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<document id="1E70C2753C12C1ED7228BCE13FCE79C2" ID-DOI="10.5281/zenodo.5404276" ID-ISSN="0374-1036" ID-Zenodo-Dep="5404276" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630132078781" checkinUser="felipe" docAuthor="Hromádka, Lubomír" docDate="2009" docId="142187F5FFC5FF8AF5AEFB86FECDFAF4" docLanguage="en" docName="ActaEntMusNatPra.2009.49.1.161-190.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 49 (1)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Philonthus aonyx Hromádka 2009, sp. nov." docType="treatment" docVersion="3" lastPageNumber="166" masterDocId="E818FF8DFFC6FF8FF464FFBCFFA1FFC7" masterDocTitle="Revision of the Afrotropical species of the Philonthus caffer species group (Coleoptera: Staphylinidae: Philonthina)" masterLastPageNumber="190" masterPageNumber="161" pageNumber="164" updateTime="1699162834734" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="7B52FBFD003F36F69CEFE1CDBDD5C334">Revision of the Afrotropical species of the Philonthus caffer species group (Coleoptera: Staphylinidae: Philonthina)</mods:title>
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<mods:namePart id="85839476F28C37C09678ED3BE0F726A3">Hromádka, Lubomír</mods:namePart>
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<mods:title id="5F2BE72398DF291F767BE2948DB6A9B1">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="C1E833C0022FBFF232CACAC2D2CA5B15">2009</mods:date>
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<mods:number id="75E0304AECAC81180D04093175DC1BE4">49</mods:number>
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<mods:identifier id="6EC3553700742C87EF388A283D7A34F6" type="ISSN">0374-1036</mods:identifier>
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<treatment id="142187F5FFC5FF8AF5AEFB86FECDFAF4" LSID="urn:lsid:plazi:treatment:142187F5FFC5FF8AF5AEFB86FECDFAF4" httpUri="http://treatment.plazi.org/id/142187F5FFC5FF8AF5AEFB86FECDFAF4" lastPageId="5" lastPageNumber="166" pageId="3" pageNumber="164">
<subSubSection id="D4926568FFC5FF8CF5AEFB86FD48FB93" box="[458,745,1082,1108]" pageId="3" pageNumber="164" type="nomenclature">
<paragraph id="9C3736E3FFC5FF8CF5AEFB86FD48FB93" blockId="3.[458,745,1082,1108]" box="[458,745,1082,1108]" pageId="3" pageNumber="164">
<heading id="C77F818FFFC5FF8CF5AEFB86FD48FB93" bold="true" box="[458,745,1082,1108]" centered="true" fontSize="8" level="2" pageId="3" pageNumber="164" reason="3">
<emphasis id="AEFCEAF1FFC5FF8CF5AEFB86FD48FB93" bold="true" box="[458,745,1082,1108]" pageId="3" pageNumber="164">
<taxonomicName id="5B884D60FFC5FF8CF5AEFB86FD2CFB94" ID-CoL="6V9VZ" authority="Hromádka, 2009" authorityName="Hromádka" authorityYear="2009" box="[458,653,1082,1107]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="164" phylum="Arthropoda" rank="species" species="aonyx" status="sp. nov.">
<emphasis id="AEFCEAF1FFC5FF8CF5AEFB86FD2CFB94" bold="true" box="[458,653,1082,1107]" italics="true" pageId="3" pageNumber="164">Philonthus aonyx</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B5CF578AFFC5FF8CF6F7FB80FD48FB93" box="[659,745,1084,1108]" pageId="3" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="D4926568FFC5FF8CF64EFBE6FD26FBAA" box="[554,647,1114,1133]" pageId="3" pageNumber="164" type="description">
<paragraph id="9C3736E3FFC5FF8CF64EFBE6FD26FBAA" blockId="3.[113,1090,1114,1301]" box="[554,647,1114,1133]" pageId="3" pageNumber="164">
(
