160 lines
19 KiB
XML
160 lines
19 KiB
XML
<document id="975B87978BDDF271503377F6945C3213" ID-DOI="10.5281/zenodo.5404276" ID-ISSN="0374-1036" ID-Zenodo-Dep="5404276" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630132078781" checkinUser="felipe" docAuthor="Hromádka, Lubomír" docDate="2009" docId="142187F5FFD1FF97F518FF66FC10FE53" docLanguage="en" docName="ActaEntMusNatPra.2009.49.1.161-190.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 49 (1)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Philonthus maridadus , Tottenham 1962" docType="treatment" docVersion="4" lastPageNumber="185" masterDocId="E818FF8DFFC6FF8FF464FFBCFFA1FFC7" masterDocTitle="Revision of the Afrotropical species of the Philonthus caffer species group (Coleoptera: Staphylinidae: Philonthina)" masterLastPageNumber="190" masterPageNumber="161" pageNumber="184" updateTime="1699162834734" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="A263F5EF3A73671BC8C2E18B4CD4AFC6">
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<mods:title id="F36CE1FA0DD21469F2FF37B0BCD6B5C2">Revision of the Afrotropical species of the Philonthus caffer species group (Coleoptera: Staphylinidae: Philonthina)</mods:title>
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</mods:titleInfo>
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<mods:name id="94607ACBE25798D34B6B044ADA3D1694" type="personal">
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<mods:roleTerm id="AAF1CDCB62990988472A56EECDECB39A">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="DB4FB28D81D27E12DD3A32B2456DA51B">Hromádka, Lubomír</mods:namePart>
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<mods:typeOfResource id="8F9DFBF5BA60DF99D42088B20E2C276E">text</mods:typeOfResource>
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<mods:title id="56C036A6A71AF0AB24962043BCF9EB24">Acta Entomologica Musei Nationalis Pragae</mods:title>
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</mods:titleInfo>
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<mods:part id="59CD905708601A1206462AC3A64749E7">
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<mods:date id="0601DBB782F1C91BE9AB17EF02F4D462">2009</mods:date>
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<mods:detail id="DBD959E3E6A293FBCBD69884C5BFC1BB" type="pubDate">
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<mods:number id="5722395B025FB574D239B02671D9FC60">2009-06-30</mods:number>
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</mods:detail>
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<mods:detail id="4CC44E79796736C3EEAB6E1BEC1ACF1F" type="volume">
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<mods:number id="D7951D7B813E2E431A824684578971CB">49</mods:number>
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</mods:detail>
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<mods:detail id="30AA32163A5ADD0D92EC952885EC61D2" type="issue">
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<mods:number id="24E38CB1CCE2BC12E9CE232B3036DD6C">1</mods:number>
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</mods:detail>
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<mods:extent id="96AA01D191FE576D0FD1A133FE084A34" unit="page">
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<mods:start id="A022C45659862CF9B572E7DB5EB3CA68">161</mods:start>
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<mods:end id="773EF7844B1DA165822C78815959C614">190</mods:end>
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</mods:extent>
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</mods:part>
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<mods:classification id="2A7C120DF8F8854CAA2228C2C526B7E2">journal article</mods:classification>
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<mods:identifier id="AA350F7EF9EA920C5BCC1B30CDBB1F63" type="DOI">10.5281/zenodo.5404276</mods:identifier>
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<mods:identifier id="57D0E9D7C76FBC6F58A0281D434014FB" type="ISSN">0374-1036</mods:identifier>
