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<document id="458DE0EC8946FB5D0371742C3E47121D" ID-CLB-Dataset="26919" ID-DOI="10.11646/zootaxa.4598.1.1" ID-GBIF-Dataset="475fd53e-c5c6-4a99-bb2e-cc2382f0171f" ID-ISSN="1175-5326" ID-Zenodo-Dep="2668063" ID-ZooBank="6BA5F086-1014-46B5-B6D5-77FE0CD07AEC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1557127411994" checkinUser="plazi" docAuthor="Seldon, David S. &amp; Buckley, Thomas R." docDate="2019" docId="03BBD0260274B77D90FBB658EFABFB4F" docLanguage="en" docName="zootaxa.4598.1.1.pdf.imf" docOrigin="Zootaxa 4598 (1)" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Mecodema tenaki Seldon &amp; Leschen 2011" docType="treatment" docVersion="11" lastPageNumber="116" masterDocId="FF82A85E0205B70E906CB267ED0BFFA4" masterDocTitle="The genus Mecodema Blanchard 1853 (Coleoptera: Carabidae: Broscini) from the North Island, New Zealand" masterLastPageNumber="148" masterPageNumber="1" pageNumber="114" updateTime="1698726038790" updateUser="ExternalLinkService">
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<mods:title id="B1A69FDCFE6E1F4910F0215EF2BF3012">The genus Mecodema Blanchard 1853 (Coleoptera: Carabidae: Broscini) from the North Island, New Zealand</mods:title>
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<mods:namePart id="3811ACF0DAF0CB075B1DF2213AF2E0A0">Seldon, David S.</mods:namePart>
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<mods:namePart id="E51501F28E56B89B7F780CB6D2851FF5">Buckley, Thomas R.</mods:namePart>
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<mods:title id="7B61961BD651F0AF4D88448E1D79B51D">Zootaxa</mods:title>
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<mods:classification id="95FC19528D0414A254BBB0EB461AC73F">journal article</mods:classification>
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<treatment id="03BBD0260274B77D90FBB658EFABFB4F" ID-DOI="http://doi.org/10.5281/zenodo.5678440" ID-GBIF-Taxon="157062703" ID-Zenodo-Dep="5678440" LSID="urn:lsid:plazi:treatment:03BBD0260274B77D90FBB658EFABFB4F" httpUri="http://treatment.plazi.org/id/03BBD0260274B77D90FBB658EFABFB4F" lastPageId="115" lastPageNumber="116" pageId="113" pageNumber="114">
<subSubSection id="C30832BB0274B77F90FBB658EFAEFBFE" box="[151,677,1087,1114]" pageId="113" pageNumber="114" type="nomenclature">
<paragraph id="8BAD61300274B77F90FBB658EFAEFBFE" blockId="113.[151,677,1087,1147]" box="[151,677,1087,1114]" pageId="113" pageNumber="114">
<heading id="D0E5D65C0274B77F90FBB658EFAEFBFE" bold="true" box="[151,677,1087,1114]" fontSize="11" level="1" pageId="113" pageNumber="114" reason="1">
<emphasis id="B966BD220274B77F90FBB658EFAEFBFE" bold="true" box="[151,677,1087,1114]" pageId="113" pageNumber="114">
<taxonomicName id="4C121AB30274B77F90FBB658EFABFBFE" ID-CoL="6QWN5" authority="Seldon &amp; Leschen, 2011" authorityName="Seldon &amp; Leschen" authorityYear="2011" box="[151,672,1087,1114]" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="113" pageNumber="114" phylum="Arthropoda" rank="species" species="tenaki">
<emphasis id="B966BD220274B77F90FBB658EC66FBFD" bold="true" box="[151,365,1087,1113]" italics="true" pageId="113" pageNumber="114">Mecodema tenaki</emphasis>
<bibRefCitation id="EF831CC10274B77F9118B627EFABFBFE" author="Seldon, D. S. &amp; Leschen, R. A. B." box="[372,672,1088,1114]" pageId="113" pageNumber="114" pagination="1 - 45" refId="ref93265" refString="Seldon, D. S. &amp; Leschen, R. A. B. (2011) Revision of the Mecodema curvidens species group (Coleoptera: Carabidae: Broscini). Zootaxa, 2829, 1 - 45. https: // doi. org / 10.11646 / zootaxa. 2829.1.1" type="journal article" year="2011">Seldon &amp; Leschen, 2011</bibRefCitation>
</taxonomicName>
.
