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<document id="9637B58D161233BDAF00B2AB05B40F2E" ID-DOI="10.5281/zenodo.5404276" ID-ISSN="0374-1036" ID-Zenodo-Dep="5404276" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630132078781" checkinUser="felipe" docAuthor="Hromádka, Lubomír" docDate="2009" docId="142187F5FFCDFF83F5F6FF66FF7CFDF3" docLanguage="en" docName="ActaEntMusNatPra.2009.49.1.161-190.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 49 (1)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Philonthus cliens , Eppelsheim 1890" docType="treatment" docVersion="3" lastPageNumber="173" masterDocId="E818FF8DFFC6FF8FF464FFBCFFA1FFC7" masterDocTitle="Revision of the Afrotropical species of the Philonthus caffer species group (Coleoptera: Staphylinidae: Philonthina)" masterLastPageNumber="190" masterPageNumber="161" pageNumber="172" updateTime="1699162834734" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="EB34783ED073EAF6E33C7430E66C5265">Revision of the Afrotropical species of the Philonthus caffer species group (Coleoptera: Staphylinidae: Philonthina)</mods:title>
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<mods:namePart id="A7766EFBFE7344B450BC171DCE150092">Hromádka, Lubomír</mods:namePart>
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<mods:title id="F3063887A44F027A224A2168E32AA5BF">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="DC0D772F80978231C53D7D1CA2DF2FD4">2009</mods:date>
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<treatment id="142187F5FFCDFF83F5F6FF66FF7CFDF3" LSID="urn:lsid:plazi:treatment:142187F5FFCDFF83F5F6FF66FF7CFDF3" httpUri="http://treatment.plazi.org/id/142187F5FFCDFF83F5F6FF66FF7CFDF3" lastPageId="12" lastPageNumber="173" pageId="11" pageNumber="172">
<subSubSection id="D4926568FFCDFF84F5F6FF66FC81FF34" box="[402,800,218,243]" pageId="11" pageNumber="172" type="nomenclature">
<paragraph id="9C3736E3FFCDFF84F5F6FF66FC81FF34" blockId="11.[402,800,218,243]" box="[402,800,218,243]" pageId="11" pageNumber="172">
<heading id="C77F818FFFCDFF84F5F6FF66FC81FF34" bold="true" box="[402,800,218,243]" centered="true" fontSize="8" level="2" pageId="11" pageNumber="172" reason="3">
<taxonomicName id="5B884D60FFCDFF84F5F6FF66FC81FF34" ID-CoL="4G9SZ" authority="Eppelsheim, 1890" authorityName=", Eppelsheim" authorityYear="1890" box="[402,800,218,243]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="172" phylum="Arthropoda" rank="species" species="cliens">
<emphasis id="AEFCEAF1FFCDFF84F5F6FF66FC81FF34" bold="true" box="[402,800,218,243]" pageId="11" pageNumber="172">
<emphasis id="AEFCEAF1FFCDFF84F5F6FF66FDF3FF34" bold="true" box="[402,594,218,243]" italics="true" pageId="11" pageNumber="172">Philonthus cliens</emphasis>
<bibRefCitation id="F8194B12FFCDFF84F63CFF67FC81FF34" author="EPPELSHEIM E." box="[600,800,219,243]" pageId="11" pageNumber="172" pagination="273 - 280" refId="ref16251" refString="EPPELSHEIM E. 1890: Staphylinidarum species novae, a Domino Dr. Bomford in India orientali collectae, descriptae. Wiener Entomologische Zeitung 9: 273 - 280." type="journal article" year="1890">Eppelsheim, 1890</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="D4926568FFCDFF84F646FF45FD31FECB" box="[546,656,249,268]" pageId="11" pageNumber="172" type="description">
<paragraph id="9C3736E3FFCDFF84F646FF45FD31FECB" blockId="11.[113,986,249,356]" box="[546,656,249,268]" pageId="11" pageNumber="172">
