treatments-xml/data/03/B5/16/03B51633FF8FFFDE5FFB58EC0572FDA7.xml

<document ID-DOI="10.11646/zootaxa.5263.2.4" ID-GBIF-Dataset="f6701c34-91f2-48c3-9ed1-4dd17fb3fb0d" ID-ISSN="1175-5326" ID-Zenodo-Dep="7801106" ID-ZooBank="4E687740-045F-44C5-810D-68C6EE9072C4" IM.illustrations_approvedBy="diego" IM.taxonomicNames_approvedBy="diego" IM.treatments_approvedBy="diego" approvalRequired="4" approvalRequired_for_textStreams="4" checkinTime="1680678286118" checkinUser="plazi" docAuthor="Qi, Zhi-Hao, He, Hao, Su, Rong-Xiang, Ai, Hong-Mu &amp; Song, Hai-Tian" docDate="2023" docId="03B51633FF8FFFDE5FFB58EC0572FDA7" docLanguage="en" docName="zootaxa.5263.2.4.pdf" docOrigin="Zootaxa 5263 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5263.2.4" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lucanus takeoi Adachi 2020" docType="treatment" docVersion="4" lastPageNumber="258" masterDocId="FF8C6E4BFF9EFFCC5F6C5E420045FFC7" masterDocTitle="Revision of the Lucanus boileaui group and notes on. takeoi Adachi, 2020 from the. maculifemoratus group (Coleoptera: Lucanidae: Lucaninae)" masterLastPageNumber="260" masterPageNumber="240" pageNumber="257" updateTime="1681158889771" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Revision of the Lucanus boileaui group and notes on. takeoi Adachi, 2020 from the. maculifemoratus group (Coleoptera: Lucanidae: Lucaninae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Qi, Zhi-Hao</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0001-8596-0913</mods:nameIdentifier>
<mods:affiliation>College of Plant Protection, Fujian Agriculture and Forestry University, Fuzhou, Fujian, 350002, China. &amp; Fujian Academy of Forestry, Fuzhou, Fujian, 350012, China. &amp; qizhihao 2021 @ 126. com; https: // orcid. org / 0000 - 0001 - 8596 - 0913</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>He, Hao</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-8871-9671</mods:nameIdentifier>
<mods:affiliation>College of Life Sciences, Yunnan Normal University, Kunming, Yunnan, 650500, China. &amp; 18835130675 @ 163. com; https: // orcid. org / 0000 - 0002 - 8871 - 9671</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Su, Rong-Xiang</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-6575-5776</mods:nameIdentifier>
<mods:affiliation>Fujian Academy of Forestry, Fuzhou, Fujian, 350012, China. &amp; surongxiang 2022 @ 126. com; https: // orcid. org / 0000 - 0002 - 6575 - 5776</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ai, Hong-Mu</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-7754-8256</mods:nameIdentifier>
<mods:affiliation>College of Plant Protection, Fujian Agriculture and Forestry University, Fuzhou, Fujian, 350002, China. &amp; aihongmu @ 163. com; https: // orcid. org / 0000 - 0002 - 7754 - 8256</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Song, Hai-Tian</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0003-1042-7959</mods:nameIdentifier>
<mods:affiliation>Fujian Academy of Forestry, Fuzhou, Fujian, 350012, China. &amp; haitiansong @ 126. com; https: // orcid. org / 0000 - 0003 - 1042 - 7959</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-04-05</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>5263</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>240</mods:start>
<mods:end>260</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.11646/zootaxa.5263.2.4</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/zootaxa.5263.2.4</mods:identifier>
<mods:identifier type="GBIF-Dataset">f6701c34-91f2-48c3-9ed1-4dd17fb3fb0d</mods:identifier>
<mods:identifier type="ISSN">1175-5326</mods:identifier>
<mods:identifier type="Zenodo-Dep">7801106</mods:identifier>
