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<document id="FB34992F0FE0309BF0144819291DC731" ID-CLB-Dataset="26919" ID-DOI="10.11646/zootaxa.4598.1.1" ID-GBIF-Dataset="475fd53e-c5c6-4a99-bb2e-cc2382f0171f" ID-ISSN="1175-5326" ID-Zenodo-Dep="2668063" ID-ZooBank="6BA5F086-1014-46B5-B6D5-77FE0CD07AEC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1557127411994" checkinUser="plazi" docAuthor="Seldon, David S. &amp; Buckley, Thomas R." docDate="2019" docId="03BBD026026FB76290FBB178EE4BF966" docLanguage="en" docName="zootaxa.4598.1.1.pdf.imf" docOrigin="Zootaxa 4598 (1)" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Mecodema simplex Castelnau 1867" docType="treatment" docVersion="11" lastPageNumber="109" masterDocId="FF82A85E0205B70E906CB267ED0BFFA4" masterDocTitle="The genus Mecodema Blanchard 1853 (Coleoptera: Carabidae: Broscini) from the North Island, New Zealand" masterLastPageNumber="148" masterPageNumber="1" pageNumber="107" updateTime="1698726038790" updateUser="ExternalLinkService">
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<mods:title id="16368B3986123583C569D0407060ADCD">The genus Mecodema Blanchard 1853 (Coleoptera: Carabidae: Broscini) from the North Island, New Zealand</mods:title>
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<mods:namePart id="E7254B4BCB5D92ADA577405ACF2B7648">Seldon, David S.</mods:namePart>
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<mods:namePart id="9B1439ACC41DECE0936E5A04438E73A9">Buckley, Thomas R.</mods:namePart>
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<mods:date id="F9C1033B0F274DE3EA6127F1A5D122B2">2019</mods:date>
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<treatment id="03BBD026026FB76290FBB178EE4BF966" ID-DOI="http://doi.org/10.5281/zenodo.5678434" ID-GBIF-Taxon="157062696" ID-Zenodo-Dep="5678434" LSID="urn:lsid:plazi:treatment:03BBD026026FB76290FBB178EE4BF966" httpUri="http://treatment.plazi.org/id/03BBD026026FB76290FBB178EE4BF966" lastPageId="108" lastPageNumber="109" pageId="106" pageNumber="107">
<subSubSection id="C30832BB026FB76490FBB178EF46FC9E" box="[151,589,799,826]" pageId="106" pageNumber="107" type="nomenclature">
<paragraph id="8BAD6130026FB76490FBB178EF46FC9E" blockId="106.[151,589,799,859]" box="[151,589,799,826]" pageId="106" pageNumber="107">
<heading id="D0E5D65C026FB76490FBB178EF46FC9E" bold="true" box="[151,589,799,826]" fontSize="11" level="1" pageId="106" pageNumber="107" reason="1">
<taxonomicName id="4C121AB3026FB76490FBB178EF46FC9E" ID-CoL="738MV" authority="Castelnau, 1867" authorityName="Castelnau" authorityYear="1867" box="[151,589,799,826]" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="106" pageNumber="107" phylum="Arthropoda" rank="species" species="simplex">
<emphasis id="B966BD22026FB76490FBB178EF46FC9E" bold="true" box="[151,589,799,826]" pageId="106" pageNumber="107">
<emphasis id="B966BD22026FB76490FBB178EC74FC9D" bold="true" box="[151,383,799,825]" italics="true" pageId="106" pageNumber="107">Mecodema simplex</emphasis>
<bibRefCitation id="EF831CC1026FB76491EAB147EF46FC9E" author="Castelnau, F." box="[390,589,800,826]" pageId="106" pageNumber="107" pagination="30 - 38" refId="ref91697" refString="Castelnau, F. Comte de (1867) Notes on Australian Coleoptera. Transactions and Proceedings of the Royal Society of Fictoria, 8, 30 - 38; 95 - 225." type="journal article" year="1867">Castelnau, 1867</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30832BB026FB76490FBB125EC01FCFF" box="[151,266,834,859]" pageId="106" pageNumber="107" type="description">
<paragraph id="8BAD6130026FB76490FBB125EC01FCFF" blockId="106.[151,589,799,859]" box="[151,266,834,859]" pageId="106" pageNumber="107">
<figureCitation id="13297DB5026FB76490FBB125EC0DFCFF" box="[151,262,834,859]" captionStart="FIGURE 53" captionStartId="107.[151,250,1855,1877]" captionTargetBox="[249,1335,199,1825]" captionTargetId="figure@107.[243,1343,193,1834]" captionTargetPageId="107" captionText="FIGURE 53. Mecodema simplex plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668171/files/figure.png" pageId="106" pageNumber="107">Figure 53</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30832BB026FB76490FBB1EDE94CFC7A" pageId="106" pageNumber="107" type="reference_group">
<paragraph id="8BAD6130026FB76490FBB1EDE922FC04" blockId="106.[151,1095,906,990]" box="[151,1065,906,928]" pageId="106" pageNumber="107">
<taxonomicName id="4C121AB3026FB76490FBB1EDEF0BFC04" ID-CoL="3YKRK" authority="Broun, 1908" authorityName="Broun" authorityYear="1908" box="[151,512,906,928]" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="106" pageNumber="107" phylum="Arthropoda" rank="species" species="acuductum">
<emphasis id="B966BD22026FB76490FBB1EDEC70FC04" box="[151,379,906,928]" italics="true" pageId="106" pageNumber="107">Mecodema acuductum</emphasis>
<bibRefCitation id="EF831CC1026FB76491EEB1ECEF0BFC04" author="Broun, T." box="[386,512,906,928]" pageId="106" pageNumber="107" pagination="543 - 547" refId="ref91387" refString="Broun, T. (1908) Descriptions of new species of New Zealand Coleoptera. Annals and Magazine of Natural History, 7 (15), 543 - 547." type="journal article" year="1908">Broun, 1908</bibRefCitation>
</taxonomicName>
(Mt Holdsworth, WN), synonymised by
<bibRefCitation id="EF831CC1026FB76493CEB1EDE92DFC04" author="Britton, E. B." box="[930,1062,906,928]" pageId="106" pageNumber="107" pagination="533 - 581" refId="ref90973" refString="Britton, E. B. (1949) The Carabidae (Coleoptera) of New Zealand part 3: A revision of the tribe Broscini. Transactions of the Royal Society of New Zealand, 77, 533 - 581." type="journal article" year="1949">Britton 1949</bibRefCitation>
.