<figureCitation id="04B32A66FFC5FF8CF654FBE6FD23FBAA" box="[560,642,1114,1133]" captionStart="Figs" captionStartId="4.[151,190,1507,1526]" captionTargetBox="[168,1121,220,1471]" captionTargetId="figure-10@4.[159,1121,210,1474]" captionTargetPageId="4" captionText="Figs. 111. 12 Philonthus aminius Tottenham, 1955; 37 P. aonyx sp. nov.; 811 P. biguttulus Fauvel, 1907 (89: Angola; 1011: Ethiopia). 1, 3, 8, 10 aedeagus, ventral view; 2 male sternite VIII, ventral view; 4 aedeagus, lateral view; 5 apex of paramere with sensory peg setae, ventral view; 6 apical portion of male sternite VIII, ventral view; 7 elytra; 9, 11 paramere with sensory peg setae, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5404278" httpUri="https://zenodo.org/record/5404278/files/figure.png" pageId="3" pageNumber="164">Figs. 37</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="D4926568FFC5FF8CF415FBC0FEC3FAD3" pageId="3" pageNumber="164" type="materials_examined">
<paragraph id="9C3736E3FFC5FF8CF415FBC0FE39FB48" blockId="3.[113,1090,1114,1301]" box="[113,408,1147,1167]" pageId="3" pageNumber="164">
<emphasis id="AEFCEAF1FFC5FF8CF415FBC0FF4BFB48" bold="true" box="[113,234,1147,1167]" pageId="3" pageNumber="164">
<typeStatus id="43338841FFC5FF8CF415FBC0FF3FFB48" box="[113,158,1148,1167]" pageId="3" pageNumber="164">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="E49F7673FFC5FF8CF48BFBC0FE9BFB48" box="[239,314,1148,1167]" name="Ethiopia" pageId="3" pageNumber="164">Ethiopia</collectingCountry>
, Bahr-Dar.
</paragraph>
<paragraph id="9C3736E3FFC5FF8CF415FB2BFD24FB18" blockId="3.[113,1090,1114,1301]" pageId="3" pageNumber="164">
<emphasis id="AEFCEAF1FFC5FF8CF415FB2BFF56FB6D" bold="true" box="[113,247,1174,1194]" pageId="3" pageNumber="164">Type material.</emphasis>
<materialsCitation id="2CE03CBEFFC5FF8CF499FB2BFCA8FB6D" box="[253,777,1170,1195]" collectingDate="1967-06-04" collectorName="P. Stys" country="Ethiopia" location="Bahr-Dar" pageId="3" pageNumber="164" specimenCount="1" typeStatus="holotype">
<typeStatus id="43338841FFC5FF8CF499FB2BFEF0FB6E" box="[253,337,1175,1194]" pageId="3" pageNumber="164" type="holotype">HOLOTYPE</typeStatus>
: J,
<collectingCountry id="E49F7673FFC5FF8CF518FB2BFE69FB6D" box="[380,456,1175,1194]" name="Ethiopia" pageId="3" pageNumber="164">Ethiopia</collectingCountry>
,
<location id="99576038FFC5FF8CF5BEFB2BFD8DFB6D" LSID="urn:lsid:plazi:treatment:142187F5FFC5FF8AF5AEFB86FECDFAF4:99576038FFC5FF8CF5BEFB2BFD8DFB6D" box="[474,556,1175,1194]" country="Ethiopia" name="Bahr-Dar" pageId="3" pageNumber="164">Bahr-Dar</location>
,
<date id="E8361023FFC5FF8CF651FB2BFD29FB6D" box="[565,648,1175,1194]" pageId="3" pageNumber="164" value="1967-06-04">
<collectingDate id="F872E9CBFFC5FF8CF651FB2BFD29FB6D" box="[565,648,1175,1194]" pageId="3" pageNumber="164" value="1967-06-04">4.vi.1967</collectingDate>
</date>
,
<collectorName id="317D5335FFC5FF8CF6F6FB2BFD6DFB6D" box="[658,716,1174,1194]" pageId="3" pageNumber="164">P. Štys</collectorName>
leg., //
</materialsCitation>
<materialsCitation id="2CE03CBEFFC5FF8CF774FB2BFD21FB03" collectionCode="NMPC" country="Ethiopia" determinerName="Hromadka" pageId="3" pageNumber="164" specimenCount="1" typeStatus="holotype">
<typeStatus id="43338841FFC5FF8CF774FB2BFC22FB6D" box="[784,899,1175,1194]" pageId="3" pageNumber="164" type="holotype">HOLOTYPE</typeStatus>
,
<taxonomicName id="5B884D60FFC5FF8CF7E8FB24FB83FB6D" authority="Hromádka, 2009" authorityName="Hromádka" authorityYear="2009" box="[908,1058,1176,1194]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="164" phylum="Arthropoda" rank="species" species="aonyx" status="sp. nov.">
<emphasis id="AEFCEAF1FFC5FF8CF7E8FB24FB83FB6D" box="[908,1058,1176,1194]" italics="true" pageId="3" pageNumber="164">Philonthus aonyx</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B5CF578AFFC5FF8CF04CFB2BFF36FB02" pageId="3" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
<determinerName id="69EBDA27FFC5FF8CF4FFFB0DFF54FB03" box="[155,245,1201,1220]" pageId="3" pageNumber="164">Hromádka</determinerName>
det.2008 [red oblong printed label] (
<collectionCode id="FA99AE26FFC5FF8CF65CFB0DFDD8FB03" box="[568,633,1201,1220]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="3" pageNumber="164">NMPC</collectionCode>
)
</materialsCitation>
.
