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<document id="385D8860C9D9472315E7750CEF155CBC" ID-CLB-Dataset="9529" ID-DOI="10.11646/zootaxa.4881.3.7" ID-GBIF-Dataset="b7622904-4534-4fc0-aebd-d4766d680eb9" ID-ISSN="1175-5326" ID-Zenodo-Dep="4283960" ID-ZooBank="003285F9-AD4C-4B33-BCFD-E15821BF26C6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1605981939210" checkinUser="plazi" docAuthor="Tian, Mingyi &amp; He, Li" docDate="2020" docId="03B5EB57AE19FFD299C47F5404321E8E" docLanguage="en" docName="zootaxa.4881.3.7.pdf" docOrigin="Zootaxa 4881 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Jujiroa zhouchaoi Tian &amp; He 2020, sp. nov." docType="treatment" docVersion="7" lastPageNumber="550" masterDocId="FF8C932FAE18FFD799537B4F01291F5F" masterDocTitle="New species and new record of cavernicolous ground beetles from Sichuan Province, China (Coleoptera: Carabidae: Pterostichini and Platynini)" masterLastPageNumber="558" masterPageNumber="545" pageNumber="546" updateTime="1698846498561" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="8E8B8BC1154BF0718ABEB984BE8FD919" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="13A509A5197A8781467A683A377C9E0A">
<mods:title id="57636CF5F17D0DEC7E9AD7D7104AB6B5">New species and new record of cavernicolous ground beetles from Sichuan Province, China (Coleoptera: Carabidae: Pterostichini and Platynini)</mods:title>
</mods:titleInfo>
<mods:name id="66575F957B0599303E37D668CEAB8CA9" type="personal">
<mods:role id="F14943D9BF33DCF3492B4B263B21DCC9">
<mods:roleTerm id="AAD05C234DCBE31BE2F3B290D12D377B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="70F9E3B3DE9C81AB61BFA61EA38F870C">Tian, Mingyi</mods:namePart>
<mods:affiliation id="080BF9EA6E2A8669595BE2D1E161C38D">Department of Entomology, College of Plant Protection, South China Agricultural University, 483 Wushan Road, Guangzhou, 510642,</mods:affiliation>
</mods:name>
<mods:name id="02A96777D412C09F34D18C5745F6EF7E" type="personal">
<mods:role id="F2E8F344829D098C48FEBC9542C5B517">
<mods:roleTerm id="2714330EB8772C55C56DC9CA9FD4800A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="043981FFEE9F168A6C420F98A45ABC00">He, Li</mods:namePart>
<mods:affiliation id="87B90DBEB5DE0A92A4FC71AB9696F433">Sichuan Cave Exploration Team, No. 66, 5 Shuangcheng Road, Chenghua District, Chengdu, 610051, China https: // orcid. org / 0000 - 0002 - 4597 - 3442</mods:affiliation>
</mods:name>
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<mods:titleInfo id="0E1D26D56E7D2CB3710B363E6D7D8FBB">
<mods:title id="735B14B4646B2446F2A9E676F7D9EF48">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="B7F3B82B9A00AE3700B614A34122ECB6">
<mods:date id="BC3021F3527BBC1CE0460F01F4094839">2020</mods:date>
<mods:detail id="D7B601440AF8BDBB23ED10129B74C109" type="pubDate">
<mods:number id="57D375077309602C38AD5AA9BD1D0C36">2020-11-20</mods:number>
</mods:detail>
<mods:detail id="36B76948AB50D5F2634AD072DE7743EB" type="volume">
<mods:number id="55586806F1E87BFA0A691829FF44CE04">4881</mods:number>
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<mods:detail id="3112E28945B997828D1E2AD74AA1E538" type="issue">
<mods:number id="654C9ECBD31B993770E3E9BC5C81643C">3</mods:number>
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</mods:extent>
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<mods:classification id="75635F3FF3B203716E569A86AE1F0614">journal article</mods:classification>
<mods:identifier id="A2EC362EC6DBC43C2750B0692052214E" type="CLB-Dataset">9529</mods:identifier>
<mods:identifier id="4717319A6D8657906296B0FD8D304F08" type="DOI">10.11646/zootaxa.4881.3.7</mods:identifier>
<mods:identifier id="A051CE655E1F3CD785D68B3933B0203B" type="GBIF-Dataset">b7622904-4534-4fc0-aebd-d4766d680eb9</mods:identifier>
<mods:identifier id="A3B5CF8DB628AD8C346538275049BEA7" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="DC0235568271EB2952EEDB8E91653634" type="Zenodo-Dep">4283960</mods:identifier>
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</mods:mods>
<treatment id="03B5EB57AE19FFD299C47F5404321E8E" ID-DOI="http://doi.org/10.5281/zenodo.4328755" ID-GBIF-Taxon="170510553" ID-Zenodo-Dep="4328755" LSID="urn:lsid:plazi:treatment:03B5EB57AE19FFD299C47F5404321E8E" httpUri="http://treatment.plazi.org/id/03B5EB57AE19FFD299C47F5404321E8E" lastPageId="5" lastPageNumber="550" pageId="1" pageNumber="546">
<subSubSection id="C30609CAAE19FFD699C47F5400D71B06" pageId="1" pageNumber="546" type="nomenclature">
<paragraph id="8BA35A41AE19FFD699C47F5400F01B6A" blockId="1.[151,510,1050,1149]" box="[151,473,1050,1077]" pageId="1" pageNumber="546">
<heading id="D0EBED2DAE19FFD699C47F5400F01B6A" bold="true" box="[151,473,1050,1077]" fontSize="11" level="1" pageId="1" pageNumber="546" reason="1">
<emphasis id="B9688653AE19FFD699C47F5400F01B6A" bold="true" box="[151,473,1050,1077]" pageId="1" pageNumber="546">
<taxonomicName id="4C1C21C2AE19FFD699C47F54005C1B6A" ID-CoL="8MH4W" authority="Tian &amp; He, 2020" authorityName="Tian &amp; He" authorityYear="2020" box="[151,373,1050,1077]" class="Insecta" family="Carabidae" genus="Jujiroa" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="546" phylum="Arthropoda" rank="species" species="zhouchaoi" status="sp. nov.">
<emphasis id="B9688653AE19FFD699C47F54005C1B6A" bold="true" box="[151,373,1050,1077]" italics="true" pageId="1" pageNumber="546">Jujiroa zhouchaoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B3B28AE19FFD698287F5400F01B6A" box="[379,473,1051,1077]" pageId="1" pageNumber="546" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA35A41AE19FFD699C47F7000D71B06" blockId="1.[151,510,1050,1149]" box="[151,510,1087,1113]" pageId="1" pageNumber="546">(Chinese name: ḄDĀṚṃm)</paragraph>