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30832BB0274B77F90FBB605EC01FBDF" box="[151,266,1122,1147]" pageId="113" pageNumber="114" type="description">
<paragraph id="8BAD61300274B77F90FBB605EC01FBDF" blockId="113.[151,677,1087,1147]" box="[151,266,1122,1147]" pageId="113" pageNumber="114">
<figureCitation id="13297DB50274B77F90FBB605EC0DFBDF" box="[151,262,1122,1147]" captionStart="FIGURE 56" captionStartId="114.[151,250,1844,1866]" captionTargetBox="[253,1331,200,1815]" captionTargetId="figure@114.[249,1337,195,1823]" captionTargetPageId="114" captionText="FIGURE 56. Mecodema tenaki plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668177/files/figure.png" pageId="113" pageNumber="114">Figure 56</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30832BB0274B77F90FBB6CDE824FAAF" pageId="113" pageNumber="114" type="diagnosis">
<paragraph id="8BAD61300274B77F90FBB6CDE824FAAF" blockId="113.[151,1437,1194,1975]" pageId="113" pageNumber="114">
<emphasis id="B966BD220274B77F90FBB6CDEC18FB67" bold="true" box="[151,275,1194,1219]" pageId="113" pageNumber="114">Diagnosis:</emphasis>
Differs from other North Island
<taxonomicName id="4C121AB30274B77F92E5B6CCEE0FFB67" authorityName="Blanchard" authorityYear="1853" box="[649,772,1195,1219]" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="113" pageNumber="114" phylum="Arthropoda" rank="genus">
<emphasis id="B966BD220274B77F92E5B6CCEE0FFB67" box="[649,772,1195,1219]" italics="true" pageId="113" pageNumber="114">Mecodema</emphasis>
</taxonomicName>
species by having:
<emphasis id="B966BD220274B77F938AB6CDEEFFFB67" bold="true" box="[998,1012,1194,1219]" pageId="113" pageNumber="114">1</emphasis>
, pronotal carina with 4-6 setae along each side (
<taxonomicName id="4C121AB30274B77F917CB6A9EC76FB42" authorityName="Broun" authorityYear="1915" box="[272,381,1230,1254]" class="Insecta" family="Carabidae" genus="Metaglymma" kingdom="Animalia" order="Coleoptera" pageId="113" pageNumber="114" phylum="Arthropoda" rank="species" species="curvidens">
<emphasis id="B966BD220274B77F917CB6A9EC76FB42" box="[272,381,1230,1254]" italics="true" pageId="113" pageNumber="114">curvidens</emphasis>
</taxonomicName>
species group);
<emphasis id="B966BD220274B77F9255B6A9EF4CFB43" bold="true" box="[569,583,1230,1255]" pageId="113" pageNumber="114">2</emphasis>
, pronotal carina smooth;
<emphasis id="B966BD220274B77F9306B6A9EE73FB43" bold="true" box="[874,888,1230,1255]" pageId="113" pageNumber="114">3</emphasis>
, pronotal disc with a small depression each side anterad pronotal foveae;
<emphasis id="B966BD220274B77F91C2B695ECB7FAAF" bold="true" box="[430,444,1266,1291]" pageId="113" pageNumber="114">4</emphasis>
, shape of the basal lobe and setal distribution of the left paramere (
<figureCitation id="13297DB50274B77F94C1B695E9F4FAAF" box="[1197,1279,1266,1291]" captionStart="FIGURE 56" captionStartId="114.[151,250,1844,1866]" captionTargetBox="[253,1331,200,1815]" captionTargetId="figure@114.[249,1337,195,1823]" captionTargetPageId="114" captionText="FIGURE 56. Mecodema tenaki plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668177/files/figure.png" pageId="113" pageNumber="114">Fig. 56</figureCitation>
LP).
</paragraph>
</subSubSection>
<subSubSection id="C30832BB0274B77D90ABB771ECA2FC4B" lastPageId="115" lastPageNumber="116" pageId="113" pageNumber="114" type="description">
<paragraph id="8BAD61300274B77F90ABB771EE14FAF7" blockId="113.[151,1437,1194,1975]" pageId="113" pageNumber="114">
<emphasis id="B966BD220274B77F90ABB771EC50FA8B" bold="true" box="[199,347,1302,1327]" pageId="113" pageNumber="114">Description:</emphasis>
Length
<quantity id="4CEACCD50274B77F91D2B771EF5AFA8A" box="[446,593,1302,1327]" metricMagnitude="-2" metricUnit="m" metricValue="2.53" metricValueMax="2.7" metricValueMin="2.36" pageId="113" pageNumber="114" unit="mm" value="25.3" valueMax="27.0" valueMin="23.6">23.627 mm</quantity>
, pronotal width
<quantity id="4CEACCD50274B77F937DB771EE95FA8A" box="[785,926,1302,1327]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" metricValueMax="7.4" metricValueMin="6.4" pageId="113" pageNumber="114" unit="mm" value="6.9" valueMax="7.4" valueMin="6.4">6.47.4 mm</quantity>
, elytral width
<quantity id="4CEACCD50274B77F9424B771E9DEFA8A" box="[1096,1237,1302,1327]" metricMagnitude="-3" metricUnit="m" metricValue="7.95" metricValueMax="8.5" metricValueMin="7.4" pageId="113" pageNumber="114" unit="mm" value="7.95" valueMax="8.5" valueMin="7.4">7.48.5 mm</quantity>
. Colour of entire dorsal body glossy black, entire ventral body matte black.