<mods:identifier type="ZooBank">4E687740-045F-44C5-810D-68C6EE9072C4</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7814615" ID-Zenodo-Dep="7814615" LSID="urn:lsid:plazi:treatment:03B51633FF8FFFDE5FFB58EC0572FDA7" httpUri="http://treatment.plazi.org/id/03B51633FF8FFFDE5FFB58EC0572FDA7" lastPageId="18" lastPageNumber="258" pageId="17" pageNumber="257">
<subSubSection box="[151,507,1709,1736]" pageId="17" pageNumber="257" type="nomenclature">
<paragraph blockId="17.[151,861,1709,1772]" box="[151,507,1709,1736]" pageId="17" pageNumber="257">
<heading bold="true" box="[151,507,1709,1736]" fontSize="11" level="1" pageId="17" pageNumber="257" reason="1">
<taxonomicName ID-CoL="7X2NK" authority="Adachi, 2020" authorityName="Adachi" authorityYear="2020" box="[151,507,1709,1736]" class="Insecta" family="Lucanidae" genus="Lucanus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="257" phylum="Arthropoda" rank="species" species="takeoi">
<emphasis bold="true" box="[151,507,1709,1736]" pageId="17" pageNumber="257">
<emphasis bold="true" box="[151,337,1709,1736]" italics="true" pageId="17" pageNumber="257">Lucanus takeoi</emphasis>
<bibRefCitation author="Adachi, N." box="[344,507,1709,1735]" pageId="17" pageNumber="257" pagination="8 - 10" refId="ref7642" refString="Adachi, N. (2020) A new species of the genus Lucanus from Szechuan province, southwest China. Kogane, 23, 8 - 10." type="journal article" year="2020">Adachi, 2020</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[151,861,1745,1772]" pageId="17" pageNumber="257" type="description">
<paragraph blockId="17.[151,861,1709,1772]" box="[151,861,1745,1772]" pageId="17" pageNumber="257">
<figureCitation box="[151,277,1745,1771]" captionStart="FIGURE 1" captionStartId="2.[151,250,1840,1864]" captionTargetBox="[158,1432,666,1804]" captionTargetId="figure-216@2.[151,1436,666,1814]" captionTargetPageId="2" captionText="FIGURE 1. Male habitus of Lucanus species. A–B, L. jiaozishanus Qi, He, Su &amp; Song, new species (33.3 mm, holotype, large sized, Kunming, Yunnan); C–D, L. ludivinae Boucher, 1998 (31.3 mm, medium sized, Gongshan, Yunnan); E–F, L. takeoi Adachi, 2020 (29.2 mm, small sized, Huidong, Sichuan); G–H, L. boileaui Planet, 1897 (41.9 mm, small sized, Mabian, Sichuan). A, C, E, G, dorsal view; B, D, F, H, ventral view. Scale bar = 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7801110" httpUri="https://zenodo.org/record/7801110/files/figure.png" pageId="17" pageNumber="257">Figs. 1E–F</figureCitation>
; 
<figureCitation box="[289,364,1745,1771]" captionStart="FIGURE 2" captionStartId="3.[151,250,1896,1920]" captionTargetBox="[151,1431,181,1873]" captionTargetId="figure-17@3.[151,1435,181,1873]" captionTargetPageId="3" captionText="FIGURE 2. Male head (A, G, M, S), clypeolabrum (B, H, N, T), canthus (C, I, O, U) and tibia (D–F, J–L, P–R, V–X) of Lucanus species in dorsal view. A–F, L. jiaozishanus Qi, He, Su &amp; Song, new species (holotype, Kunming, Yunnan); G–L, L. ludivinae Boucher, 1998 (Gongshan, Yunnan); M–R, L. takeoi Adachi, 2020 (Huidong, Sichuan); S–X, L. boileaui Planet, 1897 (Mabian, Sichuan). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7814599" httpUri="https://zenodo.org/record/7814599/files/figure.png" pageId="17" pageNumber="257">2M–R</figureCitation>
; 
<figureCitation box="[375,433,1745,1771]" captionStart="FIGURE 3" captionStartId="4.[151,250,1909,1933]" captionTargetBox="[314,1276,189,1873]" captionTargetId="figure-19@4.[307,1280,181,1885]" captionTargetPageId="4" captionText="FIGURE 3. Male abdominal tergite 8 (A, E, I, M), abdominal ventrite 8 (B, F, J, N) and abdominal segment 9 (C–D, G–H, K–L, O–P) of Lucanus species. A–D, L. jiaozishanus Qi, He, Su &amp; Song, new species (holotype, Kunming, Yunnan); E–H, L. ludivinae Boucher, 1998 (Gongshan, Yunnan); I–L, L. takeoi Adachi, 2020 (Huidong, Sichuan); M–P, L. boileaui Planet, 1897 (Mabian, Sichuan). A, D, E, H, I, L, M, P, dorsal view; B–C, F–G, J–K, N–O, ventral view. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7814603" httpUri="https://zenodo.org/record/7814603/files/figure.png" pageId="17" pageNumber="257">3I–L</figureCitation>