</subSubSection>
<subSubSection id="C30609CAAE19FFD699C47F2C01D61B22" box="[151,255,1123,1149]" pageId="1" pageNumber="546" type="description">
<paragraph id="8BA35A41AE19FFD699C47F2C01D61B22" blockId="1.[151,510,1050,1149]" box="[151,255,1123,1149]" pageId="1" pageNumber="546">
<figureCitation id="132746C4AE19FFD699C47F2C01D61B22" box="[151,255,1123,1149]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="2.[151,250,1989,2013]" captionStartId-1="3.[151,250,1702,1726]" captionStartId-2="4.[151,250,1920,1944]" captionTargetBox-1="[151,1426,993,1668]" captionTargetBox-2="[151,1435,181,1896]" captionTargetId-0="figure-20@2.[227,1361,181,1967]" captionTargetId-1="figure-376@3.[151,1435,993,1679]" captionTargetId-2="figure-20@4.[151,1435,181,1896]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURE 1. Habitus of Jujiroa zhouchaoi sp. nov., male, holotype." captionText-1="FIGURE 2. Male genitalia of Jujiroa zhouchaoi sp. nov., paratype." captionText-2="FIGURE 3. Cave Shenxian Dong, the type locality of Jujiroa zhouchaoi sp. nov. A, environs of the cave, entrance shown by arrowhead; B, C, entrance; D, individual running in cave; E, the moist and gravelly areas in cave, showing the habitat where the beetles were collected; F, Chao Zhou is collecting." figureDoi-0="http://doi.org/10.5281/zenodo.4289212" figureDoi-1="http://doi.org/10.5281/zenodo.4283962" figureDoi-2="http://doi.org/10.5281/zenodo.4283964" httpUri-0="https://zenodo.org/record/4289212/files/figure.png" httpUri-1="https://zenodo.org/record/4283962/files/figure.png" httpUri-2="https://zenodo.org/record/4283964/files/figure.png" pageId="1" pageNumber="546">Figs. 1–3</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30609CAAE19FFD699C47FE502FB1A6E" pageId="1" pageNumber="546" type="materials_examined">
<paragraph id="8BA35A41AE19FFD699C47FE502FB1A6E" blockId="1.[151,1437,1194,2013]" pageId="1" pageNumber="546">
<emphasis id="B9688653AE19FFD699C47FE5002F1B9B" bold="true" box="[151,262,1194,1220]" pageId="1" pageNumber="546">Material.</emphasis>
<materialsCitation id="3B74501CAE19FFD6985F7FE400621A52" ID-GBIF-Occurrence="2985238312" collectingDate="2020-04-11" collectionCode="SCAU" collectorName="Li He &amp; Chao Zhou" county="cave Shenxian Dong" elevation="1460" latitude="30.944223" location="Yutang" longLatPrecision="1" longitude="103.5278" municipality="Mt. Zhaogong" pageId="1" pageNumber="546" specimenCount="1" stateProvince="Sichuan" typeStatus="holotype">
<typeStatus id="54A7E4E3AE19FFD6985F7FE400501B9A" box="[268,377,1195,1221]" pageId="1" pageNumber="546" type="holotype">Holotype</typeStatus>
: male,
<collectingCounty id="62C222CDAE19FFD698957FE403861B9A" box="[454,687,1195,1221]" pageId="1" pageNumber="546">cave Shenxian Dong</collectingCounty>
,
<collectingMunicipality id="6BC7C03BAE19FFD69BEA7FE402721B9A" box="[697,859,1195,1221]" pageId="1" pageNumber="546">Mt. Zhaogong</collectingMunicipality>
,
<location id="8EC30C9AAE19FFD69A377FE4029F1B9A" LSID="urn:lsid:plazi:treatment:03B5EB57AE19FFD299C47F5404321E8E:8EC30C9AAE19FFD69A377FE4029F1B9A" box="[868,950,1195,1221]" county="cave Shenxian Dong" latitude="30.944223" longLatPrecision="1" longitude="103.5278" municipality="Mt. Zhaogong" name="Yutang" pageId="1" pageNumber="546" stateProvince="Sichuan">Yutang</location>
,
<location id="8EC30C9AAE19FFD69A927FE4056E1B9A" LSID="urn:lsid:plazi:treatment:03B5EB57AE19FFD299C47F5404321E8E:8EC30C9AAE19FFD69A927FE4056E1B9A" box="[961,1095,1195,1221]" county="cave Shenxian Dong" latitude="30.944223" longLatPrecision="1" longitude="103.5278" municipality="Mt. Zhaogong" name="Dujiangyan" pageId="1" pageNumber="546" stateProvince="Sichuan">Dujiangyan</location>
,
<location id="8EC30C9AAE19FFD69D037FE405911B9A" LSID="urn:lsid:plazi:treatment:03B5EB57AE19FFD299C47F5404321E8E:8EC30C9AAE19FFD69D037FE405911B9A" box="[1104,1208,1195,1221]" county="cave Shenxian Dong" latitude="30.944223" longLatPrecision="1" longitude="103.5278" municipality="Mt. Zhaogong" name="Chengdu" pageId="1" pageNumber="546" stateProvince="Sichuan">Chengdu</location>
,
<collectingRegion id="49D894A3AE19FFD69D907FE404341B9A" box="[1219,1309,1195,1221]" country="China" name="Sichuan" pageId="1" pageNumber="546">Sichuan</collectingRegion>
(Hnjờẁ ďḿďữûḿṮṞüŭẌƜṖIJḆ),
<geoCoordinate id="EE283C86AE19FFD69B077F9F03D61BB4" box="[596,767,1232,1259]" degrees="30" direction="north" minutes="56" orientation="latitude" pageId="1" pageNumber="546" precision="1" seconds="39.21" value="30.944223">30°56′39.21″N</geoCoordinate>
,
<geoCoordinate id="EE283C86AE19FFD69A5F7F9F02EB1BB4" box="[780,962,1232,1259]" degrees="103" direction="east" minutes="31" orientation="longitude" pageId="1" pageNumber="546" precision="1" seconds="40.07" value="103.5278">103°31′40.07″E</geoCoordinate>
,
<quantity id="4CE4F7A4AE19FFD69A9C7F8005011BB6" box="[975,1064,1231,1257]" metricMagnitude="3" metricUnit="m" metricValue="1.46" pageId="1" pageNumber="546" unit="m" value="1460.0">
<elevation id="0031BD72AE19FFD69A9C7F8005011BB6" box="[975,1064,1231,1257]" metricMagnitude="3" metricUnit="m" metricValue="1.46" pageId="1" pageNumber="546" unit="m" value="1460.0">1460 m</elevation>
</quantity>
,
<date id="FFA27C81AE19FFD69D667F8005901BB6" box="[1077,1209,1231,1257]" pageId="1" pageNumber="546" value="2020-04-11">
<collectingDate id="EFE68569AE19FFD69D667F8005901BB6" box="[1077,1209,1231,1257]" pageId="1" pageNumber="546" value="2020-04-11">IV-11-2020</collectingDate>
</date>
, leg.