</paragraph>
<paragraph id="8BAD6130026FB76490FBB1CEE90BFC1B" blockId="106.[151,1095,906,990]" box="[151,1024,937,959]" pageId="106" pageNumber="107">
<taxonomicName id="4C121AB3026FB76490FBB1CEECF9FC1B" authority="Broun, 1912" authorityName="Broun" authorityYear="1912" box="[151,498,937,959]" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="106" pageNumber="107" phylum="Arthropoda" rank="species" species="bryodium">
<emphasis id="B966BD22026FB76490FBB1CEEC66FC1B" box="[151,365,937,959]" italics="true" pageId="106" pageNumber="107">Mecodema bryodium</emphasis>
<bibRefCitation id="EF831CC1026FB7649118B1CDECF9FC1B" author="Broun, T." box="[372,498,937,959]" pageId="106" pageNumber="107" pagination="379 - 440" refId="ref91419" refString="Broun, T. (1912) Descriptions of new genera and species of Coleoptera. [Part I]. Transactions and Proceedings of the New Zealand Institute, 44, 379 - 440." type="journal article" year="1912">Broun, 1912</bibRefCitation>
</taxonomicName>
(Silverstream, WN), synonymised by
<bibRefCitation id="EF831CC1026FB7649314B1CEEEF7FC1B" author="Britton, E. B." box="[888,1020,937,959]" pageId="106" pageNumber="107" pagination="533 - 581" refId="ref90973" refString="Britton, E. B. (1949) The Carabidae (Coleoptera) of New Zealand part 3: A revision of the tribe Broscini. Transactions of the Royal Society of New Zealand, 77, 533 - 581." type="journal article" year="1949">Britton 1949</bibRefCitation>
.
</paragraph>
<paragraph id="8BAD6130026FB76490FBB1AFE94CFC7A" blockId="106.[151,1095,906,990]" box="[151,1095,968,990]" pageId="106" pageNumber="107">
<taxonomicName id="4C121AB3026FB76490FBB1AFECFAFC7A" ID-CoL="6QWMY" authority="Broun, 1912" authorityName="Broun" authorityYear="1912" box="[151,497,968,990]" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="106" pageNumber="107" phylum="Arthropoda" rank="species" species="arcuatum">
<emphasis id="B966BD22026FB76490FBB1AFEC67FC7A" box="[151,364,968,990]" italics="true" pageId="106" pageNumber="107">Mecodema arcuatum</emphasis>
<bibRefCitation id="EF831CC1026FB764911FB1AEECFAFC7A" author="Broun, T." box="[371,497,968,990]" pageId="106" pageNumber="107" pagination="379 - 440" refId="ref91419" refString="Broun, T. (1912) Descriptions of new genera and species of Coleoptera. [Part I]. Transactions and Proceedings of the New Zealand Institute, 44, 379 - 440." type="journal article" year="1912">Broun, 1912</bibRefCitation>
</taxonomicName>
(near Martinborough, WA), synonymised by
<bibRefCitation id="EF831CC1026FB76493ACB1AFE94FFC7A" author="Britton, E. B." box="[960,1092,968,990]" pageId="106" pageNumber="107" pagination="533 - 581" refId="ref90973" refString="Britton, E. B. (1949) The Carabidae (Coleoptera) of New Zealand part 3: A revision of the tribe Broscini. Transactions of the Royal Society of New Zealand, 77, 533 - 581." type="journal article" year="1949">Britton 1949</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30832BB026FB76490FBB66DE965FBCF" pageId="106" pageNumber="107" type="diagnosis">
<paragraph id="8BAD6130026FB76490FBB66DE965FBCF" blockId="106.[151,1437,1034,2034]" pageId="106" pageNumber="107">
<emphasis id="B966BD22026FB76490FBB66DEC18FB87" bold="true" box="[151,275,1034,1059]" pageId="106" pageNumber="107">Diagnosis:</emphasis>
Differing from other North Island
<taxonomicName id="4C121AB3026FB76492F1B66CEE13FB87" authorityName="Blanchard" authorityYear="1853" box="[669,792,1035,1059]" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="106" pageNumber="107" phylum="Arthropoda" rank="genus">
<emphasis id="B966BD22026FB76492F1B66CEE13FB87" box="[669,792,1035,1059]" italics="true" pageId="106" pageNumber="107">Mecodema</emphasis>
</taxonomicName>
species by having:
<emphasis id="B966BD22026FB764939AB66DE90FFB87" bold="true" box="[1014,1028,1034,1059]" pageId="106" pageNumber="107">1</emphasis>
, numerous punctures along vertexal groove and vertex;
<emphasis id="B966BD22026FB764911CB649EC75FBE3" bold="true" box="[368,382,1070,1095]" pageId="106" pageNumber="107">2</emphasis>
, pronotum carina with 5 setae along each side;
<emphasis id="B966BD22026FB76493FFB649EEAAFBE3" bold="true" box="[915,929,1070,1095]" pageId="106" pageNumber="107">3</emphasis>
, elytra with star-shaped punctures, punctures equal size entire;
<emphasis id="B966BD22026FB7649130B635EC61FBCF" bold="true" box="[348,362,1106,1131]" pageId="106" pageNumber="107">4</emphasis>
, distinctive shape of the apical portion of the penis lobe (
<figureCitation id="13297DB5026FB7649380B635E936FBCF" box="[1004,1085,1106,1131]" captionStart="FIGURE 53" captionStartId="107.[151,250,1855,1877]" captionTargetBox="[249,1335,199,1825]" captionTargetId="figure@107.[243,1343,193,1834]" captionTargetPageId="107" captionText="FIGURE 53. Mecodema simplex plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668171/files/figure.png" pageId="106" pageNumber="107">Fig. 53</figureCitation>
PL).
</paragraph>
</subSubSection>
<subSubSection id="C30832BB026FB76290ABB611ECA3FBFF" lastPageId="108" lastPageNumber="109" pageId="106" pageNumber="107" type="description">
<paragraph id="8BAD6130026FB76490ABB611E946FB17" blockId="106.[151,1437,1034,2034]" pageId="106" pageNumber="107">
<emphasis id="B966BD22026FB76490ABB611EC50FB2B" bold="true" box="[199,347,1142,1167]" pageId="106" pageNumber="107">Description:</emphasis>
Length
<quantity id="4CEACCD5026FB76491D4B611EF43FB2A" box="[440,584,1142,1167]" metricMagnitude="-2" metricUnit="m" metricValue="2.63" metricValueMax="3.0" metricValueMin="2.26" pageId="106" pageNumber="107" unit="mm" value="26.3" valueMax="30.0" valueMin="22.6">22.630 mm</quantity>
, pronotal width
<quantity id="4CEACCD5026FB7649293B611EE7FFB2A" box="[767,884,1142,1167]" metricMagnitude="-3" metricUnit="m" metricValue="7.2" metricValueMax="8.0" metricValueMin="6.4" pageId="106" pageNumber="107" unit="mm" value="7.2" valueMax="8.0" valueMin="6.4">6.48 mm</quantity>
, elytral width
<quantity id="4CEACCD5026FB7649479B610E994FB2A" box="[1045,1183,1143,1167]" metricMagnitude="-3" metricUnit="m" metricValue="8.6" metricValueMax="9.9" metricValueMin="7.3" pageId="106" pageNumber="107" unit="mm" value="8.6" valueMax="9.9" valueMin="7.3">7.39.9 mm</quantity>
. Colour of entire body matte to glossy black, except the elytra, ventral surface and legs dark reddish-brown.