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<mods:identifier id="81379931ABCF394ECCB1A18CF8C59DCA" type="Zenodo-Dep">5404276</mods:identifier>
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<treatment id="142187F5FFD1FF97F518FF66FC10FE53" ID-DOI="http://doi.org/10.5281/zenodo.5337820" ID-Zenodo-Dep="5337820" LSID="urn:lsid:plazi:treatment:142187F5FFD1FF97F518FF66FC10FE53" httpUri="http://treatment.plazi.org/id/142187F5FFD1FF97F518FF66FC10FE53" lastPageId="24" lastPageNumber="185" pageId="23" pageNumber="184">
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<subSubSection id="D4926568FFD1FF98F518FF66FC94FF34" box="[380,821,218,243]" pageId="23" pageNumber="184" type="nomenclature">
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<paragraph id="9C3736E3FFD1FF98F518FF66FC94FF34" blockId="23.[380,821,218,243]" box="[380,821,218,243]" pageId="23" pageNumber="184">
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<heading id="C77F818FFFD1FF98F518FF66FC94FF34" bold="true" box="[380,821,218,243]" centered="true" fontSize="8" level="2" pageId="23" pageNumber="184" reason="3">
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<taxonomicName id="5B884D60FFD1FF98F518FF66FC94FF34" ID-CoL="4GB8V" authority="Tottenham, 1962" authorityName=", Tottenham" authorityYear="1962" box="[380,821,218,243]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="184" phylum="Arthropoda" rank="species" species="maridadus">
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<emphasis id="AEFCEAF1FFD1FF98F518FF66FC94FF34" bold="true" box="[380,821,218,243]" pageId="23" pageNumber="184">
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<emphasis id="AEFCEAF1FFD1FF98F518FF66FDD0FF34" bold="true" box="[380,625,218,243]" italics="true" pageId="23" pageNumber="184">Philonthus maridadus</emphasis>
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<bibRefCitation id="F8194B12FFD1FF98F61CFF67FC94FF34" author="TOTTENHAM C. E." box="[632,821,219,243]" pageId="23" pageNumber="184" pagination="132 - 258" refId="ref16863" refString="TOTTENHAM C. E. 1962: Mission zoologique de l' I. R. S. A. C. en Afrique orientale (P. Basilewsky et N. Leleup, 1957). LXXVI. Coleoptera Staphylinidae Staphylininae. Annales du Musee de l'Afrique Centrale, Sciences Zoologiques, Serie 8 ° 110: 132 - 258." type="journal article" year="1962">Tottenham, 1962</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="D4926568FFD1FF98F67BFF45FD33FECB" box="[543,658,249,268]" pageId="23" pageNumber="184" type="description">
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<paragraph id="9C3736E3FFD1FF98F67BFF45FD33FECB" blockId="23.[113,658,249,303]" box="[543,658,249,268]" pageId="23" pageNumber="184">
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(
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<figureCitation id="04B32A66FFD1FF98F642FF45FD2DFECB" box="[550,652,249,268]" captionStart="Figs" captionStartId="26.[151,191,1193,1212]" captionTargetBox="[160,1118,221,1161]" captionTargetId="figure-101@26.[151,1128,214,1166]" captionTargetPageId="26" captionText="Figs. 54–61. 54–56 – Philonthus limbipennis Fauvel, 1905. 57–58 – P. maridadus Tottenham, 1962; 59–61 – P. sequens Bernhauer & Schubert, 1914; 62–63 – P. tomicus Tottenham, 1962. 54, 57, 59 – aedeagus, ventral view; 55, 58, 60 – aedeagus, lateral view; 56, 61 – apex of paramere with sensory peg setae, ventral view; 62 – aedeagus, dorsal view; 63 – paramere, ventral,view." figureDoi="http://doi.org/10.5281/zenodo.5404286" httpUri="https://zenodo.org/record/5404286/files/figure.png" pageId="23" pageNumber="184">Figs. 57–58</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="D4926568FFD1FF98F415FEA1FE54FEE8" box="[113,501,284,303]" pageId="23" pageNumber="184" type="reference_group">
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<paragraph id="9C3736E3FFD1FF98F415FEA1FE54FEE8" blockId="23.[113,658,249,303]" box="[113,501,284,303]" pageId="23" pageNumber="184">
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<treatmentCitationGroup id="BC9811CDFFD1FF98F415FEA1FE54FEE8" box="[113,501,284,303]" pageId="23" pageNumber="184">