<collectorName id="26E93F97AE19FFD69DAB7F8004131BB6" box="[1272,1338,1231,1257]" pageId="1" pageNumber="546">Li He</collectorName>
&amp;
<collectorName id="26E93F97AE19FFD69C327F8001FC1A52" pageId="1" pageNumber="546">Chao Zhou</collectorName>
, in
<collectionCode id="ED0DC284AE19FFD699AE7FBC00621A52" box="[253,331,1267,1293]" pageId="1" pageNumber="546">SCAU</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B74501CAE19FFD698067FBC02D21A52" ID-GBIF-Occurrence="2985238308" box="[341,1019,1267,1293]" collectingDate="2020-04-11" collectionCode="SCAU" collectorName="Li He &amp; Chao Zhou" county="cave Shenxian Dong" elevation="1460" latitude="30.944223" location="Yutang" longLatPrecision="1" longitude="103.5278" municipality="Mt. Zhaogong" pageId="1" pageNumber="546" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Sichuan" typeStatus="paratype">
<typeStatus id="54A7E4E3AE19FFD698067FBC00EB1A52" box="[341,450,1267,1293]" pageId="1" pageNumber="546" type="paratype">Paratypes</typeStatus>
(10):
<specimenCount id="9D1A91C8AE19FFD69B557FBC03761A52" box="[518,607,1267,1293]" pageId="1" pageNumber="546" type="male">3 males</specimenCount>
,
<specimenCount id="9D1A91C8AE19FFD69B397FBC03F01A52" box="[618,729,1267,1293]" pageId="1" pageNumber="546" type="female">2 females</specimenCount>
,
<emphasis id="B9688653AE19FFD69BB77FBC02331A52" box="[740,794,1267,1293]" italics="true" pageId="1" pageNumber="546">idem</emphasis>
<typeStatus id="54A7E4E3AE19FFD69A727FBC02AA1A52" box="[801,899,1267,1293]" pageId="1" pageNumber="546" type="holotype">holotype</typeStatus>
, in
<collectionCode id="ED0DC284AE19FFD69AFE7FBC02D21A52" box="[941,1019,1267,1293]" pageId="1" pageNumber="546">SCAU</collectionCode>
</materialsCitation>
;
<materialsCitation id="3B74501CAE19FFD69D547FBC01FA1A6E" ID-GBIF-Occurrence="2985238311" collectingDate="2020-04-11" collectionCode="SCAU" collectorName="Li He &amp; Chao Zhou" county="cave Shenxian Dong" elevation="1460" latitude="30.944223" location="Yutang" longLatPrecision="1" longitude="103.5278" municipality="Mt. Zhaogong" pageId="1" pageNumber="546" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Sichuan" typeStatus="holotype">
<specimenCount id="9D1A91C8AE19FFD69D547FBC05761A52" box="[1031,1119,1267,1293]" pageId="1" pageNumber="546" type="male">2 males</specimenCount>
,
<specimenCount id="9D1A91C8AE19FFD69D387FBC05E71A52" box="[1131,1230,1267,1293]" pageId="1" pageNumber="546" type="female">1 female</specimenCount>
,
<emphasis id="B9688653AE19FFD69D8A7FBC04261A52" box="[1241,1295,1267,1293]" italics="true" pageId="1" pageNumber="546">idem</emphasis>
<typeStatus id="54A7E4E3AE19FFD69C447FBC04501A52" box="[1303,1401,1267,1293]" pageId="1" pageNumber="546" type="holotype">holotype</typeStatus>
, in CHL
</materialsCitation>
;
<materialsCitation id="3B74501CAE19FFD6998D7E58036F1A6E" ID-GBIF-Occurrence="2985238306" box="[222,582,1303,1329]" collectingDate="2020-04-11" collectionCode="SCAU" collectorName="Li He &amp; Chao Zhou" county="cave Shenxian Dong" elevation="1460" latitude="30.944223" location="Yutang" longLatPrecision="1" longitude="103.5278" municipality="Mt. Zhaogong" pageId="1" pageNumber="546" specimenCount="2" specimenCount-male="2" stateProvince="Sichuan" typeStatus="holotype">
<specimenCount id="9D1A91C8AE19FFD6998D7E58001E1A6E" box="[222,311,1303,1329]" pageId="1" pageNumber="546" type="male">2 males</specimenCount>
,
<emphasis id="B9688653AE19FFD698117E5800511A6E" box="[322,376,1303,1329]" italics="true" pageId="1" pageNumber="546">idem</emphasis>
<typeStatus id="54A7E4E3AE19FFD698D37E5800CB1A6E" box="[384,482,1303,1329]" pageId="1" pageNumber="546" type="holotype">holotype</typeStatus>
, in CCZ
</materialsCitation>
.
