD3D" blockId="17.[113,1090,706,1562]" pageId="17" pageNumber="178">
<emphasis id="AEFCEAF1FFD7FF9EF415FD7EFEB4FD1D" bold="true" box="[113,277,706,730]" pageId="17" pageNumber="178">Redescription</emphasis>
. Body length
<quantity id="5B709B06FFD7FF9EF5D9FD7EFDE6FD1D" box="[445,583,706,730]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="9.1" metricValueMin="8.9" pageId="17" pageNumber="178" unit="mm" value="9.0" valueMax="9.1" valueMin="8.9">8.99.1 mm</quantity>
, length of fore body (to end of elytra) 4.0
<quantity id="5B709B06FFD7FF9EF41BFD5EFF75FD3D" box="[127,212,738,762]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="17" pageNumber="178" unit="mm" value="4.2">4.2 mm</quantity>
.
</paragraph>
<paragraph id="9C3736E3FFD7FF9EF4F4FCBEFCEEFC5D" blockId="17.[113,1090,706,1562]" pageId="17" pageNumber="178">Colouration. Head and abdomen black, with very slight greenish-violet hue around eyes, antennal sockets and clypeus along anterior margin very narrowly yellow-brown, maxillary and labial palpi yellow-brown, antennae black, base of antennomere 2 dark brown, pronotum black with distinct green-violet hue, scutellum and elytra black, legs black-brown, protarsomeres 15 yellow-brown, middle and posterior tarsi somewhat paler.</paragraph>
<paragraph id="9C3736E3FFD7FF9EF4F4FC1EFC11FB9D" blockId="17.[113,1090,706,1562]" pageId="17" pageNumber="178">Head quadrangular, slightly wider than long (ratio 31: 27), slightly widened posteriad. Posterior angles obtusely rounded, each with one long and several short bristles. Distance between medial interocular punctures about 3.5 times as large as distance between medial and lateral interocular puncture. Eyes flat, somewhat shorter than temples (ratio 10: 12). Posterior margin of each eye with three coarse punctures. Temporal area with several coarse punctures. Dorsal surface without microsculpture, only with micropunctation.</paragraph>
<paragraph id="9C3736E3FFD7FF9EF4F4FBDEFCAFFB7D" blockId="17.[113,1090,706,1562]" pageId="17" pageNumber="178">Antennae long, almost reaching posterior margin of pronotum when reclined. Antennomeres 18 and 11 longer than wide, antennomeres 910 as long as wide. Relative length of antennomeres: 1 = 8; 2 = 4; 3 = 5; 48 = 3.5; 910 = 3; 11 = 5.</paragraph>
<paragraph id="9C3736E3FFD7FF9EF4F4FB7FFDA0FAFD" blockId="17.[113,1090,706,1562]" pageId="17" pageNumber="178">Pronotum markedly highly convex, as long as wide, distinctly narrowed anteriad.Anterior angles each with several short bristles. Posterior angles markedly rounded. Lateral margins each with one long black bristle in anterior half. Each dorsal row with 4 equidistant punctures, each sublateral row with 2 punctures.</paragraph>
<paragraph id="9C3736E3FFD7FF9EF4F4FAFEFF4EFA5D" blockId="17.[113,1090,706,1562]" pageId="17" pageNumber="178">Posterior two thirds of scutellum finely and sparsely punctate, punctures somewhat smaller than eye-facets, separated by 2 puncture diameters in transverse direction. Anterior third impunctate.</paragraph>
<paragraph id="9C3736E3FFD7FF9EF4F4FA1EFDD9F9DD" blockId="17.[113,1090,706,1562]" pageId="17" pageNumber="178">Elytra wider than long (ratio 46: 42), parallel-sided. Anterior angles each with one long black bristle. Punctation dense and fine, punctures somewhat larger than those on scutellum, separated by distance somewhat larger than puncture diameters in transverse direction. Surface without microsculpture; setation longer, greyish.</paragraph>
