treatments-xml/data/08/5A/B2/085AB225FFE237472397FEDEFE64FEAC.xml

<document id="F5744998633ED55E5B1D97059F7CDBEC" ID-DOI="10.11646/phytotaxa.625.1.5" ID-ISSN="1179-3163" ID-Zenodo-Dep="10149605" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1700230626821" checkinUser="plazi" docAuthor="Gonzáles, James Alexander Chamaya, Hágsater, Eric, Ayala, Elizabeth Santiago, Salinas, Julian Duarte &amp; Vela, Gustavo Iberico" docDate="2023" docId="085AB225FFE237472397FEDEFE64FEAC" docLanguage="en" docName="phytotaxa.625.1.5.pdf" docOrigin="Phytotaxa 625 (1)" docSource="https://phytotaxa.mapress.com/pt/article/download/phytotaxa.625.1.5/51270" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.7:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="7" docTitle="Epidendrum bicornialpicola Hagsater, Chamaya &amp; Iberico. A. Plant 2023, sp. nov." docType="treatment" docVersion="1" lastPageNumber="86" masterDocId="F463CA5DFFE5374E231FFFF0FFF6FFC1" masterDocTitle="Three new species of Epidendrum (Orchidaceae: Laeliinae) from the cloud forests of Chota, northern Peru" masterLastPageNumber="87" masterPageNumber="77" pageNumber="84" updateTime="1700663291293" updateUser="juliana" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="FFFED78B7B6BFBC83410FA98B99D016E">Three new species of Epidendrum (Orchidaceae: Laeliinae) from the cloud forests of Chota, northern Peru</mods:title>
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<mods:namePart id="40D2E94CEFD26E04A89DA9BA93B90F9E">Gonzáles, James Alexander Chamaya</mods:namePart>
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<mods:affiliation id="77FBE847265EE1C8BD51869AEA35192A">Escuela de Posgrado, Unidad de Posgrado de la Facultad de Ciencias Agrarias, Mención en Gestión ambiental, Universidad Nacional de Cajamarca, Perú, Avenida Atahualpa Km. 3, Cajamarca, Perú &amp; chamaya 982 @ gmail. com, jchamayag _ epg 19 @ unc. edu. pe; https: // orcid. org / 0009 - 0004 - 3214 - 5420</mods:affiliation>
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<mods:namePart id="31FE12DBEA8D233D3DA83D8B73F8AC0B">Hágsater, Eric</mods:namePart>
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<mods:affiliation id="AEBC093C1F4618714C1FD711FADF54DB">Herbario AMO, Montañas Calizas 490, Miguel Hidalgo, CDMX 11000, México. Ames Herbarium, Harvard University Herbaria, Cambridge, MA 02138 &amp; erichag _ 1 @ msn. com; https: // orcid. org / 0000 - 0002 - 2371 - 9427</mods:affiliation>
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<mods:namePart id="229827EEA9F9477817DF9A01FEDE358F">Ayala, Elizabeth Santiago</mods:namePart>
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<mods:affiliation id="CC8B822DAD359C940A06593FE0AB8BB8">Herbario AMO, Montañas Calizas 490, Miguel Hidalgo, CDMX 11000, México. Ames Herbarium, Harvard University Herbaria, Cambridge, MA 02138 &amp; sayalae 2014 @ gmail. com; https: // orcid. org / 0000 - 0002 - 1368 - 9025</mods:affiliation>
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<mods:namePart id="C2BD2008217086C89521F9D3A2625470">Salinas, Julian Duarte</mods:namePart>
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<mods:affiliation id="016915873925851EF261B2F300C0AE35">Herbario AMO, Montañas Calizas 490, Miguel Hidalgo, CDMX 11000, México. Ames Herbarium, Harvard University Herbaria, Cambridge, MA 02138 &amp; julianduartesalinas @ gmail. com; https: // orcid. org / 0000 - 0003 - 4655 - 5274</mods:affiliation>
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<mods:namePart id="80DFD43744C4D61B091D26BEC906E613">Vela, Gustavo Iberico</mods:namePart>
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<mods:affiliation id="3D815622F139548A7B78B27B5C29C926">Docente y director del Herbario CPUN “ Isidoro Sánchez Vega ” de la Universidad Nacional de Cajamarca &amp; giberico @ unc. edu. pe; https: // orcid. org / 0000 - 0003 - 3301 - 5527</mods:affiliation>
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<mods:date id="D50BDD87038A92DE84CAABCC246CF2AA">2023</mods:date>
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<taxonomicName id="47F378B0FFE237492397FEDEFCECFE89" authority="Hagsater, Chamaya &amp; Iberico" authorityName="Hagsater, Chamaya &amp; Iberico. A. Plant" authorityYear="2023" box="[136,794,302,328]" class="Liliopsida" family="Orchidaceae" genus="Epidendrum" kingdom="Plantae" order="Asparagales" pageId="7" pageNumber="84" phylum="Tracheophyta" rank="species" species="bicornialpicola" status="sp. nov.">