<materialsCitation id="3B74501CAE19FFD69B027E5802E41A6E" ID-GBIF-Occurrence="2985238309" box="[593,973,1303,1329]" collectionCode="SCAU" pageId="1" pageNumber="546" specimenCount="1" specimenCount-female="1" stateProvince="Sichuan" typeStatus="paratype">
<specimenCount id="9D1A91C8AE19FFD69B027E58039C1A6E" box="[593,693,1303,1329]" pageId="1" pageNumber="546" type="female">1 female</specimenCount>
, spare in ethanol,
<collectionCode id="ED0DC284AE19FFD69A2C7E5802E41A6E" box="[895,973,1303,1329]" pageId="1" pageNumber="546">SCAU</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30609CAAE19FFD699947E75046A1A26" pageId="1" pageNumber="546" type="diagnosis">
<paragraph id="8BA35A41AE19FFD699947E75046A1A26" blockId="1.[151,1437,1194,2013]" pageId="1" pageNumber="546">
<emphasis id="B9688653AE19FFD699947E75006B1A0B" bold="true" box="[199,322,1338,1364]" pageId="1" pageNumber="546">Diagnosis.</emphasis>
A medium-sized
<taxonomicName id="4C1C21C2AE19FFD69B567E74037C1A0B" authorityName="Ueno" authorityYear="1952" box="[517,597,1339,1364]" class="Insecta" family="Carabidae" genus="Jujiroa" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="546" phylum="Arthropoda" rank="genus">
<emphasis id="B9688653AE19FFD69B567E74037C1A0B" box="[517,597,1339,1364]" italics="true" pageId="1" pageNumber="546">Jujiroa</emphasis>
</taxonomicName>
species, body depigmented, microphthalmic, head thin and nearly parallelsided, fore angles of pronotum distinctly protruded, elytral striae largely punctate, not mucronate at apices.
</paragraph>
</subSubSection>
<subSubSection id="C30609CAAE19FFD499947ECD05F61C96" lastPageId="3" lastPageNumber="548" pageId="1" pageNumber="546" type="description">
<paragraph id="8BA35A41AE19FFD699947ECD051D1AC2" blockId="1.[151,1437,1194,2013]" box="[199,1076,1410,1437]" pageId="1" pageNumber="546">
<emphasis id="B9688653AE19FFD699947ECD00701AC3" bold="true" box="[199,345,1410,1436]" pageId="1" pageNumber="546">Description.</emphasis>
Length: 13.0–15.0 mm; width: 4.0–
<quantity id="4CE4F7A4AE19FFD69BBC7ECC02611AC2" box="[751,840,1411,1437]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="1" pageNumber="546" unit="mm" value="4.4">4.4 mm</quantity>
. Habitus as in
<figureCitation id="132746C4AE19FFD69AB87ECC05071AC2" box="[1003,1070,1411,1437]" captionStart="FIGURE 1" captionStartId="2.[151,250,1989,2013]" captionTargetId="figure-20@2.[227,1361,181,1967]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of Jujiroa zhouchaoi sp. nov., male, holotype." figureDoi="http://doi.org/10.5281/zenodo.4289212" httpUri="https://zenodo.org/record/4289212/files/figure.png" pageId="1" pageNumber="546">Fig. 1</figureCitation>
.
</paragraph>
<paragraph id="8BA35A41AE19FFD699947EE800D41956" blockId="1.[151,1437,1194,2013]" pageId="1" pageNumber="546">Body concolorous reddish brown but a little darker on head, surface smooth and glabrous (though rough on sides and lateral margins of pronotum), strongly shiny. Microsculptural meshes nearly isodiametric on head, transversely on pronotum and elytra.</paragraph>
<paragraph id="8BA35A41AE19FFD699947D5C00A11882" blockId="1.[151,1437,1194,2013]" pageId="1" pageNumber="546">
Head thin and elongate, much longer than wide, HLm/HW = 2.05–2.11, HLl/HW = 1.54–1.56; widest exactly where the eyes are at about middle of head from base to labrum; genae convex, and slightly expanded at sides; frontal furrows short and shallow, ending before anterior supraorbital pores; two pairs of supraorbital pores present, anterior at little before widest point and near inner margin of eye, posterior distant from eye; eyes very small and flat; clypeus bisetose, labrum bisinuate at front margin, 6-setose; mandibles elongated, teeth reduced; labial suture clear; mentum with two setae on either side just in front of the basal pit which is very small; median tooth short, bifid at tip; submentum with three setae on each side, inner ones longest, outer ones shortest; ligula short, widened and truncated at apical margin, bisetose; palpomeres long and slender, the 2
<superScript id="7C69F709AE19FFD69AFE7C4102941843" attach="left" box="[941,957,1806,1820]" fontSize="6" pageId="1" pageNumber="546">nd</superScript>
labial palpomere bisetose on inner margin, 1.2 times as long as 3
<superScript id="7C69F709AE19FFD698D67C7D00BA181F" attach="none" box="[389,403,1842,1856]" fontSize="6" pageId="1" pageNumber="546">rd</superScript>
, the 3
<superScript id="7C69F709AE19FFD698857C7D00CD181F" attach="left" box="[470,484,1842,1856]" fontSize="6" pageId="1" pageNumber="546">rd</superScript>
maxillary palpomere as long as 4
<superScript id="7C69F709AE19FFD69A097C7D024E181F" attach="none" box="[858,871,1842,1856]" fontSize="6" pageId="1" pageNumber="546">th</superScript>
; antennae filiform, thin and very long, extended to apical 1/4 elytra, the 1
<superScript id="7C69F709AE19FFD698C07C1800B7183A" attach="left" box="[403,414,1879,1893]" fontSize="6" pageId="1" pageNumber="546">st</superScript>
to 3
<superScript id="7C69F709AE19FFD698827C1900F6183B" attach="left" box="[465,479,1878,1892]" fontSize="6" pageId="1" pageNumber="546">rd</superScript>
antennomeres glabrous, each of the 1
<superScript id="7C69F709AE19FFD69AD97C1802BC183A" attach="left" box="[906,917,1879,1893]" fontSize="6" pageId="1" pageNumber="546">st</superScript>
and 2
<superScript id="7C69F709AE19FFD69A887C1902C2183B" attach="left" box="[987,1003,1878,1892]" fontSize="6" pageId="1" pageNumber="546">nd</superScript>
with a seta near apex, pubescent from the 4
<superScript id="7C69F709AE19FFD6999D7C3501F218D7" attach="none" box="[206,219,1914,1928]" fontSize="6" pageId="1" pageNumber="546">th</superScript>
; the 2
<superScript id="7C69F709AE19FFD6984C7C35000618D7" attach="left" box="[287,303,1914,1928]" fontSize="6" pageId="1" pageNumber="546">nd</superScript>
shortest, while 5