; 4C, M–P; 5C–D; 
<figureCitation box="[642,703,1745,1771]" captionStart="FIGURE 6" captionStartId="7.[151,250,1871,1895]" captionTargetBox="[151,1433,181,1847]" captionTargetId="figure-17@7.[151,1436,181,1847]" captionTargetPageId="7" captionText="FIGURE 6. Female head (A, G, I, O), clypeolabrum (B, J, P), canthus (C, H, K, Q) and tibia (D–F, L–N, R–T) of Lucanus species in dorsal view. A–F, L. jiaozishanus Qi, He, Su &amp; Song, new species (paratype, Kunming, Yunnan); G–H (refer to p. 22 in Huang &amp; Chen, 2017; Gongshan, Yunnan); I–N, L. takeoi Adachi, 2020 (Huidong, Sichuan); O–T, L. boileaui Planet, 1897 (Mabian, Sichuan). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7814605" httpUri="https://zenodo.org/record/7814605/files/figure.png" pageId="17" pageNumber="257">6I–N</figureCitation>
; 7F–J; 13D–F
</paragraph>
</subSubSection>
<subSubSection box="[151,479,1807,1831]" pageId="17" pageNumber="257" type="reference_group">
<paragraph blockId="17.[151,479,1807,1831]" box="[151,479,1807,1831]" pageId="17" pageNumber="257">
<treatmentCitationGroup box="[151,479,1807,1831]" pageId="17" pageNumber="257">
<treatmentCitation author="Adachi, N." box="[151,475,1807,1831]" page="8" pageId="17" pageNumber="257" year="2020">
<taxonomicName ID-CoL="7X2NK" authority="Adachi, 2020: 8" authorityName="Adachi" authorityPageNumber="8" authorityYear="2020" box="[151,475,1807,1831]" class="Insecta" family="Lucanidae" genus="Lucanus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="257" phylum="Arthropoda" rank="species" species="takeoi">
<emphasis box="[151,305,1807,1831]" italics="true" pageId="17" pageNumber="257">Lucanus takeoi</emphasis>
<bibRefCitation author="Adachi, N." box="[310,475,1807,1831]" pageId="17" pageNumber="257" pagination="8 - 10" refId="ref7642" refString="Adachi, N. (2020) A new species of the genus Lucanus from Szechuan province, southwest China. Kogane, 23, 8 - 10." type="journal article" year="2020">Adachi, 2020: 8</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection pageId="17" pageNumber="257" type="materials_examined">
<paragraph blockId="17.[151,1436,1870,2041]" pageId="17" pageNumber="257">
<materialsCitation country="China" county="Liangshan Yi Autonomous Prefecture" location="Tuowu township" municipality="Mianning county" pageId="17" pageNumber="257" specimenCount="1" stateProvince="Sichuan" typeStatus="holotype">
<emphasis bold="true" box="[151,309,1870,1897]" pageId="17" pageNumber="257">
<typeStatus box="[151,210,1871,1897]" pageId="17" pageNumber="257">Type</typeStatus>
locality.
</emphasis>
<collectingCountry box="[315,385,1870,1896]" name="China" pageId="17" pageNumber="257">China</collectingCountry>
, 
<collectingRegion box="[396,595,1870,1896]" country="China" name="Sichuan" pageId="17" pageNumber="257">Sichuan Province</collectingRegion>
, 
<collectingCounty box="[606,1028,1870,1897]" pageId="17" pageNumber="257">Liangshan Yi Autonomous Prefecture</collectingCounty>
, 
<collectingMunicipality box="[1039,1231,1870,1897]" pageId="17" pageNumber="257">Mianning county</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:03B51633FF8FFFDE5FFB58EC0572FDA7:8EC3F1FEFF8FFFDD5BB6590D05DCF8AF" box="[1242,1433,1870,1897]" country="China" county="Liangshan Yi Autonomous Prefecture" municipality="Mianning county" name="Tuowu township" pageId="17" pageNumber="257" stateProvince="Sichuan">Tuowu township</location>
, 
<location LSID="urn:lsid:plazi:treatment:03B51633FF8FFFDE5FFB58EC0572FDA7:8EC3F1FEFF8FFFDD5FFB59300093F84B" box="[151,214,1906,1932]" country="China" county="Liangshan Yi Autonomous Prefecture" municipality="Mianning county" name="Baku" pageId="17" pageNumber="257" stateProvince="Sichuan">Baku</location>
</materialsCitation>
.