</paragraph>
<paragraph id="8BAD6130026FB76490ABB6D9EEFBF99B" blockId="106.[151,1437,1034,2034]" pageId="106" pageNumber="107">
<emphasis id="B966BD22026FB76490ABB6D9EC06FB72" box="[199,269,1214,1238]" italics="true" pageId="106" pageNumber="107">Head:</emphasis>
Broad and convex (
<figureCitation id="13297DB5026FB7649199B6D9EF57FB73" box="[501,604,1214,1239]" captionStart="FIGURE 14" captionStartId="20.[151,250,936,958]" captionTargetBox="[171,1412,204,911]" captionTargetId="figure@20.[160,1412,193,921]" captionTargetPageId="20" captionText="FIGURE 14. Diagrammatic cross sections that represent the width and convexity of head or deflection of pronotum (frontal view): (A) broad and flat; (B) broad and convex; (C) narrow and flat; (D) narrow and convex." httpUri="https://zenodo.org/record/2668091/files/figure.png" pageId="106" pageNumber="107">Fig. 14B</figureCitation>
). Vertex with numerous punctures and rugose wrinkles laterally; vertexal groove broad and shallow, defined by punctures the entire length (
<figureCitation id="13297DB5026FB76493CCB685EEFCFB5F" box="[928,1015,1250,1275]" captionStart="FIGURE 53" captionStartId="107.[151,250,1855,1877]" captionTargetBox="[249,1335,199,1825]" captionTargetId="figure@107.[243,1343,193,1834]" captionTargetPageId="107" captionText="FIGURE 53. Mecodema simplex plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668171/files/figure.png" pageId="106" pageNumber="107">Fig. 53</figureCitation>
); small supraorbital puncture within larger groove bearing 34 setae; 45 well-defined supraorbital grooves extended onto frons; frons medially convex with a broad and deep depression each side, transverse grooves present; frontoclypeal suture and tentorial pits indistinct; anterior area of clypeus (
<figureCitation id="13297DB5026FB7649258B729EF71FAC3" box="[564,634,1358,1383]" captionStart="FIGURE 9" captionStartId="15.[151,250,1706,1728]" captionTargetBox="[157,1433,759,1683]" captionTargetId="figure@15.[151,1436,757,1683]" captionTargetPageId="15" captionText="FIGURE 9. Dorsal view of head with detail of the morphological structures used in the species descriptions. Ƒ = vertex; SPS = supraorbital puncture setae; SP = supraorbital puncture; F = frons area; FCS = frontoclypeal suture; C = clypeus; M = mandible; LS = labrum setae; L = labrum; ACM = anterior clypeal microsculpture; CSP = clypeal setose puncture; TP = tentorial pit; SG = supraorbital grooves; E = eye; ƑG = vertexal groove." httpUri="https://zenodo.org/record/2668081/files/figure.png" pageId="106" pageNumber="107">Fig. 9</figureCitation>
) with broad and shallow grooves laterally, medial grooves narrow; 1 setose puncture each side bearing 12 setae (
<figureCitation id="13297DB5026FB76492FEB715EFEDFA2F" box="[658,742,1394,1419]" captionStart="FIGURE 53" captionStartId="107.[151,250,1855,1877]" captionTargetBox="[249,1335,199,1825]" captionTargetId="figure@107.[243,1343,193,1834]" captionTargetPageId="107" captionText="FIGURE 53. Mecodema simplex plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668171/files/figure.png" pageId="106" pageNumber="107">Fig. 53</figureCitation>
). Labrum lobate, anterior edge outwardly curved, moderately emarginated with 2 proximate central setae, 2 setae each side evenly spaced. Mentum lobes rounded (
<figureCitation id="13297DB5026FB7649546B7F1E884FA0B" box="[1322,1423,1430,1455]" captionStart="FIGURE 12" captionStartId="18.[151,250,1905,1927]" captionTargetBox="[311,1244,697,1878]" captionTargetId="figure@18.[310,1250,689,1889]" captionTargetPageId="18" captionText="FIGURE 12. Ventral view of the mentum with a number of different character states. Illustrations AD are the different forms of the indentation in the apex of the median process: (A) distinctly indentate (M. manaia); (B) moderately indentate (M. pluto); (C) slightly indentate (M. infimate); (D) indentation notched (M. parataiko). Illustrations EG are the three general forms (i.e., hashed lines) of the mentum lobes: (E) triangular; (F) rounded; (G) squared." httpUri="https://zenodo.org/record/2668087/files/figure.png" pageId="106" pageNumber="107">Fig. 12F</figureCitation>
), median process broad and short, moderately angled upward (45°), moderately indentate (
<figureCitation id="13297DB5026FB76494E8B7DDE9E0FA77" box="[1156,1259,1466,1491]" captionStart="FIGURE 12" captionStartId="18.[151,250,1905,1927]" captionTargetBox="[311,1244,697,1878]" captionTargetId="figure@18.[310,1250,689,1889]" captionTargetPageId="18" captionText="FIGURE 12. Ventral view of the mentum with a number of different character states. Illustrations AD are the different forms of the indentation in the apex of the median process: (A) distinctly indentate (M. manaia); (B) moderately indentate (M. pluto); (C) slightly indentate (M. infimate); (D) indentation notched (M. parataiko). Illustrations EG are the three general forms (i.e., hashed lines) of the mentum lobes: (E) triangular; (F) rounded; (G) squared." httpUri="https://zenodo.org/record/2668087/files/figure.png" pageId="106" pageNumber="107">Fig. 12B</figureCitation>
); mentum setae absent. Submentum sclerite (
<figureCitation id="13297DB5026FB764918CB7B9EF2FFA53" box="[480,548,1502,1527]" captionStart="FIGURE 8" captionStartId="13.[151,250,1097,1119]" captionTargetBox="[155,1432,309,1072]" captionTargetId="figure@13.[151,1436,303,1075]" captionTargetPageId="13" captionText="FIGURE 8. Detail of the ventral head with the taxonomic structures used in the species descriptions. SBMS = submentum setae; SBM = submentum sclerite; ML = mentum lobe; MP = mentum process (with indentation); MPW = mentum process width; SBS = stipes basal setae; MPL = mentum process length; MPS = mentum process setae; SBMC = submentum sclerite constriction; GP = gula pits; GS = gula suture." httpUri="https://zenodo.org/record/2668079/files/figure.png" pageId="106" pageNumber="107">Fig. 8</figureCitation>
) constriction narrow with 3 setae laterally (with a large gap medially). Stipes with 2 basal setae. Gula pits small, suture well-defined the entire length, gula slightly convex with transverse lines. Gena (
<figureCitation id="13297DB5026FB764908DB441EC28F99B" box="[225,291,1574,1599]" captionStart="FIGURE 8" captionStartId="13.[151,250,1097,1119]" captionTargetBox="[155,1432,309,1072]" captionTargetId="figure@13.[151,1436,303,1075]" captionTargetPageId="13" captionText="FIGURE 8. Detail of the ventral head with the taxonomic structures used in the species descriptions. SBMS = submentum setae; SBM = submentum sclerite; ML = mentum lobe; MP = mentum process (with indentation); MPW = mentum process width; SBS = stipes basal setae; MPL = mentum process length; MPS = mentum process setae; SBMC = submentum sclerite constriction; GP = gula pits; GS = gula suture." httpUri="https://zenodo.org/record/2668079/files/figure.png" pageId="106" pageNumber="107">Fig. 8</figureCitation>
) with rugose wrinkles forming an isodiametric pattern laterally.