<superScript id="7C69F709AE19FFD698BE7C3500D318D7" attach="left" box="[493,506,1914,1928]" fontSize="6" pageId="1" pageNumber="546">th</superScript>
longest; relative length of each antennomere compared with 2
<superScript id="7C69F709AE19FFD69DE37C3505E918D7" attach="left" box="[1200,1216,1914,1928]" fontSize="6" pageId="1" pageNumber="546">nd</superScript>
antennomere in the
<typeStatus id="54A7E4E3AE19FFD699C47CD001D018E6" box="[151,249,1951,1977]" pageId="1" pageNumber="546" type="holotype">holotype</typeStatus>
as follows: 1
<superScript id="7C69F709AE19FFD698DE7CD000B118F2" attach="left" box="[397,408,1951,1965]" fontSize="6" pageId="1" pageNumber="546">st</superScript>
(2.05), 2
<superScript id="7C69F709AE19FFD698A97CD1032318F3" attach="left" box="[506,522,1950,1964]" fontSize="6" pageId="1" pageNumber="546">nd</superScript>
(1.00), 3
<superScript id="7C69F709AE19FFD69B3C7CD1035418F3" attach="left" box="[623,637,1950,1964]" fontSize="6" pageId="1" pageNumber="546">rd</superScript>
(2.20), 4
<superScript id="7C69F709AE19FFD69BB17CD103C618F3" attach="left" box="[738,751,1950,1964]" fontSize="6" pageId="1" pageNumber="546">th</superScript>
(2.25), 5
<superScript id="7C69F709AE19FFD69A077CD1024818F3" attach="left" box="[852,865,1950,1964]" fontSize="6" pageId="1" pageNumber="546">th</superScript>
(2.40), 6
<superScript id="7C69F709AE19FFD69A907CD102F918F3" attach="left" box="[963,976,1950,1964]" fontSize="6" pageId="1" pageNumber="546">th</superScript>
(2.30), 7
<superScript id="7C69F709AE19FFD69D617CD1051618F3" attach="left" box="[1074,1087,1950,1964]" fontSize="6" pageId="1" pageNumber="546">th</superScript>
(2.25), 8
<superScript id="7C69F709AE19FFD69DF07CD1059918F3" attach="left" box="[1187,1200,1950,1964]" fontSize="6" pageId="1" pageNumber="546">th</superScript>
(2.05), 9
<superScript id="7C69F709AE19FFD69C417CD1043618F3" attach="left" box="[1298,1311,1950,1964]" fontSize="6" pageId="1" pageNumber="546">th</superScript>
(1.90), 10
<superScript id="7C69F709AE19FFD69CC37CD104B418F3" attach="none" box="[1424,1437,1950,1964]" fontSize="6" pageId="1" pageNumber="546">th</superScript>
(1.65) and 11
<superScript id="7C69F709AE19FFD6987F7C8D0010188F" attach="left" box="[300,313,1986,2000]" fontSize="6" pageId="1" pageNumber="546">th</superScript>
(1.55).
</paragraph>
<caption id="DF630AC9AE1AFFD599C47C8A026A1882" ID-DOI="http://doi.org/10.5281/zenodo.4289212" ID-Zenodo-Dep="4289212" box="[151,835,1989,2013]" httpUri="https://zenodo.org/record/4289212/files/figure.png" pageId="2" pageNumber="547" targetBox="[455,1157,253,1943]" targetPageId="2">
<paragraph id="8BA35A41AE1AFFD599C47C8A026A1882" blockId="2.[151,835,1989,2013]" box="[151,835,1989,2013]" pageId="2" pageNumber="547">
<emphasis id="B9688653AE1AFFD599C47C8A003A1882" bold="true" box="[151,275,1989,2013]" pageId="2" pageNumber="547">FIGURE 1.</emphasis>
Habitus of
<taxonomicName id="4C1C21C2AE1AFFD598D97C8903681882" authority="Tian &amp; He, 2020" authorityName="Tian &amp; He" authorityYear="2020" box="[394,577,1989,2013]" class="Insecta" family="Carabidae" genus="Jujiroa" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="547" phylum="Arthropoda" rank="species" species="zhouchaoi" status="sp. nov.">
<emphasis id="B9688653AE1AFFD598D97C8903681882" box="[394,577,1989,2013]" italics="true" pageId="2" pageNumber="547">Jujiroa zhouchaoi</emphasis>
</taxonomicName>
<emphasis id="B9688653AE1AFFD59B147C8A03B11882" bold="true" box="[583,664,1989,2013]" pageId="2" pageNumber="547">
<taxonomicNameLabel id="A25B3B28AE1AFFD59B147C8A03B11882" box="[583,664,1989,2013]" pageId="2" pageNumber="547" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male, holotype.
</paragraph>
</caption>
<paragraph id="8BA35A41AE1BFFD499947BD803061ED6" blockId="3.[151,1437,151,969]" pageId="3" pageNumber="548">Pronotum subcordate, transverse, PL/PW = 0.91–0.93; much wider but slightly shorter than head, PW/HW = 1.56–1.59, PL/HLl = 0.94; widest at about 1/3 from front, lateral margins including front and hind angles widely reflexed throughout, gently and gradually narrowed towards hind angles, basal foveae short but well-marked; only basal latero-marginal setae present, inserted just on the hind angles; fore angles extraordinarily protruded and roundly lobed, hind angles rectangular and pointed; entire lateral margins and front without borders, base finely bordered, slightly narrower than front including front angles, PbW/PfW = 0.98–0.99; both base and front nearly straight. Scutellum small, triangular.</paragraph>
<paragraph id="8BA35A41AE1BFFD499947ADC020C1DDA" blockId="3.[151,1437,151,969]" pageId="3" pageNumber="548">
Elytra elongate, amygdaloid, much longer than wide, EL/EW = 1.76–1.78; distinctly longer than forebody including mandibles, much wider than pronotum; base well-bordered (but unbordered against the 1
<superScript id="7C69F709AE1BFFD49C737AF804021E9A" attach="left" box="[1312,1323,439,453]" fontSize="6" pageId="3" pageNumber="548">st</superScript>
interval), shoulders nearly rounded; widest at about 4/7 of elytra from base, apex protruded and narrowed backwardly, not mucronate; disc convex, striae entire, impressed by large and nearly rounded punctures; scutellar striole short; basal pores absent; the 3
<superScript id="7C69F709AE1BFFD49835796D005D1D6F" attach="left" box="[358,372,546,560]" fontSize="6" pageId="3" pageNumber="548">rd</superScript>
interval with a setiferous pore at about 3/7 of elytra from apex, and presence of a preapical pore close to the 2
<superScript id="7C69F709AE1BFFD49878790900121D0B" attach="left" box="[299,315,582,596]" fontSize="6" pageId="3" pageNumber="548">nd</superScript>
stria at apical 1/8 of elytra; three pores present on 7
<superScript id="7C69F709AE1BFFD49AD2790902A71D0B" attach="left" box="[897,910,582,596]" fontSize="6" pageId="3" pageNumber="548">th</superScript>
stria posteriorly; an apical pore present; 23–25 marginal umbilicate pores present throughout, continuous.