(
<figureCitation id="04B32A66FFCDFF84F64CFF45FD28FECB" box="[552,649,249,268]" captionStart="Figs" captionStartId="9.[112,151,1487,1506]" captionTargetBox="[148,1054,223,1459]" captionTargetId="figure-13@9.[142,1059,219,1468]" captionTargetPageId="9" captionText="Figs. 1224. 1213 Philonthus biguttulus Fauvel, 1907 (Côte dIvoire); 1416 P. bucorvus sp. nov.; 1720 P. caffer Boheman, 1848; 2124 Philonthus cliens Eppelsheim, 1890. 12, 14, 17, 21 aedeagus, ventral view, 13 paramere with sensory peg setae, ventral view; 15, 18, 22 aedeagus, lateral view; 16, 20 male sternite IX, ventral view; 19, 23 aedeagus without paramere, ventral view; 24 apex of paramere with sensory peg setae, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5404280" httpUri="https://zenodo.org/record/5404280/files/figure.png" pageId="11" pageNumber="172">Figs 2124</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="D4926568FFCDFF84F415FEA1FDDCFEA3" pageId="11" pageNumber="172" type="reference_group">
<paragraph id="9C3736E3FFCDFF84F415FEA1FDDCFEA3" blockId="11.[113,986,249,356]" pageId="11" pageNumber="172">
<treatmentCitationGroup id="BC9811CDFFCDFF84F415FEA1FDDCFEA3" pageId="11" pageNumber="172">
<treatmentCitation id="1D2910F2FFCDFF84F415FEA1FE71FEE8" author="EPPELSHEIM E." box="[113,464,284,303]" page="277" pageId="11" pageNumber="172" year="1890">
<taxonomicName id="5B884D60FFCDFF84F415FEA1FE71FEE8" ID-CoL="4G9SZ" authority="Eppelsheim, 1890: 277" authorityName=", Eppelsheim" authorityPageNumber="277" authorityYear="1890" box="[113,464,284,303]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="172" phylum="Arthropoda" rank="species" species="cliens">
<emphasis id="AEFCEAF1FFCDFF84F415FEA1FEA4FEE8" box="[113,261,285,303]" italics="true" pageId="11" pageNumber="172">Philonthus cliens</emphasis>
<bibRefCitation id="F8194B12FFCDFF84F56EFEA0FE71FEE8" author="EPPELSHEIM E." box="[266,464,284,303]" pageId="11" pageNumber="172" pagination="273 - 280" refId="ref16251" refString="EPPELSHEIM E. 1890: Staphylinidarum species novae, a Domino Dr. Bomford in India orientali collectae, descriptae. Wiener Entomologische Zeitung 9: 273 - 280." type="journal article" year="1890">Eppelsheim, 1890: 277</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(resurrected from synonymy by
<treatmentCitation id="1D2910F2FFCDFF84F68CFEA0FC77FEE8" author="SCHILLHAMMER H." box="[744,982,284,303]" page="322" pageId="11" pageNumber="172" year="2004">
<bibRefCitation id="F8194B12FFCDFF84F68CFEA0FC77FEE8" author="SCHILLHAMMER H." box="[744,982,284,303]" pageId="11" pageNumber="172" pagination="319 - 325" refId="ref16707" refString="SCHILLHAMMER H. 2004: New records, synonyms and nomenclatoral changes in the tribe Staphylinini (Insecta: Coleoptera: Staphylinidae). Annalen des Naturhistorischen Museums Wien 105 B: 319 - 325." type="journal article" year="2004">SCHILLHAMMER (2004: 322))</bibRefCitation>
</treatmentCitation>
.
<taxonomicName id="5B884D60FFCDFF84F415FE8BFDEAFE8E" ID-CoL="4G9SZ" authority=": FAUVEL (1903: 159)" authorityName="FAUVEL" authorityPageNumber="159" authorityYear="1903" box="[113,587,310,330]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="172" phylum="Arthropoda" rank="variety" species="aeneipennis" variety="cliens">
<emphasis id="AEFCEAF1FFCDFF84F415FE8BFE98FE8E" box="[113,313,311,329]" italics="true" pageId="11" pageNumber="172">Philonthus aeneipennis</emphasis>
var.