<paragraph id="9C3736E3FFD4FF9DF4D1FF60FEBAFE94" blockId="18.[151,1127,219,660]" pageId="18" pageNumber="179">Legs. Metatibia somewhat longer than metatarsus (ratio 29.5: 28). Metatarsomere 1 almost as long as metatarsomeres 24 combined, metatarsomere 5 somewhat longer than metatarsomeres 3 and 4 combined. Relative length of metatarsomeres: 1 = 11; 2 = 4.5; 34 = 3; 5 = 6.5.</paragraph>
<paragraph id="9C3736E3FFD4FF9DF4D1FEE7FD0BFE73" blockId="18.[151,1127,219,660]" pageId="18" pageNumber="179">
Male. Protarsomeres 13 strongly dilated and sub-bilobed, covered with modified pale setae ventrally, protarsomere 4 distinctly narrower than preceding ones. Sternite VIII (
<figureCitation id="04B32A66FFD4FF9DF05FFEC7FF15FE73" captionStart="Figs" captionStartId="20.[151,191,1474,1493]" captionTargetBox="[202,1071,219,1393]" captionTargetId="figure-10@20.[202,1078,210,1393]" captionTargetPageId="20" captionText="Figs. 3853. 3844 Philonthus iridicollis Bernhauer, 1932; 4548 P. kenyanus Bernhauer, 1939; 4953 P. leptophilus Hromádka, 2006. 38, 45, 49 aedeagus, ventral view; 39, 46, 50 aedeagus, lateral view; 40, 51 paramere with sensory peg setae, ventral view; 41, 52 apical portion of male sternite VIII, ventral view; 42, 48, 53 male sternite IX, ventral view; 43 female tergite X, ventral view; 44 gonocoxite of female genital segment; 47 aedeagus without paramere, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5404284" httpUri="https://zenodo.org/record/5404284/files/figure.png" pageId="18" pageNumber="179">Fig. 41</figureCitation>
), sternite IX (
<figureCitation id="04B32A66FFD4FF9DF52CFE27FE39FE73" box="[328,408,411,436]" captionStart="Figs" captionStartId="20.[151,191,1474,1493]" captionTargetBox="[202,1071,219,1393]" captionTargetId="figure-10@20.[202,1078,210,1393]" captionTargetPageId="20" captionText="Figs. 3853. 3844 Philonthus iridicollis Bernhauer, 1932; 4548 P. kenyanus Bernhauer, 1939; 4953 P. leptophilus Hromádka, 2006. 38, 45, 49 aedeagus, ventral view; 39, 46, 50 aedeagus, lateral view; 40, 51 paramere with sensory peg setae, ventral view; 41, 52 apical portion of male sternite VIII, ventral view; 42, 48, 53 male sternite IX, ventral view; 43 female tergite X, ventral view; 44 gonocoxite of female genital segment; 47 aedeagus without paramere, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5404284" httpUri="https://zenodo.org/record/5404284/files/figure.png" pageId="18" pageNumber="179">Fig. 42</figureCitation>
), aedeagus (
<figureCitation id="04B32A66FFD4FF9DF67FFE27FD3FFE73" box="[539,670,411,436]" captionStart="Figs" captionStartId="20.[151,191,1474,1493]" captionTargetBox="[202,1071,219,1393]" captionTargetId="figure-10@20.[202,1078,210,1393]" captionTargetPageId="20" captionText="Figs. 3853. 3844 Philonthus iridicollis Bernhauer, 1932; 4548 P. kenyanus Bernhauer, 1939; 4953 P. leptophilus Hromádka, 2006. 38, 45, 49 aedeagus, ventral view; 39, 46, 50 aedeagus, lateral view; 40, 51 paramere with sensory peg setae, ventral view; 41, 52 apical portion of male sternite VIII, ventral view; 42, 48, 53 male sternite IX, ventral view; 43 female tergite X, ventral view; 44 gonocoxite of female genital segment; 47 aedeagus without paramere, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5404284" httpUri="https://zenodo.org/record/5404284/files/figure.png" pageId="18" pageNumber="179">Figs. 3840</figureCitation>
).