</paragraph>
<paragraph id="8BA35A41AE1BFFD4999479C000B21C02" blockId="3.[151,1437,151,969]" pageId="3" pageNumber="548">
Ventral surface smooth and glabrous. Legs slender and elongate, procoxae asetose, mesocoxae unisetose, metacoxae bisetose, without inner seta; each trochanter with a single seta; metafemur bisetose posteriorly; tibiae and tarsi smooth, without longitudinal sulci or striae externally; the 4
<superScript id="7C69F709AE1BFFD49A63799902141DBB" attach="left" box="[816,829,726,740]" fontSize="6" pageId="3" pageNumber="548">th</superScript>
tarsomere bilobed in fore and middle legs, moderately emarginated in hind ones; 1
<superScript id="7C69F709AE1BFFD4988579B400C81C56" attach="left" box="[470,481,763,777]" fontSize="6" pageId="3" pageNumber="548">st</superScript>
–3
<superScript id="7C69F709AE1BFFD49B5579B5033D1C57" attach="left" box="[518,532,762,776]" fontSize="6" pageId="3" pageNumber="548">rd</superScript>
tarsomeres longitudinally bisulcate; 1
<superScript id="7C69F709AE1BFFD49A9479B402FB1C56" attach="left" box="[967,978,763,777]" fontSize="6" pageId="3" pageNumber="548">st</superScript>
protarsomere not modified in male, but with two spongy setae on ventral surface. Each abdominal ventrite IV–VI bisetose, ventrite VII bisetose in male, quadrisetose in female.
</paragraph>
<paragraph id="8BA35A41AE1BFFD49994782805F61C96" blockId="3.[151,1437,151,969]" pageId="3" pageNumber="548">
Male genitalia (
<figureCitation id="132746C4AE1BFFD49825782800921CDE" box="[374,443,871,897]" captionStart="FIGURE 2" captionStartId="3.[151,250,1702,1726]" captionTargetBox="[151,1426,993,1668]" captionTargetId="figure-376@3.[151,1435,993,1679]" captionTargetPageId="3" captionText="FIGURE 2. Male genitalia of Jujiroa zhouchaoi sp. nov., paratype." figureDoi="http://doi.org/10.5281/zenodo.4283962" httpUri="https://zenodo.org/record/4283962/files/figure.png" pageId="3" pageNumber="548">Fig. 2</figureCitation>
): Median lobe of aedeagus (
<figureCitation id="132746C4AE1BFFD49BA6782802651CDE" box="[757,844,871,897]" captionStart="FIGURE 2" captionStartId="3.[151,250,1702,1726]" captionTargetBox="[151,1426,993,1668]" captionTargetId="figure-376@3.[151,1435,993,1679]" captionTargetPageId="3" captionText="FIGURE 2. Male genitalia of Jujiroa zhouchaoi sp. nov., paratype." figureDoi="http://doi.org/10.5281/zenodo.4283962" httpUri="https://zenodo.org/record/4283962/files/figure.png" pageId="3" pageNumber="548">Fig. 2A</figureCitation>
) slender and elongate, slightly arcuate in middle portion, then gently curved towards apex which is bluntly pointed; sagittal aileron quite small, base moderately opened, parameres developed (
<figureCitation id="132746C4AE1BFFD498C778E000C51C96" box="[404,492,943,969]" captionStart="FIGURE 2" captionStartId="3.[151,250,1702,1726]" captionTargetBox="[151,1426,993,1668]" captionTargetId="figure-376@3.[151,1435,993,1679]" captionTargetPageId="3" captionText="FIGURE 2. Male genitalia of Jujiroa zhouchaoi sp. nov., paratype." figureDoi="http://doi.org/10.5281/zenodo.4283962" httpUri="https://zenodo.org/record/4283962/files/figure.png" pageId="3" pageNumber="548">Fig. 2B</figureCitation>
). In lateral view, apical lobe thin, much longer than wide (
<figureCitation id="132746C4AE1BFFD49D2A78E005FB1C96" box="[1145,1234,943,969]" captionStart="FIGURE 2" captionStartId="3.[151,250,1702,1726]" captionTargetBox="[151,1426,993,1668]" captionTargetId="figure-376@3.[151,1435,993,1679]" captionTargetPageId="3" captionText="FIGURE 2. Male genitalia of Jujiroa zhouchaoi sp. nov., paratype." figureDoi="http://doi.org/10.5281/zenodo.4283962" httpUri="https://zenodo.org/record/4283962/files/figure.png" pageId="3" pageNumber="548">Fig. 2C</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF630AC9AE1BFFD499C47DE9026F19E0" ID-DOI="http://doi.org/10.5281/zenodo.4283962" ID-Zenodo-Dep="4283962" box="[151,838,1702,1727]" httpUri="https://zenodo.org/record/4283962/files/figure.png" pageId="3" pageNumber="548" startId="3.[151,250,1702,1726]" targetBox="[151,1426,993,1668]" targetPageId="3">
<paragraph id="8BA35A41AE1BFFD499C47DE9026F19E0" blockId="3.[151,838,1702,1727]" box="[151,838,1702,1727]" pageId="3" pageNumber="548">
<emphasis id="B9688653AE1BFFD499C47DE9003A19E1" bold="true" box="[151,275,1702,1726]" pageId="3" pageNumber="548">FIGURE 2.</emphasis>
Male genitalia of
<taxonomicName id="4C1C21C2AE1BFFD4989E7DE803AA19E1" authority="Tian &amp; He, 2020" authorityName="Tian &amp; He" authorityYear="2020" box="[461,643,1702,1726]" class="Insecta" family="Carabidae" genus="Jujiroa" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="548" phylum="Arthropoda" rank="species" species="zhouchaoi" status="sp. nov.">
<emphasis id="B9688653AE1BFFD4989E7DE803AA19E1" box="[461,643,1702,1726]" italics="true" pageId="3" pageNumber="548">Jujiroa zhouchaoi</emphasis>
</taxonomicName>
<emphasis id="B9688653AE1BFFD49BD97DE803F319E0" bold="true" box="[650,730,1703,1727]" pageId="3" pageNumber="548">
<taxonomicNameLabel id="A25B3B28AE1BFFD49BD97DE803F319E0" box="[650,730,1703,1727]" pageId="3" pageNumber="548" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, paratype.