</paragraph>
<paragraph id="8BAD6130026FB76490ABB42CEC4FF8FB" blockId="106.[151,1437,1034,2034]" pageId="106" pageNumber="107">
<emphasis id="B966BD22026FB76490ABB42CEC49F9C7" box="[199,322,1611,1635]" italics="true" pageId="106" pageNumber="107">Prothorax:</emphasis>
Prothoracic carina (
<figureCitation id="13297DB5026FB7649244B42DEF72F9C7" box="[552,633,1610,1635]" captionStart="FIGURE 11" captionStartId="17.[151,250,949,971]" captionTargetBox="[221,1358,181,923]" captionTargetId="figure@17.[221,1359,179,929]" captionTargetPageId="17" captionText="FIGURE 11. Detail of pronotum showing the taxonomic structures used in the species descriptions. PPC = posterior pronotum constriction; PS = pronotal setae; PAA = pronotum anterior angle; AE = anterior edge; PC = pronotal carina; ML = midline; PF= pronotal foveae; PPS = pronotum posterior sinuation; PE = posterior edge." httpUri="https://zenodo.org/record/2668085/files/figure.png" pageId="106" pageNumber="107">Fig. 11</figureCitation>
) broad the entire length, moderately crenulated with 5 setae each side, extended beyond anterior angle; posterior lateral sinuation evidently carinate, parallel; pronotum broad and deflected laterally, overall shape cordate (pronotal edge right angled apically); midline well-defined, anterior medial and posterior medial diamond-shaped impressions present, disc with fine transverse lines turning to welldefined wrinkles laterally (
<figureCitation id="13297DB5026FB76491A3B4BDEF2DF957" box="[463,550,1754,1779]" captionStart="FIGURE 53" captionStartId="107.[151,250,1855,1877]" captionTargetBox="[249,1335,199,1825]" captionTargetId="figure@107.[243,1343,193,1834]" captionTargetPageId="107" captionText="FIGURE 53. Mecodema simplex plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668171/files/figure.png" pageId="106" pageNumber="107">Fig. 53</figureCitation>
), may have obsolescent punctures mesad foveae; pronotal foveae broad and deep; anterior edge slightly inwardly curved, posterior edge straight, medially emarginated. Prosternum slightly convex and smooth; proepisternum densely pitted. Procoxal setae absent; protibia distally expanded and shovellike (
<figureCitation id="13297DB5026FB76490BCB521EC3CF8FB" box="[208,311,1862,1887]" captionStart="FIGURE 10" captionStartId="16.[151,250,1442,1464]" captionTargetBox="[152,1428,191,1421]" captionTargetId="figure@16.[151,1435,183,1427]" captionTargetPageId="16" captionText="FIGURE 10. The protibia form is a character used mainly to determine between genera, except for one species in Mecodema where the difference between (A) greatly distally expanded (M. antarctica) and (B) expanded distally (all other Mecodema species) is used." httpUri="https://zenodo.org/record/2668083/files/figure.png" pageId="106" pageNumber="107">Fig. 10B</figureCitation>
).
</paragraph>
<paragraph id="8BAD6130026FB76290ABB50CE9D4FF5F" blockId="106.[151,1437,1034,2034]" lastBlockId="108.[151,1437,151,1730]" lastPageId="108" lastPageNumber="109" pageId="106" pageNumber="107">
<emphasis id="B966BD22026FB76490ABB50CEC1DF827" box="[199,278,1899,1923]" italics="true" pageId="106" pageNumber="107">Elytra:</emphasis>
Broad and deflected laterally; humeral angle anteriorly convergent (
<figureCitation id="13297DB5026FB7649459B50DE998F827" box="[1077,1171,1898,1923]" captionStart="FIGURE 6" captionStartId="10.[151,250,519,541]" captionTargetBox="[156,1432,193,498]" captionTargetId="figure@10.[151,1436,193,498]" captionTargetPageId="10" captionText="FIGURE 6. Detail of the three different forms of the elytral humeral angle: (A) = anteriorly convergent (M. parataiko), (B) = evenly convex (M. oconnori), (C) = subangulate (M. atuanui)." httpUri="https://zenodo.org/record/2668075/files/figure.png" pageId="106" pageNumber="107">Fig. 6A</figureCitation>
); basal margin slightly curved and gently sloped to base, interval 1 extended to scutellum; lateral carina narrow the entire length, broadened apically, extended to humeral angle; humerus with 3 setose punctures closely spaced, 4
<superScript id="7C67CC78026FB764948BB5D5E9F8F864" attach="left" box="[1255,1267,1970,1984]" fontSize="6" pageId="106" pageNumber="107">th</superScript>
double-spaced posterad; suture well-defined; striae with regularly sized and spaced, asetose punctures striae 14 extended beyond basal margin; intervals 3, 5 and 7 moderately convex, interval 9 strongly convex, especially anterior ⅓; interval microsculpture in form of fine lines radiating outwardly from punctures (star-shaped); 7
<superScript id="7C67CC780269B762941FB2DCE974FF6D" attach="left" box="[1139,1151,187,201]" fontSize="6" pageId="108" pageNumber="109">th</superScript>
strial setal pattern with 3 setose punctures in anterior ½, 4 setose punctures in posterior ½ (
<figureCitation id="13297DB50269B7629318B285EECCFF5F" box="[884,967,226,251]" captionStart="FIGURE 53" captionStartId="107.[151,250,1855,1877]" captionTargetBox="[249,1335,199,1825]" captionTargetId="figure@107.[243,1343,193,1834]" captionTargetPageId="107" captionText="FIGURE 53. Mecodema simplex plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668171/files/figure.png" pageId="108" pageNumber="109">Fig. 53</figureCitation>
), setose punctures small.
</paragraph>
<caption id="DF6D31B8026EB76590FBB558E808F836" httpUri="https://zenodo.org/record/2668171/files/figure.png" pageId="107" pageNumber="108" startId="107.[151,250,1855,1877]" targetBox="[249,1335,199,1825]" targetPageId="107">
<paragraph id="8BAD6130026EB76590FBB558E808F836" blockId="107.[151,1436,1855,1938]" pageId="107" pageNumber="108">
<emphasis id="B966BD22026EB76590FBB558EC29F8F1" bold="true" box="[151,290,1855,1877]" pageId="107" pageNumber="108">FIGURE 53.</emphasis>
<taxonomicName id="4C121AB3026EB7659146B558ECE4F8F1" authorityName="Castelnau" authorityYear="1867" box="[298,495,1855,1877]" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="107" pageNumber="108" phylum="Arthropoda" rank="species" species="simplex">
<emphasis id="B966BD22026EB7659146B558ECE4F8F1" box="[298,495,1855,1877]" italics="true" pageId="107" pageNumber="108">Mecodema simplex</emphasis>
</taxonomicName>
plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe.