<emphasis id="B287DF21FFE237492397FEDEFE30FE89" box="[136,454,302,328]" italics="true" pageId="7" pageNumber="84">Epidendrum bicornialpicola</emphasis>
Hágsater, Chamaya &amp; Iberico
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, 
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<taxonomicNameLabel id="A9B4625AFFE237492037FEDFFC8BFE89" box="[808,893,303,328]" pageId="7" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
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(
<figureCitation id="18C81FB6FFE237492094FEDEFC00FE89" box="[907,1014,302,328]" captionStart="FIGURE 1" captionStartId="2.[136,229,1953,1975]" captionTargetBox="[196,1392,194,1923]" captionTargetId="figure-25@2.[189,1398,187,1925]" captionTargetPageId="2" captionText="FIGURE 1. Distribution map of new species of Epidendrum. Prepared by James Chamaya." figureDoi="http://doi.org/10.5281/zenodo.10149607" httpUri="https://zenodo.org/record/10149607/files/figure.png" pageId="7" pageNumber="84">Figures 1</figureCitation>
, 
<figureCitation id="18C81FB6FFE23749271BFEDEFBE5FE89" box="[1028,1043,302,328]" captionStart="FIGURE 4" captionStartId="8.[136,229,1952,1974]" captionTargetBox="[195,1393,190,1928]" captionTargetId="figure-25@8.[195,1393,190,1928]" captionTargetPageId="8" captionText="FIGURE 4. Lankester Composite Dissection Plate (LCDP) of Epidendrum bicornialpicola Hágsater, Chamaya &amp; Iberico. A. Plant habit. B. Inflorescence. C. Flower, lateral view. D. Dissected perianth. E. Ovary with lip and column; upper and lateral views. F. Pollinarium. Prepared by Anaís Cisneros from the images by J. A. Chamaya taken from the specimen that served as the holotype." figureDoi="http://doi.org/10.5281/zenodo.10149614" httpUri="https://zenodo.org/record/10149614/files/figure.png" pageId="7" pageNumber="84">4</figureCitation>
)
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<emphasis id="B287DF21FFE237492397FE8AFF4CFE4E" bold="true" box="[136,186,378,399]" pageId="7" pageNumber="84">Type</emphasis>
: 
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<materialsCitation id="309B096EFFE2374923C6FE8AFAFEFE4E" box="[217,1288,378,400]" collectingDate="2022-07-19" collectionCode="CPUN" collectorName="J. A. Chamaya" country="Peru" elevation="2778" location="Bosque La Palma" municipality="Chota" pageId="7" pageNumber="84" specimenCount="1" stateProvince="Cajamarca" typeStatus="holotype">
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: 
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: 
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: 
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, 
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, 
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, 
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G. 104
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(
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: 
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!)
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.
</paragraph>
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<taxonomicName id="47F378B0FFE237492397FE4EFE3FFE19" authorityName="Hagsater, Chamaya &amp; Iberico. A. Plant" authorityYear="2023" box="[136,457,446,472]" class="Liliopsida" family="Orchidaceae" genus="Epidendrum" kingdom="Plantae" order="Asparagales" pageId="7" pageNumber="84" phylum="Tracheophyta" rank="species" species="bicornialpicola">
<emphasis id="B287DF21FFE237492397FE4EFE3FFE19" box="[136,457,446,472]" italics="true" pageId="7" pageNumber="84">Epidendrum bicornialpicola</emphasis>
</taxonomicName>
is similar to 
<taxonomicName id="47F378B0FFE23749217CFE4FFBE8FE19" authority="Hagsater &amp; Cisneros (2020" authorityName="Hagsater &amp; Cisneros" authorityYear="2020" box="[611,1054,446,472]" class="Liliopsida" family="Orchidaceae" genus="Epidendrum" kingdom="Plantae" order="Asparagales" pageId="7" pageNumber="84" phylum="Tracheophyta" rank="species" species="delsyae">