<emphasis id="AEFCEAF1FFCDFF84F507FE8BFE34FE8E" box="[355,405,311,329]" italics="true" pageId="11" pageNumber="172">cliens</emphasis>
: FAUVEL (1903: 159)
</taxonomicName>
.
<treatmentCitation id="1D2910F2FFCDFF84F415FEEEFDDBFEA3" author="SCHEERPELTZ O." box="[113,634,337,356]" page="5" pageId="11" pageNumber="172" year="1960">
<taxonomicName id="5B884D60FFCDFF84F415FEEEFDDBFEA3" ID-CoL="76ZC6" authority="Scheerpeltz, 1960: 5" authorityName="Scheerpeltz" authorityPageNumber="5" authorityYear="1960" box="[113,634,337,356]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="172" phylum="Arthropoda" rank="variety" species="aeneipennis" variety="lindemanni">
<emphasis id="AEFCEAF1FFCDFF84F415FEEEFE98FEA3" box="[113,313,338,356]" italics="true" pageId="11" pageNumber="172">Philonthus aeneipennis</emphasis>
var.
<emphasis id="AEFCEAF1FFCDFF84F507FEEEFE62FEA3" box="[355,451,338,356]" italics="true" pageId="11" pageNumber="172">lindemanni</emphasis>
<bibRefCitation id="F8194B12FFCDFF84F5ACFEEDFDDBFEA3" author="SCHEERPELTZ O." box="[456,634,337,356]" pageId="11" pageNumber="172" pagination="1 - 7" refId="ref16670" refString="SCHEERPELTZ O. 1960: Die von Dr. Chr. Lindemann gelegentlich ihrer Reise 1955 / 56 in Westpakistan aufgesammelten Staphyliniden {Col.) Opuscula Zoologica (Munchen) 51: 1 - 7." type="journal article" year="1960">Scheerpeltz, 1960: 5</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="D4926568FFCDFF84F415FE3AFEE0FD93" pageId="11" pageNumber="172" type="materials_examined">
<paragraph id="9C3736E3FFCDFF84F415FE3AFED7FE5D" blockId="11.[113,1090,389,596]" box="[113,374,389,410]" pageId="11" pageNumber="172">
<emphasis id="AEFCEAF1FFCDFF84F415FE3AFF4BFE5E" bold="true" box="[113,234,389,409]" pageId="11" pageNumber="172">
<typeStatus id="43338841FFCDFF84F415FE3AFF3FFE5E" box="[113,158,390,409]" pageId="11" pageNumber="172">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="E49F7673FFCDFF84F48BFE3AFEBCFE5E" box="[239,285,390,409]" name="India" pageId="11" pageNumber="172">India</collectingCountry>
, Nagpore.
</paragraph>
<paragraph id="9C3736E3FFCDFF84F415FE1DFE47FDC3" blockId="11.[113,1090,389,596]" pageId="11" pageNumber="172">
<emphasis id="AEFCEAF1FFCDFF84F415FE1DFEF3FE73" bold="true" box="[113,338,416,436]" pageId="11" pageNumber="172">Type material examined.</emphasis>
<typeStatus id="43338841FFCDFF84F533FE1DFE0AFE74" box="[343,427,417,436]" pageId="11" pageNumber="172" type="holotype">HOLOTYPE</typeStatus>
: J, Nagpore,
<collectingCountry id="E49F7673FFCDFF84F64FFE1DFDF6FE73" box="[555,599,417,436]" name="India" pageId="11" pageNumber="172">India</collectingCountry>
centr., Dr. D. Bomford [white oblong printed label] //
<taxonomicName id="5B884D60FFCDFF84F040FE1DFF6AFE08" authority="Eppel." authorityName=", Eppelsheim" authorityYear="1890" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="172" phylum="Arthropoda" rank="species" species="cliens">cliens Eppel.</taxonomicName>
spec. propria. det Gridelli [white oblong label, handwritten] / /
<taxonomicName id="5B884D60FFCDFF84F6BBFE01FCD5FE08" authority="Boh." authorityName="Boh." box="[735,884,444,463]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="172" phylum="Arthropoda" rank="species" species="aeneipennis">
<emphasis id="AEFCEAF1FFCDFF84F6BBFE01FCE4FE08" box="[735,837,445,463]" italics="true" pageId="11" pageNumber="172">aeneipennis</emphasis>
Boh.