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<treatmentCitation id="1D2910F2FFD1FF98F415FEA1FE53FEE8" author="TOTTENHAM C. E." box="[113,498,284,303]" page="223" pageId="23" pageNumber="184" year="1962">
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<taxonomicName id="5B884D60FFD1FF98F415FEA1FE53FEE8" ID-CoL="4GB8V" authority="Tottenham, 1962: 223" authorityName=", Tottenham" authorityPageNumber="223" authorityYear="1962" box="[113,498,284,303]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="184" phylum="Arthropoda" rank="species" species="maridadus">
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<emphasis id="AEFCEAF1FFD1FF98F415FEA1FE8FFEE8" box="[113,302,285,303]" italics="true" pageId="23" pageNumber="184">Philonthus maridadus</emphasis>
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<bibRefCitation id="F8194B12FFD1FF98F557FEA0FE53FEE8" author="TOTTENHAM C. E." box="[307,498,284,303]" pageId="23" pageNumber="184" pagination="132 - 258" refId="ref16863" refString="TOTTENHAM C. E. 1962: Mission zoologique de l' I. R. S. A. C. en Afrique orientale (P. Basilewsky et N. Leleup, 1957). LXXVI. Coleoptera Staphylinidae Staphylininae. Annales du Musee de l'Afrique Centrale, Sciences Zoologiques, Serie 8 ° 110: 132 - 258." type="journal article" year="1962">Tottenham, 1962: 223</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="D4926568FFD1FF98F415FEEDFC0CFE5E" pageId="23" pageNumber="184" type="materials_examined">
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<paragraph id="9C3736E3FFD1FF98F415FEEDFEDBFEA3" blockId="23.[113,1090,336,410]" box="[113,378,336,356]" pageId="23" pageNumber="184">
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<emphasis id="AEFCEAF1FFD1FF98F415FEEDFF4BFEA3" bold="true" box="[113,234,336,356]" pageId="23" pageNumber="184">
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<typeStatus id="43338841FFD1FF98F415FEEDFF3FFEA3" box="[113,158,337,356]" pageId="23" pageNumber="184">Type</typeStatus>
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locality.
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</emphasis>
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<collectingCountry id="E49F7673FFD1FF98F48BFEEDFE8BFEA3" box="[239,298,337,356]" name="Kenya" pageId="23" pageNumber="184">Kenya</collectingCountry>
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: Nairobi.
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</paragraph>
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<paragraph id="9C3736E3FFD1FF98F415FED0FC0CFE5E" blockId="23.[113,1090,336,410]" pageId="23" pageNumber="184">
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<emphasis id="AEFCEAF1FFD1FF98F415FED0FEF5FEB8" bold="true" box="[113,340,363,383]" pageId="23" pageNumber="184">Type material examined.</emphasis>
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<materialsCitation id="2CE03CBEFFD1FF98F53EFED0FC0CFE5E" collectingDate="1954-12-27" collectionCode="BMNH" collectorName="Nairobi & V. F. Eastop. & C. E. Tottenham & C. H. BOHEMAN" pageId="23" pageNumber="184" specimenCount="1" typeStatus="holotype">
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<typeStatus id="43338841FFD1FF98F53EFED0FE0CFEBA" box="[346,429,364,383]" pageId="23" pageNumber="184" type="holotype">HOLOTYPE</typeStatus>
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: J, ‛
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<collectorName id="317D5335FFD1FF98F5BFFED0FDBFFEB8" box="[475,542,364,383]" pageId="23" pageNumber="184">Nairobi</collectorName>
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,
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<date id="E8361023FFD1FF98F643FED0FD2BFEB8" box="[551,650,364,383]" pageId="23" pageNumber="184" value="1954-12-27">
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<collectingDate id="F872E9CBFFD1FF98F643FED0FD2BFEB8" box="[551,650,364,383]" pageId="23" pageNumber="184" value="1954-12-27">27.xii.1954</collectingDate>
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</date>
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,