<emphasis id="B287DF21FFE23749217CFE4FFD2EFE19" box="[611,728,446,472]" italics="true" pageId="7" pageNumber="84">E. delsyae</emphasis>
<bibRefCitation id="E4627EC2FFE2374921FEFE4EFBE8FE19" author="Hagsater, E. &amp; Cisneros, A." box="[737,1054,446,472]" pageId="7" pageNumber="84" pagination="1809" refId="ref6069" refString="Hagsater, E. &amp; Cisneros, A. (2020) Epidendrum delsyae. In: Hagsater, E. &amp; Santiago, E. (Eds.) Icones Orchidacearum, Fascicle 18 (1) The Genus Epidendrum, Part 14. Herbario Amo, t. 1809." type="book chapter" year="2020">Hágsater &amp; Cisneros (2020</bibRefCitation>
</taxonomicName>
: t. 1809) but the leaves are half as long, (3.2–6.0 cm long 
<emphasis id="B287DF21FFE237492290FE13FE5BFE3D" box="[399,429,483,508]" italics="true" pageId="7" pageNumber="84">vs.</emphasis>
<quantity id="470BAED6FFE2374922ABFE12FDC9FE3C" box="[436,575,482,509]" metricMagnitude="-1" metricUnit="m" metricValue="1.04" metricValueMax="1.34" metricValueMin="0.74" pageId="7" pageNumber="84" unit="cm" value="10.4" valueMax="13.4" valueMin="7.4">7.4–13.4 cm</quantity>
long), pale yellow-green flowers with a lip ochre (
<emphasis id="B287DF21FFE237492764FE13FB6CFE3D" box="[1147,1178,483,508]" italics="true" pageId="7" pageNumber="84">vs.</emphasis>
ochre flowers with a lip ochre-orange), smaller sepals (
<quantity id="470BAED6FFE2374922FFFDF6FD9FFDE0" box="[480,617,518,545]" metricMagnitude="-3" metricUnit="m" metricValue="7.65" metricValueMax="8.6" metricValueMin="6.7" pageId="7" pageNumber="84" unit="mm" value="7.65" valueMax="8.6" valueMin="6.7">6.7–8.6 mm</quantity>
long 
<emphasis id="B287DF21FFE2374921B7FDF7FD31FDE1" box="[680,711,519,544]" italics="true" pageId="7" pageNumber="84">vs.</emphasis>
<quantity id="470BAED6FFE2374921D1FDF6FCAAFDE0" box="[718,860,518,545]" metricMagnitude="-2" metricUnit="m" metricValue="1.14" metricValueMax="1.28" metricValueMin="1.0" pageId="7" pageNumber="84" unit="mm" value="11.4" valueMax="12.8" valueMin="10.0">10–12.8 mm</quantity>
long), and the lip with calli upright, acute, divergent (
<emphasis id="B287DF21FFE23749238EFDDBFF46FD85" box="[145,176,555,580]" italics="true" pageId="7" pageNumber="84">vs.</emphasis>
calli non upright, rounded, slightly paralell).
</paragraph>
</subSubSection>
<subSubSection id="C8E950B8FFE2374923A2FDBEFAF1FA64" pageId="7" pageNumber="84" type="description">
<paragraph id="804C0333FFE2374923A2FDBEFAF1FA64" blockId="7.[136,1452,446,2021]" pageId="7" pageNumber="84">
<emphasis id="B287DF21FFE2374923A2FDBEFE9BFDA9" bold="true" box="[189,365,590,616]" pageId="7" pageNumber="84">Description:—</emphasis>
Lithophytic and epiphytic, sympodial, caespitose 
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, ca. 
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tall (including inflorescence). 
<emphasis id="B287DF21FFE237492397FD83FF3EFD4D" box="[136,200,627,652]" italics="true" pageId="7" pageNumber="84">Roots</emphasis>
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in diameter, basal, thick, white. 
<emphasis id="B287DF21FFE2374921A6FD83FD0CFD4C" box="[697,762,627,653]" italics="true" pageId="7" pageNumber="84">Stems</emphasis>
44 × 
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, simple, cane-like, terete below, laterally compressed above, erect to arching; basal 1/3 covered by papyraceous sheaths ca. 4.0 × 
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, light brown. 
<emphasis id="B287DF21FFE237492641FD67FA5AFD71" box="[1374,1452,663,688]" italics="true" pageId="7" pageNumber="84">Leaves</emphasis>
7, articulate, alternate, sub-coriaceous and lower leaves smaller, spreading; sheaths 2.0–3.0 × 
<quantity id="470BAED6FFE2374927D4FD4AFABDFD14" box="[1227,1355,698,725]" metricMagnitude="-3" metricUnit="m" metricValue="3" metricValueMax="4.0" metricValueMin="2.0" pageId="7" pageNumber="84" unit="cm" value="0.3" valueMax="0.4" valueMin="0.2">0.2–0.4 cm</quantity>
tubular, somewhat laterally compressed, glabrous; blades 3.2–6.0 × 
<quantity id="470BAED6FFE23749202FFD2EFC47FD38" box="[816,945,734,761]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.2" metricValueMin="0.8" pageId="7" pageNumber="84" unit="cm" value="1.0" valueMax="1.2" valueMin="0.8">0.8–1.2 cm</quantity>
, 5:1, linear-lanceolate, acuminate, sub-erect, margin entire, spreading, medium green. 