<typeStatus id="43338841FFC5FF8CF6ECFB0EFD41FB04" box="[648,736,1202,1221]" pageId="3" pageNumber="164" type="paratype">PARATYPES</typeStatus>
: 1 J
<specimenCount id="8A8EFD6AFFC5FF8CF776FB0EFC93FB03" box="[786,818,1195,1221]" pageId="3" pageNumber="164" type="female">1 ♀</specimenCount>
, same label data as in
<typeStatus id="43338841FFC5FF8CF792FB0DFBE0FB03" box="[1014,1089,1201,1220]" pageId="3" pageNumber="164" type="holotype">holotype</typeStatus>
[both specimens with red oblong printed type-labels] (
<collectionCode id="FA99AE26FFC5FF8CF65BFB70FDDBFB18" box="[575,634,1228,1247]" pageId="3" pageNumber="164">LHPC</collectionCode>
).
</paragraph>
<paragraph id="9C3736E3FFC5FF8CF415FB5AFEC3FAD3" blockId="3.[113,1090,1114,1301]" pageId="3" pageNumber="164">
<emphasis id="AEFCEAF1FFC5FF8CF415FB5AFDA3FB3D" bold="true" box="[113,514,1254,1274]" pageId="3" pageNumber="164">
Additional material examined.
<collectingCountry id="E49F7673FFC5FF8CF5F4FB5AFE5FFB3D" box="[400,510,1254,1274]" name="Ethiopia" pageId="3" pageNumber="164">ETHIOPIA</collectingCountry>
:
</emphasis>
<specimenCount id="8A8EFD6AFFC5FF8CF66CFB5BFDE9FB3D" box="[520,584,1255,1274]" pageId="3" pageNumber="164" type="generic">5 spec.</specimenCount>
, Ethiopia, Arsi, Zasela,
<date id="E8361023FFC5FF8CF773FB5BFCCEFB3D" box="[791,879,1255,1274]" pageId="3" pageNumber="164" value="1988-12-07">7.xii.1988</date>
,
<quantity id="5B709B06FFC5FF8CF71EFB5BFC16FB3D" box="[890,951,1255,1274]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="3" pageNumber="164" unit="m" value="2400.0">2400m</quantity>
, cow dung, leg. S. Persson (
<collectionCode id="FA99AE26FFC5FF8CF4B1FABEFEAEFAD2" box="[213,271,1282,1301]" pageId="3" pageNumber="164">LHPC</collectionCode>
,
<collectionCode id="FA99AE26FFC5FF8CF57CFABDFEF9FAD3" box="[280,344,1281,1300]" country="Sweden" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34994" lsid="urn:lsid:biocol.org:col:34994" name="Lund University" pageId="3" pageNumber="164">MZLU</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="D4926568FFC5FF8AF415FA91FEFDFBD4" lastPageId="5" lastPageNumber="166" pageId="3" pageNumber="164" type="description">
<paragraph id="9C3736E3FFC5FF8CF415FA91FB9CFA83" blockId="3.[113,1089,1325,1605]" box="[113,1085,1325,1349]" pageId="3" pageNumber="164">
<emphasis id="AEFCEAF1FFC5FF8CF415FA91FF5DFA82" bold="true" box="[113,252,1325,1349]" pageId="3" pageNumber="164">Description.</emphasis>
Body length
<quantity id="5B709B06FFC5FF8CF5E8FA91FDBDFA83" box="[396,540,1325,1349]" metricMagnitude="-2" metricUnit="m" metricValue="1.005" metricValueMax="1.02" metricValueMin="0.99" pageId="3" pageNumber="164" unit="mm" value="10.05" valueMax="10.2" valueMin="9.9">9.910.2 mm</quantity>
, length of fore body (to end of elytra) 4.65.0 mm.