</taxonomicName>
var. [white oblong label handwritten] //
<emphasis id="AEFCEAF1FFCDFF84F493FE6BFEC9FE2E" box="[247,360,471,489]" italics="true" pageId="11" pageNumber="172">quisquilarius</emphasis>
<taxonomicName id="5B884D60FFCDFF84F50AFE6BFE7FFE2E" box="[366,478,471,490]" pageId="11" pageNumber="172" rank="variety" variety="inquinatus">
v.
<emphasis id="AEFCEAF1FFCDFF84F5E0FE6BFE7FFE2E" box="[388,478,471,489]" italics="true" pageId="11" pageNumber="172">inquinatus</emphasis>
</taxonomicName>
[white oblongblabel] //
<taxonomicName id="5B884D60FFCDFF84F6D5FE6BFC19FE2E" authority=", Eppelsheim" authorityName=", Eppelsheim" authorityYear="1890" box="[689,952,470,490]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="172" phylum="Arthropoda" rank="species" species="cliens">
<emphasis id="AEFCEAF1FFCDFF84F6D5FE6BFCE7FE2E" box="[689,838,471,489]" italics="true" pageId="11" pageNumber="172">Philonthus cliens</emphasis>
, Eppelsheim
</taxonomicName>
,
<typeStatus id="43338841FFCDFF84F7A6FE6AFBE0FE2E" box="[962,1089,470,489]" pageId="11" pageNumber="172" type="holotype">HOLOTYPUS</typeStatus>
[red oblong label, handwritten] (
<collectionCode id="FA99AE26FFCDFF84F5F5FE4DFE7DFDC3" box="[401,476,497,516]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="11" pageNumber="172" type="Museum">NHMW</collectionCode>
).
</paragraph>
<paragraph id="9C3736E3FFCDFF84F415FDB7FEE0FD93" blockId="11.[113,1090,389,596]" pageId="11" pageNumber="172">
<emphasis id="AEFCEAF1FFCDFF84F415FDB7FDB1FDD8" bold="true" box="[113,528,523,543]" pageId="11" pageNumber="172">
Additional material examined.
<collectingCountry id="E49F7673FFCDFF84F5EAFDB7FDAAFDD8" box="[398,523,523,543]" name="Botswana" pageId="11" pageNumber="172">BOTSWANA</collectingCountry>
:
</emphasis>
<specimenCount id="8A8EFD6AFFCDFF84F671FDB0FDF5FDD8" box="[533,596,524,543]" pageId="11" pageNumber="172" type="generic">6 spec.</specimenCount>
,
<quantity id="5B709B06FFCDFF84F605FDB0FD2EFDD8" box="[609,655,524,543]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="11" pageNumber="172" unit="km" value="6.0">6 km</quantity>
E Kakfontein,
<date id="E8361023FFCDFF84F777FDB0FCC7FDD8" box="[787,870,524,543]" pageId="11" pageNumber="172" value="1993-03-06">6.iii.1993</date>
,
<geoCoordinate id="F9BC5024FFCDFF84F714FDB0FC70FDD8" box="[880,977,523,543]" degrees="20" direction="south" minutes="04" orientation="latitude" pageId="11" pageNumber="172" precision="15" seconds="24" value="-20.073334">20°0424″S</geoCoordinate>
/
<geoCoordinate id="F9BC5024FFCDFF84F7BCFDB0FB9CFDD8" box="[984,1085,523,543]" degrees="20" direction="east" minutes="50" orientation="longitude" pageId="11" pageNumber="172" precision="15" seconds="05" value="20.834723">20°5005″E</geoCoordinate>
, lux, M. Uhlig, leg. (
<collectionCode id="FA99AE26FFCDFF84F54CFD9AFECBFDFE" box="[296,362,550,569]" pageId="11" pageNumber="172">ZMHB</collectionCode>
).