<emphasis id="B287DF21FFE237492144FCF2FD44FCDD" box="[603,690,770,796]" italics="true" pageId="7" pageNumber="84">Spathes</emphasis>
1–2, 3.0–6.0 × 
<quantity id="470BAED6FFE237492077FCF2FC1FFCDC" box="[872,1001,770,797]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="7" pageNumber="84" unit="cm" value="0.25" valueMax="0.3" valueMin="0.2">0.2–0.3 cm</quantity>
, tubular, acuminate, lower one leaf-like towards apex, slightly imbricating. 
<emphasis id="B287DF21FFE237492107FCD6FD58FC81" box="[536,686,806,832]" italics="true" pageId="7" pageNumber="84">Inflorescence</emphasis>
<quantity id="470BAED6FFE2374921AAFCD6FD0DFC80" box="[693,763,806,833]" metricMagnitude="-1" metricUnit="m" metricValue="1.7" pageId="7" pageNumber="84" unit="cm" value="17.0">17 cm</quantity>
long, apical, a sub-densely flowered raceme, arching nutant; peduncle 
<quantity id="470BAED6FFE2374923E9FCBAFEB5FCA4" box="[246,323,842,869]" metricMagnitude="-2" metricUnit="m" metricValue="8.5" pageId="7" pageNumber="84" unit="cm" value="8.5">8.5 cm</quantity>
long, erect, straight, nearly totally hidden by spathes, rachis 
<quantity id="470BAED6FFE2374920E4FCBAFBC5FCA4" box="[1019,1075,842,869]" metricMagnitude="-2" metricUnit="m" metricValue="8.0" pageId="7" pageNumber="84" unit="cm" value="8.0">8 cm</quantity>
long, strongly arched and nutant at base, nearly straight thereafter. 
<emphasis id="B287DF21FFE23749210AFC9EFD5BFC49" box="[533,685,878,904]" italics="true" pageId="7" pageNumber="84">Floral bracts</emphasis>
<quantity id="470BAED6FFE2374921A7FC9EFD0BFC48" box="[696,765,878,905]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="7" pageNumber="84" unit="mm" value="4.0">4 mm</quantity>
long, prominent, nearly as long as ovary, linear-triangular, acuminate, embracing. 
<emphasis id="B287DF21FFE237492290FC62FE1DFC6D" box="[399,491,914,940]" italics="true" pageId="7" pageNumber="84">Flowers</emphasis>
ca. 24, simultaneous, resupinate, apical lip always pointing to axis of rachis, flowers pale yellow-green, lip ochre, column white; fragrance reminiscent of lavender during daytime. 
<emphasis id="B287DF21FFE2374927A6FC47FB09FC10" box="[1209,1279,951,977]" italics="true" pageId="7" pageNumber="84">Ovary</emphasis>
4.5–9.0 × 1.0– 
<quantity id="470BAED6FFE237492397FC2AFF29FC34" box="[136,223,986,1013]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="7" pageNumber="84" unit="mm" value="1.1">1.1 mm</quantity>
, terete, thin, arching, not inflated, unornamented, furrowed, green. 
<emphasis id="B287DF21FFE2374920D6FC2AFBE4FC35" box="[969,1042,986,1012]" italics="true" pageId="7" pageNumber="84">Sepals</emphasis>
free, fleshy, 3-veined, margin entire, spreading; dorsal sepal 6.4–6.7 × 
<quantity id="470BAED6FFE23749211BFC0EFD78FBD8" box="[516,654,1022,1049]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" metricValueMax="2.7" metricValueMin="2.5" pageId="7" pageNumber="84" unit="mm" value="2.6" valueMax="2.7" valueMin="2.5">2.5–2.7 mm</quantity>
, somewhat concave, obovate-elliptic, acuminate; lateral sepals 8.4–8.6 × 
<quantity id="470BAED6FFE237492380FBD2FED1FBFC" box="[159,295,1058,1085]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" metricValueMax="2.9" metricValueMin="2.7" pageId="7" pageNumber="84" unit="mm" value="2.8" valueMax="2.9" valueMin="2.7">2.7–2.9 mm</quantity>
, partly concave, ovate-elliptic, acuminate, oblique, aristate, with low dorsal keel that continues apically to form an aristate tip, margin of keel entire. 