</paragraph>
</caption>
<paragraph id="8BAD61300269B76290ABB360EC5EFE53" blockId="108.[151,1437,151,1730]" pageId="108" pageNumber="109">
<emphasis id="B966BD220269B76290ABB360EC77FEBB" box="[199,380,263,287]" italics="true" pageId="108" pageNumber="109">Ventral surface:</emphasis>
Mesepisternum and metepisternum (
<figureCitation id="13297DB50269B7629349B361EE61FEBB" box="[805,874,262,287]" captionStart="FIGURE 2" captionStartId="6.[151,250,800,822]" captionTargetBox="[159,1432,197,773]" captionTargetId="figure@6.[151,1436,193,779]" captionTargetPageId="6" captionText="FIGURE 2. Ventral view of a Mecodema specimen showing specific morphological structures, excluding taxonomic structures indicated in detail figures, used in the species descriptions. ƑLF = ventrite lateral foveae; ƑSP = ventrite setose punctures; MTC = metacoxa; MTƑP = metaventrite process; MSC = mesocoxa; PC = procoxa; PS = prosternum; G = gena; PES = proepisternum; MSE = mesepisternum; MTE = metepisternum; Ƒ1Ƒ6 = ventrites 1-6 (ventrites 1-3 may be fused); M = midline (dashed line, not a taxonomic structure)." httpUri="https://zenodo.org/record/2668067/files/figure.png" pageId="108" pageNumber="109">Fig. 2</figureCitation>
) with numerous punctures across entire area and spilling onto latero-posterior edges of mesosternum; setose punctures present on mesocoxa (2) and metacoxa (2). All abdominal ventrites lineate, ventrite 1 sparsely micropunctate; ventrites 35 with 1 setose puncture each side of midline; ventrite 6 setae present:
<specimenCount id="9D14AAB90269B7629266B314EF14FE2E" box="[522,543,371,394]" pageId="108" pageNumber="109" type="male"></specimenCount>
with a single setose puncture each side near junction of straight apical edge and curve anterad;
<specimenCount id="9D14AAB90269B762912DB3FFEC5DFE0B" box="[321,342,408,431]" pageId="108" pageNumber="109" type="female"></specimenCount>
with 2 setose punctures each side, apical edge straight between first pair; ventrites 36 foveate laterally. Anterior metaventrite process (
<figureCitation id="13297DB50269B7629237B3DDEF94FE77" box="[603,671,442,467]" captionStart="FIGURE 2" captionStartId="6.[151,250,800,822]" captionTargetBox="[159,1432,197,773]" captionTargetId="figure@6.[151,1436,193,779]" captionTargetPageId="6" captionText="FIGURE 2. Ventral view of a Mecodema specimen showing specific morphological structures, excluding taxonomic structures indicated in detail figures, used in the species descriptions. ƑLF = ventrite lateral foveae; ƑSP = ventrite setose punctures; MTC = metacoxa; MTƑP = metaventrite process; MSC = mesocoxa; PC = procoxa; PS = prosternum; G = gena; PES = proepisternum; MSE = mesepisternum; MTE = metepisternum; Ƒ1Ƒ6 = ventrites 1-6 (ventrites 1-3 may be fused); M = midline (dashed line, not a taxonomic structure)." httpUri="https://zenodo.org/record/2668067/files/figure.png" pageId="108" pageNumber="109">Fig. 2</figureCitation>
) a short and sharp triangle with a very broad and pronounced carina the entire length.
</paragraph>
<paragraph id="8BAD61300269B76290ABB064E880FC4B" blockId="108.[151,1437,151,1730]" pageId="108" pageNumber="109">
<emphasis id="B966BD220269B76290ABB064EC70FDBF" box="[199,379,515,539]" italics="true" pageId="108" pageNumber="109">Male genitalia:</emphasis>
Apical portion of penis lobe asymmetrically hooked (almost symmetrical) with a slight deflection to right of vertical axis (VV) (
<figureCitation id="13297DB50269B7629206B041EFD9FD9B" box="[618,722,550,575]" captionStart="FIGURE 15" captionStartId="20.[151,250,1943,1965]" captionTargetBox="[162,1424,1039,1914]" captionTargetId="figure@20.[162,1424,1038,1921]" captionTargetPageId="20" captionText="FIGURE 15. The penis lobe in ventral view showing the relative deflection to right of vertical: (A) distinctly deflected (M. regulus); (B) moderately deflected (M. haunoho); (C) slightly deflected (M. oregoides). Scale bars = 1 mm. Structures of penis lobe indicated are: base (B); lateral edge (LE); apical portion (AP); ventral edge (ƑE)." httpUri="https://zenodo.org/record/2668093/files/figure.png" pageId="108" pageNumber="109">Fig. 15C</figureCitation>
); ventral process curved from shaft to form a rounded point at centre of vertical axis, apex is broadly and symmetrically curved upwards to form dorsal process (
<figureCitation id="13297DB50269B7629484B02DE833FDC7" box="[1256,1336,586,611]" captionStart="FIGURE 53" captionStartId="107.[151,250,1855,1877]" captionTargetBox="[249,1335,199,1825]" captionTargetId="figure@107.[243,1343,193,1834]" captionTargetPageId="107" captionText="FIGURE 53. Mecodema simplex plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668171/files/figure.png" pageId="108" pageNumber="109">Fig. 53</figureCitation>
PL) with a more rounded point (behind ventral point) that recurves to shaft, both relatively the same depth (LV); shaft of penis lobe narrow but gradually broadened toward base (
<figureCitation id="13297DB50269B762937CB0F5EE6BFD0F" box="[784,864,658,683]" captionStart="FIGURE 53" captionStartId="107.[151,250,1855,1877]" captionTargetBox="[249,1335,199,1825]" captionTargetId="figure@107.[243,1343,193,1834]" captionTargetPageId="107" captionText="FIGURE 53. Mecodema simplex plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668171/files/figure.png" pageId="108" pageNumber="109">Fig. 53</figureCitation>
PL); ventral edge of penis lobe shaft with moderate curve to left, basal to apical portion curve to right (VV); overall length of penis lobe with dictinct curve ventrally (LV) (
<figureCitation id="13297DB50269B76290B3B0BDEC42FD57" box="[223,329,730,755]" captionStart="FIGURE 16" captionStartId="21.[151,250,897,919]" captionTargetBox="[151,1436,181,869]" captionTargetId="figure@21.[151,1436,181,876]" captionTargetPageId="21" captionText="FIGURE 16. Relative strength of the ventral curve over the length of the penis lobe (lateral view): (A) slightly curved (M. oblongum); (B) moderately curved (M. ponaiti); (C) distinctly curved (M. manaia). Scale bar = 1 mm." httpUri="https://zenodo.org/record/2668095/files/figure.png" pageId="108" pageNumber="109">Fig. 16C</figureCitation>
). Structures of the endophallus (