<emphasis id="AEFCEAF1FFCDFF84F51FFD99FE60FDFE" bold="true" box="[379,449,549,569]" pageId="11" pageNumber="172">
<collectingCountry id="E49F7673FFCDFF84F51FFD99FE1CFDFE" box="[379,445,549,569]" name="India" pageId="11" pageNumber="172">INDIA</collectingCountry>
:
</emphasis>
1 J, Nagpore, //
<taxonomicName id="5B884D60FFCDFF84F603FD9BFD62FDFE" authority="Hromádka, 2009" authorityName="Hromádka" authorityYear="2009" box="[615,707,551,569]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="172" phylum="Arthropoda" rank="genus" status="sp. nov.">
<emphasis id="AEFCEAF1FFCDFF84F603FD9BFD62FDFE" box="[615,707,551,569]" italics="true" pageId="11" pageNumber="172">Philonthus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B5CF578AFFCDFF84F6AEFD9BFD5AFDFD" box="[714,763,551,570]" pageId="11" pageNumber="172" rank="species">n. sp.</taxonomicNameLabel>
Epp. //
<taxonomicName id="5B884D60FFCDFF84F720FD9BFBB2FDFE" authority=", Epp." authorityName=", Eppelsheim" authorityYear="1890" box="[836,1043,550,570]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="172" phylum="Arthropoda" rank="species" species="cliens">
<emphasis id="AEFCEAF1FFCDFF84F720FD9BFC78FDFE" box="[836,985,551,569]" italics="true" pageId="11" pageNumber="172">Philonthus cliens</emphasis>
, Epp.
</taxonomicName>
, det. Schillhammer (
<collectionCode id="FA99AE26FFCDFF84F490FDFDFE97FD93" box="[244,310,577,596]" pageId="11" pageNumber="172">ZMHB</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="D4926568FFCDFF83F415FDD0FE23FE94" lastPageId="12" lastPageNumber="173" pageId="11" pageNumber="172" type="description">
<paragraph id="9C3736E3FFCDFF84F415FDD0FC4CFD42" blockId="11.[113,1089,620,1572]" box="[113,1005,620,645]" pageId="11" pageNumber="172">
<emphasis id="AEFCEAF1FFCDFF84F415FDD0FEB6FD43" bold="true" box="[113,279,620,644]" pageId="11" pageNumber="172">Redescription.</emphasis>
Body length
<quantity id="5B709B06FFCDFF84F5C3FDD0FE5AFD43" box="[423,507,620,644]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="11" pageNumber="172" unit="mm" value="7.5">7.5 mm</quantity>
, length of fore body (to end of elytra
<quantity id="5B709B06FFCDFF84F7EFFDD0FC40FD42" box="[907,993,620,645]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="11" pageNumber="172" unit="mm" value="3.5">3.5 mm</quantity>
).