<emphasis id="B287DF21FFE237492167FBB6FD48FBA1" box="[632,702,1094,1120]" italics="true" pageId="7" pageNumber="84">Petals</emphasis>
4.5–5.0 × 
<quantity id="470BAED6FFE23749202BFBB6FC4AFBA0" box="[820,956,1094,1121]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="7" pageNumber="84" unit="mm" value="0.55" valueMax="0.6" valueMin="0.5">0.5–0.6 mm</quantity>
, free, spreading, linear oblanceolate-oblong, acuminate, 1-veined, margin entire, spreading. 
<emphasis id="B287DF21FFE23749218CFB9BFD4EFB45" box="[659,696,1131,1156]" italics="true" pageId="7" pageNumber="84">Lip</emphasis>
4.3–4.5 × 
<quantity id="470BAED6FFE237492030FB9AFC43FB44" box="[815,949,1130,1157]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" metricValueMax="3.8" metricValueMin="3.6" pageId="7" pageNumber="84" unit="mm" value="3.7" valueMax="3.8" valueMin="3.6">3.6–3.8 mm</quantity>
(with spread lateral lobes), united to column, fleshy, 3-lobed, base cordate; bicallose, calli prominent, laminar, short, acute, upright, divergent; disc with a very short thick broad mid-rib, extending beyond calli to base of mid-lobe; lateral lobes ca. 1.0–1.2 × 2.0– 
<quantity id="470BAED6FFE2374927D0FB42FADEFB0C" box="[1231,1320,1202,1229]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="7" pageNumber="84" unit="mm" value="2.2">2.2 mm</quantity>
, embracing apex of column without covering it, transversely elliptic, margin slightly erose; mid-lobe 2.0–2.3 × 
<quantity id="470BAED6FFE237492605FB26FA51FB30" box="[1306,1447,1238,1265]" metricMagnitude="-3" metricUnit="m" metricValue="1.55" metricValueMax="1.6" metricValueMin="1.5" pageId="7" pageNumber="84" unit="mm" value="1.55" valueMax="1.6" valueMin="1.5">1.5–1.6 mm</quantity>
, slightly arched upwards in natural position, triangular, acuminate, margin entire. 
<emphasis id="B287DF21FFE237492719FB0AFBA9FAD5" box="[1030,1119,1274,1300]" italics="true" pageId="7" pageNumber="84">Column</emphasis>
<quantity id="470BAED6FFE23749277BFB0AFB4FFAD4" box="[1124,1209,1274,1301]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="7" pageNumber="84" unit="mm" value="2.3">2.3 mm</quantity>
long including lateral wings, dorsally 
<quantity id="470BAED6FFE23749225DFAEEFE6DFAF8" box="[322,411,1310,1337]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="7" pageNumber="84" unit="mm" value="1.3">1.3 mm</quantity>
to edge of clinandrium-hood, dorsally clinandrium hood somewhat upturned, thick, apex oblique, with pair of rounded wings that surpass clinandrium-hood. 
<emphasis id="B287DF21FFE237492068FAB2FBB1FA9D" box="[887,1095,1346,1372]" italics="true" pageId="7" pageNumber="84">Clinandrium-hood</emphasis>
short, slightly upturned, margin erose. 
<taxonomicName id="47F378B0FFE2374923CEFA96FEB7FA41" authorityName="Montfort" authorityYear="1810" box="[209,321,1382,1408]" class="Gastropoda" family="Rostellariidae" genus="Rostellum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="7" pageNumber="84" phylum="Mollusca" rank="genus">
<emphasis id="B287DF21FFE2374923CEFA96FEB7FA41" box="[209,321,1382,1408]" italics="true" pageId="7" pageNumber="84">Rostellum</emphasis>
</taxonomicName>
sub-apical, slit; viscarium semi-liquid. 
<emphasis id="B287DF21FFE2374921E0FA96FBF5FA41" box="[767,1027,1382,1408]" italics="true" pageId="7" pageNumber="84">Lateral lobes of stigma</emphasis>
, 
<emphasis id="B287DF21FFE23749270EFA96FB8AFA41" box="[1041,1148,1382,1408]" italics="true" pageId="7" pageNumber="84">cuniculus</emphasis>
not seen. 
<emphasis id="B287DF21FFE2374927F0FA96FACDFA41" box="[1263,1339,1382,1408]" italics="true" pageId="7" pageNumber="84">Anther</emphasis>
reniform, 4-celled. 