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<collectorName id="317D5335FFD1FF98F6F7FED0FCA3FEB8" box="[659,770,364,383]" pageId="23" pageNumber="184">V. F. Eastop.</collectorName>
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//
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<taxonomicName id="5B884D60FFD1FF98F77CFED1FB9CFEB8" authority=", Tottenham" authorityName=", Tottenham" authorityYear="1962" box="[792,1085,364,383]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="184" phylum="Arthropoda" rank="species" species="maridadus">
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<emphasis id="AEFCEAF1FFD1FF98F77CFED1FC74FEB8" box="[792,981,365,383]" italics="true" pageId="23" pageNumber="184">Philonthus maridadus</emphasis>
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, Tottenham
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</taxonomicName>
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, TYPE [white oblong label handwritten] //
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<collectorName id="317D5335FFD1FF98F5BFFE3AFDC7FE5E" box="[475,614,390,410]" pageId="23" pageNumber="184">C. E. Tottenham</collectorName>
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collection,
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<collectorName id="317D5335FFD1FF98F6AEFE3BFD56FE5D" box="[714,759,391,410]" name="C. H. BOHEMAN" pageId="23" pageNumber="184">B.M.</collectorName>
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1974-587’ (
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<collectionCode id="FA99AE26FFD1FF98F73AFE3AFC02FE5E" box="[862,931,390,409]" pageId="23" pageNumber="184">BMNH</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="D4926568FFD1FF97F415FE0EFE0BFF34" lastPageId="24" lastPageNumber="185" pageId="23" pageNumber="184" type="description">
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<paragraph id="9C3736E3FFD1FF98F415FE0EFBB5FE0E" blockId="23.[113,1089,434,1578]" box="[113,1044,434,458]" pageId="23" pageNumber="184">
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<emphasis id="AEFCEAF1FFD1FF98F415FE0EFEB6FE0D" bold="true" box="[113,279,434,458]" pageId="23" pageNumber="184">Redescription.</emphasis>
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Body length
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<quantity id="5B709B06FFD1FF98F5C3FE0EFE5AFE0E" box="[423,507,434,458]" metricMagnitude="-3" metricUnit="m" metricValue="8.8" pageId="23" pageNumber="184" unit="mm" value="8.8">8.8 mm</quantity>
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, length of fore body (to the end of elytra)
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<quantity id="5B709B06FFD1FF98F7DFFE0EFBB1FE0E" box="[955,1040,434,458]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="23" pageNumber="184" unit="mm" value="4.1">4.1 mm</quantity>
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.
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</paragraph>
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<paragraph id="9C3736E3FFD1FF98F4DBFE6EFE47FD6D" blockId="23.[113,1089,434,1578]" pageId="23" pageNumber="184">Colouration. Head black, pronotum and abdomen brown. Elytra yellowish brown, anterior part of each elytron dark, border between anterior dark and posterior pale colouration rather weak, going in a straight line from posterior apex of scutellum to posterior third of elytral length at lateral margin when observed from above; deflexed margins of elytra yellow. Maxillary and labial palpi brown, distal palpomeres slightly paler, mandibles brown, antennae brown, antennomere 1 and base of antennomere 2 of maxillary and labial palpi paler, legs yellow, all tibiae somewhat darker.</paragraph>
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<paragraph id="9C3736E3FFD1FF98F4DEFD0EFD81FCAD" blockId="23.[113,1089,434,1578]" pageId="23" pageNumber="184">Head inconspicuously longer than wide (ratio 25: 23), lateral margins behind eyes subparallel to posterior angles. Eyes slightly projecting, vaguely shorter than temples (ratio 10: 12). Distance between medial interocular punctures about 4 times as large as distance between medial and lateral interocular punctures. Lateral interocular punctures slightly shifted anteriad. Posterior margin of each eye with 3 coarse punctures. Surface with irregular, almost indistinct microsculpture here and there.</paragraph>