</paragraph>
<paragraph id="9C3736E3FFCDFF84F4F4FD30FEE1FCC3" blockId="11.[113,1089,620,1572]" pageId="11" pageNumber="172">Colouration. Head black, pronotum, scutellum and abdomen black-brown, elytra red, shoulders narrowly smoky black, maxillary and labial palpi yellow-brown, antennomeres 12 and anterior half of antennomere 3 yellow-brown, remaining antennomeres brown-black, legs yellow-brown.</paragraph>
<paragraph id="9C3736E3FFCDFF84F4F4FCB0FDE9FC03" blockId="11.[113,1089,620,1572]" pageId="11" pageNumber="172">Head as wide as long, narrowed posteriad from posterior margin of eyes, posterior angles each with one long and several short black bristles. Eyes as long as temples, slightly convex. Four punctures between eyes, distance between medial interocular punctures about 4 times as large as distance between medial and lateral interocular punctures. Posterior margin of eyes with 4 coarse punctures. Temporal areal almost impunctate. Dorsal surface with very fine irregular microsculpture here and there.</paragraph>
<paragraph id="9C3736E3FFCDFF84F4F4FC71FBB1FBE3" blockId="11.[113,1089,620,1572]" pageId="11" pageNumber="172">Antennae long and slender, reaching posterior fifth of pronotum when reclined. Antennomere 1 longer than antennomere 11, antennomere 2 shorter than antennomere 3. Relative length of antennomeres: 1 = 7.5; 2 = 4; 3 = 5; 46 = 3.5; 78 = 3; 910 = 2.5; 11 = 4.5.</paragraph>
<paragraph id="9C3736E3FFCDFF84F4F4FB91FCDBFB03" blockId="11.[113,1089,620,1572]" pageId="11" pageNumber="172">Pronotum highly convex, somewhat longer than wide (ratio 28: 25), slightly narrowed anteriad, posterior angles markedly rounded. Each dorsal row with 5 fine punctures, punctures 14 equidistant, distance between punctures 45 somewhat larger than distance between punctures 14. Each sublateral row with 2 punctures, puncture 1 situated approximately behind level of puncture 3 of dorsal row. Microsculpture similar to that on head.</paragraph>
<paragraph id="9C3736E3FFCDFF84F4F4FB70FE2DFAC3" blockId="11.[113,1089,620,1572]" pageId="11" pageNumber="172">Scutellum very coarsely and densely punctate, punctures larger than eye-facets, slightly contiguous here and there.</paragraph>
<paragraph id="9C3736E3FFCDFF84F4F4FAB0FDACFAA3" blockId="11.[113,1089,620,1572]" pageId="11" pageNumber="172">Elytra combined as wide as long, slightly widened posteriad. Punctation somewhat coarser and sparser than that on scutellum, separated by a puncture diameter, or somewhat smaller here and there; setation yellow-brown.</paragraph>
<paragraph id="9C3736E3FFCDFF84F4F4FAD1FD68FA63" blockId="11.[113,1089,620,1572]" pageId="11" pageNumber="172">Legs. Metatibia as long as metatarsus. Metatarsomere 1 as long as metatarsomere 5. Relative length of metatarsomeres: 1 = 6; 2 = 2.5; 34 = 2; 5 = 6.</paragraph>
<paragraph id="9C3736E3FFCDFF84F4F4FA10FC3CF9E3" blockId="11.[113,1089,620,1572]" pageId="11" pageNumber="172">Abdomen parallel-sided anteriorly, very gradually narrowed from visible tergite III towards apex. Elevated area between two basal lines on first three visible tergites impunctate. Punctation at base of all tergites finer and denser than that on elytra, gradually becoming finer and much sparser towards apex of each tergite. Setation similar to that on elytra.</paragraph>
<paragraph id="9C3736E3FFCAFF83F4D1FF67FF4BFEF3" blockId="12.[151,1127,219,564]" pageId="12" pageNumber="173">
Male. Protarsomeres 13 markedly dilated, sub-bilobed, covered with modified pale setae ventrally, protarsomere 4 distinctly narrower than preceding ones, triangular.Aedeagus (
<figureCitation id="04B32A66FFCAFF83F053FF47FF7FFEF3" captionStart="Figs" captionStartId="9.[112,151,1487,1506]" captionTargetBox="[148,1054,223,1459]" captionTargetId="figure-13@9.[142,1059,219,1468]" captionTargetPageId="9" captionText="Figs. 1224. 1213 Philonthus biguttulus Fauvel, 1907 (Côte dIvoire); 1416 P. bucorvus sp. nov.; 1720 P. caffer Boheman, 1848; 2124 Philonthus cliens Eppelsheim, 1890. 12, 14, 17, 21 aedeagus, ventral view, 13 paramere with sensory peg setae, ventral view; 15, 18, 22 aedeagus, lateral view; 16, 20 male sternite IX, ventral view; 19, 23 aedeagus without paramere, ventral view; 24 apex of paramere with sensory peg setae, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5404280" httpUri="https://zenodo.org/record/5404280/files/figure.png" pageId="12" pageNumber="173">Figs 2124</figureCitation>
).