<taxonomicName id="47F378B0FFE2374923EFFA7AFEBCFA65" box="[240,330,1418,1444]" class="Liliopsida" family="Poaceae" genus="Pollinia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Poales" pageId="7" pageNumber="84" phylum="Tracheophyta" rank="genus">
<emphasis id="B287DF21FFE2374923EFFA7AFEBCFA65" box="[240,330,1418,1444]" italics="true" pageId="7" pageNumber="84">Pollinia</emphasis>
</taxonomicName>
4, ovoid, laterally compressed, caudicles soft and granulose, short. 
<emphasis id="B287DF21FFE23749275EFA7AFB6BFA65" box="[1089,1181,1418,1444]" italics="true" pageId="7" pageNumber="84">Capsule</emphasis>
not seen.
</paragraph>
</subSubSection>
<subSubSection id="C8E950B8FFE2374923A2FA5EFAECFA2D" pageId="7" pageNumber="84" type="distribution">
<paragraph id="804C0333FFE2374923A2FA5EFAECFA2D" blockId="7.[136,1452,446,2021]" pageId="7" pageNumber="84">
<emphasis id="B287DF21FFE2374923A2FA5EFE05FA09" bold="true" box="[189,499,1454,1480]" pageId="7" pageNumber="84">Distribution and ecology:</emphasis>
—Presently known only from the 
<typeStatus id="5F48BD91FFE23749209EFA5FFC47FA08" box="[897,945,1455,1481]" pageId="7" pageNumber="84">type</typeStatus>
in northern 
<collectingCountry id="F8E443A3FFE23749275BFA5FFB8AFA08" box="[1092,1148,1455,1481]" name="Peru" pageId="7" pageNumber="84">Peru</collectingCountry>
: 
<collectingRegion id="4237CDD1FFE237492796FA5EFAF2FA09" box="[1161,1284,1454,1480]" country="Peru" name="Cajamarca" pageId="7" pageNumber="84">Cajamarca</collectingRegion>
, Chota; grows lithophytic and epiphytic at 
<quantity id="470BAED6FFE2374922DDFA22FDE1FA2C" box="[450,535,1490,1517]" metricMagnitude="3" metricUnit="m" metricValue="2.778" pageId="7" pageNumber="84" unit="m" value="2778.0">2778 m</quantity>
elevation in well illuminated places. Flowering in July to November
</paragraph>
</subSubSection>
<subSubSection id="C8E950B8FFE2374923A2FA06FE63F999" pageId="7" pageNumber="84" type="etymology">
<paragraph id="804C0333FFE2374923A2FA06FE63F999" blockId="7.[136,1452,446,2021]" pageId="7" pageNumber="84">
<emphasis id="B287DF21FFE2374923A2FA06FEBEF9D1" bold="true" box="[189,328,1526,1552]" pageId="7" pageNumber="84">Etymology:</emphasis>
—From the Latin 
<emphasis id="B287DF21FFE237492106FA06FD9CF9D1" box="[537,618,1526,1552]" italics="true" pageId="7" pageNumber="84">bicorni</emphasis>
-, who horned, and 
<emphasis id="B287DF21FFE237492053FA06FC5EF9D1" box="[844,936,1526,1552]" italics="true" pageId="7" pageNumber="84">alpicola</emphasis>
, Alpine, found in mountains, the species has unusual calli which are triangular, upright and divergent, when all other species have the calli parallel to the lip and more or less finger-like.