</paragraph>
<paragraph blockId="17.[151,1436,1870,2041]" pageId="17" pageNumber="257">
<emphasis bold="true" box="[199,433,1942,1968]" pageId="17" pageNumber="257">Material examined.</emphasis>
<materialsCitation box="[441,908,1942,1971]" collectionCode="CZHQ, CRXS" country="China" location="CHINA" pageId="17" pageNumber="257" specimenCount="1">
<emphasis bold="true" box="[441,544,1942,1968]" pageId="17" pageNumber="257">
<collectingCountry box="[441,539,1942,1968]" name="China" pageId="17" pageNumber="257">CHINA</collectingCountry>
:
</emphasis>
4♁♁ (3♁♁ 
<collectionCode box="[689,768,1943,1969]" pageId="17" pageNumber="257">CZHQ</collectionCode>
, 1♁ 
<collectionCode box="[823,900,1943,1969]" pageId="17" pageNumber="257">CRXS</collectionCode>
)
</materialsCitation>
, 
<materialsCitation box="[920,1062,1943,1969]" collectionCode="CZHQ" country="China" pageId="17" pageNumber="257" specimenCount="1" specimenCount-female="1">
<specimenCount box="[920,955,1943,1969]" pageId="17" pageNumber="257" type="female">1♀</specimenCount>
(
<collectionCode box="[971,1054,1943,1969]" pageId="17" pageNumber="257">CZHQ</collectionCode>
)
</materialsCitation>
: 
<materialsCitation collectingDate="2022-07-15" collectingDateMax="2022-07-20" collectingDateMin="2022-07-15" country="China" county="Huidong County" elevation="2450" location="Jiangzhou Town" municipality="Liangshan Yi Autonomous Prefecture" pageId="17" pageNumber="257" specimenCount="1" stateProvince="Sichuan">
<collectingRegion box="[1074,1274,1942,1968]" country="China" name="Sichuan" pageId="17" pageNumber="257">Sichuan Province</collectingRegion>
, 
<collectingMunicipality pageId="17" pageNumber="257">Liangshan Yi Autonomous Prefecture</collectingMunicipality>
, 
<collectingCounty box="[437,626,1978,2005]" pageId="17" pageNumber="257">Huidong County</collectingCounty>
[
<emphasis bold="true" box="[645,729,1979,2002]" pageId="17" pageNumber="257">ĤŤB</emphasis>
], 
<location LSID="urn:lsid:plazi:treatment:03B51633FF8FFFDE5FFB58EC0572FDA7:8EC3F1FEFF8FFFDD5D9F59F803E9F812" box="[755,940,1978,2005]" country="China" county="Huidong County" municipality="Liangshan Yi Autonomous Prefecture" name="Jiangzhou Town" pageId="17" pageNumber="257" stateProvince="Sichuan">Jiangzhou Town</location>
[
<emphasis bold="true" box="[959,1043,1981,2004]" pageId="17" pageNumber="257">ƊNJDz</emphasis>
], 
<location LSID="urn:lsid:plazi:treatment:03B51633FF8FFFDE5FFB58EC0572FDA7:8EC3F1FEFF8FFFDD5B4059F804EDF813" box="[1068,1192,1978,2004]" country="China" county="Huidong County" municipality="Liangshan Yi Autonomous Prefecture" name="Matoushan" pageId="17" pageNumber="257" stateProvince="Sichuan">Matoushan</location>
[
<emphasis bold="true" box="[1212,1296,1979,2002]" pageId="17" pageNumber="257">ḢĿƜ</emphasis>
]; alt. 