<figureCitation id="13297DB50269B76292AFB0BDEE07FD57" box="[707,780,730,755]" captionStart="FIGURE 4" captionStartId="7.[151,250,973,995]" captionTargetBox="[354,1232,193,952]" captionTargetId="figure@7.[354,1232,193,952]" captionTargetPageId="7" captionText="FIGURE 4. An Environmental Scanning Electron Microscope (ESEM) photograph of the four main structures of the apical plate of the endophallus within the penis lobe. This is an example from Mecodema haunoho showing the central spicule (c.s), left (l.s.f) and the right setose flange (r.s.f), and microtrichial field (m.f). These structures are variable among species but within a species, they are very consistent, even over distant geographical ranges. The microtrichial field can be covered in flattened scales as in photo above, or with elongated setae-like scales." httpUri="https://zenodo.org/record/2668071/files/figure.png" pageId="108" pageNumber="109">Fig. 4</figureCitation>
): lateral form of the apex of the central spicule pointed; dorsal form of the apex of the central spicule greatly expanded without projection; setal (scales) coverage of the apical plate extensive (76100%); size of left setose flange small; size of right setose flange large. Left paramere articulated at right angle from penis lobe, basal lobe rectangular with large dorsal hump (almost making lobe triangular) that forms steeply (60°) curved slope to arm (
<figureCitation id="13297DB50269B762937DB10DEE69FC27" box="[785,866,874,899]" captionStart="FIGURE 53" captionStartId="107.[151,250,1855,1877]" captionTargetBox="[249,1335,199,1825]" captionTargetId="figure@107.[243,1343,193,1834]" captionTargetPageId="107" captionText="FIGURE 53. Mecodema simplex plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668171/files/figure.png" pageId="108" pageNumber="109">Fig. 53</figureCitation>
LP); arm short and narrow; terminal lobe short and broad with apical tuft of long setae, short setae extended along edge, longer comb prior to arm junction; setae extended along apical ½ of straight ventral edge (
<figureCitation id="13297DB50269B76292D7B1D5EE00FC6F" box="[699,779,946,971]" captionStart="FIGURE 53" captionStartId="107.[151,250,1855,1877]" captionTargetBox="[249,1335,199,1825]" captionTargetId="figure@107.[243,1343,193,1834]" captionTargetPageId="107" captionText="FIGURE 53. Mecodema simplex plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668171/files/figure.png" pageId="108" pageNumber="109">Fig. 53</figureCitation>
LP). Right paramere narrowly triangular with a double row of long to medium-lengthed setae extended along apical ¾ of ventral edge (
<figureCitation id="13297DB50269B7629389B1B1E93EFC4B" box="[997,1077,982,1007]" captionStart="FIGURE 53" captionStartId="107.[151,250,1855,1877]" captionTargetBox="[249,1335,199,1825]" captionTargetId="figure@107.[243,1343,193,1834]" captionTargetPageId="107" captionText="FIGURE 53. Mecodema simplex plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668171/files/figure.png" pageId="108" pageNumber="109">Fig. 53</figureCitation>
RP), curved inwardly apically.
</paragraph>
<paragraph id="8BAD61300269B76290ABB19CECA3FBFF" blockId="108.[151,1437,151,1730]" pageId="108" pageNumber="109">
<emphasis id="B966BD220269B76290ABB19CEC9AFBB7" box="[199,401,1019,1043]" italics="true" pageId="108" pageNumber="109">Female genitalia:</emphasis>
Basal gonocoxite 1 short and narrow, apically broadened, ventral surface smooth, internal dorso-lateral carina (
<figureCitation id="13297DB50269B76291ECB679ECC8FB93" box="[384,451,1054,1079]" captionStart="FIGURE 5" captionStartId="9.[151,250,1714,1736]" captionTargetBox="[167,1436,790,1674]" captionTargetId="figure@9.[151,1436,790,1693]" captionTargetPageId="9" captionText="FIGURE 5. The entire female reproductive tract (ventral view) including taxonomic structures (for all other abbreviated structure names see Seldon et al. 2012). However, for this revision only the following structures are important: HS = helminthoid sclerite; G1 = gonocoxite 1; G2 = gonocoxite 2; TDC = transverse dorsal carina; R = ramus." httpUri="https://zenodo.org/record/2668073/files/figure.png" pageId="108" pageNumber="109">Fig. 5</figureCitation>
) with 2 setae. Gonocoxite 2 bluntly rounded triangle, slightly reflexed apically. Ramus (
<figureCitation id="13297DB50269B76290F3B625EDE9FBFF" box="[159,226,1090,1115]" captionStart="FIGURE 5" captionStartId="9.[151,250,1714,1736]" captionTargetBox="[167,1436,790,1674]" captionTargetId="figure@9.[151,1436,790,1693]" captionTargetPageId="9" captionText="FIGURE 5. The entire female reproductive tract (ventral view) including taxonomic structures (for all other abbreviated structure names see Seldon et al. 2012). However, for this revision only the following structures are important: HS = helminthoid sclerite; G1 = gonocoxite 1; G2 = gonocoxite 2; TDC = transverse dorsal carina; R = ramus." httpUri="https://zenodo.org/record/2668073/files/figure.png" pageId="108" pageNumber="109">Fig. 5</figureCitation>
) short and broad.
</paragraph>
</subSubSection>
<subSubSection id="C30832BB0269B76290ABB601E943FB6E" pageId="108" pageNumber="109" type="discussion">
<paragraph id="8BAD61300269B76290ABB601E943FB6E" blockId="108.[151,1437,151,1730]" pageId="108" pageNumber="109">
<emphasis id="B966BD220269B76290ABB601EC59FBDB" bold="true" box="[199,338,1126,1151]" pageId="108" pageNumber="109">Comments:</emphasis>
The
<typeStatus id="54A9DF920269B76291E6B601ECE7FBDB" box="[394,492,1126,1151]" pageId="108" pageNumber="109" type="holotype">holotype</typeStatus>
locality is
<collectingRegion id="49D6AFD20269B762920AB601EFDDFBDB" box="[614,726,1126,1151]" country="New Zealand" name="Auckland" pageId="108" pageNumber="109">Auckland</collectingRegion>
, but we suggest that this refers to the
<collectingRegion id="49D6AFD20269B7629414B601E9EEFBDB" box="[1144,1253,1126,1151]" country="New Zealand" name="Auckland" pageId="108" pageNumber="109">Auckland</collectingRegion>
Province that in the 1860s extended from Cape Reinga to the 39
<superScript id="7C67CC780269B76292A9B6EDEFDAFB3C" attach="left" box="[709,721,1162,1176]" fontSize="6" pageId="108" pageNumber="109">th</superScript>
latitude. This southern boundary would be the most northern range of
<taxonomicName id="4C121AB30269B7629096B6D6ECDAFB6D" authorityName="Castelnau" authorityYear="1867" box="[250,465,1201,1225]" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="108" pageNumber="109" phylum="Arthropoda" rank="species" species="simplex">
<emphasis id="B966BD220269B7629096B6D6ECDAFB6D" box="[250,465,1201,1225]" italics="true" pageId="108" pageNumber="109">Mecodema simplex</emphasis>
</taxonomicName>
, which is much more prevalent in the Wairarapa region.