</paragraph>
</subSubSection>
<subSubSection id="C8E950B8FFE2374923A2F992FBBAF9BC" box="[189,1100,1634,1661]" pageId="7" pageNumber="84" type="conservation">
<paragraph id="804C0333FFE2374923A2F992FBBAF9BC" blockId="7.[136,1452,446,2021]" box="[189,1100,1634,1661]" pageId="7" pageNumber="84">
<emphasis id="B287DF21FFE2374923A2F992FE42F9BD" bold="true" box="[189,436,1634,1660]" pageId="7" pageNumber="84">Conservation status:</emphasis>
—DD. Data deficient. Presently known only from the 
<typeStatus id="5F48BD91FFE23749270BF993FBBEF9BC" box="[1044,1096,1635,1661]" pageId="7" pageNumber="84">type</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C8E950B8FFE2374723A2F976FE64FEAC" lastPageId="9" lastPageNumber="86" pageId="7" pageNumber="84" type="discussion">
<paragraph id="804C0333FFE2374723A2F976FE03FF1D" blockId="7.[136,1452,446,2021]" lastBlockId="9.[136,1451,158,365]" lastPageId="9" lastPageNumber="86" pageId="7" pageNumber="84">
<emphasis id="B287DF21FFE2374923A2F976FE39F961" bold="true" box="[189,463,1670,1696]" pageId="7" pageNumber="84">Taxonomic Discussion:</emphasis>
— 
<taxonomicName id="47F378B0FFE2374922F4F976FCDFF961" authorityName="Hagsater, Chamaya &amp; Iberico. A. Plant" authorityYear="2023" box="[491,809,1670,1696]" class="Liliopsida" family="Orchidaceae" genus="Epidendrum" kingdom="Plantae" order="Asparagales" pageId="7" pageNumber="84" phylum="Tracheophyta" rank="species" species="bicornialpicola">
<emphasis id="B287DF21FFE2374922F4F976FCDFF961" box="[491,809,1670,1696]" italics="true" pageId="7" pageNumber="84">Epidendrum bicornialpicola</emphasis>
</taxonomicName>
belongs to the Alpicola Group, which is characterized by the simple stems, the long, narrow spathes, numerous flowers, the fleshy lips, the triangular mid-lobe and roundish lateral lobes, and the lip with two laminar calli and a rounded process in between. The new species is recognized by the peduncle of the inflorescence, straight, erect, about as long as apical leaves, the spathes of the peduncle tubular, closely appressed, the rachis sharply aching nutant, nearly straight but nutant, the yellow-green flowers, ochre lip and white column, the lateral sepals 8.4–8.6 × 
<quantity id="470BAED6FFE237492178F8CAFD07F894" box="[615,753,1850,1877]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" metricValueMax="2.9" metricValueMin="2.7" pageId="7" pageNumber="84" unit="mm" value="2.8" valueMax="2.9" valueMin="2.7">2.7–2.9 mm</quantity>
, the petals 4.5–5.0 × 
<quantity id="470BAED6FFE2374920FCF8CAFB9AF894" box="[995,1132,1850,1877]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="7" pageNumber="84" unit="mm" value="0.55" valueMax="0.6" valueMin="0.5">0.5–0.6 mm</quantity>
, linear oblanceolate-oblong, acuminate, and the lip 4.3–4.5 × 
<quantity id="470BAED6FFE23749211EF8AEFD7CF8B8" box="[513,650,1886,1913]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" metricValueMax="3.8" metricValueMin="3.6" pageId="7" pageNumber="84" unit="mm" value="3.7" valueMax="3.8" valueMin="3.6">3.6–3.8 mm</quantity>
when spread, bicallose, calli prominent, laminar, short, acute, upright, divergent; disc with a very short thick wide mid-rib, extending beyond calli to base of mid-lobe. 
<taxonomicName id="47F378B0FFE2374927D9F872FA5DF85D" authorityName="Hagsater &amp; Cisneros" authorityYear="2020" box="[1222,1451,1922,1948]" class="Liliopsida" family="Orchidaceae" genus="Epidendrum" kingdom="Plantae" order="Asparagales" pageId="7" pageNumber="84" phylum="Tracheophyta" rank="species" species="delsyae">
<emphasis id="B287DF21FFE2374927D9F872FA5DF85D" box="[1222,1451,1922,1948]" italics="true" pageId="7" pageNumber="84">Epidendrum delsyae</emphasis>
</taxonomicName>
is larger overall, the leaves 7.4–13.4 × 1.0– 
<quantity id="470BAED6FFE23749216BF856FD35F800" box="[628,707,1958,1985]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="7" pageNumber="84" unit="cm" value="1.2">1.2 cm</quantity>
, the peduncle of the inflorescence shorter than the apical leaf, the flowers relatively large, the sepals and petals ochre, apex greenish orange, lip ochre-orange, column green, the sepals 
<quantity id="470BAED6FFEC37472397FF6EFEEEFF78" box="[136,280,158,185]" metricMagnitude="-2" metricUnit="m" metricValue="1.14" metricValueMax="1.28" metricValueMin="1.0" pageId="9" pageNumber="86" unit="mm" value="11.4" valueMax="12.8" valueMin="10.0">10–12.8 mm</quantity>
long,, and the lip with a mid-lobe 3.8 × 
<quantity id="470BAED6FFEC374721E5FF6EFCA2FF78" box="[762,852,158,185]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="9" pageNumber="86" unit="mm" value="1.8">1.8 mm</quantity>
, sub-trilobed in general form, acute, attenuate at the middle, the apical half ensiform.