</paragraph>
</caption>
<subSubSection id="C30609CAAE1BFFD499947DA203941810" pageId="3" pageNumber="548" type="discussion">
<paragraph id="8BA35A41AE1BFFD499947DA203941810" blockId="3.[151,1437,1773,2015]" pageId="3" pageNumber="548">
<emphasis id="B9688653AE1BFFD499947DA200121858" bold="true" box="[199,315,1773,1799]" pageId="3" pageNumber="548">Remarks.</emphasis>
Similar to
<taxonomicName id="4C1C21C2AE1BFFD498EB7DA103FB1858" authority="Ueno, 2007" authorityName="Ueno" authorityYear="2007" box="[440,722,1773,1799]" class="Insecta" family="Carabidae" genus="Jujiroa" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="548" phylum="Arthropoda" rank="species" species="satoi">
<emphasis id="B9688653AE1BFFD498EB7DA1036C1858" box="[440,581,1774,1799]" italics="true" pageId="3" pageNumber="548">Jujiroa satoi</emphasis>
<bibRefCitation id="EF8D27B0AE1BFFD49B1F7DA203FB1858" author="Ueno, S. I." box="[588,722,1773,1799]" pageId="3" pageNumber="548" pagination="21 - 26" refId="ref6281" refString="Ueno, S. I. (2007) Occurrence of a new cave species of Jujiroa (Coleoptera: Carabidae: Platyninae) from Central Sichuan, Southwest China. Elytra, 35 (1), 21 - 26." type="journal article" year="2007">Uéno, 2007</bibRefCitation>
</taxonomicName>
, but it is easily distinguished by its slenderer body, thinner head which is almost parallel-sided, coarsely punctate striae on elytra, presence of an additional short seta on each outer side of submentum, and slenderer male genitalia.
</paragraph>
</subSubSection>
<subSubSection id="C30609CAAE1BFFD499947C16033018C8" pageId="3" pageNumber="548" type="etymology">
<paragraph id="8BA35A41AE1BFFD499947C16033018C8" blockId="3.[151,1437,1773,2015]" pageId="3" pageNumber="548">
<emphasis id="B9688653AE1BFFD499947C160067182C" bold="true" box="[199,334,1881,1907]" pageId="3" pageNumber="548">Etymology.</emphasis>
The new species is dedicated to Mr. Chao Zhou, an enthusiastic amateur entomologist who is one of the collectors of the
<typeStatus id="54A7E4E3AE1BFFD498C97C3200E318C8" box="[410,458,1917,1943]" pageId="3" pageNumber="548">type</typeStatus>
series.
</paragraph>
</subSubSection>
<subSubSection id="C30609CAAE1BFFD299947CEF04321E8E" lastPageId="5" lastPageNumber="550" pageId="3" pageNumber="548" type="distribution">
<paragraph id="8BA35A41AE1BFFD499947CEF00E81880" blockId="3.[151,1437,1773,2015]" pageId="3" pageNumber="548">
<emphasis id="B9688653AE1BFFD499947CEF004818E5" bold="true" box="[199,353,1952,1978]" pageId="3" pageNumber="548">Distribution.</emphasis>
<collectingCountry id="F30B1AD1AE1BFFD4983B7CEE008218E4" box="[360,427,1953,1979]" name="China" pageId="3" pageNumber="548">China</collectingCountry>
(
<collectingRegion id="49D894A3AE1BFFD498E87CEE033518E3" box="[443,540,1953,1980]" country="China" name="Sichuan" pageId="3" pageNumber="548">Sichuan</collectingRegion>
). Known from the cave Shenxian Dong and two other anonymous caves in Dujiangyan,
<collectingRegion id="49D894A3AE1BFFD498557C8A00491880" box="[262,352,1989,2015]" country="China" name="Sichuan" pageId="3" pageNumber="548">Sichuan</collectingRegion>
(
<figureCitation id="132746C4AE1BFFD4983C7C8A009D1880" box="[367,436,1989,2015]" captionStart="FIGURE 3" captionStartId="4.[151,250,1920,1944]" captionTargetBox="[151,1435,181,1896]" captionTargetId="figure-20@4.[151,1435,181,1896]" captionTargetPageId="4" captionText="FIGURE 3. Cave Shenxian Dong, the type locality of Jujiroa zhouchaoi sp. nov. A, environs of the cave, entrance shown by arrowhead; B, C, entrance; D, individual running in cave; E, the moist and gravelly areas in cave, showing the habitat where the beetles were collected; F, Chao Zhou is collecting." figureDoi="http://doi.org/10.5281/zenodo.4283964" httpUri="https://zenodo.org/record/4283964/files/figure.png" pageId="3" pageNumber="548">Fig. 3</figureCitation>
).
</paragraph>
<caption id="DF630AC9AE1CFFD399C47CCF03B318BF" ID-DOI="http://doi.org/10.5281/zenodo.4283964" ID-Zenodo-Dep="4283964" httpUri="https://zenodo.org/record/4283964/files/figure.png" pageId="4" pageNumber="549" startId="4.[151,250,1920,1944]" targetBox="[151,1435,181,1896]" targetPageId="4">
<paragraph id="8BA35A41AE1CFFD399C47CCF03B318BF" blockId="4.[151,1436,1920,2016]" pageId="4" pageNumber="549">
<emphasis id="B9688653AE1CFFD399C47CCF003D18C7" bold="true" box="[151,276,1920,1944]" pageId="4" pageNumber="549">FIGURE 3.</emphasis>
Cave Shenxian Dong, the type locality of
<taxonomicName id="4C1C21C2AE1CFFD39B9A7CCE02A818C7" authority="Tian &amp; He, 2020" authorityName="Tian &amp; He" authorityYear="2020" box="[713,897,1920,1944]" class="Insecta" family="Carabidae" genus="Jujiroa" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="549" phylum="Arthropoda" rank="species" species="zhouchaoi" status="sp. nov.">
<emphasis id="B9688653AE1CFFD39B9A7CCE02A818C7" box="[713,897,1920,1944]" italics="true" pageId="4" pageNumber="549">Jujiroa zhouchaoi</emphasis>
</taxonomicName>
<emphasis id="B9688653AE1CFFD39ADB7CCF02DA18C7" bold="true" box="[904,1011,1920,1944]" pageId="4" pageNumber="549">
<taxonomicNameLabel id="A25B3B28AE1CFFD39ADB7CCF02F018C7" box="[904,985,1920,1944]" pageId="4" pageNumber="549" rank="species">sp. nov.</taxonomicNameLabel>
A
</emphasis>
, environs of the cave, entrance shown by arrowhead;
<emphasis id="B9688653AE1CFFD3985C7CEB000918E3" bold="true" box="[271,288,1956,1980]" pageId="4" pageNumber="549">B</emphasis>
,
<emphasis id="B9688653AE1CFFD3987F7CEB001718E3" bold="true" box="[300,318,1956,1980]" pageId="4" pageNumber="549">C</emphasis>
, entrance;
<emphasis id="B9688653AE1CFFD398FF7CEB009718E3" bold="true" box="[428,446,1956,1980]" pageId="4" pageNumber="549">D</emphasis>
, individual running in cave;
<emphasis id="B9688653AE1CFFD39BB37CEB03D818E3" bold="true" box="[736,753,1956,1980]" pageId="4" pageNumber="549">E</emphasis>
, the moist and gravelly areas in cave, showing the habitat where the beetles were collected;
<emphasis id="B9688653AE1CFFD398D77C8700BA18BF" bold="true" box="[388,403,1992,2016]" pageId="4" pageNumber="549">F</emphasis>
, Chao Zhou is collecting.