<quantity metricMagnitude="3" metricUnit="m" metricValue="2.45" metricValueMax="2.5" metricValueMin="2.4" pageId="17" pageNumber="257" unit="m" value="2450.0" valueMax="2500.0" valueMin="2400.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="2.45" metricValueMax="2.5" metricValueMin="2.4" pageId="17" pageNumber="257" unit="m" value="2450.0" valueMax="2500.0" valueMin="2400.0">2400– 2500 m</elevation>
</quantity>
, 
<date box="[250,431,2014,2041]" pageId="17" pageNumber="257" value="2022-07-15" valueMax="2022-07-20" valueMin="2022-07-15">
<collectingDate box="[250,431,2014,2041]" pageId="17" pageNumber="257" value="2022-07-15" valueMax="2022-07-20" valueMin="2022-07-15">15–20.VII.2022</collectingDate>
</date>
, local collector leg
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection box="[199,550,150,176]" pageId="18" pageNumber="258" type="distribution">
<paragraph blockId="18.[151,1437,150,608]" box="[199,550,150,176]" pageId="18" pageNumber="258">
<emphasis bold="true" box="[199,353,150,176]" pageId="18" pageNumber="258">Distribution.</emphasis>
<collectingCountry box="[359,426,150,176]" name="China" pageId="18" pageNumber="258">China</collectingCountry>
(
<collectingRegion box="[441,538,150,176]" country="China" name="Sichuan" pageId="18" pageNumber="258">Sichuan</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="258" type="discussion">
<paragraph blockId="18.[151,1437,150,608]" pageId="18" pageNumber="258">
<emphasis bold="true" box="[199,315,186,212]" pageId="18" pageNumber="258">Remarks.</emphasis>
This species should be assigned to the 
<taxonomicName authorityName="Motschulsky" authorityYear="1861" box="[747,1036,186,212]" class="Insecta" family="Lucanidae" genus="Lucanus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="258" phylum="Arthropoda" rank="species" species="maculifemoratus">
<emphasis box="[747,1036,186,212]" italics="true" pageId="18" pageNumber="258">Lucanus maculifemoratus</emphasis>
</taxonomicName>
group (sensu 
<bibRefCitation author="Huang, H. &amp; Chen, C. - C." box="[1190,1425,186,213]" pageId="18" pageNumber="258" refId="ref7824" refString="Huang, H. &amp; Chen, C. - C. (2010) Stag Beetles of China I. Formosa Ecological Company, Taipei, 288 pp." type="book" year="2010">Huang &amp; Chen 2010</bibRefCitation>
). 
<bibRefCitation author="Adachi, N." box="[151,316,222,248]" pageId="18" pageNumber="258" pagination="8 - 10" refId="ref7642" refString="Adachi, N. (2020) A new species of the genus Lucanus from Szechuan province, southwest China. Kogane, 23, 8 - 10." type="journal article" year="2020">Adachi (2020)</bibRefCitation>
described the species on a single male specimen from Baku, Mianning county, 
<collectingRegion box="[1231,1433,222,248]" country="China" name="Sichuan" pageId="18" pageNumber="258">Sichuan Province</collectingRegion>
. The female and host plant of 
<taxonomicName authority="Adachi, 2020" authorityName="Adachi" authorityYear="2020" box="[492,755,258,285]" class="Insecta" family="Lucanidae" genus="Lucanus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="258" phylum="Arthropoda" rank="species" species="takeoi">
<emphasis box="[492,508,259,284]" italics="true" pageId="18" pageNumber="258">L</emphasis>
. 
<emphasis box="[524,592,258,284]" italics="true" pageId="18" pageNumber="258">takeoi</emphasis>
<bibRefCitation author="Adachi, N." box="[602,755,258,284]" pageId="18" pageNumber="258" pagination="8 - 10" refId="ref7642" refString="Adachi, N. (2020) A new species of the genus Lucanus from Szechuan province, southwest China. Kogane, 23, 8 - 10." type="journal article" year="2020">Adachi, 2020</bibRefCitation>
</taxonomicName>
are unknown. In 2021, the natives collected this species in Huidong County (
<collectingRegion box="[353,553,294,320]" country="China" name="Sichuan" pageId="18" pageNumber="258">Sichuan Province</collectingRegion>
) for the first time.