</paragraph>
<paragraph id="9C3736E3FFC5FF8CF4F4FAF1FE02FA22" blockId="3.[113,1089,1325,1605]" pageId="3" pageNumber="164">
Colouration. Head black, antennal sockets and clypeus along anterior margin narrowly yellow-brown, mandibles and pronotum black, maxillary and labial palpi brown-yellow, antennae dark brown, scutellum black, sides in anterior half very narrowly red, elytra black, each with red spot (
<figureCitation id="04B32A66FFC5FF8CF522FA11FE29FA02" box="[326,392,1453,1477]" captionStart="Figs" captionStartId="4.[151,190,1507,1526]" captionTargetBox="[168,1121,220,1471]" captionTargetId="figure-10@4.[159,1121,210,1474]" captionTargetPageId="4" captionText="Figs. 111. 12 Philonthus aminius Tottenham, 1955; 37 P. aonyx sp. nov.; 811 P. biguttulus Fauvel, 1907 (89: Angola; 1011: Ethiopia). 1, 3, 8, 10 aedeagus, ventral view; 2 male sternite VIII, ventral view; 4 aedeagus, lateral view; 5 apex of paramere with sensory peg setae, ventral view; 6 apical portion of male sternite VIII, ventral view; 7 elytra; 9, 11 paramere with sensory peg setae, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5404278" httpUri="https://zenodo.org/record/5404278/files/figure.png" pageId="3" pageNumber="164">Fig. 7</figureCitation>
), abdomen black with violet-green reflex, femora yellow-brown, tibiae and tarsi black-brown.
</paragraph>
<paragraph id="9C3736E3FFC5FF8AF4F4FA51FC90FEF4" blockId="3.[113,1089,1325,1605]" lastBlockId="5.[113,1090,219,1331]" lastPageId="5" lastPageNumber="166" pageId="3" pageNumber="164">Head rounded quadrangular, wider than long (ratio 35: 25), parallel-sided. Posterior angles obtusely rounded, each with several short bristles. Eyes flat, conspicuously shorter than temples (ratio 10: 13). Punctures between eyes arranged in straight line, distance between medial interocular punctures about 4 times as long as distance between medial and lateral interocular punctures. Temporal area and area along posterior margin of head with many punctures of variable size. Dorsal surface without microsculpture.</paragraph>
<caption id="C8F7666BFFC2FF8BF4F3FA5FFC96F981" ID-DOI="http://doi.org/10.5281/zenodo.5404278" ID-Zenodo-Dep="5404278" httpUri="https://zenodo.org/record/5404278/files/figure.png" pageId="4" pageNumber="165" startId="4.[151,190,1507,1526]" targetBox="[168,1121,220,1471]" targetPageId="4">
<paragraph id="9C3736E3FFC2FF8BF4F3FA5FFC96F981" blockId="4.[151,1129,1507,1606]" pageId="4" pageNumber="165">
Figs. 111. 12
<taxonomicName id="5B884D60FFC2FF8BF555FA58FDCEFA31" authority="Tottenham, 1955" authorityName=", Tottenham" authorityYear="1955" box="[305,623,1507,1526]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="165" phylum="Arthropoda" rank="species" species="aminius">
<emphasis id="AEFCEAF1FFC2FF8BF555FA58FE76FA31" box="[305,471,1508,1526]" italics="true" pageId="4" pageNumber="165">Philonthus aminius</emphasis>
<bibRefCitation id="F8194B12FFC2FF8BF5B8FA5FFDCEFA31" author="TOTTENHAM C. E." box="[476,623,1507,1526]" pageId="4" pageNumber="165" pagination="153 - 195" refId="ref16822" refString="TOTTENHAM C. E. 1955: Studies in the genus Philonthus Stephens (Coleoptera: Staphylinidae). Parts II, III, and IV. Transactions of the Royal Entomological Society of London 106: 153 - 195." type="journal article" year="1955">Tottenham, 1955</bibRefCitation>
</taxonomicName>
; 37
<taxonomicName id="5B884D60FFC2FF8BF6CAFA58FD57FA31" authority="Hromádka, 2009" authorityName="Hromádka" authorityYear="2009" box="[686,758,1508,1526]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="165" phylum="Arthropoda" rank="species" species="aonyx" status="sp. nov.">
