treatments-xml/data/14/21/87/142187F5FFC3FF89F5F2FAE6FEE2F9E8.xml
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<document id="1964CFADE7F072FDDC0133F078D15F2A" ID-DOI="10.5281/zenodo.5404276" ID-ISSN="0374-1036" ID-Zenodo-Dep="5404276" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630132078781" checkinUser="felipe" docAuthor="Hromádka, Lubomír" docDate="2009" docId="142187F5FFC3FF89F5F2FAE6FEE2F9E8" docLanguage="en" docName="ActaEntMusNatPra.2009.49.1.161-190.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 49 (1)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Philonthus biguttulus Fauvel 1907" docType="treatment" docVersion="3" lastPageNumber="167" masterDocId="E818FF8DFFC6FF8FF464FFBCFFA1FFC7" masterDocTitle="Revision of the Afrotropical species of the Philonthus caffer species group (Coleoptera: Staphylinidae: Philonthina)" masterLastPageNumber="190" masterPageNumber="161" pageNumber="166" updateTime="1699162834734" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="E9002F7DBF917F002FE4C807157E9B4A">Revision of the Afrotropical species of the Philonthus caffer species group (Coleoptera: Staphylinidae: Philonthina)</mods:title>
</mods:titleInfo>
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<mods:namePart id="D29AA8F0BA1C5BB1FC9AF7273D926ABB">Hromádka, Lubomír</mods:namePart>
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<mods:title id="7485F3DBC9925134DFFFB38F6505C914">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="34D343C9FCC52F1766F86F850D0B4C82">2009</mods:date>
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<mods:number id="2407BD1BB143DEE1FBEAA00939251174">2009-06-30</mods:number>
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<mods:number id="E8C1E2EB334220838F87127E46516E1F">49</mods:number>
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<mods:number id="C8B04C3C91DD5E3BCF3AEDDA7A5E48C6">1</mods:number>
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<mods:identifier id="B6C684DB6415B15177C08D7A95CCA18B" type="ISSN">0374-1036</mods:identifier>
<mods:identifier id="FAC5227EFE02203C9B6445B215F70C97" type="Zenodo-Dep">5404276</mods:identifier>
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<treatment id="142187F5FFC3FF89F5F2FAE6FEE2F9E8" LSID="urn:lsid:plazi:treatment:142187F5FFC3FF89F5F2FAE6FEE2F9E8" httpUri="http://treatment.plazi.org/id/142187F5FFC3FF89F5F2FAE6FEE2F9E8" lastPageId="6" lastPageNumber="167" pageId="5" pageNumber="166">
<subSubSection id="D4926568FFC3FF8AF5F2FAE6FCBAFAB4" box="[406,795,1370,1396]" pageId="5" pageNumber="166" type="nomenclature">
<paragraph id="9C3736E3FFC3FF8AF5F2FAE6FCBAFAB4" blockId="5.[406,795,1370,1396]" box="[406,795,1370,1396]" pageId="5" pageNumber="166">
<heading id="C77F818FFFC3FF8AF5F2FAE6FCBAFAB4" bold="true" box="[406,795,1370,1396]" centered="true" fontSize="8" level="2" pageId="5" pageNumber="166" reason="3">
<taxonomicName id="5B884D60FFC3FF8AF5F2FAE6FCBAFAB4" ID-CoL="76XHH" authority="Fauvel, 1907" authorityName="Fauvel" authorityYear="1907" box="[406,795,1370,1396]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="166" phylum="Arthropoda" rank="species" species="biguttulus">
<emphasis id="AEFCEAF1FFC3FF8AF5F2FAE6FCBAFAB4" bold="true" box="[406,795,1370,1396]" pageId="5" pageNumber="166">
<emphasis id="AEFCEAF1FFC3FF8AF5F2FAE6FD24FAB4" bold="true" box="[406,645,1370,1395]" italics="true" pageId="5" pageNumber="166">Philonthus biguttulus</emphasis>
<bibRefCitation id="F8194B12FFC3FF8AF6E8FAE0FCBAFAB4" author="FAUVEL A." box="[652,795,1371,1396]" pageId="5" pageNumber="166" pagination="10 - 70" refId="ref16384" refString="FAUVEL A. 1907: Voyage de M. Ch. Alluaud dans l'Afrique Orientale. Staphylinidae. Revue d'Entomologie 26: 10 - 70." type="journal article" year="1907">Fauvel, 1907</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="D4926568FFC3FF8AF641FAC6FD2CFA4A" box="[549,653,1402,1421]" pageId="5" pageNumber="166" type="description">