</paragraph>
<paragraph blockId="18.[151,1437,150,608]" pageId="18" pageNumber="258">
In 2022, we obtained some specimens of this species from the local collector in Huidong County, which enabled us to illustrate the female (
<figureCitation box="[460,596,366,393]" captionStart="FIGURE 5" captionStartId="6.[151,250,1756,1780]" captionTargetBox="[164,1418,194,1720]" captionTargetId="figure-19@6.[151,1435,181,1732]" captionTargetPageId="6" captionText="FIGURE 5. Female habitus of Lucanus species. A–B, L. jiaozishanus Qi, He, Su &amp; Song, new species (24.8 mm, paratype, Kunming, Yunnan); C–D, L. takeoi Adachi, 2020 (27.4 mm, Huidong, Sichuan); E–F, L. boileaui Planet, 1897 (35.0 mm, Mabian, Sichuan). A, C, E, dorsal view; B, D, F, ventral view. Scale bar = 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7801118" httpUri="https://zenodo.org/record/7801118/files/figure.png" pageId="18" pageNumber="258">Figs. 5C–D</figureCitation>
; 
<figureCitation box="[609,670,366,392]" captionStart="FIGURE 6" captionStartId="7.[151,250,1871,1895]" captionTargetBox="[151,1433,181,1847]" captionTargetId="figure-17@7.[151,1436,181,1847]" captionTargetPageId="7" captionText="FIGURE 6. Female head (A, G, I, O), clypeolabrum (B, J, P), canthus (C, H, K, Q) and tibia (D–F, L–N, R–T) of Lucanus species in dorsal view. A–F, L. jiaozishanus Qi, He, Su &amp; Song, new species (paratype, Kunming, Yunnan); G–H (refer to p. 22 in Huang &amp; Chen, 2017; Gongshan, Yunnan); I–N, L. takeoi Adachi, 2020 (Huidong, Sichuan); O–T, L. boileaui Planet, 1897 (Mabian, Sichuan). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7814605" httpUri="https://zenodo.org/record/7814605/files/figure.png" pageId="18" pageNumber="258">6I–N</figureCitation>
), the males of different sizes (
<figureCitation box="[1018,1147,366,392]" captionStart="FIGURE 1" captionStartId="2.[151,250,1840,1864]" captionTargetBox="[158,1432,666,1804]" captionTargetId="figure-216@2.[151,1436,666,1814]" captionTargetPageId="2" captionText="FIGURE 1. Male habitus of Lucanus species. A–B, L. jiaozishanus Qi, He, Su &amp; Song, new species (33.3 mm, holotype, large sized, Kunming, Yunnan); C–D, L. ludivinae Boucher, 1998 (31.3 mm, medium sized, Gongshan, Yunnan); E–F, L. takeoi Adachi, 2020 (29.2 mm, small sized, Huidong, Sichuan); G–H, L. boileaui Planet, 1897 (41.9 mm, small sized, Mabian, Sichuan). A, C, E, G, dorsal view; B, D, F, H, ventral view. Scale bar = 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7801110" httpUri="https://zenodo.org/record/7801110/files/figure.png" pageId="18" pageNumber="258">Figs. 1E–F</figureCitation>
; 
<figureCitation box="[1161,1236,366,392]" captionStart="FIGURE 2" captionStartId="3.[151,250,1896,1920]" captionTargetBox="[151,1431,181,1873]" captionTargetId="figure-17@3.[151,1435,181,1873]" captionTargetPageId="3" captionText="FIGURE 2. Male head (A, G, M, S), clypeolabrum (B, H, N, T), canthus (C, I, O, U) and tibia (D–F, J–L, P–R, V–X) of Lucanus species in dorsal view. A–F, L. jiaozishanus Qi, He, Su &amp; Song, new species (holotype, Kunming, Yunnan); G–L, L. ludivinae Boucher, 1998 (Gongshan, Yunnan); M–R, L. takeoi Adachi, 2020 (Huidong, Sichuan); S–X, L. boileaui Planet, 1897 (Mabian, Sichuan). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7814599" httpUri="https://zenodo.org/record/7814599/files/figure.png" pageId="18" pageNumber="258">2M–R</figureCitation>
; 13D–F), and the genitalia of both sexes (
<figureCitation box="[426,549,402,428]" captionStart="FIGURE 3" captionStartId="4.[151,250,1909,1933]" captionTargetBox="[314,1276,189,1873]" captionTargetId="figure-19@4.[307,1280,181,1885]" captionTargetPageId="4" captionText="FIGURE 3. Male abdominal tergite 8 (A, E, I, M), abdominal ventrite 8 (B, F, J, N) and abdominal segment 9 (C–D, G–H, K–L, O–P) of Lucanus species. A–D, L. jiaozishanus Qi, He, Su &amp; Song, new species (holotype, Kunming, Yunnan); E–H, L. ludivinae Boucher, 1998 (Gongshan, Yunnan); I–L, L. takeoi Adachi, 2020 (Huidong, Sichuan); M–P, L. boileaui Planet, 1897 (Mabian, Sichuan). A, D, E, H, I, L, M, P, dorsal view; B–C, F–G, J–K, N–O, ventral view. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7814603" httpUri="https://zenodo.org/record/7814603/files/figure.png" pageId="18" pageNumber="258">Figs. 3I–L</figureCitation>