</paragraph>
<paragraph id="9C3736E3FFD4FF9DF4D1FE07FF5DFDD3" blockId="18.[151,1127,219,660]" pageId="18" pageNumber="179">
Female. Protarsomeres 13 less dilated than in male, covered with modified pale setae ventrally, protarsomere 4 small. Tergite X (
<figureCitation id="04B32A66FFD4FF9DF601FE67FD17FE33" box="[613,694,475,500]" captionStart="Figs" captionStartId="20.[151,191,1474,1493]" captionTargetBox="[202,1071,219,1393]" captionTargetId="figure-10@20.[202,1078,210,1393]" captionTargetPageId="20" captionText="Figs. 3853. 3844 Philonthus iridicollis Bernhauer, 1932; 4548 P. kenyanus Bernhauer, 1939; 4953 P. leptophilus Hromádka, 2006. 38, 45, 49 aedeagus, ventral view; 39, 46, 50 aedeagus, lateral view; 40, 51 paramere with sensory peg setae, ventral view; 41, 52 apical portion of male sternite VIII, ventral view; 42, 48, 53 male sternite IX, ventral view; 43 female tergite X, ventral view; 44 gonocoxite of female genital segment; 47 aedeagus without paramere, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5404284" httpUri="https://zenodo.org/record/5404284/files/figure.png" pageId="18" pageNumber="179">Fig. 43</figureCitation>
), gonocoxites of female genital segment (
<figureCitation id="04B32A66FFD4FF9DF4FBFE47FF51FDD3" box="[159,240,507,532]" captionStart="Figs" captionStartId="20.[151,191,1474,1493]" captionTargetBox="[202,1071,219,1393]" captionTargetId="figure-10@20.[202,1078,210,1393]" captionTargetPageId="20" captionText="Figs. 3853. 3844 Philonthus iridicollis Bernhauer, 1932; 4548 P. kenyanus Bernhauer, 1939; 4953 P. leptophilus Hromádka, 2006. 38, 45, 49 aedeagus, ventral view; 39, 46, 50 aedeagus, lateral view; 40, 51 paramere with sensory peg setae, ventral view; 41, 52 apical portion of male sternite VIII, ventral view; 42, 48, 53 male sternite IX, ventral view; 43 female tergite X, ventral view; 44 gonocoxite of female genital segment; 47 aedeagus without paramere, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5404284" httpUri="https://zenodo.org/record/5404284/files/figure.png" pageId="18" pageNumber="179">Fig. 44</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="D4926568FFD4FF9DF4F3FDA7FC81FDB4" pageId="18" pageNumber="179" type="diagnosis">
<paragraph id="9C3736E3FFD4FF9DF4F3FDA7FBA3FD94" blockId="18.[151,1127,219,660]" pageId="18" pageNumber="179">
<emphasis id="AEFCEAF1FFD4FF9DF4F3FDA7FE31FDF4" bold="true" box="[151,400,539,563]" pageId="18" pageNumber="179">Differential diagnosis.</emphasis>
<taxonomicName id="5B884D60FFD4FF9DF5F1FDA1FDD7FDF3" authorityName="M. Bernhauer" authorityYear="1932" box="[405,630,541,564]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="179" phylum="Arthropoda" rank="species" species="iridicollis">
<emphasis id="AEFCEAF1FFD4FF9DF5F1FDA1FDD7FDF3" box="[405,630,541,564]" italics="true" pageId="18" pageNumber="179">Philonthus iridicollis</emphasis>
</taxonomicName>
may be distinguished from the similar
<taxonomicName id="5B884D60FFD4FF9DF06AFDA1FBC6FDF3" authority="Hromádka, 2009" authorityName="Hromádka" authorityYear="2009" box="[1038,1127,541,564]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="179" phylum="Arthropoda" rank="species" species="aonyx" status="sp. nov.">
<emphasis id="AEFCEAF1FFD4FF9DF06AFDA1FBC6FDF3" box="[1038,1127,541,564]" italics="true" pageId="18" pageNumber="179">P. aonyx</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B5CF578AFFD4FF9DF4F3FD80FF4BFD94" box="[151,234,572,595]" pageId="18" pageNumber="179" rank="species">sp. nov.</taxonomicNameLabel>
by unicoloured elytra and legs and by the different shape of the aedeagus.
</paragraph>
<paragraph id="9C3736E3FFD4FF9DF4F3FDE7FC81FDB4" blockId="18.[151,1127,219,660]" box="[151,800,603,628]" pageId="18" pageNumber="179">
<emphasis id="AEFCEAF1FFD4FF9DF4F3FDE7FEABFDB4" bold="true" box="[151,266,603,627]" pageId="18" pageNumber="179">Bionomic.</emphasis>
Some specimens were collected in grass cuttings.
</paragraph>
</subSubSection>
<subSubSection id="D4926568FFD4FF9DF4F3FDC7FC35FD54" box="[151,916,635,660]" pageId="18" pageNumber="179" type="distribution">
<paragraph id="9C3736E3FFD4FF9DF4F3FDC7FC35FD54" blockId="18.[151,1127,219,660]" box="[151,916,635,660]" pageId="18" pageNumber="179">
<emphasis id="AEFCEAF1FFD4FF9DF4F3FDC7FE8BFD54" bold="true" box="[151,298,635,659]" pageId="18" pageNumber="179">Distribution.</emphasis>
<collectingCountry id="E49F7673FFD4FF9DF554FDC7FD09FD53" box="[304,680,635,660]" name="Democratic Republic of the Congo" pageId="18" pageNumber="179">Democratic Republic of the Congo</collectingCountry>
,
<collectingCountry id="E49F7673FFD4FF9DF6D6FDC7FCAEFD54" box="[690,783,635,659]" name="Ethiopia" pageId="18" pageNumber="179">Ethiopia</collectingCountry>
,
<collectingCountry id="E49F7673FFD4FF9DF77EFDC7FC31FD54" box="[794,912,635,659]" name="Zimbabwe" pageId="18" pageNumber="179">Zimbabwe</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>