</paragraph>
</subSubSection>
<subSubSection id="C30832BB0269B76290ABB6B2EC11FAB6" pageId="108" pageNumber="109" type="distribution">
<paragraph id="8BAD61300269B76290ABB6B2EC11FAB6" blockId="108.[151,1437,151,1730]" pageId="108" pageNumber="109">
<emphasis id="B966BD220269B76290ABB6B2EC69FB4A" bold="true" box="[199,354,1237,1262]" pageId="108" pageNumber="109">Distribution:</emphasis>
<collectingCountry id="F30521A00269B7629103B6B1EF05FB4A" box="[367,526,1237,1262]" name="New Zealand" pageId="108" pageNumber="109">New Zealand</collectingCountry>
, North Island, eastern Tararua Ranges, Hawkes Bay, Rangitikei, Wairarapa, Wellington.
</paragraph>
</subSubSection>
<subSubSection id="C30832BB0269B76290ABB77AEE4BF966" pageId="108" pageNumber="109" type="materials_examined">
<paragraph id="8BAD61300269B76290ABB77AECBCFADA" blockId="108.[151,1437,151,1730]" pageId="108" pageNumber="109">
<materialsCitation id="3B7A6B6D0269B76290ABB77AEF8CFAFE" ID-GBIF-Occurrence="2237851803" collectionCode="MCNG" collectorName="Coll. Castelnau" location="Auckland" pageId="108" pageNumber="109" specimenCount="1" stateProvince="Auckland" typeStatus="holotype">
<emphasis id="B966BD220269B76290ABB77AEC36FA92" bold="true" box="[199,317,1309,1334]" pageId="108" pageNumber="109">
<typeStatus id="54A9DF920269B76290ABB77AEC33FA92" box="[199,312,1309,1334]" pageId="108" pageNumber="109" type="holotype">Holotype</typeStatus>
:
</emphasis>
<collectionCode id="ED03F9F50269B7629124B779EC97FA92" box="[328,412,1310,1334]" country="Venezuela" httpUri="http://grbio.org/cool/tyz9-grz8" name="Museo de Ciencias Naturales de la UNELLEZ en Guanare" pageId="108" pageNumber="109">MCNG</collectionCode>
specimen labelled.
<collectingRegion id="49D6AFD20269B76292E4B77AEFFEFA92" box="[648,757,1309,1334]" country="New Zealand" name="Auckland" pageId="108" pageNumber="109">Auckland</collectingRegion>
<collectorName id="26E704E60269B762936CB77AEEBBFA92" box="[768,944,1309,1334]" pageId="108" pageNumber="109">Coll. Castelnau</collectorName>
[hw, black border] / Typus [red font, red border] /
<taxonomicName id="4C121AB30269B762916CB726EC91FAFE" authority="Cast." authorityName="Castelnau" authorityYear="1867" box="[256,410,1345,1370]" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="108" pageNumber="109" phylum="Arthropoda" rank="species" species="simplex">simplex Cast.</taxonomicName>
[hw] /
<taxonomicName id="4C121AB30269B7629180B726EF8CFAFE" authority="Cast." authorityName="Castelnau" authorityYear="1867" box="[492,647,1345,1370]" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="108" pageNumber="109" phylum="Arthropoda" rank="species" species="simplex">Simplex Cast</taxonomicName>
</materialsCitation>
. /
<materialsCitation id="3B7A6B6D0269B76292CDB725ECB8FADA" ID-GBIF-Occurrence="2237851814" location="Museo Civico di Genova" pageId="108" pageNumber="109" specimenCount="1" stateProvince="Auckland" typeStatus="holotype">
<typeStatus id="54A9DF920269B76292CDB725EE4CFAFE" box="[673,839,1346,1370]" pageId="108" pageNumber="109" type="holotype">HOLOTYPUS</typeStatus>
<taxonomicName id="4C121AB30269B7629323B726E9E9FAFE" authority="Castelnau 1867" authorityName="Castelnau" authorityYear="1867" box="[847,1250,1345,1370]" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="108" pageNumber="109" phylum="Arthropoda" rank="species" species="simplex">
Mecodema simplex
<bibRefCitation id="EF831CC10269B7629458B726E9E9FAFE" author="Castelnau, F." box="[1076,1250,1345,1370]" pageId="108" pageNumber="109" pagination="30 - 38" refId="ref91697" refString="Castelnau, F. Comte de (1867) Notes on Australian Coleoptera. Transactions and Proceedings of the Royal Society of Fictoria, 8, 30 - 38; 95 - 225." type="journal article" year="1867">Castelnau 1867</bibRefCitation>
</taxonomicName>
[hw, red label] /
<location id="8ECD37EB0269B76290FBB701ECB8FADA" LSID="urn:lsid:plazi:treatment:03BBD026026FB76290FBB178EE4BF966:8ECD37EB0269B76290FBB701ECB8FADA" box="[151,435,1381,1406]" name="Museo Civico di Genova" pageId="108" pageNumber="109" stateProvince="Auckland">Museo Civico di Genova</location>
</materialsCitation>
.