; 4C, M–P; 7F–J). In addition, we found that the female of the species has an almost completely sclerotized spermathecal duct (
<figureCitation box="[724,816,438,464]" captionStart="FIGURE 7" captionStartId="8.[151,250,970,994]" captionTargetBox="[177,1411,181,946]" captionTargetId="figure-19@8.[177,1411,181,946]" captionTargetPageId="8" captionText="FIGURE 7. Female abdominal tergite 8 (A, F, K), abdominal ventrite 8 (B, G, L) and genitalia (C, H, M) including details of hemisternite (D, I, N), spermatheca and spermathecal gland (E, J, O) of Lucanus species. A–E, L. jiaozishanus Qi, He, Su &amp; Song, new species (paratype, Kunming, Yunnan); F–J, L. takeoi Adachi, 2020 (Huidong, Sichuan); K–O, L. boileaui Planet, 1897 (Mabian, Sichuan). A, F, K, dorsal view; B–E, G–J, L–O, ventral view. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7801120" httpUri="https://zenodo.org/record/7801120/files/figure.png" pageId="18" pageNumber="258">Fig. 7H</figureCitation>
), which is a feature distinct from other species of the same species-group. The adults of 
<taxonomicName authorityName="Adachi" authorityYear="2020" box="[541,639,474,501]" class="Insecta" family="Lucanidae" genus="Lucanus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="258" phylum="Arthropoda" rank="species" species="takeoi">
<emphasis box="[541,557,475,500]" italics="true" pageId="18" pageNumber="258">L</emphasis>
. 
<emphasis box="[571,639,474,500]" italics="true" pageId="18" pageNumber="258">takeoi</emphasis>
</taxonomicName>
mainly inhabit the 
<taxonomicName box="[858,1006,475,501]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="18" pageNumber="258" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis box="[858,952,475,501]" italics="true" pageId="18" pageNumber="258">Quercus</emphasis>
spp.
</taxonomicName>
(
<taxonomicName authorityName="Dumortier" authorityYear="1829" box="[1020,1129,474,500]" class="Magnoliopsida" family="Fagaceae" kingdom="Plantae" order="Fagales" pageId="18" pageNumber="258" phylum="Tracheophyta" rank="family">Fagaceae</taxonomicName>
, 
<figureCitation box="[1140,1276,474,500]" captionStart="FIGURE 14" captionStartId="16.[151,250,1897,1921]" captionTargetBox="[158,1430,181,1873]" captionTargetId="figure-19@16.[158,1430,181,1873]" captionTargetPageId="16" captionText="FIGURE 14. Host plants of Lucanus species (L. takeoi Adachi, 2020, L. moae Qi, 2021, L. delavayi Fairmaire, 1887, L. thibetanus thibetanus Planet, 1898 and L. kraatzi kraatzi Nagel, 1926) at an altitude of about 2400 m in Huidong, Sichuan (Photo by local collector). A, C, Quercus spp. in the wild; B, D, details of Quercus spp. leaves (mating L. kraatzi kraatzi in red circle)." figureDoi="http://doi.org/10.5281/zenodo.7801131" httpUri="https://zenodo.org/record/7801131/files/figure.png" pageId="18" pageNumber="258">Fig. 14A–D</figureCitation>
) at an altitude about 
<quantity box="[220,305,510,537]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="18" pageNumber="258" unit="m" value="2400.0">2400 m</quantity>
in Huidong County and can be found from June to August. These plants are also the host plants of 
<taxonomicName authority="Fairmaire, 1887" authorityName="Fairmaire" authorityYear="1887" class="Insecta" family="Lucanidae" genus="Lucanus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="258" phylum="Arthropoda" rank="species" species="delavayi">
<emphasis box="[1414,1430,511,536]" italics="true" pageId="18" pageNumber="258">L</emphasis>
. 