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<paragraph id="9C3736E3FFD1FF98F4CCFCCEFEF9FC0D" blockId="23.[113,1089,434,1578]" pageId="23" pageNumber="184">Antennae long, reaching posterior margin of pronotum when reclined, all antennomeres longer than wide. Relative length of antennomeres: 1 = 8; 2 = 4.5; 3 = 2.5; 4–5 = 4.5; 6–7 = 4; 8–10 = 3.5; 11 = 5.</paragraph>
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<paragraph id="9C3736E3FFD1FF98F4CCFC6EFC7BFB8D" blockId="23.[113,1089,434,1578]" pageId="23" pageNumber="184">Pronotum highly convex, hardly narrowed anteriad, longer than wide (ratio 32: 28). Each dorsal row with 5 coarse punctures, punctures 2–4 equidistant, distance between punctures 1–2 and 4–5 twice as long as distance between punctures 2–4. Each sublateral row with 2 punctures, puncture 2 slightly shifted toward lateral margin. Surface without microsculpture.</paragraph>
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<paragraph id="9C3736E3FFD1FF98F4D2FBEEFC9DFB4D" blockId="23.[113,1089,434,1578]" pageId="23" pageNumber="184">Entire scutellum very densely and coarsely punctate, punctures somewhat larger than eye-facets, separated by distance much smaller than their diameter.</paragraph>
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<paragraph id="9C3736E3FFD1FF98F4CBFB2EFDA9FACD" blockId="23.[113,1089,434,1578]" pageId="23" pageNumber="184">Elytra combined slightly wider than long (ratio 42: 40), parallel-sided, distinctly widened posteriad. Punctation fine and relatively sparse. Punctures hardly larger than eye-facets, separated by 1.5 or 2 puncture diameters in transverse direction. Surface between punctures without microsculpture; setation dark.</paragraph>
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<paragraph id="9C3736E3FFD1FF98F4A0FAAEFC89FAAD" blockId="23.[113,1089,434,1578]" pageId="23" pageNumber="184">Legs. Metatibia vaguely longer than metatarsus (ratio 25.5: 24), metatarsomere 1 almost as long as metatarsomere 5, metatarsomere 2 hardly longer than metatarsomere 3. Relative length of metatarsomeres: 1 = 7; 2 = 3; 3–4 = 2.5; 5 = 8.</paragraph>
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<paragraph id="9C3736E3FFD1FF98F4D8FACEFC3EFA2D" blockId="23.[113,1089,434,1578]" pageId="23" pageNumber="184">Abdomen slightly narrowed toward apex. Elevated area between two basal lines on first three visible tergites densely punctate. Punctation of visible tergites finer and denser than that on elytra, becoming somewhat sparser toward posterior margin of each tergite. Surface between punctures without microsculpture; setation similar to that on elytra.</paragraph>
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<paragraph id="9C3736E3FFD1FF98F4D2FA4EFD36F9ED" blockId="23.[113,1089,434,1578]" pageId="23" pageNumber="184">
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Male. Protarsomeres 1–3 strongly dilated, sub-bilobed, protarsomere 4 distinctly narrower than preceding ones. Aedeagus (
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<figureCitation id="04B32A66FFD1FF98F66BF9AEFD2DF9ED" box="[527,652,1554,1578]" captionStart="Figs" captionStartId="26.[151,191,1193,1212]" captionTargetBox="[160,1118,221,1161]" captionTargetId="figure-101@26.[151,1128,214,1166]" captionTargetPageId="26" captionText="Figs. 54–61. 54–56 – Philonthus limbipennis Fauvel, 1905. 57–58 – P. maridadus Tottenham, 1962; 59–61 – P. sequens Bernhauer & Schubert, 1914; 62–63 – P. tomicus Tottenham, 1962. 54, 57, 59 – aedeagus, ventral view; 55, 58, 60 – aedeagus, lateral view; 56, 61 – apex of paramere with sensory peg setae, ventral view; 62 – aedeagus, dorsal view; 63 – paramere, ventral,view." figureDoi="http://doi.org/10.5281/zenodo.5404286" httpUri="https://zenodo.org/record/5404286/files/figure.png" pageId="23" pageNumber="184">Figs 57–58</figureCitation>
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).