<paragraph id="9C3736E3FFC3FF8AF641FAC6FD2CFA4A" blockId="5.[113,1090,1402,1570]" box="[549,653,1402,1421]" pageId="5" pageNumber="166">
(
<figureCitation id="04B32A66FFC3FF8AF64FFAC6FD26FA4A" box="[555,647,1402,1421]" captionStart-0="Figs" captionStart-1="Figs" captionStartId-0="4.[151,190,1507,1526]" captionStartId-1="9.[112,151,1487,1506]" captionTargetBox-0="[168,1121,220,1471]" captionTargetBox-1="[148,1054,223,1459]" captionTargetId-0="figure-10@4.[159,1121,210,1474]" captionTargetId-1="figure-13@9.[142,1059,219,1468]" captionTargetPageId-0="4" captionTargetPageId-1="9" captionText-0="Figs. 111. 12 Philonthus aminius Tottenham, 1955; 37 P. aonyx sp. nov.; 811 P. biguttulus Fauvel, 1907 (89: Angola; 1011: Ethiopia). 1, 3, 8, 10 aedeagus, ventral view; 2 male sternite VIII, ventral view; 4 aedeagus, lateral view; 5 apex of paramere with sensory peg setae, ventral view; 6 apical portion of male sternite VIII, ventral view; 7 elytra; 9, 11 paramere with sensory peg setae, ventral view." captionText-1="Figs. 1224. 1213 Philonthus biguttulus Fauvel, 1907 (Côte dIvoire); 1416 P. bucorvus sp. nov.; 1720 P. caffer Boheman, 1848; 2124 Philonthus cliens Eppelsheim, 1890. 12, 14, 17, 21 aedeagus, ventral view, 13 paramere with sensory peg setae, ventral view; 15, 18, 22 aedeagus, lateral view; 16, 20 male sternite IX, ventral view; 19, 23 aedeagus without paramere, ventral view; 24 apex of paramere with sensory peg setae, ventral view." figureDoi-0="http://doi.org/10.5281/zenodo.5404278" figureDoi-1="http://doi.org/10.5281/zenodo.5404280" httpUri-0="https://zenodo.org/record/5404278/files/figure.png" httpUri-1="https://zenodo.org/record/5404280/files/figure.png" pageId="5" pageNumber="166">Figs. 813</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="D4926568FFC3FF8AF415FA21FE65FA68" box="[113,452,1436,1455]" pageId="5" pageNumber="166" type="reference_group">
<paragraph id="9C3736E3FFC3FF8AF415FA21FE65FA68" blockId="5.[113,1090,1402,1570]" box="[113,452,1436,1455]" pageId="5" pageNumber="166">
<treatmentCitationGroup id="BC9811CDFFC3FF8AF415FA21FE65FA68" box="[113,452,1436,1455]" pageId="5" pageNumber="166">
<treatmentCitation id="1D2910F2FFC3FF8AF415FA21FE61FA68" author="FAUVEL A." box="[113,448,1436,1455]" page="44" pageId="5" pageNumber="166" year="1907">
<taxonomicName id="5B884D60FFC3FF8AF415FA21FE61FA68" ID-CoL="76XHH" authority="Fauvel, 1907: 44" authorityName="Fauvel" authorityPageNumber="44" authorityYear="1907" box="[113,448,1436,1455]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="166" phylum="Arthropoda" rank="species" species="biguttulus">
<emphasis id="AEFCEAF1FFC3FF8AF415FA21FE89FA68" box="[113,296,1437,1455]" italics="true" pageId="5" pageNumber="166">Philonthus biguttulus</emphasis>
<bibRefCitation id="F8194B12FFC3FF8AF54AFA20FE61FA68" author="FAUVEL A." box="[302,448,1436,1455]" pageId="5" pageNumber="166" pagination="10 - 70" refId="ref16384" refString="FAUVEL A. 1907: Voyage de M. Ch. Alluaud dans l'Afrique Orientale. Staphylinidae. Revue d'Entomologie 26: 10 - 70." type="journal article" year="1907">Fauvel, 1907: 44</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="D4926568FFC3FF89F415FA02FB98FEF8" lastPageId="6" lastPageNumber="167" pageId="5" pageNumber="166" type="materials_examined">
<paragraph id="9C3736E3FFC3FF8AF415FA02FE1BFA15" blockId="5.[113,1090,1402,1570]" box="[113,442,1470,1490]" pageId="5" pageNumber="166">
<emphasis id="AEFCEAF1FFC3FF8AF415FA02FF4BFA15" bold="true" box="[113,234,1470,1490]" pageId="5" pageNumber="166">
<typeStatus id="43338841FFC3FF8AF415FA02FF3FFA16" box="[113,158,1470,1489]" pageId="5" pageNumber="166">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="E49F7673FFC3FF8AF48AFA02FEE7FA16" box="[238,326,1470,1489]" name="Ethiopia" pageId="5" pageNumber="166">Abyssinie</collectingCountry>
, Escarpment.