</paragraph>
<caption id="D48C53BBFFED37462397F850FB12F83E" ID-DOI="http://doi.org/10.5281/zenodo.10149614" ID-Zenodo-Dep="10149614" httpUri="https://zenodo.org/record/10149614/files/figure.png" pageId="8" pageNumber="85" startId="8.[136,229,1952,1974]" targetBox="[195,1393,190,1928]" targetPageId="8" targetType="figure">
<paragraph id="804C0333FFED37462397F850FB12F83E" blockId="8.[136,1453,1952,2047]" pageId="8" pageNumber="85">
<emphasis id="B287DF21FFED37462397F850FF0BF874" bold="true" box="[136,253,1952,1974]" pageId="8" pageNumber="85">FIGURE 4.</emphasis>
Lankester Composite Dissection Plate (LCDP) of 
<taxonomicName id="47F378B0FFED374621FCF851FA99F876" authority="Hagsater, Chamaya &amp; Iberico. A. Plant" authorityName="Hagsater, Chamaya &amp; Iberico. A. Plant" authorityYear="2023" box="[739,1391,1952,1975]" class="Liliopsida" family="Orchidaceae" genus="Epidendrum" kingdom="Plantae" order="Asparagales" pageId="8" pageNumber="85" phylum="Tracheophyta" rank="species" species="bicornialpicola">
<emphasis id="B287DF21FFED374621FCF851FC05F877" box="[739,1011,1953,1974]" italics="true" pageId="8" pageNumber="85">Epidendrum bicornialpicola</emphasis>
Hágsater, Chamaya &amp; Iberico. 
<emphasis id="B287DF21FFED3746263EF850FACFF874" bold="true" box="[1313,1337,1952,1973]" pageId="8" pageNumber="85">A.</emphasis>
Plant
</taxonomicName>
habit. 
<emphasis id="B287DF21FFED37462397F835FF68F81B" bold="true" box="[136,158,1989,2010]" pageId="8" pageNumber="85">B.</emphasis>
Inflorescence. 
<emphasis id="B287DF21FFED3746222CF834FEBCF81B" bold="true" box="[307,330,1988,2010]" pageId="8" pageNumber="85">C.</emphasis>
Flower, lateral view. 
<emphasis id="B287DF21FFED37462101F835FDC0F81B" bold="true" box="[542,566,1989,2010]" pageId="8" pageNumber="85">D.</emphasis>
Dissected perianth. 
<emphasis id="B287DF21FFED374621E3F835FCE4F81B" bold="true" box="[764,786,1989,2010]" pageId="8" pageNumber="85">E.</emphasis>
Ovary with lip and column; upper and lateral views. 
<emphasis id="B287DF21FFED37462604F835FADBF81B" bold="true" box="[1307,1325,1989,2010]" pageId="8" pageNumber="85">F.</emphasis>
Pollinarium. Prepared by Anaís Cisneros from the images by J. A. Chamaya taken from the specimen that served as the holotype.
</paragraph>
</caption>
<paragraph id="804C0333FFEC374723A2FF16FE64FEAC" blockId="9.[136,1451,158,365]" pageId="9" pageNumber="86">
The Ecuadorian 
<taxonomicName id="47F378B0FFEC3747226BFF16FC72FEC1" authority="Hagsater &amp; Dodson" authorityName="Hagsater &amp; Dodson" box="[372,900,230,256]" class="Liliopsida" family="Orchidaceae" genus="Epidendrum" kingdom="Plantae" order="Asparagales" pageId="9" pageNumber="86" phylum="Tracheophyta" rank="species" species="aenigmaticum">
<emphasis id="B287DF21FFEC3747226BFF16FD55FEC1" box="[372,675,230,256]" italics="true" pageId="9" pageNumber="86">Epidendrum aenigmaticum</emphasis>
Hágsater &amp; Dodson
</taxonomicName>
(2007: t. 902) is recognized by the leaves oblongelliptic, obtuse, 9.5–15 × 
<quantity id="470BAED6FFEC374722B6FEFAFDDCFEE4" box="[425,554,266,293]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" metricValueMax="2.1" metricValueMin="1.5" pageId="9" pageNumber="86" unit="cm" value="1.8" valueMax="2.1" valueMin="1.5">1.5–2.1 cm</quantity>
, flowers brown-orange, the dorsal sepal 5.4–6.0 mm long, and the lip 3.6 × 
<quantity id="470BAED6FFEC37472697FEFAFF40FE88" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="9" pageNumber="86" unit="mm" value="3.5">3.5 mm</quantity>
, the calli laminar, rounded, with a thick, elliptic, short mid-keel, the lateral lobes transversely ovate-elliptic, the mid-lobe 1.7 × 
<quantity id="470BAED6FFEC3747222AFEA2FE78FEAC" box="[309,398,338,365]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="9" pageNumber="86" unit="mm" value="1.2">1.2 mm</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>