</paragraph>
<paragraph id="8BAD61300269B76290ABB7EEE862F9F2" blockId="108.[151,1437,151,1730]" pageId="108" pageNumber="109">
<emphasis id="B966BD220269B76290ABB7EEEC34FA06" bold="true" box="[199,319,1417,1442]" pageId="108" pageNumber="109">
<typeStatus id="54A9DF920269B76290ABB7EEEC34FA06" box="[199,319,1417,1442]" pageId="108" pageNumber="109" type="paratype">Paratypes</typeStatus>
</emphasis>
(
<taxonomicName id="4C121AB30269B7629122B7EEEF2DFA05" authorityName="Castelnau" authorityYear="1867" box="[334,550,1417,1441]" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="108" pageNumber="109" phylum="Arthropoda" rank="species" species="simplex">
<emphasis id="B966BD220269B7629122B7EEEF2DFA05" box="[334,550,1417,1441]" italics="true" pageId="108" pageNumber="109">Mecodema simplex</emphasis>
</taxonomicName>
): BMNH card-mounted male labelled. Type [round label with red border glued to card] / 2602
<specimenCount id="9D14AAB90269B7629151B7CAEC5BFA62" box="[317,336,1453,1478]" pageId="108" pageNumber="109" type="male"></specimenCount>
[hw] / New Zeal. [red underline] Broun Coll. Brit. Mus. 1922-482 / Mt. Holdsworth Tararua [hw] /
<taxonomicName id="4C121AB30269B76290CBB7B6ECACFA4E" authorityName="Broun" authorityYear="1908" box="[167,423,1489,1514]" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="108" pageNumber="109" phylum="Arthropoda" rank="species" species="acuductum">Mecodema acuductum</taxonomicName>
[hw]. BMNH male labelled. Type [round label with red border] / 3115 [hw] / New Zeal. [red underline] Broun Coll. Brit. Mus. 1922-482 / Silverstream
<collectingRegion id="49D6AFD20269B7629301B792EEE2F9AA" box="[877,1001,1525,1550]" country="New Zealand" name="Wellington" pageId="108" pageNumber="109">Wellington</collectingRegion>
[hw] /
<taxonomicName id="4C121AB30269B7629453B792E83AF9AA" authorityName="Broun" authorityYear="1912" box="[1087,1329,1525,1550]" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="108" pageNumber="109" phylum="Arthropoda" rank="species" species="bryobium">Mecodema bryobium</taxonomicName>
<specimenCount id="9D14AAB90269B7629556B791E844F9A9" box="[1338,1359,1526,1549]" pageId="108" pageNumber="109" type="male"></specimenCount>
[hw] / card-mounted male genitalia; BMNH female labelled. Type [round label with red border] / 3168 [hw] / New Zeal. [red underline] Broun Coll. Brit. Mus. 1922-482 / Martinboro
<collectingRegion id="49D6AFD20269B762933DB45AEEC6F9F2" box="[849,973,1597,1622]" country="New Zealand" name="Wellington" pageId="108" pageNumber="109">Wellington</collectingRegion>
[hw] /
<taxonomicName id="4C121AB30269B7629472B45AE801F9F1" authorityName="Broun" authorityYear="1912" box="[1054,1290,1597,1622]" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="108" pageNumber="109" phylum="Arthropoda" rank="species" species="arcuatum">Mecodema arcuatum</taxonomicName>
<specimenCount id="9D14AAB90269B762957DB459E82DF9F1" box="[1297,1318,1598,1621]" pageId="108" pageNumber="109" type="female"></specimenCount>
[hw].
</paragraph>
<paragraph id="8BAD61300269B76290ABB406EE4BF966" blockId="108.[151,1437,151,1730]" pageId="108" pageNumber="109">
<emphasis id="B966BD220269B76290ABB406ECB9F9DE" bold="true" box="[199,434,1633,1658]" pageId="108" pageNumber="109">Material examined:</emphasis>
<materialsCitation id="3B7A6B6D0269B76291D4B405EF84F93A" ID-GBIF-Occurrence="2237851682" collectingDate="1992-02-23" collectionCode="LOVEI, COLLECTION" collectorName="Bellance Reserve, RI &amp; G. Lovei" location="Manawatu Gorge" municipality="Bellance Reserve" pageId="108" pageNumber="109" specimenCode="AMNZ 503340" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D14AAB90269B76291D4B405ECD6F9DE" box="[440,477,1634,1658]" pageId="108" pageNumber="109" type="male">1♂</specimenCount>
,
<location id="8ECD37EB0269B7629185B405EFA5F9DE" LSID="urn:lsid:plazi:treatment:03BBD026026FB76290FBB178EE4BF966:8ECD37EB0269B7629185B405EFA5F9DE" box="[489,686,1634,1658]" municipality="Bellance Reserve" name="Manawatu Gorge" pageId="108" pageNumber="109">Manawatu Gorge</location>
,
<collectorName id="26E704E60269B76292D4B406EEAFF9DE" box="[696,932,1633,1658]" pageId="108" pageNumber="109">
<collectingMunicipality id="6BC9FB4A0269B76292D4B406EE75F9DD" box="[696,894,1633,1658]" pageId="108" pageNumber="109">Bellance Reserve</collectingMunicipality>
, RI
</collectorName>
/WN, forest pitfall 3/2,
<date id="FFAC47F00269B76294C5B405E9F4F9DE" box="[1193,1279,1634,1658]" pageId="108" pageNumber="109" value="1992-02-23">
<collectingDate id="EFE8BE180269B76294C5B405E9F4F9DE" box="[1193,1279,1634,1658]" pageId="108" pageNumber="109" value="1992-02-23">23/2/92</collectingDate>
</date>
coll.
<collectorName id="26E704E60269B7629550B405E897F9DE" box="[1340,1436,1633,1658]" pageId="108" pageNumber="109">G. Lovei</collectorName>
/ G.
<collectionCode id="ED03F9F50269B76290AFB4E1ECC5F93A" box="[195,462,1670,1694]" pageId="108" pageNumber="109">LOVEI COLLECTION</collectionCode>
/
<specimenCode id="DBB4C94B0269B762918EB4E1EF84F93A" box="[482,655,1669,1694]" collectionCode="AMNZ" country="New Zealand" httpUri="http://grbio.org/cool/76et-m1kt" name="Auckland Institute and Museum" pageId="108" pageNumber="109" type="Museum">AMNZ 503340</specimenCode>
</materialsCitation>
[green label];
<materialsCitation id="3B7A6B6D0269B762935EB4E1EE37F966" ID-GBIF-Occurrence="2237851821" collectingDate="1976-04-11" collectionCode="JNPC" collectorName="WA Puketoi SF &amp; J. Nunn" country="New Zealand" determinerName="J. Nunn" location="NEW ZEALAND" pageId="108" pageNumber="109" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D14AAB90269B762935EB4E1EE5EF93A" box="[818,853,1670,1694]" pageId="108" pageNumber="109" type="female">1♀</specimenCount>
,
<collectingCountry id="F30521A00269B762930EB4E1E927F93A" box="[866,1068,1670,1694]" name="New Zealand" pageId="108" pageNumber="109">NEW ZEALAND</collectingCountry>
<collectorName id="26E704E60269B762945FB4E1E836F939" box="[1075,1341,1669,1694]" pageId="108" pageNumber="109">WA Puketoi SF J. Nunn</collectorName>
2411
<date id="FFAC47F00269B76295EDB4E1EDEEF966" pageId="108" pageNumber="109" value="1976-04-11">
<collectingDate id="EFE8BE180269B76295EDB4E1EDEEF966" pageId="108" pageNumber="109" value="1976-04-11">11 Apr 76</collectingDate>
</date>
[hw] /
<taxonomicName id="4C121AB30269B762915BB4CEEF18F966" authorityName="Castelnau" authorityYear="1867" box="[311,531,1705,1730]" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="108" pageNumber="109" phylum="Arthropoda" rank="species" species="simplex">Mecodema simplex</taxonomicName>
det.
<determinerName id="7E718DF40269B7629226B4CDEFAAF965" box="[586,673,1706,1729]" pageId="108" pageNumber="109">J. Nunn</determinerName>
[hw] (
<collectionCode id="ED03F9F50269B7629280B4CDEE3FF966" box="[748,820,1706,1730]" pageId="108" pageNumber="109">JNPC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>