</paragraph>
<paragraph id="9C3736E3FFCAFF83F4D1FE87FE23FE94" blockId="12.[151,1127,219,564]" box="[181,386,315,339]" pageId="12" pageNumber="173">Female. Unknown.</paragraph>
</subSubSection>
<subSubSection id="D4926568FFCAFF83F4F3FEE7FE2DFE34" pageId="12" pageNumber="173" type="diagnosis">
<paragraph id="9C3736E3FFCAFF83F4F3FEE7FE0EFE14" blockId="12.[151,1127,219,564]" pageId="12" pageNumber="173">
<emphasis id="AEFCEAF1FFCAFF83F4F3FEE7FE32FEB4" bold="true" box="[151,403,347,371]" pageId="12" pageNumber="173">Differential diagnosis.</emphasis>
<taxonomicName id="5B884D60FFCAFF83F5FFFEE1FDF4FEB3" authorityName=", Eppelsheim" authorityYear="1890" box="[411,597,349,372]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="173" phylum="Arthropoda" rank="species" species="cliens">
<emphasis id="AEFCEAF1FFCAFF83F5FFFEE1FDF4FEB3" box="[411,597,349,372]" italics="true" pageId="12" pageNumber="173">Philonthus cliens</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="5B884D60FFCAFF83F68CFEE1FCC6FEB3" authorityName="M. Bernhauer" authorityYear="1939" box="[744,871,349,372]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="173" phylum="Arthropoda" rank="species" species="kenyanus">
<emphasis id="AEFCEAF1FFCAFF83F68CFEE1FCC6FEB3" box="[744,871,349,372]" italics="true" pageId="12" pageNumber="173">P. kenyanus</emphasis>
</taxonomicName>
in most characters, but differens as follows: head from posterior margin of eyes distinctly narrowed posteriad, pronotum longer, punctation of elytra coarser and somewhat sparser. Both species also differ by the shape of the aedeagus.
</paragraph>
<paragraph id="9C3736E3FFCAFF83F4F3FE67FE2DFE34" blockId="12.[151,1127,219,564]" box="[151,396,475,499]" pageId="12" pageNumber="173">
<emphasis id="AEFCEAF1FFCAFF83F4F3FE67FEB4FE34" bold="true" box="[151,277,475,499]" pageId="12" pageNumber="173">Bionomics.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="D4926568FFCAFF83F4F3FE47FF7CFDF3" pageId="12" pageNumber="173" type="distribution">
<paragraph id="9C3736E3FFCAFF83F4F3FE47FF7CFDF3" blockId="12.[151,1127,219,564]" pageId="12" pageNumber="173">
<emphasis id="AEFCEAF1FFCAFF83F4F3FE47FE83FDD4" bold="true" box="[151,290,507,531]" pageId="12" pageNumber="173">Distribution</emphasis>
.
<collectingCountry id="E49F7673FFCAFF83F54BFE40FE3AFDD4" box="[303,411,508,531]" name="Botswana" pageId="12" pageNumber="173">Botswana</collectingCountry>
,
<collectingCountry id="E49F7673FFCAFF83F5C2FE47FE7CFDD4" box="[422,477,507,531]" name="India" pageId="12" pageNumber="173">India</collectingCountry>
(this paper);
<collectingCountry id="E49F7673FFCAFF83F60EFE47FD0DFDD4" box="[618,684,507,531]" name="Nepal" pageId="12" pageNumber="173">Nepal</collectingCountry>
,
<collectingCountry id="E49F7673FFCAFF83F6D3FE47FCABFDD4" box="[695,778,507,531]" name="Nigeria" pageId="12" pageNumber="173">Nigeria</collectingCountry>
,
<collectingCountry id="E49F7673FFCAFF83F770FE47FCCAFDD4" box="[788,875,507,531]" name="Senegal" pageId="12" pageNumber="173">Senegal</collectingCountry>
, Tanganyika (HERMAN 2001).
</paragraph>
</subSubSection>
</treatment>
</document>