</paragraph>
</caption>
<paragraph id="8BA35A41AE1DFFD299947BD8032C1EF2" blockId="5.[151,1436,151,465]" pageId="5" pageNumber="550">
The calcareous conglomerate cave Shenxian Dong is located on the mountainside of Mt. Zhaogong, the main peak of Mt. Qingcheng. It is about a
<quantity id="4CE4F7A4AE1DFFD29B6A7BF503A11F8A" box="[569,648,186,213]" metricMagnitude="3" metricUnit="m" metricValue="5.5" pageId="5" pageNumber="550" unit="km" value="5.5">5.5 km</quantity>
walk along the hiking trails from Longfeng Village to the entrance of the cave, which is located at the bottom of a hill (
<figureCitation id="132746C4AE1DFFD29BE97B90023D1FA6" box="[698,788,223,249]" captionStart="FIGURE 3" captionStartId="4.[151,250,1920,1944]" captionTargetBox="[151,1435,181,1896]" captionTargetId="figure-20@4.[151,1435,181,1896]" captionTargetPageId="4" captionText="FIGURE 3. Cave Shenxian Dong, the type locality of Jujiroa zhouchaoi sp. nov. A, environs of the cave, entrance shown by arrowhead; B, C, entrance; D, individual running in cave; E, the moist and gravelly areas in cave, showing the habitat where the beetles were collected; F, Chao Zhou is collecting." figureDoi="http://doi.org/10.5281/zenodo.4283964" httpUri="https://zenodo.org/record/4283964/files/figure.png" pageId="5" pageNumber="550">Fig. 3A</figureCitation>
). The opening of the entrance is small and there is a statue of Magu (a goddess in Chinese Taoist mythology) outside the entrance (
<figureCitation id="132746C4AE1DFFD29AED7A4C051E1E42" box="[958,1079,259,285]" captionStart="FIGURE 3" captionStartId="4.[151,250,1920,1944]" captionTargetBox="[151,1435,181,1896]" captionTargetId="figure-20@4.[151,1435,181,1896]" captionTargetPageId="4" captionText="FIGURE 3. Cave Shenxian Dong, the type locality of Jujiroa zhouchaoi sp. nov. A, environs of the cave, entrance shown by arrowhead; B, C, entrance; D, individual running in cave; E, the moist and gravelly areas in cave, showing the habitat where the beetles were collected; F, Chao Zhou is collecting." figureDoi="http://doi.org/10.5281/zenodo.4283964" httpUri="https://zenodo.org/record/4283964/files/figure.png" pageId="5" pageNumber="550">Fig. 3B, C</figureCitation>
). The cave is narrow and moist, with some narrower branches at both sides of the main passage. Since the passage ahead is narrow, we do not know how long the cave is. Individuals of
<taxonomicName id="4C1C21C2AE1DFFD29B7E7A0403931E3A" authority="Tian &amp; He, 2020" authorityName="Tian &amp; He" authorityYear="2020" box="[557,698,331,357]" class="Insecta" family="Carabidae" genus="Jujiroa" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="550" phylum="Arthropoda" rank="species" species="zhouchaoi" status="sp. nov.">
<emphasis id="B9688653AE1DFFD29B7E7A0403931E3A" box="[557,698,331,357]" italics="true" pageId="5" pageNumber="550">J. zhouchaoi</emphasis>
</taxonomicName>
<emphasis id="B9688653AE1DFFD29B927A0402331E3A" bold="true" box="[705,794,331,357]" pageId="5" pageNumber="550">
<taxonomicNameLabel id="A25B3B28AE1DFFD29B927A0402331E3A" box="[705,794,331,357]" pageId="5" pageNumber="550" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
were collected under stone or deadwood in the moist and gravelly areas about 10–70 meters from the entrance (
<figureCitation id="132746C4AE1DFFD29BAD7A2002691ED6" box="[766,832,367,393]" captionStart="FIGURE 3" captionStartId="4.[151,250,1920,1944]" captionTargetBox="[151,1435,181,1896]" captionTargetId="figure-20@4.[151,1435,181,1896]" captionTargetPageId="4" captionText="FIGURE 3. Cave Shenxian Dong, the type locality of Jujiroa zhouchaoi sp. nov. A, environs of the cave, entrance shown by arrowhead; B, C, entrance; D, individual running in cave; E, the moist and gravelly areas in cave, showing the habitat where the beetles were collected; F, Chao Zhou is collecting." figureDoi="http://doi.org/10.5281/zenodo.4283964" httpUri="https://zenodo.org/record/4283964/files/figure.png" pageId="5" pageNumber="550">Fig. 3</figureCitation>
D–F). Other cave animals living in the cave are bats, crickets, spiders and amphipods.
</paragraph>
<paragraph id="8BA35A41AE1DFFD299947AF804321E8E" blockId="5.[151,1436,151,465]" box="[199,1307,439,465]" pageId="5" pageNumber="550">Elytral debris of this new species was also collected in two anonymous caves near Shenxian Dong.</paragraph>
</subSubSection>
</treatment>
</document>
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