</paragraph>
<paragraph id="9C3736E3FFC3FF8AF415FA65FEC4FA2B" blockId="5.[113,1090,1402,1570]" box="[113,357,1496,1516]" pageId="5" pageNumber="166">
<emphasis id="AEFCEAF1FFC3FF8AF415FA65FF57FA2B" bold="true" box="[113,246,1496,1516]" pageId="5" pageNumber="166">
<typeStatus id="43338841FFC3FF8AF415FA65FF3FFA2B" box="[113,158,1497,1516]" pageId="5" pageNumber="166">Type</typeStatus>
material.
</emphasis>
<collectingRegion id="5E4CF801FFC3FF8AF49FFA65FEBAFA2B" box="[251,283,1497,1516]" country="Cameroon" name="North" pageId="5" pageNumber="166">Not</collectingRegion>
studied.
</paragraph>
<paragraph id="9C3736E3FFC3FF8AF415FA4FFBE1F9E6" blockId="5.[113,1090,1402,1570]" pageId="5" pageNumber="166">
<emphasis id="AEFCEAF1FFC3FF8AF415FA4FFEDEF9C0" bold="true" box="[113,383,1523,1543]" pageId="5" pageNumber="166">Additional material examined.</emphasis>
<materialsCitation id="2CE03CBEFFC3FF8AF5E6FA4FFCFBF9E6" collectingDate="1961-01" collectionCode="MRAC" collectorName="Rives &amp; Calundo &amp; Aug. &amp; A. de Barros Machado &amp; Mus. Tervuren" country="Angola" determinerName="C. E. Tottenham" location="ANGOLA" pageId="5" pageNumber="166" specimenCount="1">
<emphasis id="AEFCEAF1FFC3FF8AF5E6FA4FFE44F9C0" bold="true" box="[386,485,1523,1543]" pageId="5" pageNumber="166">
<collectingCountry id="E49F7673FFC3FF8AF5E6FA4FFE43F9C0" box="[386,482,1523,1543]" name="Angola" pageId="5" pageNumber="166">ANGOLA</collectingCountry>
:
</emphasis>
1J.,
<collectorName id="317D5335FFC3FF8AF67CFA48FDEAF9C0" box="[536,587,1524,1543]" pageId="5" pageNumber="166">Rives</collectorName>
du lac
<collectorName id="317D5335FFC3FF8AF6E7FA48FD6BF9C0" box="[643,714,1524,1543]" pageId="5" pageNumber="166">Calundo</collectorName>
lumiere,
<collectorName id="317D5335FFC3FF8AF771FA48FCE1F9C0" box="[789,832,1524,1543]" pageId="5" pageNumber="166">Aug.</collectorName>
1.1955,
<collectorName id="317D5335FFC3FF8AF7E7FA48FB9CF9C0" box="[899,1085,1524,1543]" pageId="5" pageNumber="166">A. de Barros Machado</collectorName>
, coll.
<collectorName id="317D5335FFC3FF8AF4FFF9B3FEB6F9E5" box="[155,279,1551,1570]" pageId="5" pageNumber="166">Mus.Tervuren</collectorName>
//
<taxonomicName id="5B884D60FFC3FF8AF54FF9B3FDB7F9E5" authority="Fauv." authorityName="Fauvel" authorityYear="1907" box="[299,534,1551,1570]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="166" phylum="Arthropoda" rank="species" species="biguttulus">
<emphasis id="AEFCEAF1FFC3FF8AF54FF9B3FE40F9E6" box="[299,481,1551,1569]" italics="true" pageId="5" pageNumber="166">Philonthus biguttulus</emphasis>
Fauv.
</taxonomicName>
det.
<determinerName id="69EBDA27FFC3FF8AF659F9B2FD67F9E6" box="[573,710,1550,1570]" pageId="5" pageNumber="166">C. E. Tottenham</determinerName>
<date id="E8361023FFC3FF8AF6AEF9B2FCA3F9E6" box="[714,770,1550,1569]" pageId="5" pageNumber="166" value="1961-01">
<collectingDate id="F872E9CBFFC3FF8AF6AEF9B2FCA3F9E6" box="[714,770,1550,1569]" pageId="5" pageNumber="166" value="1961-01">i.1961</collectingDate>
</date>
(
<collectionCode id="FA99AE26FFC3FF8AF774F9B2FCF5F9E6" box="[784,852,1550,1569]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de lAfrique Centrale" pageId="5" pageNumber="166" type="Museum">MRAC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="2CE03CBEFFC3FF8AF705F9B2FBE1F9E6" box="[865,1088,1544,1570]" collectorName="de Canavezes" country="Angola" location="Marco de Canavezes" pageId="5" pageNumber="166" specimenCount="1" specimenCount-female="1">
<specimenCount id="8A8EFD6AFFC3FF8AF705F9B2FCDEF9E6" box="[865,895,1544,1569]" pageId="5" pageNumber="166" type="female">1 ♀</specimenCount>
,
<location id="99576038FFC3FF8AF7E9F9B2FBE1F9E6" LSID="urn:lsid:plazi:treatment:142187F5FFC3FF89F5F2FAE6FEE2F9E8:99576038FFC3FF8AF7E9F9B2FBE1F9E6" box="[909,1088,1550,1569]" country="Angola" name="Marco de Canavezes" pageId="5" pageNumber="166">
Marco
<collectorName id="317D5335FFC3FF8AF7A9F9B2FBE1F9E6" box="[973,1088,1550,1569]" pageId="5" pageNumber="166">de Canavezes</collectorName>
</location>
</materialsCitation>
</paragraph>
<paragraph id="9C3736E3FFC0FF89F4F3FF67FB98FEF8" blockId="6.[151,1127,219,319]" pageId="6" pageNumber="167">
(distr.Benguela)
<materialsCitation id="2CE03CBEFFC0FF89F542FF67FBC2FF28" box="[294,1123,219,239]" collectingDate="1956-03" collectingDateMax="1961-02" collectingDateMin="1956-03" collectionCode="MRAC" collectorName="E. Luna de Caravalho" determinerName="C. E. Tottenham" location="Luna de Caravalho" pageId="6" pageNumber="167" specimenCount="1">
<date id="E8361023FFC0FF89F542FF67FECDFF29" box="[294,364,219,238]" pageId="6" pageNumber="167" value="1956-03">
<collectingDate id="F872E9CBFFC0FF89F542FF67FECDFF29" box="[294,364,219,238]" pageId="6" pageNumber="167" value="1956-03">iii. 1956</collectingDate>
</date>
,
<collectorName id="317D5335FFC0FF89F517FF60FD86FF29" box="[371,551,219,239]" pageId="6" pageNumber="167">
E.
<location id="99576038FFC0FF89F5EDFF60FD86FF29" LSID="urn:lsid:plazi:treatment:142187F5FFC3FF89F5F2FAE6FEE2F9E8:99576038FFC0FF89F5EDFF60FD86FF29" box="[393,551,219,239]" name="Luna de Caravalho" pageId="6" pageNumber="167">Luna de Caravalho</location>
</collectorName>
//
<taxonomicName id="5B884D60FFC0FF89F652FF60FCBDFF28" authority="Fauv." authorityName="Fauvel" authorityYear="1907" box="[566,796,220,239]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="167" phylum="Arthropoda" rank="species" species="biguttulus">
<emphasis id="AEFCEAF1FFC0FF89F652FF60FD48FF29" box="[566,745,220,238]" italics="true" pageId="6" pageNumber="167">Philonthus biguttulus</emphasis>
Fauv.
</taxonomicName>
det.