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</paragraph>
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<paragraph id="9C3736E3FFDEFF97F4B9FF67FE0BFF34" blockId="24.[151,1130,219,404]" box="[221,426,219,243]" pageId="24" pageNumber="185">Female. Unknown.</paragraph>
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</subSubSection>
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<subSubSection id="D4926568FFDEFF97F4F3FF47FE20FEB4" pageId="24" pageNumber="185" type="diagnosis">
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<paragraph id="9C3736E3FFDEFF97F4F3FF47FDC1FE94" blockId="24.[151,1130,219,404]" pageId="24" pageNumber="185">
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<emphasis id="AEFCEAF1FFDEFF97F4F3FF47FE35FED4" bold="true" box="[151,404,251,275]" pageId="24" pageNumber="185">Differential diagnosis.</emphasis>
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<taxonomicName id="5B884D60FFDEFF97F5F9FF41FD2DFED3" authorityName=", Tottenham" authorityYear="1962" box="[413,652,253,276]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="185" phylum="Arthropoda" rank="species" species="maridadus">
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<emphasis id="AEFCEAF1FFDEFF97F5F9FF41FD2DFED3" box="[413,652,253,276]" italics="true" pageId="24" pageNumber="185">Philonthus maridadus</emphasis>
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</taxonomicName>
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is similar in most external characters to
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<taxonomicName id="5B884D60FFDEFF97F037FF41FF58FEF3" authority="Hromádka, 2009" authorityName="Hromádka" authorityYear="2009" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="185" phylum="Arthropoda" rank="species" species="bucorvus" status="sp. nov.">
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<emphasis id="AEFCEAF1FFDEFF97F037FF41FF58FEF3" italics="true" pageId="24" pageNumber="185">P. bucorvus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="B5CF578AFFDEFF97F564FEA0FEF2FEF4" box="[256,339,284,307]" pageId="24" pageNumber="185" rank="species">sp. nov.</taxonomicNameLabel>
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It differs by shorter antennae, black elytra each with red triangular patch, and by the different shape of the aedeagus.
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</paragraph>
|
||
<paragraph id="9C3736E3FFDEFF97F4F3FEE7FE20FEB4" blockId="24.[151,1130,219,404]" box="[151,385,347,371]" pageId="24" pageNumber="185">
|
||
<emphasis id="AEFCEAF1FFDEFF97F4F3FEE7FEABFEB4" bold="true" box="[151,266,347,371]" pageId="24" pageNumber="185">Bionomic.</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D4926568FFDEFF97F4F3FEC7FC10FE53" box="[151,945,379,404]" pageId="24" pageNumber="185" type="distribution">
|
||
<paragraph id="9C3736E3FFDEFF97F4F3FEC7FC10FE53" blockId="24.[151,1130,219,404]" box="[151,945,379,404]" pageId="24" pageNumber="185">
|
||
<emphasis id="AEFCEAF1FFDEFF97F4F3FEC7FE8BFE54" bold="true" box="[151,298,379,403]" pageId="24" pageNumber="185">Distribution.</emphasis>
|
||
<collectingCountry id="E49F7673FFDEFF97F554FEC0FED7FE53" box="[304,374,380,404]" name="Kenya" pageId="24" pageNumber="185">Kenya</collectingCountry>
|
||
(this paper); ‘Tanganyika’,
|
||
<collectingCountry id="E49F7673FFDEFF97F6FBFEC7FD5EFE54" box="[671,767,379,403]" name="Tanzania" pageId="24" pageNumber="185">Tanzania</collectingCountry>
|
||
(HERMAN 2001).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |