treatments-xml/data/03/EB/BB/03EBBB3FFFC5F261EBE3FA14F6E2FBFA.xml
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<document id="515A67A642FF2A11CF5AAF0B19CCAB2B" ID-CLB-Dataset="20500" ID-DOI="10.5169/seals-787079" ID-GBIF-Dataset="3ba6188e-2e5e-4162-b055-905cfc2764ac" ID-ISSN="2624-9170" ID-Zenodo-Dep="6283928" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1637935189009" checkinUser="carolina" docAuthor="Balkenohl, Michael &amp; Aston, Paul" docDate="2015" docId="03EBBB3FFFC5F261EBE3FA14F6E2FBFA" docLanguage="en" docName="ContribNatHist.31.1-12.pdf" docOrigin="Contributions to Natural History 31" docStyle="DocumentStyle:D319E9290ED630111D2A11900DAFC11F.4:ContribNatHist.2011-.journal_article" docStyleId="D319E9290ED630111D2A11900DAFC11F" docStyleName="ContribNatHist.2011-.journal_article" docStyleVersion="4" docTitle="Asioreicheia guenardi Balkenohl &amp; Aston 2015, sp. nov." docType="treatment" docVersion="12" lastPageNumber="10" masterDocId="FFD2C347FFC7F268EB3EFFA2F777FFA3" masterDocTitle="Asioreicheia guenardi sp. nov. (Insecta: Coleoptera: Carabidae: Clivinini), a new peculiar species from Hong Kong" masterLastPageNumber="12" masterPageNumber="1" pageNumber="3" updateTime="1699470509822" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="7E89915761E29EBFA02A8562A6177DD1">Asioreicheia guenardi sp. nov. (Insecta: Coleoptera: Carabidae: Clivinini), a new peculiar species from Hong Kong</mods:title>
</mods:titleInfo>
<mods:name id="C52A802A1821047B116C12E41C2DE62C" type="personal">
<mods:role id="A49A533E0557E8BC4674F2B6E2A77D47">
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<mods:namePart id="1962D926F6821DF3353BB120C8574943">Balkenohl, Michael</mods:namePart>
</mods:name>
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<mods:role id="F36865FFB7A6B37C777F90BE357EDD3F">
<mods:roleTerm id="2BDB970397E9485832BA7A0658F8EA9B">Author</mods:roleTerm>
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<mods:namePart id="5C2ADC26A0354D716796DFD5A4017AFD">Aston, Paul</mods:namePart>
</mods:name>
<mods:typeOfResource id="FE95944FFD070A9CE9954429D2B1BAF5">text</mods:typeOfResource>
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<mods:title id="9F705FBEAE9435D6FF02103A906F3C00">Contributions to Natural History</mods:title>
</mods:titleInfo>
<mods:part id="43FD937E9170682AFA3ABF76E0E47809">
<mods:date id="489D821CB927EE97B508DF0880A97992">2015</mods:date>
<mods:detail id="5185089929F503ABF99799CAFCD01CF2" type="pubDate">
<mods:number id="706ADBBD330478CAADB3DCA5C9EF09AE">2015-12-11</mods:number>
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<mods:number id="D5B4858916938DD945B66F52CFD9AB72">31</mods:number>
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<mods:identifier id="F6C17838C084159C119D82B874E725AD" type="DOI">10.5169/seals-787079</mods:identifier>
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<treatment id="03EBBB3FFFC5F261EBE3FA14F6E2FBFA" ID-DOI="http://doi.org/10.5281/zenodo.6315562" ID-GBIF-Taxon="193585911" ID-Zenodo-Dep="6315562" LSID="urn:lsid:plazi:treatment:03EBBB3FFFC5F261EBE3FA14F6E2FBFA" httpUri="http://treatment.plazi.org/id/03EBBB3FFFC5F261EBE3FA14F6E2FBFA" lastPageId="9" lastPageNumber="10" pageId="2" pageNumber="3">
<subSubSection id="C35859A2FFC5F26AEBE3FA14F544FA6C" box="[221,563,1462,1487]" pageId="2" pageNumber="3" type="nomenclature">
<paragraph id="8BFD0A29FFC5F26AEBE3FA14F544FA6C" blockId="2.[221,563,1462,1487]" box="[221,563,1462,1487]" pageId="2" pageNumber="3">
<heading id="D0B5BD45FFC5F26AEBE3FA14F544FA6C" box="[221,563,1462,1487]" centered="true" fontSize="36" level="2" pageId="2" pageNumber="3" reason="4">
<emphasis id="B936D63BFFC5F26AEBE3FA14F544FA6C" bold="true" box="[221,563,1462,1487]" pageId="2" pageNumber="3">
<taxonomicName id="4C4271AAFFC5F26AEBE3FA14F6A2FA6C" ID-CoL="5VTWC" authority="Balkenohl &amp; Aston, 2015" authorityName="Balkenohl &amp; Aston" authorityYear="2015" box="[221,469,1462,1487]" class="Insecta" family="Carabidae" genus="Asioreicheia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="guenardi" status="sp. nov.">
<emphasis id="B936D63BFFC5F26AEBE3FA14F6A2FA6C" bold="true" box="[221,469,1462,1487]" italics="true" pageId="2" pageNumber="3">Asioreicheia guenardi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2056B40FFC5F26AEAE2FA14F544FA6C" box="[476,563,1462,1487]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35859A2FFC5F26BEBE3F9A3F540FE7D" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3" type="materials_examined">
<paragraph id="8BFD0A29FFC5F26BEBE3F9A3F66EFE87" blockId="2.[221,1106,1534,1636]" lastBlockId="3.[184,1069,156,478]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
Type material:
<materialsCitation id="3B2A0074FFC5F26BEABFF9A3F66EFE87" ID-GBIF-Occurrence="3496976303" collectingDate="2014-11-25" collectionCode="NMBE" collectorName="Dr. Benoit Guenard &amp; M. Wong." country="Hong Kong" determinerName="Paul Aston" elevation="156" lastPageId="3" lastPageNumber="4" location="Adjacent to Hong Kong University Campus" municipality="Determined Paul Aston" pageId="2" pageNumber="3" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="54F9B48BFFC5F26AEABFF9A3F690F9BA" box="[385,487,1537,1561]" pageId="2" pageNumber="3" type="holotype">Holotype</typeStatus>
<specimenCount id="9D44C1A0FFC5F26AEAD3FA5CF576F9BA" box="[493,513,1534,1561]" pageId="2" pageNumber="3" type="male"></specimenCount>
, with printed label &quot;Locality: Lung Fu shan C.P.,
<collectingCountry id="F3554AB9FFC5F26AEF2BF9A3F66FF99D" name="Hong Kong" pageId="2" pageNumber="3">Hong Kong</collectingCountry>
(
<location id="8E9D5CF2FFC5F26AEA17F984F445F99D" LSID="urn:lsid:plazi:treatment:03EBBB3FFFC5F261EBE3FA14F6E2FBFA:8E9D5CF2FFC5F26AEA17F984F445F99D" box="[297,818,1574,1598]" country="Hong Kong" municipality="Determined Paul Aston" name="Adjacent to Hong Kong University Campus" pageId="2" pageNumber="3">Adjacent to Hong Kong University Campus</location>
)&quot;; second printed label &quot;Leg.
<collectorName id="26B76FFFFFC5F26AEA1CF9EEF68AF9C7" box="[290,509,1612,1636]" pageId="2" pageNumber="3">Dr. Benoit Guénard</collectorName>
&amp;
<collectorName id="26B76FFFFFC5F26AE91FF9EEF5F8F9C7" box="[545,655,1612,1636]" pageId="2" pageNumber="3">M. Wong.</collectorName>
Hong Kong University&quot;; blue handwritten label &quot;12 of
<date id="FFFC2CE9FFC4F26BEA41FF3EF6AFFF17" box="[383,472,156,180]" pageId="3" pageNumber="4" value="2014-11-25">
<collectingDate id="EFB8D501FFC4F26BEA41FF3EF6AFFF17" box="[383,472,156,180]" pageId="3" pageNumber="4" value="2014-11-25">25.xi.14</collectingDate>
</date>
H.K.U. campus, alt
<quantity id="4CBAA7CCFFC4F26BE985FF3EF475FF17" box="[699,770,156,180]" metricMagnitude="2" metricUnit="m" metricValue="1.56" pageId="3" pageNumber="4" unit="m" value="156.0">
<elevation id="006FED1AFFC4F26BE985FF3EF475FF17" box="[699,770,156,180]" metricMagnitude="2" metricUnit="m" metricValue="1.56" pageId="3" pageNumber="4" unit="m" value="156.0">156 m</elevation>
</quantity>
HKG winkler&quot;; backside black printed &quot;coll: B. Guénard Hong Kong, China&quot;; second blue handwritten label &quot;Clivini [sic!] sp I&quot; backside black printed &quot;Determined
<determinerName id="7E21E6EDFFC4F26BE8A0FF44F369FF5D" box="[926,1054,230,254]" pageId="3" pageNumber="4">Paul Aston</determinerName>
&quot; (
<collectionCode id="ED5392ECFFC4F26BEB80FEAEF678FE87" box="[190,271,268,292]" collectionName="Switzerland, Bern, Naturhistorische Museums" country="Switzerland" httpUri="http://grbio.org/cool/5hwi-0wgz" name="Naturhistorisches Museum der Burgergemeinde Bern" pageId="3" pageNumber="4">NMBE</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BFD0A29FFC4F26BEB86FE93F540FE7D" blockId="3.[184,1069,156,478]" pageId="3" pageNumber="4">
<materialsCitation id="3B2A0074FFC4F26BEB86FE93F5A3FEEA" ID-GBIF-Occurrence="3496976301" box="[184,724,302,329]" collectionCode="CBB" collectorName=" Dr. Benoit Guenard &amp; M. Wong" country="Hong Kong" elevation="156" location="Clivini" municipality="Determined Paul Aston" pageId="3" pageNumber="4" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<typeStatus id="54F9B48BFFC4F26BEB86FE93F646FEEA" box="[184,305,305,329]" pageId="3" pageNumber="4" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D44C1A0FFC4F26BEA03FE93F61FFEEA" box="[317,360,302,329]" pageId="3" pageNumber="4" type="female">1 ♀</specimenCount>
: same data as holotype (
<collectionCode id="ED5392ECFFC4F26BE9ADFE93F5BAFEEA" box="[659,717,305,329]" collectionName="CBB" pageId="3" pageNumber="4">CBB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B2A0074FFC4F26BE9DEFE93F544FE7D" ID-GBIF-Occurrence="3496976302" collectingDate="2014-11-24" collectionCode="CAHK" collectorName="B. Guenard" country="Hong Kong" determinerName="Paul Aston" elevation="295" location="Hong Kong" municipality="Determined Paul Aston" pageId="3" pageNumber="4" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D44C1A0FFC4F26BE9DEFE93F47BFEEA" box="[736,780,302,329]" pageId="3" pageNumber="4" type="male">1 ♂</specimenCount>
, same data as holotype, but with blue handwritten label data &quot;2 of
<date id="FFFC2CE9FFC4F26BE9B1FEF4F590FECD" box="[655,743,342,366]" pageId="3" pageNumber="4" value="2014-11-24">
<collectingDate id="EFB8D501FFC4F26BE9B1FEF4F590FECD" box="[655,743,342,366]" pageId="3" pageNumber="4" value="2014-11-24">24.xi.14</collectingDate>
</date>
HKU campus, HKG alt
<quantity id="4CBAA7CCFFC4F26BE8E1FEF4F35BFECD" box="[991,1068,342,366]" metricMagnitude="2" metricUnit="m" metricValue="2.95" pageId="3" pageNumber="4" unit="m" value="295.0">
<elevation id="006FED1AFFC4F26BE8E1FEF4F35BFECD" box="[991,1068,342,366]" metricMagnitude="2" metricUnit="m" metricValue="2.95" pageId="3" pageNumber="4" unit="m" value="295.0">295 m</elevation>
</quantity>
winkler extractor&quot;; backside black printed: &quot;coll:
<collectorName id="26B76FFFFFC4F26BE824FEDEF4EBFE37" box="[794,924,380,404]" pageId="3" pageNumber="4">B. Guénard</collectorName>
<collectingCountry id="F3554AB9FFC4F26BE89BFEDEF35EFE37" box="[933,1065,380,404]" name="Hong Kong" pageId="3" pageNumber="4">Hong Kong</collectingCountry>
,
<collectingCountry id="F3554AB9FFC4F26BEB86FE03F781FE1A" box="[184,246,417,441]" name="China" pageId="3" pageNumber="4">China</collectingCountry>
&quot;; second blue handwritten label &quot;
<taxonomicName id="4C4271AAFFC4F26BE9BDFE03F5AAFE1A" box="[643,733,417,441]" class="Insecta" family="Carabidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="tribe" tribe="Clivinini">Clivinini</taxonomicName>
sp I.&quot; backside black printed &quot;Determined
<determinerName id="7E21E6EDFFC4F26BEA6EFE64F6BFFE7D" box="[336,456,454,478]" pageId="3" pageNumber="4">Paul Aston</determinerName>
&quot; (
<collectionCode id="ED5392ECFFC4F26BEADCFE64F559FE7D" box="[482,558,454,478]" collectionName="CAHK" pageId="3" pageNumber="4">CAHK</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C35859A2FFC4F26FEB86FD94F619FCDA" lastPageId="7" lastPageNumber="8" pageId="3" pageNumber="4" type="description">
<paragraph id="8BFD0A29FFC4F26BEB86FD94F5EFFDED" blockId="3.[184,664,566,591]" box="[184,664,566,591]" pageId="3" pageNumber="4">
<emphasis id="B936D63BFFC4F26BEB86FD94F64CFDEC" bold="true" box="[184,315,566,591]" pageId="3" pageNumber="4">Description</emphasis>
(
<figureCitation id="137916ACFFC4F26BEA76FD94F6C2FDED" box="[328,437,566,590]" captionStart-0="Figs" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="4.[221,260,160,178]" captionStartId-1="5.[856,885,160,178]" captionStartId-2="6.[221,249,1430,1448]" captionStartId-3="8.[221,250,160,178]" captionTargetBox-0="[439,1143,89,949]" captionTargetBox-1="[163,815,90,605]" captionTargetBox-2="[171,1172,39,1373]" captionTargetBox-3="[431,1136,87,1078]" captionTargetId-0="figure-215@4.[477,1108,150,559]" captionTargetId-1="figure-284@5.[180,573,128,528]" captionTargetId-2="figure-17@6.[221,1105,144,755]" captionTargetId-3="figure-216@8.[462,922,153,1049]" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="8" captionText-0="Figs. 1a, b: Asioreicheia guenardi sp. nov., holotype, male, a: dorsal view, b: lateral view." captionText-1="Fig. 2: Asioreicheia guenardi sp. nov., holotype, male, head with mouthparts, dorsal view." captionText-2="Fig. 3ac: Asioreicheia guenardi sp. nov., holotype, male. a: aedeagus with median lobe and parameres, ventral view; b: apical part of median lobe, ventral view; c: genital ring." captionText-3="Fig. 4: Asioreicheia guenardi sp. nov., paratype female, left coxostylus, dorsal view." figureDoi-0="http://doi.org/10.5281/zenodo.6283932" figureDoi-1="http://doi.org/10.5281/zenodo.6283936" figureDoi-2="http://doi.org/10.5281/zenodo.6283948" figureDoi-3="http://doi.org/10.5281/zenodo.6283954" httpUri-0="https://zenodo.org/record/6283932/files/figure.png" httpUri-1="https://zenodo.org/record/6283936/files/figure.png" httpUri-2="https://zenodo.org/record/6283948/files/figure.png" httpUri-3="https://zenodo.org/record/6283954/files/figure.png" pageId="3" pageNumber="4">Figs. 14</figureCitation>
; habitus
<figureCitation id="137916ACFFC4F26BE925FD94F5F8FDED" box="[539,655,566,590]" captionStart="Figs" captionStartId="4.[221,260,160,178]" captionTargetBox="[439,1143,89,949]" captionTargetId="figure-215@4.[477,1108,150,559]" captionText="Figs. 1a, b: Asioreicheia guenardi sp. nov., holotype, male, a: dorsal view, b: lateral view." figureDoi="http://doi.org/10.5281/zenodo.6283932" httpUri="https://zenodo.org/record/6283932/files/figure.png" pageId="3" pageNumber="4">Figs. 1a, b</figureCitation>
)
</paragraph>
<paragraph id="8BFD0A29FFC4F26BEB86FD23F41CFD47" blockId="3.[184,1068,641,1636]" pageId="3" pageNumber="4">
Measurements (n = 3): Body length
<quantity id="4CBAA7CCFFC4F26BE94BFD23F46BFD3A" box="[629,796,641,665]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" metricValueMax="2.15" metricValueMin="1.95" pageId="3" pageNumber="4" unit="mm" value="2.05" valueMax="2.15" valueMin="1.95">1.952.15 mm</quantity>
(X =
<quantity id="4CBAA7CCFFC4F26BE858FD23F4A1FD3A" box="[870,982,641,665]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" pageId="3" pageNumber="4" unit="mm" value="2.05">2.05 mm</quantity>
), width
<quantity id="4CBAA7CCFFC4F26BEB86FD04F616FD1D" box="[184,353,678,702]" metricMagnitude="-4" metricUnit="m" metricValue="7.35" metricValueMax="7.7" metricValueMin="7.0" pageId="3" pageNumber="4" unit="mm" value="0.735" valueMax="0.77" valueMin="0.7">0.700.77 mm</quantity>
(X =
<quantity id="4CBAA7CCFFC4F26BEA99FD04F563FD1D" box="[423,532,678,702]" metricMagnitude="-4" metricUnit="m" metricValue="7.199999999999999" pageId="3" pageNumber="4" unit="mm" value="0.72">0.72 mm</quantity>
), ratio length/width of pronotum 0.980.1.04 (X = 1.01), ratio length/width of elytra 1.531.55 (X = 1.55).
</paragraph>
<paragraph id="8BFD0A29FFC4F26BEBE3FD53F56CFC67" blockId="3.[184,1068,641,1636]" pageId="3" pageNumber="4">Colour: Head, pronotum, dorsal and ventral surface middle to dark reddish brown. Vault of supraantennal plates, lateral teeth of clypeus light brown. Mandibles middle brown with carinae and apices darkened; mandibular and maxillar palpi yellowish-beige, transparent at apical tip; femora and tibiae yellowish red brown; tarsomeres, scapus and pedicellus light brown; antennomeres 311 yellowish-beige.</paragraph>
<paragraph id="8BFD0A29FFC4F26CEBE3FC73F40FFB07" blockId="3.[184,1068,641,1636]" lastBlockId="4.[221,1106,1014,1636]" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
Head (
<figureCitation id="137916ACFFC4F26BEA16FC73F61DFC4A" box="[296,362,977,1001]" captionStart="Fig" captionStartId="5.[856,885,160,178]" captionTargetBox="[163,815,90,605]" captionTargetId="figure-284@5.[180,573,128,528]" captionTargetPageId="5" captionText="Fig. 2: Asioreicheia guenardi sp. nov., holotype, male, head with mouthparts, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.6283936" httpUri="https://zenodo.org/record/6283936/files/figure.png" pageId="3" pageNumber="4">Fig. 2</figureCitation>
): A third narrower than pronotum. Clypeus and lateral teeth finely margined. Clypeus straight at anterior margin, lateral teeth moderately projecting, obtuse at tip, divided from supraantennal plates by indistinct obtuse-angular notches; clypeal field pentagonal, prolonged to frons as triangular elevation with sharp keel; frons moderately convex, with transverse rugae laterally to keel; clypeal field and frons with distinctly meshed reticulation. Supraantennal plates convex, smooth, laterally and posteriorly with distinct carina. Frontal furrows deep, diverging anteriorly and posteriorly of keel level. Anterior supraorbital setae situated at the end of supraorbital vault, the posterior ones at level of the end of genae; bilateral fovea bearing the clypeofrontal setae situated laterally and anteriorly to base of keel. Eyes reduced to a single convex to slightly globally protruding ommatidium, pointing anteriorly, surrounded by dark pigment; genae distinctly enlarged, higher than eyes, smooth. Labrum slightly trilobed anteriorly, with irregular reticulation, 5-setose. Mandible distinctly curved, acute at apex. Apical segment of maxillary palpomere slender, securiform, pointed at apex; apical segment of labial palpomere bottle-like, somewhat broadened, penultimate segment bisetose. Mentum bordered at base, lobes broadly rounded-off, apex of lobes with transverse rugae, median tooth not extending over lateral lobes; ligula with moderately long seta; antenna moderately long, extending up to posterior setigerous puncture of pronotum, scapus about as long as pedicellus, with one dorsoapical seta, third segment 1.5 times longer than wide, segments 410 moniliform, fully pubescent from segment 4 onwards.
</paragraph>
<caption id="DF3D5AA1FFC3F26CEBE3FF02F617FEBE" ID-DOI="http://doi.org/10.5281/zenodo.6283932" ID-Zenodo-Dep="6283932" httpUri="https://zenodo.org/record/6283932/files/figure.png" pageId="4" pageNumber="5" startId="4.[221,260,160,178]" targetBox="[439,1143,89,949]" targetPageId="4">
<paragraph id="8BFD0A29FFC3F26CEBE3FF02F617FEBE" blockId="4.[221,430,160,285]" pageId="4" pageNumber="5">
<emphasis id="B936D63BFFC3F26CEBE3FF02F617FEBE" bold="true" pageId="4" pageNumber="5">
Figs. 1a, b:
<emphasis id="B936D63BFFC3F26CEA7CFF02F65DFF6E" bold="true" italics="true" pageId="4" pageNumber="5">
<taxonomicName id="4C4271AAFFC3F26CEA7CFF02F6D9FF11" authority="Balkenohl &amp; Aston, 2015" authorityName="Balkenohl &amp; Aston" authorityYear="2015" box="[322,430,160,178]" pageId="4" pageNumber="5" status="sp. nov.">Asioreicheia</taxonomicName>
guenardi
</emphasis>
<taxonomicNameLabel id="A2056B40FFC3F26CEA11FF19F602FF6E" box="[303,373,187,205]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
,
<typeStatus id="54F9B48BFFC3F26CEBE3FF77F659FF44" box="[221,302,213,231]" pageId="4" pageNumber="5" type="holotype">holotype</typeStatus>
, male, a: dorsal view, b: lateral view.
</emphasis>
</paragraph>
</caption>
<paragraph id="8BFD0A29FFC3F26CEA3DFB13F461F9C7" blockId="4.[221,1106,1014,1636]" pageId="4" pageNumber="5">Pronotum: Outline subrectangular, in lateral view moderately convex, moderately to slightly convex to base. As wide as long, maximum width at middle. Lateral border slightly convex in middle part, convexity more distinct from posterior half to base. Reflexed lateral margin distinct, reaching from rounded anterior angles up to base, not joining anterior transverse line. Lateral channel narrow but distinct. Proepisternum tumid posterolaterally, well visible from above. Median line small, sharp, distinct, deeper at base, joining finely basal constriction (seen best at 140 x magnification), not joining anterior transverse line; anterior transverse line intimated, indistinct, visible in some specimens by its pigmentation; surface shiny, with irregular reticulation (seen best at 160 x magnification). Ring-like flange moderately convex (lateral view), constriction formed by row of rough foveae.</paragraph>
<caption id="DF3D5AA1FFC2F26DE866FF02F4B5FEBE" ID-DOI="http://doi.org/10.5281/zenodo.6283936" ID-Zenodo-Dep="6283936" httpUri="https://zenodo.org/record/6283936/files/figure.png" pageId="5" pageNumber="6" startId="5.[856,885,160,178]" targetBox="[163,815,90,605]" targetPageId="5">
<paragraph id="8BFD0A29FFC2F26DE866FF02F4B5FEBE" blockId="5.[856,1056,160,285]" pageId="5" pageNumber="6">
Fig. 2:
<taxonomicName id="4C4271AAFFC2F26DE8ADFF02F4D2FF6E" authority="Balkenohl &amp; Aston, 2015" authorityName="Balkenohl &amp; Aston" authorityYear="2015" class="Insecta" family="Carabidae" genus="Asioreicheia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="guenardi" status="sp. nov.">
<emphasis id="B936D63BFFC2F26DE8ADFF02F4D2FF6E" bold="true" italics="true" pageId="5" pageNumber="6">Asioreicheia guenardi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2056B40FFC2F26DE894FF19F487FF6E" box="[938,1008,187,205]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, male,
<emphasis id="B936D63BFFC2F26DE866FF52F4B5FEBE" bold="true" pageId="5" pageNumber="6">head with mouthparts, dorsal view.</emphasis>
</paragraph>
</caption>
<paragraph id="8BFD0A29FFC2F26DEBE3FD04F406FB6A" blockId="5.[184,1069,678,1636]" pageId="5" pageNumber="6">Elytra: Moderately convex on disc, distinctly convex in basal third (lateral view). Outline regularly long-oval, maximum width at middle, margined from pedunculus to apex; margin serrate up to apical third, teeth of serration sharp basally, becoming indistinct apically; base nearly straight, with fine and smooth reflexed margin; humeral angle visible, obtuse; lateral channel broad from level of humerus to apex, narrow at middle, with fold-like carina near apex crossing lateral channel, with broad setigerous tubercles; setae robust, long. Suture at base somewhat impressed. Basal granula absent; basal setigerous puncture with distinct tubercle, situated in projected extension of first interval. No scutellar stria. Striae 13 consisting of partly connected punctures, 15 reaching base but not up to basal declivity, all becoming less impressed towards apex and laterally. First interval slightly to moderately convex, others flattened. Interval 3 with 1012 and interval 5 with 78 fine setigerous punctures, located with nearly equal distance to each other but with irregular distance to striae, setae pili-like, short, fine, upright.</paragraph>
<paragraph id="8BFD0A29FFC2F26DEBE3FB74F513FB4D" blockId="5.[184,1069,678,1636]" box="[221,612,1238,1262]" pageId="5" pageNumber="6">Hind wings: Completely atrophied.</paragraph>
<paragraph id="8BFD0A29FFC2F26DEBE3FB5EF667F9BA" blockId="5.[184,1069,678,1636]" pageId="5" pageNumber="6">Ventral surface: Submarginal furrow of proepisternum visible from level of anterior setigerous puncture to middle of pronotum where it is turning to the disc due to convexity of the proepisternum, surface with some fine transverse wrinkles. Furrow between prosternum and proepisternum narrow, distinct. Abdominal sternites nearly smooth; terminal segment with an indistinct and irregular reticulation in male (seen best at 160 x magnification), with irregular transverse reticulation in female; two apical setigerous punctures distant.</paragraph>
<paragraph id="8BFD0A29FFC2F26FEBE3F984F485FF7A" blockId="5.[184,1069,678,1636]" lastBlockId="7.[184,1070,156,1636]" lastPageId="7" lastPageNumber="8" pageId="5" pageNumber="6">Legs: Protibia: Surface with distinct longitudinal reticulation; lateral upper spine ventrally curved. Movable spur smaller than spine, nearly straight, ventrally curved at apex. Praeapical lateral denticle robust, sharp, second one smaller. Hind legs: Tarsomere 1 longer than tarsomeres 2+3 (ratio 12:10).</paragraph>
<caption id="DF3D5AA1FFC1F26EEBE3FA34F432FA61" ID-DOI="http://doi.org/10.5281/zenodo.6283948" ID-Zenodo-Dep="6283948" httpUri="https://zenodo.org/record/6283948/files/figure.png" pageId="6" pageNumber="7" startId="6.[221,249,1430,1448]" targetBox="[171,1172,39,1373]" targetPageId="6">
<paragraph id="8BFD0A29FFC1F26EEBE3FA34F432FA61" blockId="6.[221,1107,1430,1474]" pageId="6" pageNumber="7">
Fig. 3ac:
<taxonomicName id="4C4271AAFFC1F26EEA0AFA34F699FA0B" authority="Balkenohl &amp; Aston, 2015" authorityName="Balkenohl &amp; Aston" authorityYear="2015" box="[308,494,1430,1448]" class="Insecta" family="Carabidae" genus="Asioreicheia" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="guenardi" status="sp. nov.">
<emphasis id="B936D63BFFC1F26EEA0AFA34F699FA0B" bold="true" box="[308,494,1430,1448]" italics="true" pageId="6" pageNumber="7">Asioreicheia guenardi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2056B40FFC1F26EEACCFA34F540FA0B" box="[498,567,1430,1448]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, male. a: aedeagus with median lobe and parameres, ventral view; b: apical part of median lobe, ventral view; c: genital ring.
</paragraph>
</caption>
<paragraph id="8BFD0A29FFC0F26FEBE3FF44F6F7FD8A" blockId="7.[184,1070,156,1636]" pageId="7" pageNumber="8">
Male genitalia (
<figureCitation id="137916ACFFC0F26FEAB0FF44F57CFF5D" box="[398,523,230,254]" captionStart="Fig" captionStartId="6.[221,249,1430,1448]" captionTargetBox="[171,1172,39,1373]" captionTargetId="figure-17@6.[221,1105,144,755]" captionTargetPageId="6" captionText="Fig. 3ac: Asioreicheia guenardi sp. nov., holotype, male. a: aedeagus with median lobe and parameres, ventral view; b: apical part of median lobe, ventral view; c: genital ring." figureDoi="http://doi.org/10.5281/zenodo.6283948" httpUri="https://zenodo.org/record/6283948/files/figure.png" pageId="7" pageNumber="8">Figs. 3ac</figureCitation>
): Median lobe moderately sclerotized, shape in ventral view about rectangularly arcuate in middle part, slightly flattened in apical half, somewhat distorted, apex formed by asymmetric rounded small spatula. Oroficium half as long as lobe, closing lips less sclerotized. Endophallus with numerous wrinkles, no spines or bristles visible (seen best at 500 x magnification). Parameres asymmetrically, ventral one much more slender and one third shorter than dorsal one, dorsal one twisted, each with two long setae at apex arising closely together. Genital ring wide, closed, oval-shaped, slightly distorted.
</paragraph>
<paragraph id="8BFD0A29FFC0F26FEBE3FD94F4A7FD47" blockId="7.[184,1070,156,1636]" pageId="7" pageNumber="8">
Female genitalia: Coxostylus slender (
<figureCitation id="137916ACFFC0F26FE9A2FD94F597FDED" box="[668,736,566,590]" captionStart="Fig" captionStartId="8.[221,250,160,178]" captionTargetBox="[431,1136,87,1078]" captionTargetId="figure-216@8.[462,922,153,1049]" captionTargetPageId="8" captionText="Fig. 4: Asioreicheia guenardi sp. nov., paratype female, left coxostylus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.6283954" httpUri="https://zenodo.org/record/6283954/files/figure.png" pageId="7" pageNumber="8">Fig. 4</figureCitation>
), dorsoventrally moderately broadened (oval-funnel like), distinctly curved apically, slightly carinate nearly up to apex; with two broader moderately long ensiform setae in middle part arising from knob-like tubercles, and one conspicuously long lateral ensiform seta at base; with two long and two medium sized nematiforme setae.
</paragraph>
<paragraph id="8BFD0A29FFC0F26FEBE3FD53F619FCDA" blockId="7.[184,1070,156,1636]" pageId="7" pageNumber="8">
Variation: In the
<specimenCount id="9D44C1A0FFC0F26FEAA5FD53F521FCAA" box="[411,598,753,777]" pageId="7" pageNumber="8" type="generic">three specimens</specimenCount>
examined, the anterior transverse line of the pronotum is more or less distinctly visible as a clear line. The punctures of the stria on the elytra show high variation in strenght of impression and also vary in position.
</paragraph>
</subSubSection>
<subSubSection id="C35859A2FFC0F26FEBE3FC24F6A5FC4A" pageId="7" pageNumber="8" type="diagnosis">
<paragraph id="8BFD0A29FFC0F26FEBE3FC24F6A5FC4A" blockId="7.[184,1070,156,1636]" pageId="7" pageNumber="8">
Differential diagnosis:
<taxonomicName id="4C4271AAFFC0F26FEAE0FC24F5A3FC3D" authority="Balkenohl &amp; Aston, 2015" authorityName="Balkenohl &amp; Aston" authorityYear="2015" box="[478,724,902,926]" class="Insecta" family="Carabidae" genus="Asioreicheia" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="guenardi" status="sp. nov.">
<emphasis id="B936D63BFFC0F26FEAE0FC24F5A3FC3D" box="[478,724,902,926]" italics="true" pageId="7" pageNumber="8">Asioreicheia guenardi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2056B40FFC0F26FE9E2FC24F442FC3D" box="[732,821,902,926]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
differs from the most related species
<taxonomicName id="4C4271AAFFC0F26FEA55FC0EF41BFC67" authority="(BULIRSCH, MAGRINI &amp; JIA, 2013)" baseAuthorityName="BULIRSCH, MAGRINI &amp; JIA" baseAuthorityYear="2013" box="[363,876,940,964]" class="Insecta" family="Carabidae" genus="Asioreicheia" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="chinensis">
<emphasis id="B936D63BFFC0F26FEA55FC0EF573FC67" box="[363,516,940,964]" italics="true" pageId="7" pageNumber="8">A. chinensis (</emphasis>
<bibRefCitation id="EFD377D8FFC0F26FE93BFC0EF411FC67" author="Bulirsch, P. &amp; Magrini, P. &amp; Jia, F." box="[517,870,940,964]" pageId="7" pageNumber="8" pagination="59 - 64" refId="ref3804" refString="Bulirsch, P., Magrini, P. &amp; Jia, F. (2013): Antireicheia chinensis sp. nov. of the subtribe Reicheiina (Coleoptera: Carabidae: Scaritiae) from the south-eastern China. - Acta Entomologica Musei Nationalis Prague 53: 59 - 64." type="journal article" year="2013">BULIRSCH, MAGRINI &amp; JIA, 2013</bibRefCitation>
)
</taxonomicName>
by the character states provided in
<tableCitation id="C6C03F92FFC0F26FEAB6FC73F6BCFC4A" box="[392,459,977,1001]" captionStart="Tab" captionStartId="9.[184,218,843,861]" captionTargetPageId="9" captionText="Tab. 1: Dividing character states of Asioreicheia chinensis (BULIRSCH, MAGRINI &amp; JIA, 2013) and A. guenardi sp. nov." httpUri="http://table.plazi.org/id/DF3D5AA1FFCEF261EB86FCE9F63AFCDB" pageId="7" pageNumber="8" tableUuid="DF3D5AA1FFCEF261EB86FCE9F63AFCDB">Tab. 1</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C35859A2FFC0F26FEBE3FC54F37FFA9A" pageId="7" pageNumber="8" type="discussion">
<paragraph id="8BFD0A29FFC0F26FEBE3FC54F37FFA9A" blockId="7.[184,1070,156,1636]" pageId="7" pageNumber="8">
Other
<taxonomicName id="4C4271AAFFC0F26FEA1BFC54F57CFBAD" box="[293,523,1014,1038]" class="Insecta" family="Carabidae" genus="Asioreicheia" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B936D63BFFC0F26FEA1BFC54F6D9FBAD" box="[293,430,1014,1038]" italics="true" pageId="7" pageNumber="8">Asioreicheia</emphasis>
species
</taxonomicName>
differ from
<taxonomicName id="4C4271AAFFC0F26FE9ABFC54F46DFBAD" authorityName="Balkenohl &amp; Aston" authorityYear="2015" box="[661,794,1014,1038]" class="Insecta" family="Carabidae" genus="Asioreicheia" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="guenardi">
<emphasis id="B936D63BFFC0F26FE9ABFC54F46DFBAD" box="[661,794,1014,1038]" italics="true" pageId="7" pageNumber="8">A. guenardi</emphasis>
</taxonomicName>
as follows:
<taxonomicName id="4C4271AAFFC0F26FE899FC54F6DAFB97" authority="(BALKENOHL, 2005)" baseAuthorityName="BALKENOHL" baseAuthorityYear="2005" class="Insecta" family="Carabidae" genus="Asioreicheia" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="margolata">
<emphasis id="B936D63BFFC0F26FE899FC54F7B9FB97" italics="true" pageId="7" pageNumber="8">A. margolata</emphasis>
(
<bibRefCitation id="EFD377D8FFC0F26FEBE1FBBEF6DFFB97" author="Balkenohl, M." box="[223,424,1052,1076]" pageId="7" pageNumber="8" pagination="1 - 10" refId="ref3699" refString="Balkenohl, M. (2005): First records of Reicheiina species from the Oriental Region and Japan (Coleoptera, Carabidae, Scarititae, Clivinini). - Coleoptera 9: 1 - 10." type="journal article" year="2005">BALKENOHL, 2005</bibRefCitation>
)
</taxonomicName>
does not have setigerous punctures on interval 5 of the elytra, and has an ovate shaped elytra. In
<taxonomicName id="4C4271AAFFC0F26FE9A5FBE3F787FBDD" authority="BULIRSCH &amp; MAGRINI, 2014" authorityName="BULIRSCH &amp; MAGRINI" authorityYear="2014" class="Insecta" family="Carabidae" genus="Asioreicheia" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="vietnamica">
<emphasis id="B936D63BFFC0F26FE9A5FBE3F443FBFA" box="[667,820,1089,1113]" italics="true" pageId="7" pageNumber="8">A. vietnamica</emphasis>
<bibRefCitation id="EFD377D8FFC0F26FE802FBE3F787FBDD" author="Bulirsch, P. &amp; Magrini, P." pageId="7" pageNumber="8" pagination="41 - 52" refId="ref3736" refString="Bulirsch, P. &amp; Magrini, P. (2014): A new genus Asioreicheia gen. nov. and two new species of the subtribes Reicheiina (Coleoptera: Carabidae: Scaritinae) from the eastern Asia and re-assessment of next two species from the same region. - Studies and reports of District Museum Prague-East, Taxonomical Series 10: 41 - 52." type="book chapter" year="2014">BULIRSCH &amp; MAGRINI, 2014</bibRefCitation>
</taxonomicName>
and in
<taxonomicName id="4C4271AAFFC0F26FEA79FBC4F46AFBDD" authority="BULIRSCH &amp; MAGRINI, 2014" authorityName="BULIRSCH &amp; MAGRINI" authorityYear="2014" box="[327,797,1126,1150]" class="Insecta" family="Carabidae" genus="Asioreicheia" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="fedorenkoi">
<emphasis id="B936D63BFFC0F26FEA79FBC4F697FBDD" box="[327,480,1126,1150]" italics="true" pageId="7" pageNumber="8">A. fedorenkoi</emphasis>
<bibRefCitation id="EFD377D8FFC0F26FEAD6FBC4F46AFBDD" author="Bulirsch, P. &amp; Magrini, P." box="[488,797,1126,1150]" pageId="7" pageNumber="8" pagination="41 - 52" refId="ref3736" refString="Bulirsch, P. &amp; Magrini, P. (2014): A new genus Asioreicheia gen. nov. and two new species of the subtribes Reicheiina (Coleoptera: Carabidae: Scaritinae) from the eastern Asia and re-assessment of next two species from the same region. - Studies and reports of District Museum Prague-East, Taxonomical Series 10: 41 - 52." type="book chapter" year="2014">BULIRSCH &amp; MAGRINI, 2014</bibRefCitation>
</taxonomicName>
, the pronotum is longer than wide.
<taxonomicName id="4C4271AAFFC0F26FEA0FFB2EF595FB07" authority="BALKENOHL, 2005" authorityName="BALKENOHL" authorityYear="2005" box="[305,738,1164,1188]" class="Insecta" family="Carabidae" genus="Laoreicheia" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="bulirschi">
<emphasis id="B936D63BFFC0F26FEA0FFB2EF56FFB07" box="[305,536,1164,1188]" italics="true" pageId="7" pageNumber="8">Laoreicheia bulirschi</emphasis>
<bibRefCitation id="EFD377D8FFC0F26FE920FB2EF595FB07" author="Balkenohl, M." box="[542,738,1164,1188]" pageId="7" pageNumber="8" pagination="1 - 10" refId="ref3699" refString="Balkenohl, M. (2005): First records of Reicheiina species from the Oriental Region and Japan (Coleoptera, Carabidae, Scarititae, Clivinini). - Coleoptera 9: 1 - 10." type="journal article" year="2005">BALKENOHL, 2005</bibRefCitation>
</taxonomicName>
exhibits two pairs of paramedian setae on the pronotum, and the elytra are completely serrate. In
<taxonomicName id="4C4271AAFFC0F26FE8F0FB13F6A1FB4D" authority="BALKENOHL, 2005" authorityName="BALKENOHL" authorityYear="2005" class="Insecta" family="Carabidae" genus="Reicheia" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="moritai">
<emphasis id="B936D63BFFC0F26FE8F0FB13F67FFB4D" italics="true" pageId="7" pageNumber="8">Reicheia moritai</emphasis>
<bibRefCitation id="EFD377D8FFC0F26FEA31FB74F6A1FB4D" author="Balkenohl, M." box="[271,470,1238,1262]" pageId="7" pageNumber="8" pagination="1 - 10" refId="ref3699" refString="Balkenohl, M. (2005): First records of Reicheiina species from the Oriental Region and Japan (Coleoptera, Carabidae, Scarititae, Clivinini). - Coleoptera 9: 1 - 10." type="journal article" year="2005">BALKENOHL, 2005</bibRefCitation>
</taxonomicName>
the proepisternum is not visible dorsally, the pronotum is distinctly cup-shaped, and the lateral channel of the elytra is smaller. For further differences compare
<bibRefCitation id="EFD377D8FFC0F26FEACCFA83F5C7FA9A" author="Balkenohl, M." box="[498,688,1313,1337]" pageId="7" pageNumber="8" pagination="1 - 10" refId="ref3699" refString="Balkenohl, M. (2005): First records of Reicheiina species from the Oriental Region and Japan (Coleoptera, Carabidae, Scarititae, Clivinini). - Coleoptera 9: 1 - 10." type="journal article" year="2005">Balkenohl (2005)</bibRefCitation>
and
<bibRefCitation id="EFD377D8FFC0F26FE9DBFA83F377FA9A" author="Bulirsch, P. &amp; Magrini, P." box="[741,1024,1313,1337]" pageId="7" pageNumber="8" pagination="41 - 52" refId="ref3736" refString="Bulirsch, P. &amp; Magrini, P. (2014): A new genus Asioreicheia gen. nov. and two new species of the subtribes Reicheiina (Coleoptera: Carabidae: Scaritinae) from the eastern Asia and re-assessment of next two species from the same region. - Studies and reports of District Museum Prague-East, Taxonomical Series 10: 41 - 52." type="book chapter" year="2014">Bulirsch &amp; Magrini (2014)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C35859A2FFC0F26FEBE3FAE4F6A7FA27" pageId="7" pageNumber="8" type="etymology">
<paragraph id="8BFD0A29FFC0F26FEBE3FAE4F6A7FA27" blockId="7.[184,1070,156,1636]" pageId="7" pageNumber="8">
Etymology: The name is a patronym of Dr. Benoit Guénard,
<collectingCountry id="F3554AB9FFC0F26FE84BFAE4F485FAFD" box="[885,1010,1350,1374]" name="Hong Kong" pageId="7" pageNumber="8">Hong Kong</collectingCountry>
, who collected the specimens.
</paragraph>
</subSubSection>
<subSubSection id="C35859A2FFC0F26FEBE3FA33F5A6F9C7" pageId="7" pageNumber="8" type="distribution">
<paragraph id="8BFD0A29FFC0F26FEBE3FA33F42CFA0A" blockId="7.[184,1070,156,1636]" box="[221,859,1425,1449]" pageId="7" pageNumber="8">
Distribution: Known from the
<typeStatus id="54F9B48BFFC0F26FE912FA33F528FA0A" box="[556,607,1425,1449]" pageId="7" pageNumber="8">type</typeStatus>
locality in
<collectingCountry id="F3554AB9FFC0F26FE9E6FA33F420FA0A" box="[728,855,1425,1449]" name="Hong Kong" pageId="7" pageNumber="8">Hong Kong</collectingCountry>
.
</paragraph>
<paragraph id="8BFD0A29FFC0F26FEBE3FA14F5A6F9C7" blockId="7.[184,1070,156,1636]" pageId="7" pageNumber="8">
Zoogeographic considerations: Between the records of
<taxonomicName id="4C4271AAFFC0F26FE86DFA14F4AAFA6D" baseAuthorityName="BULIRSCH, MAGRINI &amp; JIA" baseAuthorityYear="2013" box="[851,989,1462,1486]" class="Insecta" family="Carabidae" genus="Asioreicheia" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="chinensis">
<emphasis id="B936D63BFFC0F26FE86DFA14F4AAFA6D" box="[851,989,1462,1486]" italics="true" pageId="7" pageNumber="8">A. chinensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4271AAFFC0F26FEF2BFA14F65EFA57" authority="Balkenohl &amp; Aston, 2015" authorityName="Balkenohl &amp; Aston" authorityYear="2015" class="Insecta" family="Carabidae" genus="Asioreicheia" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="guernardi" status="sp. nov.">
<emphasis id="B936D63BFFC0F26FEF2BFA14F65EFA57" italics="true" pageId="7" pageNumber="8">A. guernardi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2056B40FFC0F26FEA0FFA7EF6FAFA57" box="[305,397,1500,1524]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
there are three geographic barriers formed by: (1) the Xi and Bei river sections, (2) the Pearl delta rivers, (3) the sea. The specimens were found on
<collectingCountry id="F3554AB9FFC0F26FEA5DF984F547F99D" box="[355,560,1574,1598]" name="Hong Kong Island" pageId="7" pageNumber="8">Hong Kong Island</collectingCountry>
, historically separated from the mainland by the sea forming a very deep
<quantity id="4CBAA7CCFFC0F26FEAC9F9EEF559F9C7" box="[503,558,1612,1636]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="8" unit="km" value="1.0">1 km</quantity>
wide channel.
</paragraph>
</subSubSection>
<caption id="DF3D5AA1FFCFF260EBE3FF02F63FFEBF" ID-DOI="http://doi.org/10.5281/zenodo.6283954" ID-Zenodo-Dep="6283954" httpUri="https://zenodo.org/record/6283954/files/figure.png" pageId="8" pageNumber="9" startId="8.[221,250,160,178]" targetBox="[431,1136,87,1078]" targetPageId="8">
<paragraph id="8BFD0A29FFCFF260EBE3FF02F63FFEBF" blockId="8.[221,388,160,284]" pageId="8" pageNumber="9">
Fig. 4:
<taxonomicName id="4C4271AAFFCFF260EA26FF02F6F3FF11" box="[280,388,160,178]" class="Insecta" family="Carabidae" genus="Asioreicheia" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">Asioreicheia</taxonomicName>
<emphasis id="B936D63BFFCFF260EBE3FF18F63FFEBF" bold="true" pageId="8" pageNumber="9">
<taxonomicName id="4C4271AAFFCFF260EBE3FF18F65DFF6F" authority="Balkenohl &amp; Aston, 2015" authorityName="Balkenohl &amp; Aston" authorityYear="2015" box="[221,298,186,204]" class="Insecta" family="Carabidae" genus="Asioreicheia" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="guenardi" status="sp. nov.">
<emphasis id="B936D63BFFCFF260EBE3FF18F65DFF6F" bold="true" box="[221,298,186,204]" italics="true" pageId="8" pageNumber="9">guenardi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2056B40FFCFF260EA11FF18F602FF6F" box="[303,373,186,204]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
,
<typeStatus id="54F9B48BFFCFF260EBE3FF77F65BFF44" box="[221,300,213,231]" pageId="8" pageNumber="9" type="paratype">paratype</typeStatus>
female, left coxostylus, dorsal view.
</emphasis>
</paragraph>
</caption>
<subSubSection id="C35859A2FFCFF260EA3DFBC4F60CFA57" pageId="8" pageNumber="9" type="discussion">
<paragraph id="8BFD0A29FFCFF260EA3DFBC4F60CFA57" blockId="8.[221,1107,1126,1636]" pageId="8" pageNumber="9">
However, during the last ice age the sea receded for
<quantity id="4CBAA7CCFFCFF260E851FBC4F4C0FBDD" box="[879,951,1126,1150]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="9" unit="km" value="50.0">50 km</quantity>
(B. Guénard, personal communication), and we have to assume that during this period this channel would have been a deep river. Heishiding National Reserve (the record of
<taxonomicName id="4C4271AAFFCFF260EA74FB74F6A3FB4D" baseAuthorityName="BULIRSCH, MAGRINI &amp; JIA" baseAuthorityYear="2013" box="[330,468,1238,1262]" class="Insecta" family="Carabidae" genus="Asioreicheia" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="chinensis">
<emphasis id="B936D63BFFCFF260EA74FB74F6A3FB4D" box="[330,468,1238,1262]" italics="true" pageId="8" pageNumber="9">A. chinensis</emphasis>
</taxonomicName>
) is located between the He, Xi, and Bei river. All the rivers to the west of and including the Xi and Bei river sections are deeply cut rivers and will not have moved in 100,000's of years except for the point where the Xi meets the Bei which is on the Pearl river flood plain. The Pearl delta rivers are all massive, and there are a lot of them. But as they are on a flood plain, they will have all moved back and forth. So, it is supposed to be unlikely for a flightless terricol species to extend its distribution across these barriers and the delta.
</paragraph>
</subSubSection>
<subSubSection id="C35859A2FFCFF260EA3DF9A3F503F9C7" pageId="8" pageNumber="9" type="biology_ecology">
<paragraph id="8BFD0A29FFCFF260EA3DF9A3F503F9C7" blockId="8.[221,1107,1126,1636]" pageId="8" pageNumber="9">Habitat: The specimens were collected by soil sieving in a mature disturbed secondary forest with closed canopy and very dense leaf litter at altitudes between 156 and 295 meters.</paragraph>
</subSubSection>
<paragraph id="8BFD0A29FFCEF261EB87FF04F41DFC82" pageId="9" pageNumber="10">
<table id="F942F889FFCE0D97EB87FF04F354FC82" box="[185,1059,166,801]" gridcols="3" gridrows="9" pageId="9" pageNumber="10">
<tr id="3572086BFFCE0D97EB87FF04F354FF19" box="[185,1059,166,186]" gridrow="0" pageId="9" pageNumber="10">
<th id="76A36117FFCE0D97EB87FF04F6B5FF19" box="[185,450,166,186]" gridcol="0" gridrow="0" pageId="9" pageNumber="10">
<emphasis id="B936D63BFFCEF261EB87FF04F633FF19" bold="true" box="[185,324,166,186]" pageId="9" pageNumber="10">Character state</emphasis>
</th>
<th id="76A36117FFCE0D97EAD3FF04F59EFF19" box="[493,745,166,186]" gridcol="1" gridrow="0" pageId="9" pageNumber="10">
<taxonomicName id="4C4271AAFFCEF261EAD3FF04F52CFF19" baseAuthorityName="BULIRSCH, MAGRINI &amp; JIA" baseAuthorityYear="2013" box="[493,603,166,186]" class="Insecta" family="Carabidae" genus="Asioreicheia" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="chinensis">
<emphasis id="B936D63BFFCEF261EAD3FF04F52CFF19" bold="true" box="[493,603,166,186]" italics="true" pageId="9" pageNumber="10">A. chinensis</emphasis>
</taxonomicName>
</th>
<th id="76A36117FFCE0D97E82AFF04F354FF19" box="[788,1059,166,186]" gridcol="2" gridrow="0" pageId="9" pageNumber="10">
<emphasis id="B936D63BFFCEF261E82AFF04F4BEFF19" bold="true" box="[788,969,166,186]" pageId="9" pageNumber="10">
<taxonomicName id="4C4271AAFFCEF261E82AFF04F40AFF19" authority="Balkenohl &amp; Aston, 2015" authorityName="Balkenohl &amp; Aston" authorityYear="2015" box="[788,893,166,186]" class="Insecta" family="Carabidae" genus="Asioreicheia" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="guenardi" status="sp. nov.">
<emphasis id="B936D63BFFCEF261E82AFF04F40AFF19" bold="true" box="[788,893,166,186]" italics="true" pageId="9" pageNumber="10">A. guenardi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2056B40FFCEF261E8BCFF04F4BEFF19" box="[898,969,166,186]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
</tr>
<tr id="3572086BFFCE0D97EB87FF6AF354FEA8" box="[185,1059,200,267]" gridrow="1" pageId="9" pageNumber="10">
<th id="76A36117FFCE0D97EB87FF6AF6B5FEA8" box="[185,450,200,267]" gridcol="0" gridrow="1" pageId="9" pageNumber="10">
<emphasis id="B936D63BFFCEF261EB87FF6AF79EFF7F" bold="true" box="[185,233,200,220]" pageId="9" pageNumber="10">Head</emphasis>
terminal maxillary palpomere eyes: ommatidium
</th>
<td id="76A36117FFCE0D97EAD3FF6AF59EFEA8" box="[493,745,200,267]" gridcol="1" gridrow="1" pageId="9" pageNumber="10">shorter flattened</td>
<td id="76A36117FFCE0D97E82AFF6AF354FEA8" box="[788,1059,200,267]" gridcol="2" gridrow="1" pageId="9" pageNumber="10">more slender,elongated to apex convex, protruding</td>
</tr>
<tr id="3572086BFFCE0D97EB87FE82F354FEC0" box="[185,1059,288,355]" gridrow="2" pageId="9" pageNumber="10">
<th id="76A36117FFCE0D97EB87FE82F6B5FEC0" box="[185,450,288,355]" gridcol="0" gridrow="2" pageId="9" pageNumber="10">
<emphasis id="B936D63BFFCEF261EB87FE82F665FE97" bold="true" box="[185,274,288,308]" pageId="9" pageNumber="10">Pronotum</emphasis>
lateral view before constriction anterior angles
</th>
<td id="76A36117FFCE0D97EAD3FE82F59EFEC0" box="[493,745,288,355]" gridcol="1" gridrow="2" pageId="9" pageNumber="10">distinctly convex angled</td>
<td id="76A36117FFCE0D97E82AFE82F354FEC0" box="[788,1059,288,355]" gridcol="2" gridrow="2" pageId="9" pageNumber="10">flatter, slightly convex rounded off</td>
</tr>
<tr id="3572086BFFCE0D97EB87FECBF354FE08" box="[185,1059,361,427]" gridrow="3" pageId="9" pageNumber="10">
<th id="76A36117FFCE0D97EB87FECBF6B5FE08" box="[185,450,361,427]" gridcol="0" gridrow="3" pageId="9" pageNumber="10">median line proepisternum surface</th>
<td id="76A36117FFCE0D97EAD3FECBF59EFE08" box="[493,745,361,427]" gridcol="1" gridrow="3" pageId="9" pageNumber="10">not joining basal constriction just visible dorsally smooth</td>
<td id="76A36117FFCE0D97E82AFECBF354FE08" box="[788,1059,361,427]" gridcol="2" gridrow="3" pageId="9" pageNumber="10">joining basal constriction distinctly tumid laterally irregular reticulation</td>
</tr>
<tr id="3572086BFFCE0D97EB87FE1DF354FD90" box="[185,1059,447,563]" gridrow="4" pageId="9" pageNumber="10">
<th id="76A36117FFCE0D97EB87FE1DF6B5FD90" box="[185,450,447,563]" gridcol="0" gridrow="4" pageId="9" pageNumber="10">
<emphasis id="B936D63BFFCEF261EB87FE1DF798FE70" bold="true" box="[185,239,447,467]" pageId="9" pageNumber="10">Elytra</emphasis>
lateral view, disk outline humerus lateral channel
</th>
<td id="76A36117FFCE0D97EAD3FE1DF59EFD90" box="[493,745,447,563]" gridcol="1" gridrow="4" pageId="9" pageNumber="10">flattened on disk elongated oval, more slender intimated equally developed</td>
<td id="76A36117FFCE0D97E82AFE1DF354FD90" box="[788,1059,447,563]" gridcol="2" gridrow="4" pageId="9" pageNumber="10">regularly convex regularly long-oval distinct narrow at middle</td>
</tr>
<tr id="3572086BFFCE0D97EB87FD9AF354FD30" box="[185,1059,568,659]" gridrow="5" pageId="9" pageNumber="10">
<th id="76A36117FFCE0D97EB87FD9AF6B5FD30" box="[185,450,568,659]" gridcol="0" gridrow="5" pageId="9" pageNumber="10">reflexed lateral margin teeth of serration number of setae on intervals carina in lateral channel at apex</th>
<td id="76A36117FFCE0D97EAD3FD9AF59EFD30" box="[493,745,568,659]" gridcol="1" gridrow="5" pageId="9" pageNumber="10">serrate up to middle indistinct, slightly rounded off 3.: 8; 5.: 5 missing</td>
<td id="76A36117FFCE0D97E82AFD9AF354FD30" box="[788,1059,568,659]" gridcol="2" gridrow="5" pageId="9" pageNumber="10">serrate up to apical third sharp up to middle 3.: 1012; 5.: 78 present</td>
</tr>
<tr id="3572086BFFCE0D97EB87FD05F354FD49" box="[185,1059,679,746]" gridrow="6" pageId="9" pageNumber="10">
<th id="76A36117FFCE0D97EB87FD05F6B5FD49" box="[185,450,679,746]" gridcol="0" gridrow="6" pageId="9" pageNumber="10">
<emphasis id="B936D63BFFCEF261EB87FD05F6D7FD18" bold="true" box="[185,416,679,699]" pageId="9" pageNumber="10">Median lobe of aedeagus</emphasis>
Shape/outline
</th>
<td id="76A36117FFCE0D97EAD3FD05F59EFD49" box="[493,745,679,746]" gridcol="1" gridrow="6" pageId="9" pageNumber="10">conspicuously curved, &lt;90°, turned differently,Fig. 5a</td>
<td id="76A36117FFCE0D97E82AFD05F354FD49" box="[788,1059,679,746]" gridcol="2" gridrow="6" pageId="9" pageNumber="10">less curved, rectangular,turned slightly, Fig. 3a</td>
</tr>
<tr id="3572086BFFCE0D97EB87FD52F354FCA1" box="[185,1059,752,770]" gridrow="7" pageId="9" pageNumber="10">
<th id="76A36117FFCE0D97EB87FD52F6B5FCA1" box="[185,450,752,770]" gridcol="0" gridrow="7" pageId="9" pageNumber="10">spatula at apex</th>
<td id="76A36117FFCE0D97EAD3FD52F59EFCA1" box="[493,745,752,770]" gridcol="1" gridrow="7" pageId="9" pageNumber="10">elongated, Fig. 5b</td>
<td id="76A36117FFCE0D97E82AFD52F354FCA1" box="[788,1059,752,770]" gridcol="2" gridrow="7" pageId="9" pageNumber="10">short, Fig. 3b</td>
</tr>
<tr id="3572086BFFCE0D97EB87FCADF354FC82" box="[185,1059,783,801]" gridrow="8" pageId="9" pageNumber="10">
<th id="76A36117FFCE0D97EB87FCADF6B5FC82" box="[185,450,783,801]" gridcol="0" gridrow="8" pageId="9" pageNumber="10">lips of orificium</th>
<td id="76A36117FFCE0D97EAD3FCADF354FC82" box="[493,1059,783,801]" colspan="2" colspanRight="1" gridcol="1" gridrow="8" pageId="9" pageNumber="10">different closing pattern</td>
</tr>
</table>
</paragraph>
<caption id="DF3D5AA1FFCEF261EB86FCE9F63AFCDB" ID-Table-UUID="DF3D5AA1FFCEF261EB86FCE9F63AFCDB" httpUri="http://table.plazi.org/id/DF3D5AA1FFCEF261EB86FCE9F63AFCDB" pageId="9" pageNumber="10" startId="9.[184,218,843,861]" targetBox="[185,1059,166,801]" targetIsTable="true" targetPageId="9">
<paragraph id="8BFD0A29FFCEF261EB86FCE9F63AFCDB" blockId="9.[184,1068,843,888]" pageId="9" pageNumber="10">
Tab. 1: Dividing character states of
<taxonomicName id="4C4271AAFFCEF261EAC7FCE9F49BFCFE" authority="(BULIRSCH, MAGRINI &amp; JIA, 2013)" baseAuthorityName="BULIRSCH, MAGRINI &amp; JIA" baseAuthorityYear="2013" box="[505,1004,843,861]" class="Insecta" family="Carabidae" genus="Asioreicheia" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="chinensis">
<emphasis id="B936D63BFFCEF261EAC7FCE9F5BAFCFE" bold="true" box="[505,717,843,861]" italics="true" pageId="9" pageNumber="10">Asioreicheia chinensis (</emphasis>
<bibRefCitation id="EFD377D8FFCEF261E9F0FCE9F490FCFE" author="Bulirsch, P. &amp; Magrini, P. &amp; Jia, F." box="[718,999,843,861]" pageId="9" pageNumber="10" pagination="59 - 64" refId="ref3804" refString="Bulirsch, P., Magrini, P. &amp; Jia, F. (2013): Antireicheia chinensis sp. nov. of the subtribe Reicheiina (Coleoptera: Carabidae: Scaritiae) from the south-eastern China. - Acta Entomologica Musei Nationalis Prague 53: 59 - 64." type="journal article" year="2013">BULIRSCH, MAGRINI &amp; JIA, 2013</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4C4271AAFFCEF261EF24FCE9F672FCDB" authority="Balkenohl &amp; Aston, 2015" authorityName="Balkenohl &amp; Aston" authorityYear="2015" class="Insecta" family="Carabidae" genus="Asioreicheia" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="guenardi" status="sp. nov.">
<emphasis id="B936D63BFFCEF261EF24FCE9F672FCDB" bold="true" italics="true" pageId="9" pageNumber="10">A. guenardi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2056B40FFCEF261EA34FCC4F63AFCDB" box="[266,333,870,888]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<subSubSection id="C35859A2FFCEF261EBE3FC73F6E2FBFA" pageId="9" pageNumber="10" type="discussion">
<paragraph id="8BFD0A29FFCEF261EBE3FC73F6E2FBFA" blockId="9.[184,1068,977,1113]" pageId="9" pageNumber="10">
Remark: According to the label of the holo- and
<typeStatus id="54F9B48BFFCEF261E832FC73F405FC4A" box="[780,882,977,1001]" pageId="9" pageNumber="10" type="paratype">paratype</typeStatus>
of
<emphasis id="B936D63BFFCEF261E8A4FC73F35BFC4A" box="[922,1068,977,1001]" italics="true" pageId="9" pageNumber="10">
<taxonomicName id="4C4271AAFFCEF261E8A4FC73F35FFC4A" baseAuthorityName="BULIRSCH, MAGRINI &amp; JIA" baseAuthorityYear="2013" box="[922,1064,977,1001]" class="Insecta" family="Carabidae" genus="Asioreicheia" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="chinensis">A. chinensis</taxonomicName>
,
</emphasis>
this species was collected by &quot;sifting of moist leaf litter in the dried-up stream beds and along the streams in the primary lowland forest&quot;, and at an altitude of 190260 meters.
</paragraph>
</subSubSection>
</treatment>
</document>