<emphasis box="[151,246,546,572]" italics="true" pageId="18" pageNumber="258">delavayi</emphasis>
Fairmaire, 1887
</taxonomicName>
, 
<taxonomicName authority="Nagel, 1926" authorityName="Nagel" authorityYear="1926" box="[445,780,546,573]" class="Insecta" family="Lucanidae" genus="Lucanus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="258" phylum="Arthropoda" rank="subSpecies" species="kraatzi" subSpecies="kraatzi">
<emphasis box="[445,461,547,572]" italics="true" pageId="18" pageNumber="258">L</emphasis>
. 
<emphasis box="[474,637,546,572]" italics="true" pageId="18" pageNumber="258">kraatzi kraatzi</emphasis>
Nagel, 1926
</taxonomicName>
(
<figureCitation box="[795,898,546,572]" captionStart="FIGURE 14" captionStartId="16.[151,250,1897,1921]" captionTargetBox="[158,1430,181,1873]" captionTargetId="figure-19@16.[158,1430,181,1873]" captionTargetPageId="16" captionText="FIGURE 14. Host plants of Lucanus species (L. takeoi Adachi, 2020, L. moae Qi, 2021, L. delavayi Fairmaire, 1887, L. thibetanus thibetanus Planet, 1898 and L. kraatzi kraatzi Nagel, 1926) at an altitude of about 2400 m in Huidong, Sichuan (Photo by local collector). A, C, Quercus spp. in the wild; B, D, details of Quercus spp. leaves (mating L. kraatzi kraatzi in red circle)." figureDoi="http://doi.org/10.5281/zenodo.7801131" httpUri="https://zenodo.org/record/7801131/files/figure.png" pageId="18" pageNumber="258">Fig. 14D</figureCitation>
), 
<taxonomicName authority="Qi, 2021" authorityName="Qi" authorityYear="2021" box="[918,1113,546,573]" class="Insecta" family="Lucanidae" genus="Lucanus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="258" phylum="Arthropoda" rank="species" species="moae">
<emphasis box="[918,934,547,572]" italics="true" pageId="18" pageNumber="258">L</emphasis>
. 
<emphasis box="[948,1009,547,572]" italics="true" pageId="18" pageNumber="258">moae</emphasis>
<bibRefCitation author="Qi, Z. - H." box="[1015,1113,546,572]" pageId="18" pageNumber="258" pagination="1 - 7" refId="ref8174" refString="Qi, Z. - H. (2021) Lucanus moae sp. nov., a new species from Sichuan, China (Coleoptera: Lucanidae: Lucaninae). Faunitaxys, 9 (23), 1 - 7." type="journal article" year="2021">Qi, 2021</bibRefCitation>
</taxonomicName>
and 
<taxonomicName authority="Planet, 1898" authorityName="Planet" authorityYear="1898" class="Insecta" family="Lucanidae" genus="Lucanus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="258" phylum="Arthropoda" rank="subSpecies" species="thibetanus" subSpecies="thibetanus">
<emphasis box="[1167,1183,547,572]" italics="true" pageId="18" pageNumber="258">L</emphasis>
. 
<emphasis box="[1196,1437,546,572]" italics="true" pageId="18" pageNumber="258">thibetanus thibetanus</emphasis>
Planet, 
<quantity box="[235,291,582,608]" metricMagnitude="1" metricUnit="m" metricValue="4.820919999999999" pageId="18" pageNumber="258" unit="in" value="1898.0">1898</quantity>
</taxonomicName>
in Huidong County (Zhi-Hao Qi’s personal communication with the local collector in 2022).
</paragraph>
</subSubSection>
</treatment>
</document>