<emphasis id="AEFCEAF1FFC2FF8BF6CAFA58FD57FA31" box="[686,758,1508,1526]" italics="true" pageId="4" pageNumber="165">P. aonyx</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B5CF578AFFC2FF8BF69FFA5FFCE0FA31" box="[763,833,1507,1526]" pageId="4" pageNumber="165" rank="species">sp. nov.</taxonomicNameLabel>
; 811
<taxonomicName id="5B884D60FFC2FF8BF7ECFA58FBC9FA31" authority="Fauvel, 1907" authorityName="Fauvel" authorityYear="1907" box="[904,1128,1507,1526]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="165" phylum="Arthropoda" rank="species" species="biguttulus">
<emphasis id="AEFCEAF1FFC2FF8BF7ECFA58FC52FA31" box="[904,1011,1508,1526]" italics="true" pageId="4" pageNumber="165">P. biguttulus</emphasis>
<bibRefCitation id="F8194B12FFC2FF8BF79DFA5FFBC9FA31" author="FAUVEL A." box="[1017,1128,1507,1526]" pageId="4" pageNumber="165" pagination="10 - 70" refId="ref16384" refString="FAUVEL A. 1907: Voyage de M. Ch. Alluaud dans l'Afrique Orientale. Staphylinidae. Revue d'Entomologie 26: 10 - 70." type="journal article" year="1907">Fauvel, 1907</bibRefCitation>
</taxonomicName>
(89: Angola; 1011: Ethiopia). 1, 3, 8, 10 aedeagus, ventral view; 2 male sternite VIII, ventral view; 4 aedeagus, lateral view; 5 apex of paramere with sensory peg setae, ventral view; 6 apical portion of male sternite VIII, ventral view; 7 elytra; 9, 11 paramere with sensory peg setae, ventral view.
</paragraph>
</caption>
<paragraph id="9C3736E3FFC3FF8AF4F4FE80FCD5FE54" blockId="5.[113,1090,219,1331]" pageId="5" pageNumber="166">Antennae reaching posterior third of pronotum when reclined. Antennomere 1 somewhat shorter than antennomeres 23 combined, antennomere 11 as long as antennomere 2. Relative length of antennomeres: 1 = 9; 2 = 5; 3 = 6; 47 = 4.5; 810 = 4; 11 = 5.</paragraph>
<paragraph id="9C3736E3FFC3FF8AF4F4FE20FCF2FDD4" blockId="5.[113,1090,219,1331]" pageId="5" pageNumber="166">Pronotum highly convex, slightly transverse (ratio 42: 40), distinctly narrowed anteriad. Anterior angles bluntly rounded, with several short bristles, posterior angles markedly rounded. Each dorsal row with 4 equidistant punctures. Each sublateral row with 2 punctures, puncture 2 distinctly shifted to lateral margin. Surface without microsculpture.</paragraph>
<paragraph id="9C3736E3FFC3FF8AF4F4FDA7FCC2FD94" blockId="5.[113,1090,219,1331]" pageId="5" pageNumber="166">Entire scutellum finely and sparsely punctate, punctures smaller than eye-facets, separated by 2 puncture diameters in transverse direction. Setation longer, black.</paragraph>
<paragraph id="9C3736E3FFC3FF8AF4F4FDE7FCE9FD74" blockId="5.[113,1090,219,1331]" pageId="5" pageNumber="166">
Elytra combined (
<figureCitation id="04B32A66FFC3FF8AF52AFDE7FE2FFDB3" box="[334,398,603,628]" captionStart="Figs" captionStartId="4.[151,190,1507,1526]" captionTargetBox="[168,1121,220,1471]" captionTargetId="figure-10@4.[159,1121,210,1474]" captionTargetPageId="4" captionText="Figs. 111. 12 Philonthus aminius Tottenham, 1955; 37 P. aonyx sp. nov.; 811 P. biguttulus Fauvel, 1907 (89: Angola; 1011: Ethiopia). 1, 3, 8, 10 aedeagus, ventral view; 2 male sternite VIII, ventral view; 4 aedeagus, lateral view; 5 apex of paramere with sensory peg setae, ventral view; 6 apical portion of male sternite VIII, ventral view; 7 elytra; 9, 11 paramere with sensory peg setae, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5404278" httpUri="https://zenodo.org/record/5404278/files/figure.png" pageId="5" pageNumber="166">Fig. 7</figureCitation>
) distinctly wider than long (ratio 50: 42), parallel-sided. Punctation very fine and sparse, punctures somewhat larger than those on scutellum, separated by 2 puncture diameters in transverse direction; setation brown-yellow.
</paragraph>
<paragraph id="9C3736E3FFC3FF8AF4F4FD00FEC3FCD4" blockId="5.[113,1090,219,1331]" pageId="5" pageNumber="166">Legs. Metatibia somewhat shorter than metatarsus (ratio 27: 30). Metatarsomere 1 as long as metatarsomeres 45 combined. Relative length of metatarsomeres: 1 = 10; 2 = 4.4; 3 = 4; 4 = 3.5; 5 = 6.5.</paragraph>
<paragraph id="9C3736E3FFC3FF8AF4F4FCA7FED3FC54" blockId="5.[113,1090,219,1331]" pageId="5" pageNumber="166">Abdomen wide, punctation of tergites finer and denser than that on elytra, becoming sparser and finer towards posterior margin of each tergite. Elevated area between two basal lines on first three visible tergites relatively densely punctate. Surface without microsculpture; setation similar to that on elytra.</paragraph>
<paragraph id="9C3736E3FFC3FF8AF4F4FC27FEF7FC34" blockId="5.[113,1090,219,1331]" pageId="5" pageNumber="166">
Male. Protarsomeres 13 dilated and sub-bilobed, densely covered with modified pale setae ventrally, protarsomere 4 distinctly narrower than preceding ones. Sternite VIII (
<figureCitation id="04B32A66FFC3FF8AF79DFC07FB95FC14" box="[1017,1076,955,979]" captionStart="Figs" captionStartId="4.[151,190,1507,1526]" captionTargetBox="[168,1121,220,1471]" captionTargetId="figure-10@4.[159,1121,210,1474]" captionTargetPageId="4" captionText="Figs. 111. 12 Philonthus aminius Tottenham, 1955; 37 P. aonyx sp. nov.; 811 P. biguttulus Fauvel, 1907 (89: Angola; 1011: Ethiopia). 1, 3, 8, 10 aedeagus, ventral view; 2 male sternite VIII, ventral view; 4 aedeagus, lateral view; 5 apex of paramere with sensory peg setae, ventral view; 6 apical portion of male sternite VIII, ventral view; 7 elytra; 9, 11 paramere with sensory peg setae, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5404278" httpUri="https://zenodo.org/record/5404278/files/figure.png" pageId="5" pageNumber="166">Fig.6</figureCitation>
), aedeagus (
<figureCitation id="04B32A66FFC3FF8AF486FC67FEEBFC34" box="[226,330,987,1011]" captionStart="Figs" captionStartId="4.[151,190,1507,1526]" captionTargetBox="[168,1121,220,1471]" captionTargetId="figure-10@4.[159,1121,210,1474]" captionTargetPageId="4" captionText="Figs. 111. 12 Philonthus aminius Tottenham, 1955; 37 P. aonyx sp. nov.; 811 P. biguttulus Fauvel, 1907 (89: Angola; 1011: Ethiopia). 1, 3, 8, 10 aedeagus, ventral view; 2 male sternite VIII, ventral view; 4 aedeagus, lateral view; 5 apex of paramere with sensory peg setae, ventral view; 6 apical portion of male sternite VIII, ventral view; 7 elytra; 9, 11 paramere with sensory peg setae, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5404278" httpUri="https://zenodo.org/record/5404278/files/figure.png" pageId="5" pageNumber="166">Figs. 35</figureCitation>
).
</paragraph>
<paragraph id="9C3736E3FFC3FF8AF4F4FC47FEFDFBD4" blockId="5.[113,1090,219,1331]" box="[144,348,1019,1043]" pageId="5" pageNumber="166">Female. Unknown.</paragraph>
</subSubSection>
<subSubSection id="D4926568FFC3FF8AF415FBA7FE5FFB74" pageId="5" pageNumber="166" type="diagnosis">
<paragraph id="9C3736E3FFC3FF8AF415FBA7FE5FFB74" blockId="5.[113,1090,219,1331]" pageId="5" pageNumber="166">
<emphasis id="AEFCEAF1FFC3FF8AF415FBA7FECCFBF4" bold="true" box="[113,365,1051,1075]" pageId="5" pageNumber="166">Differential diagnosis.</emphasis>
The new species is similar to
<taxonomicName id="5B884D60FFC3FF8AF6DDFBA1FC9CFBF3" authorityName="M. Bernhauer" authorityYear="1932" box="[697,829,1053,1076]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="166" phylum="Arthropoda" rank="species" species="iridicollis">
<emphasis id="AEFCEAF1FFC3FF8AF6DDFBA1FC9CFBF3" box="[697,829,1053,1076]" italics="true" pageId="5" pageNumber="166">P. iridicollis</emphasis>
</taxonomicName>
but it differs by the red spot on each elytron, by the yellow-brown femora and black- brown tibiae and tarsi. It differs from
<taxonomicName id="5B884D60FFC3FF8AF4C8FBE1FE9FFBB3" authorityName=", Levasseur" authorityYear="1966" box="[172,318,1117,1140]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="166" phylum="Arthropoda" rank="species" species="gabonensis">
<emphasis id="AEFCEAF1FFC3FF8AF4C8FBE1FE9FFBB3" box="[172,318,1117,1140]" italics="true" pageId="5" pageNumber="166">P. gabonensis</emphasis>
</taxonomicName>
by the red spot on each elytron, the wider head, the differently coloured antennomere 1 and by the coarser punctation of the abdomen. It differs from both above species by the shape of the aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="D4926568FFC3FF8AF415FB00FEC7FAD4" pageId="5" pageNumber="166" type="etymology">
<paragraph id="9C3736E3FFC3FF8AF415FB00FDF0FB33" blockId="5.[113,1090,219,1331]" pageId="5" pageNumber="166">
<emphasis id="AEFCEAF1FFC3FF8AF415FB00FF53FB13" bold="true" box="[113,242,1212,1236]" pageId="5" pageNumber="166">Etymology.</emphasis>
The name of this species, a noun in apposition, is the Latin generic name of the African otter
<taxonomicName id="5B884D60FFC3FF8AF565FB61FEE5FB33" authorityName="Hromádka" authorityYear="2009" box="[257,324,1245,1268]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="166" phylum="Arthropoda" rank="species" species="aonyx">
<emphasis id="AEFCEAF1FFC3FF8AF565FB61FEE5FB33" box="[257,324,1245,1268]" italics="true" pageId="5" pageNumber="166">Aonyx</emphasis>
</taxonomicName>
<taxonomicName id="5B884D60FFC3FF8AF52EFB61FDECFB33" authority="(Schinz, 1821)" baseAuthorityName="Schinz" baseAuthorityYear="1821" box="[330,589,1243,1268]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="166" phylum="Arthropoda" rank="species" species="capensis">
<emphasis id="AEFCEAF1FFC3FF8AF52EFB61FE07FB33" box="[330,422,1245,1268]" italics="true" pageId="5" pageNumber="166">capensis</emphasis>
(Schinz, 1821)
</taxonomicName>
.
</paragraph>
<paragraph id="9C3736E3FFC3FF8AF415FB47FEC7FAD4" blockId="5.[113,1090,219,1331]" box="[113,358,1275,1299]" pageId="5" pageNumber="166">
<emphasis id="AEFCEAF1FFC3FF8AF415FB47FF4EFAD4" bold="true" box="[113,239,1275,1299]" pageId="5" pageNumber="166">Bionomics.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="D4926568FFC3FF8AF415FAA7FECDFAF4" box="[113,364,1307,1331]" pageId="5" pageNumber="166" type="distribution">
<paragraph id="9C3736E3FFC3FF8AF415FAA7FECDFAF4" blockId="5.[113,1090,219,1331]" box="[113,364,1307,1331]" pageId="5" pageNumber="166">
<emphasis id="AEFCEAF1FFC3FF8AF415FAA7FEA5FAF4" bold="true" box="[113,260,1307,1331]" pageId="5" pageNumber="166">Distribution.</emphasis>
<collectingCountry id="E49F7673FFC3FF8AF56EFAA7FEC9FAF4" box="[266,360,1307,1331]" name="Ethiopia" pageId="5" pageNumber="166">Ethiopia</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>