<determinerName id="69EBDA27FFC0FF89F725FF60FC6BFF29" box="[833,970,219,239]" pageId="6" pageNumber="167">C. E. Tottenham</determinerName>
,
<date id="E8361023FFC0FF89F7B4FF67FBACFF29" box="[976,1037,219,238]" pageId="6" pageNumber="167" value="1961-02">
<collectingDate id="F872E9CBFFC0FF89F7B4FF67FBACFF29" box="[976,1037,219,238]" pageId="6" pageNumber="167" value="1961-02">ii.1961</collectingDate>
</date>
(
<collectionCode id="FA99AE26FFC0FF89F07EFF60FBFCFF28" box="[1050,1117,220,239]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de lAfrique Centrale" pageId="6" pageNumber="167" type="Museum">MRAC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="2CE03CBEFFC0FF89F4F3FF49FCC8FECE" box="[151,873,241,266]" collectingDate="1998-04-27" collectionCode="LHPC" country="Ivory Coast" latitude="8.748334" location="Komoe" longLatPrecision="21" longitude="-3.8133333" pageId="6" pageNumber="167" specimenCount="1" stateProvince="Comoe">
<emphasis id="AEFCEAF1FFC0FF89F4F3FF49FE9FFECE" bold="true" box="[151,318,245,265]" pageId="6" pageNumber="167">
<collectingCountry id="E49F7673FFC0FF89F4F3FF49FE9BFECE" box="[151,314,245,265]" name="Ivory Coast" pageId="6" pageNumber="167">CÔTE dIVOIRE</collectingCountry>
:
</emphasis>
1J,
<collectingRegion id="5E4CF801FFC0FF89F508FF4AFE0FFECE" box="[364,430,246,265]" country="Ivory Coast" name="Comoe" pageId="6" pageNumber="167">Komoe</collectingRegion>
NP,
<date id="E8361023FFC0FF89F5B0FF4AFD8EFECE" box="[468,559,246,265]" pageId="6" pageNumber="167" value="1998-04-27">
<collectingDate id="F872E9CBFFC0FF89F5B0FF4AFD8EFECE" box="[468,559,246,265]" pageId="6" pageNumber="167" value="1998-04-27">27.iv.1998</collectingDate>
</date>
, lux,
<geoCoordinate id="F9BC5024FFC0FF89F63EFF4AFD1BFECE" box="[602,698,245,265]" degrees="3" direction="west" minutes="48" orientation="longitude" pageId="6" pageNumber="167" precision="15" seconds="48" value="-3.8133333">3°4848″W</geoCoordinate>
/
<geoCoordinate id="F9BC5024FFC0FF89F6A5FF4AFCBFFECE" box="[705,798,245,265]" degrees="8" direction="north" minutes="44" orientation="latitude" pageId="6" pageNumber="167" precision="15" seconds="54" value="8.748334">8°4454″N</geoCoordinate>
(
<collectionCode id="FA99AE26FFC0FF89F74CFF4AFCC2FECE" box="[808,867,246,265]" pageId="6" pageNumber="167">LHPC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="2CE03CBEFFC0FF89F714FF49FB94FEF8" collectingDate="1967-06-04" collectingDateMax="1993-11-27" collectingDateMin="1967-06-04" collectionCode="LHPC, MYLU" collectorName="P. Stys &amp; Lund University Sierra Leone Expedition &amp; L. Cederholm &amp; R. Danielsson" country="Ethiopia" latitude="8.883333" location="Makeni" longLatPrecision="1300" longitude="-12.05" municipality="Bahr-Dar" pageId="6" pageNumber="167" specimenCount="62">
<emphasis id="AEFCEAF1FFC0FF89F714FF49FC40FECE" bold="true" box="[880,993,245,265]" pageId="6" pageNumber="167">
<collectingCountry id="E49F7673FFC0FF89F714FF49FC7CFECE" box="[880,989,245,265]" name="Ethiopia" pageId="6" pageNumber="167">ETHIOPIA</collectingCountry>
:
</emphasis>
1J,
<collectingMunicipality id="7C53AC99FFC0FF89F06BFF4AFBC5FECE" box="[1039,1124,246,265]" pageId="6" pageNumber="167">Bahr-Dar</collectingMunicipality>
,
<date id="E8361023FFC0FF89F4F3FEADFF4BFEE3" box="[151,234,273,292]" pageId="6" pageNumber="167" value="1967-06-04">
<collectingDate id="F872E9CBFFC0FF89F4F3FEADFF4BFEE3" box="[151,234,273,292]" pageId="6" pageNumber="167" value="1967-06-04">4.vi.1967</collectingDate>
</date>
,
<collectorName id="317D5335FFC0FF89F495FEADFE89FEE3" box="[241,296,272,292]" pageId="6" pageNumber="167">P.Štys</collectorName>
leg. (
<collectionCode id="FA99AE26FFC0FF89F53EFEADFE34FEE3" box="[346,405,273,292]" pageId="6" pageNumber="167">LHPC</collectionCode>
),
<emphasis id="AEFCEAF1FFC0FF89F5C6FEACFDE6FEE3" bold="true" box="[418,583,272,292]" pageId="6" pageNumber="167">
<collectingCountry id="E49F7673FFC0FF89F5C6FEACFDE2FEE3" box="[418,579,272,292]" name="Sierra Leone" pageId="6" pageNumber="167">SIERRA LEONE</collectingCountry>
:
</emphasis>
<specimenCount id="8A8EFD6AFFC0FF89F62EFEADFD33FEE3" box="[586,658,273,292]" pageId="6" pageNumber="167" type="generic">62 spec.</specimenCount>
,
<location id="99576038FFC0FF89F6FCFEADFD7AFEE3" LSID="urn:lsid:plazi:treatment:142187F5FFC3FF89F5F2FAE6FEE2F9E8:99576038FFC0FF89F6FCFEADFD7AFEE3" box="[664,731,273,292]" country="Ethiopia" latitude="8.883333" longLatPrecision="1300" longitude="-12.05" municipality="Bahr-Dar" name="Makeni" pageId="6" pageNumber="167">Makeni</location>
,
<geoCoordinate id="F9BC5024FFC0FF89F686FEADFC8CFEE3" box="[738,813,272,292]" degrees="12" direction="west" minutes="03" orientation="longitude" pageId="6" pageNumber="167" precision="925" value="-12.05">12°03W</geoCoordinate>
,
<geoCoordinate id="F9BC5024FFC0FF89F751FEADFCD2FEE3" box="[821,883,272,292]" degrees="8" direction="north" minutes="53" orientation="latitude" pageId="6" pageNumber="167" precision="925" value="8.883333">8°53N</geoCoordinate>
,
<date id="E8361023FFC0FF89F71EFEADFC76FEE3" box="[890,983,273,292]" pageId="6" pageNumber="167" value="1993-11-27">
<collectingDate id="F872E9CBFFC0FF89F71EFEADFC76FEE3" box="[890,983,273,292]" pageId="6" pageNumber="167" value="1993-11-27">27.xi.1993</collectingDate>
</date>
, loc., 9,
<collectingMethod id="45C94EF4FFC0FF89F07FFEADFBC7FEE3" box="[1051,1126,273,292]" pageId="6" pageNumber="167">light trap</collectingMethod>
18-21,
<collectorName id="317D5335FFC0FF89F4B0FE97FD97FEF9" box="[212,566,299,319]" pageId="6" pageNumber="167">Lund University Sierra Leone Expedition</collectorName>
1993, leg.,
<collectorName id="317D5335FFC0FF89F6FFFE90FCB0FEF9" box="[667,785,299,319]" pageId="6" pageNumber="167">L. Cederholm</collectorName>
<collectorName id="317D5335FFC0FF89F743FE90FC3FFEF9" box="[807,926,299,319]" pageId="6" pageNumber="167">R. Danielsson</collectorName>
(
<collectionCode id="FA99AE26FFC0FF89F7CDFE90FC4AFEF8" box="[937,1003,300,319]" pageId="6" pageNumber="167">MYLU</collectionCode>
,
<collectionCode id="FA99AE26FFC0FF89F790FE90FB8FFEF8" box="[1012,1070,300,319]" pageId="6" pageNumber="167">LHPC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="D4926568FFC0FF89F4F3FEEBFE23FAA8" pageId="6" pageNumber="167" type="description">
<paragraph id="9C3736E3FFC0FF89F4F3FEEBFF5CFE49" blockId="6.[151,1127,343,1583]" pageId="6" pageNumber="167">
<emphasis id="AEFCEAF1FFC0FF89F4F3FEEBFE9AFEA8" bold="true" box="[151,315,343,367]" pageId="6" pageNumber="167">Redescription</emphasis>
. Body length 7.88.0 mm, length of fore body (to end of elytra) 3.7
<quantity id="5B709B06FFC0FF89F4C1FECBFF5BFE49" box="[165,250,375,399]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="6" pageNumber="167" unit="mm" value="3.9">3.9 mm</quantity>
.
</paragraph>
<paragraph id="9C3736E3FFC0FF89F4D1FE2BFCFDFE28" blockId="6.[151,1127,343,1583]" pageId="6" pageNumber="167">Colouration. Head, pronotum, scutellum, elytra and abdomen black, only posterior fourth of elytra red, maxillary and labial palpi, antennomere 1 and base of antennomere 2 brownyellow, remaining antennomeres black-brown, legs yellow-brown.</paragraph>
<paragraph id="9C3736E3FFC0FF89F4D1FE4BFC18FD48" blockId="6.[151,1127,343,1583]" pageId="6" pageNumber="167">Head longer than wide (ratio 24: 21), posterior angles obtusely rounded, with one long, black bristle. Eyes slightly projecting, vaguely shorter than temples (ratio 9: 10.5). Distance between medial interocular punctures about 4 times as long as distance between medial and lateral interocular punctures. Lateral interocular punctures slightly shifted anteriad. Temporal area with many punctures of variable size. Surface without microsculpture.</paragraph>
<paragraph id="9C3736E3FFC0FF89F4D1FD2BFD61FD28" blockId="6.[151,1127,343,1583]" pageId="6" pageNumber="167">Antennae reaching posterior fourth of pronotum when reclined. Antennomere 1 somewhat longer than antennomere 11, antennomere 2 shorter than antennomere 3. Relative length of antennomeres: 1 = 6; 2 = 4; 3 = 5; 410 = 3; 11 = 5.</paragraph>
<paragraph id="9C3736E3FFC0FF89F4D1FD4BFCB3FC68" blockId="6.[151,1127,343,1583]" pageId="6" pageNumber="167">Pronotum highly convex, distinctly longer than wide (ratio 30: 24), parallel-sided.Anterior angles and lateral margins each with several short bristles. Posterior angles markedly rounded. Each dorsal row with 5 punctures, punctures 14 equidistant, interval between punctures 4 and 5 about 1.5 times as large as distance between punctures 1 and 2. Each sublateral row with 2 punctures, puncture 1 situated behind level of punctures 23 of dorsal rows, puncture 2 distinctly shifted laterad. Surface without microsculpture.</paragraph>
<paragraph id="9C3736E3FFC0FF89F4D1FC0BFC8BFC28" blockId="6.[151,1127,343,1583]" pageId="6" pageNumber="167">Scutellum very densely and coarsely punctate, punctures somewhat larger than eye-facets, distance between punctures much smaller than their diameter.</paragraph>
<paragraph id="9C3736E3FFC0FF89F4D1FC4BFD2AFBE8" blockId="6.[151,1127,343,1583]" pageId="6" pageNumber="167">Elytra combined as long as wide, very slightly widened posteriad. Punctation similar to that on scutellum. Setation fine, brown-yellow.</paragraph>
<paragraph id="9C3736E3FFC0FF89F4D1FB8BFC0DFB48" blockId="6.[151,1127,343,1583]" pageId="6" pageNumber="167">Legs. Metatibia longer than metatarsus (ratio 26: 21). Metatarsomere 1 somewhat longer than metatarsomeres 23 combined, metatarsomere 5 as long as metatarsomeres 34 combined. Relative length of metatarsomeres: 1 = 7; 2 = 3.5; 3 = 3; 4 = 3; 5 = 6.</paragraph>
<paragraph id="9C3736E3FFC0FF89F4D1FB2BFDABFAC8" blockId="6.[151,1127,343,1583]" pageId="6" pageNumber="167">Abdomen parallel-sided, slightly narrowed from tergite V posteriad. Elevated area between two basal lines on first four visible tergites almost impunctate. Entire visible tergites very densely and coarsely punctate, most punctures of raindrop shape. Distance between punctures much smaller than their diameters.</paragraph>
<paragraph id="9C3736E3FFC0FF89F4D1FAABFCC6FA88" blockId="6.[151,1127,343,1583]" pageId="6" pageNumber="167">
Male. Protarsomeres 13 strongly dilated, each densely covered with modified pale setae ventrally, protarsomere 4 narrow and small. Aedeagus (
<figureCitation id="04B32A66FFC0FF89F682FA8BFCFBFA88" box="[742,858,1335,1359]" captionStart-0="Figs" captionStart-1="Figs" captionStartId-0="4.[151,190,1507,1526]" captionStartId-1="9.[112,151,1487,1506]" captionTargetBox-0="[168,1121,220,1471]" captionTargetBox-1="[148,1054,223,1459]" captionTargetId-0="figure-10@4.[159,1121,210,1474]" captionTargetId-1="figure-13@9.[142,1059,219,1468]" captionTargetPageId-0="4" captionTargetPageId-1="9" captionText-0="Figs. 111. 12 Philonthus aminius Tottenham, 1955; 37 P. aonyx sp. nov.; 811 P. biguttulus Fauvel, 1907 (89: Angola; 1011: Ethiopia). 1, 3, 8, 10 aedeagus, ventral view; 2 male sternite VIII, ventral view; 4 aedeagus, lateral view; 5 apex of paramere with sensory peg setae, ventral view; 6 apical portion of male sternite VIII, ventral view; 7 elytra; 9, 11 paramere with sensory peg setae, ventral view." captionText-1="Figs. 1224. 1213 Philonthus biguttulus Fauvel, 1907 (Côte dIvoire); 1416 P. bucorvus sp. nov.; 1720 P. caffer Boheman, 1848; 2124 Philonthus cliens Eppelsheim, 1890. 12, 14, 17, 21 aedeagus, ventral view, 13 paramere with sensory peg setae, ventral view; 15, 18, 22 aedeagus, lateral view; 16, 20 male sternite IX, ventral view; 19, 23 aedeagus without paramere, ventral view; 24 apex of paramere with sensory peg setae, ventral view." figureDoi-0="http://doi.org/10.5281/zenodo.5404278" figureDoi-1="http://doi.org/10.5281/zenodo.5404280" httpUri-0="https://zenodo.org/record/5404278/files/figure.png" httpUri-1="https://zenodo.org/record/5404280/files/figure.png" pageId="6" pageNumber="167">Figs. 813</figureCitation>
).
</paragraph>
<paragraph id="9C3736E3FFC0FF89F4D1FAEBFE23FAA8" blockId="6.[151,1127,343,1583]" box="[181,386,1367,1391]" pageId="6" pageNumber="167">Female. Unknown.</paragraph>
</subSubSection>
<subSubSection id="D4926568FFC0FF89F4F3FACBFE2DFA28" pageId="6" pageNumber="167" type="diagnosis">
<paragraph id="9C3736E3FFC0FF89F4F3FACBFD92FA08" blockId="6.[151,1127,343,1583]" pageId="6" pageNumber="167">
<emphasis id="AEFCEAF1FFC0FF89F4F3FACBFE2DFA48" bold="true" box="[151,396,1399,1423]" pageId="6" pageNumber="167">Differential diagnosis.</emphasis>
<taxonomicName id="5B884D60FFC0FF89F5F4FAC4FDCEFA48" authorityName="Fauvel" authorityYear="1907" box="[400,623,1400,1423]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="167" phylum="Arthropoda" rank="species" species="biguttulus">
<emphasis id="AEFCEAF1FFC0FF89F5F4FAC4FDCEFA48" box="[400,623,1400,1423]" italics="true" pageId="6" pageNumber="167">Philonthus biguttulus</emphasis>
</taxonomicName>
may be distinguished from the habitually similar
<taxonomicName id="5B884D60FFC0FF89F4F3FA24FEA4FA68" authorityName="Tottenham" authorityYear="1962" box="[151,261,1432,1455]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="167" phylum="Arthropoda" rank="species" species="tomicus">
<emphasis id="AEFCEAF1FFC0FF89F4F3FA24FEA4FA68" box="[151,261,1432,1455]" italics="true" pageId="6" pageNumber="167">P. tomicus</emphasis>
</taxonomicName>
by longer eyes, shorter antennae, elytra without red spot, yellow-brown legs and by the different shape of the aedeagus.
</paragraph>
<paragraph id="9C3736E3FFC0FF89F4F3FA6BFE2DFA28" blockId="6.[151,1127,343,1583]" box="[151,396,1495,1519]" pageId="6" pageNumber="167">
<emphasis id="AEFCEAF1FFC0FF89F4F3FA6BFEB4FA28" bold="true" box="[151,277,1495,1519]" pageId="6" pageNumber="167">Bionomics.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="D4926568FFC0FF89F4F3FA4BFEE2F9E8" pageId="6" pageNumber="167" type="distribution">
<paragraph id="9C3736E3FFC0FF89F4F3FA4BFEE2F9E8" blockId="6.[151,1127,343,1583]" pageId="6" pageNumber="167">
<emphasis id="AEFCEAF1FFC0FF89F4F3FA4BFE8BF9C8" bold="true" box="[151,298,1527,1551]" pageId="6" pageNumber="167">Distribution.</emphasis>
<collectingCountry id="E49F7673FFC0FF89F555FA4BFE23F9C8" box="[305,386,1527,1551]" name="Angola" pageId="6" pageNumber="167">Angola</collectingCountry>
,
<collectingCountry id="E49F7673FFC0FF89F5EBFA4BFD83F9C8" box="[399,546,1527,1551]" name="Ivory Coast" pageId="6" pageNumber="167">Côte dIvoire</collectingCountry>
,
<collectingCountry id="E49F7673FFC0FF89F64BFA4BFD2DF9C8" box="[559,652,1527,1551]" name="Ethiopia" pageId="6" pageNumber="167">Ethiopia</collectingCountry>
,
<collectingCountry id="E49F7673FFC0FF89F6FDFA4BFC84F9C9" box="[665,805,1527,1551]" name="Sierra Leone" pageId="6" pageNumber="167">Sierra Leone</collectingCountry>
(this paper);
<collectingCountry id="E49F7673FFC0FF89F7DEFA4BFBA3F9C8" box="[954,1026,1527,1551]" name="Kenya" pageId="6" pageNumber="167">Kenya</collectingCountry>
,
<collectingCountry id="E49F7673FFC0FF89F06BFA4BFBC7F9C8" box="[1039,1126,1527,1551]" name="Rwanda" pageId="6" pageNumber="167">Rwanda</collectingCountry>
(HERMAN 2001).
</paragraph>
</subSubSection>
</treatment>
</document>