</paragraph>
<paragraph id="8BAD61300274B77F90ABB739EE1FF91E" blockId="113.[151,1437,1194,1975]" pageId="113" pageNumber="114">
<emphasis id="B966BD220274B77F90ABB739EC06FAD2" box="[199,269,1374,1398]" italics="true" pageId="113" pageNumber="114">Head:</emphasis>
Broad and flat (
<figureCitation id="13297DB50274B77F91A6B739EF3FFAD3" box="[458,564,1374,1399]" captionStart="FIGURE 14" captionStartId="20.[151,250,936,958]" captionTargetBox="[171,1412,204,911]" captionTargetId="figure@20.[160,1412,193,921]" captionTargetPageId="20" captionText="FIGURE 14. Diagrammatic cross sections that represent the width and convexity of head or deflection of pronotum (frontal view): (A) broad and flat; (B) broad and convex; (C) narrow and flat; (D) narrow and convex." httpUri="https://zenodo.org/record/2668091/files/figure.png" pageId="113" pageNumber="114">Fig. 14A</figureCitation>
). Vertex smooth; vertexal groove narrow, defined entire length with V-shape medially (
<figureCitation id="13297DB50274B77F9164B7E5EC57FA3F" box="[264,348,1410,1435]" captionStart="FIGURE 56" captionStartId="114.[151,250,1844,1866]" captionTargetBox="[253,1331,200,1815]" captionTargetId="figure@114.[249,1337,195,1823]" captionTargetPageId="114" captionText="FIGURE 56. Mecodema tenaki plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668177/files/figure.png" pageId="113" pageNumber="114">Fig. 56</figureCitation>
); large supraorbital puncture bearing 3 setae; 2 shallow supraorbital grooves; frons smooth with a large and very shallow depression each side of midline; frontoclypeal suture (
<figureCitation id="13297DB50274B77F9470B7C1E96AFA1B" box="[1052,1121,1446,1471]" captionStart="FIGURE 9" captionStartId="15.[151,250,1706,1728]" captionTargetBox="[157,1433,759,1683]" captionTargetId="figure@15.[151,1436,757,1683]" captionTargetPageId="15" captionText="FIGURE 9. Dorsal view of head with detail of the morphological structures used in the species descriptions. Ƒ = vertex; SPS = supraorbital puncture setae; SP = supraorbital puncture; F = frons area; FCS = frontoclypeal suture; C = clypeus; M = mandible; LS = labrum setae; L = labrum; ACM = anterior clypeal microsculpture; CSP = clypeal setose puncture; TP = tentorial pit; SG = supraorbital grooves; E = eye; ƑG = vertexal groove." httpUri="https://zenodo.org/record/2668081/files/figure.png" pageId="113" pageNumber="114">Fig. 9</figureCitation>
) indistinct, except laterally where it meets deep tentorial pits; anterior area of clypeus with indistinct or without microsculpture; clypeus with 1 large setose puncture on each side bearing 23 setae (
<figureCitation id="13297DB50274B77F9293B789EE5DF9A3" box="[767,854,1518,1543]" captionStart="FIGURE 56" captionStartId="114.[151,250,1844,1866]" captionTargetBox="[253,1331,200,1815]" captionTargetId="figure@114.[249,1337,195,1823]" captionTargetPageId="114" captionText="FIGURE 56. Mecodema tenaki plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668177/files/figure.png" pageId="113" pageNumber="114">Fig. 56</figureCitation>
). Labrum rounded, anterior edge outwardly curved with 2 proximate central setae, 2 setae each side evenly spaced. Mentum lobes squared (
<figureCitation id="13297DB50274B77F941FB475E9D0F98F" box="[1139,1243,1554,1579]" captionStart="FIGURE 12" captionStartId="18.[151,250,1905,1927]" captionTargetBox="[311,1244,697,1878]" captionTargetId="figure@18.[310,1250,689,1889]" captionTargetPageId="18" captionText="FIGURE 12. Ventral view of the mentum with a number of different character states. Illustrations AD are the different forms of the indentation in the apex of the median process: (A) distinctly indentate (M. manaia); (B) moderately indentate (M. pluto); (C) slightly indentate (M. infimate); (D) indentation notched (M. parataiko). Illustrations EG are the three general forms (i.e., hashed lines) of the mentum lobes: (E) triangular; (F) rounded; (G) squared." httpUri="https://zenodo.org/record/2668087/files/figure.png" pageId="113" pageNumber="114">Fig. 12G</figureCitation>
), median process very short and broad, slightly angled upward (15°), strongly indentate (
<figureCitation id="13297DB50274B77F939CB451E955F9EB" box="[1008,1118,1590,1615]" captionStart="FIGURE 12" captionStartId="18.[151,250,1905,1927]" captionTargetBox="[311,1244,697,1878]" captionTargetId="figure@18.[310,1250,689,1889]" captionTargetPageId="18" captionText="FIGURE 12. Ventral view of the mentum with a number of different character states. Illustrations AD are the different forms of the indentation in the apex of the median process: (A) distinctly indentate (M. manaia); (B) moderately indentate (M. pluto); (C) slightly indentate (M. infimate); (D) indentation notched (M. parataiko). Illustrations EG are the three general forms (i.e., hashed lines) of the mentum lobes: (E) triangular; (F) rounded; (G) squared." httpUri="https://zenodo.org/record/2668087/files/figure.png" pageId="113" pageNumber="114">Fig. 12A</figureCitation>
); setose punctures present. Submentum sclerite constriction narrow with 46 setae clustered laterally. Stipes with 2 basal setae. Gula pits large within defined suture, gula (
<figureCitation id="13297DB50274B77F91B0B419EF2BF933" box="[476,544,1662,1687]" captionStart="FIGURE 8" captionStartId="13.[151,250,1097,1119]" captionTargetBox="[155,1432,309,1072]" captionTargetId="figure@13.[151,1436,303,1075]" captionTargetPageId="13" captionText="FIGURE 8. Detail of the ventral head with the taxonomic structures used in the species descriptions. SBMS = submentum setae; SBM = submentum sclerite; ML = mentum lobe; MP = mentum process (with indentation); MPW = mentum process width; SBS = stipes basal setae; MPL = mentum process length; MPS = mentum process setae; SBMC = submentum sclerite constriction; GP = gula pits; GS = gula suture." httpUri="https://zenodo.org/record/2668079/files/figure.png" pageId="113" pageNumber="114">Fig. 8</figureCitation>
) weakly convex with very fine and closely packed transverse wrinkles. Gena with well-defined isodiametric pattern across entire area.
</paragraph>
<paragraph id="8BAD61300274B77F90ABB4A1EE8EF813" blockId="113.[151,1437,1194,1975]" pageId="113" pageNumber="114">
<emphasis id="B966BD220274B77F90ABB4A1EC49F97A" box="[199,322,1734,1758]" italics="true" pageId="113" pageNumber="114">Prothorax:</emphasis>
Prothoracic carina narrow the entire length, slightly crenulated with 46 setae each side (
<figureCitation id="13297DB50274B77F9557B4A1E884F97B" box="[1339,1423,1734,1759]" captionStart="FIGURE 56" captionStartId="114.[151,250,1844,1866]" captionTargetBox="[253,1331,200,1815]" captionTargetId="figure@114.[249,1337,195,1823]" captionTargetPageId="114" captionText="FIGURE 56. Mecodema tenaki plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668177/files/figure.png" pageId="113" pageNumber="114">Fig. 56</figureCitation>
), extended beyond anterior angle; posterior lateral sinuation indistinctly carinate, slightly angled inward; pronotum broad and flattened but deflected to carina, overall shape cordate (
<figureCitation id="13297DB50274B77F93E2B569EEEFF883" box="[910,996,1806,1831]" captionStart="FIGURE 56" captionStartId="114.[151,250,1844,1866]" captionTargetBox="[253,1331,200,1815]" captionTargetId="figure@114.[249,1337,195,1823]" captionTargetPageId="114" captionText="FIGURE 56. Mecodema tenaki plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668177/files/figure.png" pageId="113" pageNumber="114">Fig. 56</figureCitation>
); midline well-defined, fine transverse wrinkles may be present laterally on disc, small shallow dents present either side of midline; pronotal foveae deep and narrow; anterior edge slightly inwardly curved, posterior edge straight, slightly emarginated medially. Prosternum flat with fine, evenly spaced, transverse lines; proepisternum without microsculpture. Procoxal (
<figureCitation id="13297DB50274B77F953DB51DE89FF837" box="[1361,1428,1914,1939]" captionStart="FIGURE 2" captionStartId="6.[151,250,800,822]" captionTargetBox="[159,1432,197,773]" captionTargetId="figure@6.[151,1436,193,779]" captionTargetPageId="6" captionText="FIGURE 2. Ventral view of a Mecodema specimen showing specific morphological structures, excluding taxonomic structures indicated in detail figures, used in the species descriptions. ƑLF = ventrite lateral foveae; ƑSP = ventrite setose punctures; MTC = metacoxa; MTƑP = metaventrite process; MSC = mesocoxa; PC = procoxa; PS = prosternum; G = gena; PES = proepisternum; MSE = mesepisternum; MTE = metepisternum; Ƒ1Ƒ6 = ventrites 1-6 (ventrites 1-3 may be fused); M = midline (dashed line, not a taxonomic structure)." httpUri="https://zenodo.org/record/2668067/files/figure.png" pageId="113" pageNumber="114">Fig. 2</figureCitation>
) setae absent; protibia distally expanded and shovel-like (
<figureCitation id="13297DB50274B77F937DB5F9EE73F813" box="[785,888,1950,1975]" captionStart="FIGURE 10" captionStartId="16.[151,250,1442,1464]" captionTargetBox="[152,1428,191,1421]" captionTargetId="figure@16.[151,1435,183,1427]" captionTargetPageId="16" captionText="FIGURE 10. The protibia form is a character used mainly to determine between genera, except for one species in Mecodema where the difference between (A) greatly distally expanded (M. antarctica) and (B) expanded distally (all other Mecodema species) is used." httpUri="https://zenodo.org/record/2668083/files/figure.png" pageId="113" pageNumber="114">Fig. 10B</figureCitation>
).
</paragraph>
<caption id="DF6D31B80277B77C90FBB553E808F82C" httpUri="https://zenodo.org/record/2668177/files/figure.png" pageId="114" pageNumber="115" startId="114.[151,250,1844,1866]" targetBox="[253,1331,200,1815]" targetPageId="114">
<paragraph id="8BAD61300277B77C90FBB553E808F82C" blockId="114.[151,1436,1844,1928]" pageId="114" pageNumber="115">
<emphasis id="B966BD220277B77C90FBB553EC2BF8EE" bold="true" box="[151,288,1844,1866]" pageId="114" pageNumber="115">FIGURE 56.</emphasis>
<taxonomicName id="4C121AB30277B77C914AB553ECD2F8EE" authorityName="Seldon &amp; Leschen" authorityYear="2011" box="[294,473,1844,1866]" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="tenaki">
<emphasis id="B966BD220277B77C914AB553ECD2F8EE" box="[294,473,1844,1866]" italics="true" pageId="114" pageNumber="115">Mecodema tenaki</emphasis>
</taxonomicName>
plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe.
</paragraph>
</caption>
<paragraph id="8BAD61300276B77D90ABB2FFE919FEC3" blockId="115.[151,1437,151,1259]" pageId="115" pageNumber="116">
<emphasis id="B966BD220276B77D90ABB2FFEC1DFF14" box="[199,278,152,176]" italics="true" pageId="115" pageNumber="116">Elytra:</emphasis>
Broad and flattened but deflected to carina; humeral angle anteriorly convergent (
<figureCitation id="13297DB50276B77D9486B2F0E842FF14" box="[1258,1353,151,176]" captionStart="FIGURE 6" captionStartId="10.[151,250,519,541]" captionTargetBox="[156,1432,193,498]" captionTargetId="figure@10.[151,1436,193,498]" captionTargetPageId="10" captionText="FIGURE 6. Detail of the three different forms of the elytral humeral angle: (A) = anteriorly convergent (M. parataiko), (B) = evenly convex (M. oconnori), (C) = subangulate (M. atuanui)." httpUri="https://zenodo.org/record/2668075/files/figure.png" pageId="115" pageNumber="116">Fig. 6A</figureCitation>
); basal margin sharply angled (
<figureCitation id="13297DB50276B77D91CAB2DBECF2FF71" box="[422,505,188,213]" captionStart="FIGURE 56" captionStartId="114.[151,250,1844,1866]" captionTargetBox="[253,1331,200,1815]" captionTargetId="figure@114.[249,1337,195,1823]" captionTargetPageId="114" captionText="FIGURE 56. Mecodema tenaki plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668177/files/figure.png" pageId="115" pageNumber="116">Fig. 56</figureCitation>
) and bevelled to base, interval 1 extended to scutellum, other intervals and striae truncated at margin; humerus with 12 setose punctures (
<figureCitation id="13297DB50276B77D9370B2B8EE7BFF5C" box="[796,880,223,248]" captionStart="FIGURE 56" captionStartId="114.[151,250,1844,1866]" captionTargetBox="[253,1331,200,1815]" captionTargetId="figure@114.[249,1337,195,1823]" captionTargetPageId="114" captionText="FIGURE 56. Mecodema tenaki plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668177/files/figure.png" pageId="115" pageNumber="116">Fig. 56</figureCitation>
); lateral carina (
<figureCitation id="13297DB50276B77D944BB2B8E97CFF5C" box="[1063,1143,223,248]" captionStart="FIGURE 11" captionStartId="17.[151,250,949,971]" captionTargetBox="[221,1358,181,923]" captionTargetId="figure@17.[221,1359,179,929]" captionTargetPageId="17" captionText="FIGURE 11. Detail of pronotum showing the taxonomic structures used in the species descriptions. PPC = posterior pronotum constriction; PS = pronotal setae; PAA = pronotum anterior angle; AE = anterior edge; PC = pronotal carina; ML = midline; PF= pronotal foveae; PPS = pronotum posterior sinuation; PE = posterior edge." httpUri="https://zenodo.org/record/2668085/files/figure.png" pageId="115" pageNumber="116">Fig. 11</figureCitation>
) narrow the entire length, extended to humeral angle; suture well-defined; striae with small, regularly distributed, asetose punctures; intervals weakly convex; interval microsculpture absent; 7
<superScript id="7C67CC780276B77D92AEB340EFC5FE91" attach="left" box="[706,718,295,309]" fontSize="6" pageId="115" pageNumber="116">th</superScript>
strial setal pattern with 01 setose punctures in anterior ½, 34 evenly spaced setose punctures in posterior ½ (
<figureCitation id="13297DB50276B77D92CBB329EFF1FEC3" box="[679,762,334,359]" captionStart="FIGURE 56" captionStartId="114.[151,250,1844,1866]" captionTargetBox="[253,1331,200,1815]" captionTargetId="figure@114.[249,1337,195,1823]" captionTargetPageId="114" captionText="FIGURE 56. Mecodema tenaki plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668177/files/figure.png" pageId="115" pageNumber="116">Fig. 56</figureCitation>
), setose punctures small.
</paragraph>
<paragraph id="8BAD61300276B77D90ABB314E92CFD9B" blockId="115.[151,1437,151,1259]" pageId="115" pageNumber="116">
<emphasis id="B966BD220276B77D90ABB314EC8DFE2F" box="[199,390,371,395]" italics="true" pageId="115" pageNumber="116">Ventral surface:</emphasis>
Mesepisternum with rugose wrinkles; metepisternum without microsculpture; setose punctures present on mesocoxae (1) and metacoxae (1). All abdominal ventrites (
<figureCitation id="13297DB50276B77D9405B3F1E9B9FE0B" box="[1129,1202,406,431]" captionStart="FIGURE 2" captionStartId="6.[151,250,800,822]" captionTargetBox="[159,1432,197,773]" captionTargetId="figure@6.[151,1436,193,779]" captionTargetPageId="6" captionText="FIGURE 2. Ventral view of a Mecodema specimen showing specific morphological structures, excluding taxonomic structures indicated in detail figures, used in the species descriptions. ƑLF = ventrite lateral foveae; ƑSP = ventrite setose punctures; MTC = metacoxa; MTƑP = metaventrite process; MSC = mesocoxa; PC = procoxa; PS = prosternum; G = gena; PES = proepisternum; MSE = mesepisternum; MTE = metepisternum; Ƒ1Ƒ6 = ventrites 1-6 (ventrites 1-3 may be fused); M = midline (dashed line, not a taxonomic structure)." httpUri="https://zenodo.org/record/2668067/files/figure.png" pageId="115" pageNumber="116">Fig. 2</figureCitation>
) wrinkled laterally; ventrites 13 without setose punctures, ventrites 4 and 5 setose punctures present (12) each side of midline; lateral foveae absent; ventrite 6 setae present:
<specimenCount id="9D14AAB90276B77D9234B387EF66FE53" box="[600,621,480,503]" pageId="115" pageNumber="116" type="male"></specimenCount>
with 24 setose punctures each side of midline, apical edge straight,
<specimenCount id="9D14AAB90276B77D95EBB387E897FE53" box="[1415,1436,480,503]" pageId="115" pageNumber="116" type="female"></specimenCount>
with 24 setose punctures each side of midline, apical edge bluntly pointed. Anterior metaventrite process (
<figureCitation id="13297DB50276B77D953CB065E89FFDBF" box="[1360,1428,514,539]" captionStart="FIGURE 2" captionStartId="6.[151,250,800,822]" captionTargetBox="[159,1432,197,773]" captionTargetId="figure@6.[151,1436,193,779]" captionTargetPageId="6" captionText="FIGURE 2. Ventral view of a Mecodema specimen showing specific morphological structures, excluding taxonomic structures indicated in detail figures, used in the species descriptions. ƑLF = ventrite lateral foveae; ƑSP = ventrite setose punctures; MTC = metacoxa; MTƑP = metaventrite process; MSC = mesocoxa; PC = procoxa; PS = prosternum; G = gena; PES = proepisternum; MSE = mesepisternum; MTE = metepisternum; Ƒ1Ƒ6 = ventrites 1-6 (ventrites 1-3 may be fused); M = midline (dashed line, not a taxonomic structure)." httpUri="https://zenodo.org/record/2668067/files/figure.png" pageId="115" pageNumber="116">Fig. 2</figureCitation>
) an apically broad triangle with carina poorly defined apically, narrowed posterad.
</paragraph>
<paragraph id="8BAD61300276B77D90ABB02CEEC1FC27" blockId="115.[151,1437,151,1259]" pageId="115" pageNumber="116">
<emphasis id="B966BD220276B77D90ABB02CEC7CFDC7" box="[199,375,587,611]" italics="true" pageId="115" pageNumber="116">Male genitalia:</emphasis>
Apical portion of penis lobe symmetrically rounded (
<figureCitation id="13297DB50276B77D9386B02DE936FDC7" box="[1002,1085,586,611]" captionStart="FIGURE 56" captionStartId="114.[151,250,1844,1866]" captionTargetBox="[253,1331,200,1815]" captionTargetId="figure@114.[249,1337,195,1823]" captionTargetPageId="114" captionText="FIGURE 56. Mecodema tenaki plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668177/files/figure.png" pageId="115" pageNumber="116">Fig. 56</figureCitation>
PL) with a slight deflection to right of vertical axis (VV) (
<figureCitation id="13297DB50276B77D91BDB009EF32FD23" box="[465,569,622,647]" captionStart="FIGURE 15" captionStartId="20.[151,250,1943,1965]" captionTargetBox="[162,1424,1039,1914]" captionTargetId="figure@20.[162,1424,1038,1921]" captionTargetPageId="20" captionText="FIGURE 15. The penis lobe in ventral view showing the relative deflection to right of vertical: (A) distinctly deflected (M. regulus); (B) moderately deflected (M. haunoho); (C) slightly deflected (M. oregoides). Scale bars = 1 mm. Structures of penis lobe indicated are: base (B); lateral edge (LE); apical portion (AP); ventral edge (ƑE)." httpUri="https://zenodo.org/record/2668093/files/figure.png" pageId="115" pageNumber="116">Fig. 15C</figureCitation>
); ventral edge of penis lobe straight (VV); overall length of shaft penis lobe broad and short, slightly curved ventrally (LV) (
<figureCitation id="13297DB50276B77D92C0B0F5EE18FD0F" box="[684,787,658,683]" captionStart="FIGURE 16" captionStartId="21.[151,250,897,919]" captionTargetBox="[151,1436,181,869]" captionTargetId="figure@21.[151,1436,181,876]" captionTargetPageId="21" captionText="FIGURE 16. Relative strength of the ventral curve over the length of the penis lobe (lateral view): (A) slightly curved (M. oblongum); (B) moderately curved (M. ponaiti); (C) distinctly curved (M. manaia). Scale bar = 1 mm." httpUri="https://zenodo.org/record/2668095/files/figure.png" pageId="115" pageNumber="116">Fig. 16A</figureCitation>
). Structures of the endophallus (
<figureCitation id="13297DB50276B77D9416B0F5E9B4FD0F" box="[1146,1215,658,683]" captionStart="FIGURE 4" captionStartId="7.[151,250,973,995]" captionTargetBox="[354,1232,193,952]" captionTargetId="figure@7.[354,1232,193,952]" captionTargetPageId="7" captionText="FIGURE 4. An Environmental Scanning Electron Microscope (ESEM) photograph of the four main structures of the apical plate of the endophallus within the penis lobe. This is an example from Mecodema haunoho showing the central spicule (c.s), left (l.s.f) and the right setose flange (r.s.f), and microtrichial field (m.f). These structures are variable among species but within a species, they are very consistent, even over distant geographical ranges. The microtrichial field can be covered in flattened scales as in photo above, or with elongated setae-like scales." httpUri="https://zenodo.org/record/2668071/files/figure.png" pageId="115" pageNumber="116">Fig. 4</figureCitation>
): lateral form of the apex of the central spicule rounded; dorsal form of the apex of the central spicule short and narrow; seta (scales) coverage of the apical plate moderate (2675%); size of left setose flange small; size of right setose flange large. Left paramere basal lobe ovate, narrowed slightly toward arm (
<figureCitation id="13297DB50276B77D9336B099EEA0FCB3" box="[858,939,766,791]" captionStart="FIGURE 56" captionStartId="114.[151,250,1844,1866]" captionTargetBox="[253,1331,200,1815]" captionTargetId="figure@114.[249,1337,195,1823]" captionTargetPageId="114" captionText="FIGURE 56. Mecodema tenaki plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668177/files/figure.png" pageId="115" pageNumber="116">Fig. 56</figureCitation>
LP), short slope to arm; arm long and broad, narrowed to indistinct terminal lobe; apical tuft of sparse, medium-lengthed setae; setae extended along apical ¾ of ventral edge (
<figureCitation id="13297DB50276B77D9151B121EC9EFCFB" box="[317,405,838,863]" captionStart="FIGURE 56" captionStartId="114.[151,250,1844,1866]" captionTargetBox="[253,1331,200,1815]" captionTargetId="figure@114.[249,1337,195,1823]" captionTargetPageId="114" captionText="FIGURE 56. Mecodema tenaki plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668177/files/figure.png" pageId="115" pageNumber="116">Fig. 56</figureCitation>
LP), ventral edge curved. Right paramere narrowly rectangular, gradually narrowed to terminus with setae extended along apical ¾ of ventral edge (
<figureCitation id="13297DB50276B77D932AB10DEE93FC27" box="[838,920,874,899]" captionStart="FIGURE 56" captionStartId="114.[151,250,1844,1866]" captionTargetBox="[253,1331,200,1815]" captionTargetId="figure@114.[249,1337,195,1823]" captionTargetPageId="114" captionText="FIGURE 56. Mecodema tenaki plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668177/files/figure.png" pageId="115" pageNumber="116">Fig. 56</figureCitation>
RP).
</paragraph>
<paragraph id="8BAD61300276B77D90ABB1E8ECA2FC4B" blockId="115.[151,1437,151,1259]" pageId="115" pageNumber="116">
<emphasis id="B966BD220276B77D90ABB1E8EC85FC03" box="[199,398,911,935]" italics="true" pageId="115" pageNumber="116">Female genitalia:</emphasis>
Basal gonocoxite 1 (
<figureCitation id="13297DB50276B77D9216B1E9EFB7FC03" box="[634,700,910,935]" captionStart="FIGURE 5" captionStartId="9.[151,250,1714,1736]" captionTargetBox="[167,1436,790,1674]" captionTargetId="figure@9.[151,1436,790,1693]" captionTargetPageId="9" captionText="FIGURE 5. The entire female reproductive tract (ventral view) including taxonomic structures (for all other abbreviated structure names see Seldon et al. 2012). However, for this revision only the following structures are important: HS = helminthoid sclerite; G1 = gonocoxite 1; G2 = gonocoxite 2; TDC = transverse dorsal carina; R = ramus." httpUri="https://zenodo.org/record/2668073/files/figure.png" pageId="115" pageNumber="116">Fig. 5</figureCitation>
) short and broad, ventral surface with a few longitudinal grooves, internal dorso-lateral carina with 2 setae. Gonocoxite 2 a short, rounded triangle with apical edge slightly reflexed. Ramus long and narrow.
</paragraph>
</subSubSection>
<subSubSection id="C30832BB0276B77D90ABB19DEE66FB93" pageId="115" pageNumber="116" type="discussion">
<paragraph id="8BAD61300276B77D90ABB19DEE66FB93" blockId="115.[151,1437,151,1259]" pageId="115" pageNumber="116">
<emphasis id="B966BD220276B77D90ABB19DEC5AFBB7" bold="true" box="[199,337,1018,1043]" pageId="115" pageNumber="116">Comments:</emphasis>
<taxonomicName id="4C121AB30276B77D9134B19CEF15FBB7" authorityName="Seldon &amp; Leschen" authorityYear="2011" box="[344,542,1019,1043]" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="115" pageNumber="116" phylum="Arthropoda" rank="species" species="tenaki">
<emphasis id="B966BD220276B77D9134B19CEF15FBB7" box="[344,542,1019,1043]" italics="true" pageId="115" pageNumber="116">Mecodema tenaki</emphasis>
</taxonomicName>
has a very limited range, residing in degraded native kanuka (
<taxonomicName id="4C121AB30276B77D94B6B19CE882FBB7" box="[1242,1417,1019,1043]" class="Magnoliopsida" family="Myrtaceae" genus="Kunzea" higherTaxonomySource="CoL" kingdom="Plantae" order="Myrtales" pageId="115" pageNumber="116" phylum="Tracheophyta" rank="species" species="robusta">
<emphasis id="B966BD220276B77D94B6B19CE882FBB7" box="[1242,1417,1019,1043]" italics="true" pageId="115" pageNumber="116">Kunzea robusta</emphasis>
</taxonomicName>
)- broadleaf forest patches in the North Cape to Cape Reinga areas.
</paragraph>
</subSubSection>
<subSubSection id="C30832BB0276B77D90ABB625EC5FFBDB" pageId="115" pageNumber="116" type="distribution">
<paragraph id="8BAD61300276B77D90ABB625EC5FFBDB" blockId="115.[151,1437,151,1259]" pageId="115" pageNumber="116">
<emphasis id="B966BD220276B77D90ABB625EC69FBFF" bold="true" box="[199,354,1090,1115]" pageId="115" pageNumber="116">Distribution:</emphasis>
<collectingCountry id="F30521A00276B77D9100B624EF0CFBFF" box="[364,519,1090,1115]" name="New Zealand" pageId="115" pageNumber="116">New Zealand</collectingCountry>
, North Island, Northland, restricted to the Cape Reinga to North Cape area (see
<bibRefCitation id="EF831CC10276B77D90FBB601EC42FBDB" author="Ball, O. J. &amp; Whaley, P. T. &amp; Booth, A. M. &amp; Hartley, S." box="[151,329,1126,1151]" pageId="115" pageNumber="116" pagination="84 - 94" refId="ref90712" refString="Ball, O. J. - P, Whaley, P. T., Booth, A. M. &amp; Hartley, S. (2013) Habitat associations and detectability of the endemic Te Paki ground beetle Mecodema tenaki (Coleoptera: Carabidae). New Zealand Journal of Ecology, 37 (1), 84 - 94." type="journal article" year="2013">
Ball
<emphasis id="B966BD220276B77D90A0B60FEC0EFBDB" box="[204,261,1127,1151]" italics="true" pageId="115" pageNumber="116">et al.</emphasis>
2013
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30832BB0276B77D90ABB6EDEFABFB4F" pageId="115" pageNumber="116" type="materials_examined">
<paragraph id="8BAD61300276B77D90ABB6EDE95AFB63" blockId="115.[151,1437,151,1259]" pageId="115" pageNumber="116">
<materialsCitation id="3B7A6B6D0276B77D90ABB6EDE880FB07" ID-GBIF-Occurrence="2237851782" box="[199,1419,1162,1187]" collectingDate="2006-11-13" collectionCode="NZAC, ND" collectorName="Cape Reinga Te Paki &amp; O. Ball" country="New Zealand" location="NEW ZEALAND" pageId="115" pageNumber="116" specimenCount="1" typeStatus="holotype">
<emphasis id="B966BD220276B77D90ABB6EDEC36FB07" bold="true" box="[199,317,1162,1187]" pageId="115" pageNumber="116">
<typeStatus id="54A9DF920276B77D90ABB6EDEC33FB07" box="[199,312,1162,1187]" pageId="115" pageNumber="116" type="holotype">Holotype</typeStatus>
:
</emphasis>
<collectionCode id="ED03F9F50276B77D912AB6ECEC99FB07" box="[326,402,1163,1187]" country="New Zealand" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34070" lsid="urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="115" pageNumber="116" type="Other institutional type">NZAC</collectionCode>
male labelled.
<collectingCountry id="F30521A00276B77D922EB6ECEE04FB07" box="[578,783,1163,1187]" name="New Zealand" pageId="115" pageNumber="116">NEW ZEALAND</collectingCountry>
<collectionCode id="ED03F9F50276B77D9374B6ECEE4BFB06" box="[792,832,1163,1186]" country="USA" lsid="urn:lsid:biocol.org:col:15197" name="University of Notre Dame" pageId="115" pageNumber="116" type="Herbarium">ND</collectionCode>
<collectorName id="26E704E60276B77D9325B6ECE931FB07" box="[841,1082,1162,1187]" pageId="115" pageNumber="116">Cape Reinga Te Paki</collectorName>
RBB-7
<date id="FFAC47F00276B77D94F6B6ECE83BFB07" box="[1178,1328,1162,1187]" pageId="115" pageNumber="116" value="2006-11-13">
<collectingDate id="EFE8BE180276B77D94F6B6ECE83BFB07" box="[1178,1328,1162,1187]" pageId="115" pageNumber="116" value="2006-11-13">13 Nov 2006</collectingDate>
</date>
<collectorName id="26E704E60276B77D9555B6ECE880FB07" box="[1337,1419,1162,1187]" pageId="115" pageNumber="116">O. Ball</collectorName>
</materialsCitation>
/
<typeStatus id="54A9DF920276B77D90FBB6C8EC20FB63" box="[151,299,1199,1223]" pageId="115" pageNumber="116" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4C121AB30276B77D915EB6C9ECF1FB63" authority="Seldon &amp; Buckley, 2019" authorityName="Seldon &amp; Buckley" authorityYear="2019" box="[306,506,1198,1223]" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="115" pageNumber="116" phylum="Arthropoda" rank="species" species="tenaki" status="sp. nov.">Mecodema tenaki</taxonomicName>
<emphasis id="B966BD220276B77D926CB6C9EF34FB63" bold="true" box="[512,575,1198,1223]" pageId="115" pageNumber="116">
<taxonomicNameLabel id="A25500590276B77D926CB6C9EF34FB63" box="[512,575,1198,1223]" pageId="115" pageNumber="116" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
design. by Seldon &amp; Leschen 2010 [red label].
</paragraph>
<paragraph id="8BAD61300276B77D90ABB6B5EFABFB4F" blockId="115.[151,1437,151,1259]" box="[199,672,1234,1259]" pageId="115" pageNumber="116">
<emphasis id="B966BD220276B77D90ABB6B5EC43FB4F" bold="true" box="[199,328,1234,1259]" pageId="115" pageNumber="116">
<typeStatus id="54A9DF920276B77D90ABB6B5EC48FB4F" box="[199,323,1234,1259]" pageId="115" pageNumber="116" type="paratype">Paratypes</typeStatus>
:
</emphasis>
(see
<bibRefCitation id="EF831CC10276B77D91EFB6B5EF9EFB4F" author="Seldon, D. S. &amp; Leschen, R. A. B." box="[387,661,1234,1259]" pageId="115" pageNumber="116" pagination="1 - 45" refId="ref93265" refString="Seldon, D. S. &amp; Leschen, R. A. B. (2011) Revision of the Mecodema curvidens species group (Coleoptera: Carabidae: Broscini). Zootaxa, 2829, 1 - 45. https: // doi. org / 10.11646 / zootaxa. 2829.1.1" type="journal article" year="2011">Seldon &amp; Leschen 2011</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>