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<document id="CD8615825F11DD8FDF27BA69E778F3C5" ID-CLB-Dataset="46589" ID-DOI="10.11646/zootaxa.3755.4.1" ID-GBIF-Dataset="5106994e-9a64-477c-8340-340b42733531" ID-ISSN="1175-5326" ID-Zenodo-Dep="251910" ID-ZooBank="220B94FD-F265-40B5-BEE8-53DB0194AB35" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460373966255" checkinUser="plazi" docAuthor="Konishi, Kazuhiko" docDate="2014" docId="03AD7C3FD47F4337B6D8FE27FD4B255E" docLanguage="en" docName="zt03755p346.pdf" docOrigin="Zootaxa 3755 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Netelia (Bessobates) longipad Konishi, 2014, n. sp." docType="treatment" docVersion="9" lastPageNumber="322" masterDocId="FF940447D46A4322B64FFFB3FFC32231" masterDocTitle="A revision of the subgenus Bessobates of the genus Netelia from Japan (Hymenoptera, Ichneumonidae, Tryphoninae)" masterLastPageNumber="346" masterPageNumber="301" pageNumber="322" updateTime="1698296137341" updateUser="ExternalLinkService">
<mods:mods id="D19B06DE065E810AAAF2F91F5597AB1F" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="7ED959F05F56B08062D6229F99A41E11">
<mods:title id="AFE56129990383453FFBBC051D811A0D">A revision of the subgenus Bessobates of the genus Netelia from Japan (Hymenoptera, Ichneumonidae, Tryphoninae)</mods:title>
</mods:titleInfo>
<mods:name id="3753DA6F284DBD28D9DAE2092F1438D2" type="personal">
<mods:role id="8573AED32240D915177EEBAD2242D4EF">
<mods:roleTerm id="48634799F1E837EC38197B60E7837B03">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EBBAFC02FFA3844993D8252AE1991095">Konishi, Kazuhiko</mods:namePart>
</mods:name>
<mods:typeOfResource id="6E727354F3140E061B336C6ECC3E7F74">text</mods:typeOfResource>
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<mods:title id="D3CBC26987E3F96A5430D3C15F819277">Zootaxa</mods:title>
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<mods:part id="F17F37B1B760DEED71B9CDFB16154EA5">
<mods:date id="F515DE08B17FF6F907AB77D58C4CD65F">2014</mods:date>
<mods:detail id="C54898383B89F865214981D8C8077566" type="volume">
<mods:number id="E4DD5F928DB2743E7CBF11BB499F4835">3755</mods:number>
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<mods:detail id="CE0256B8C9FD353A6493EC76F062E61C" type="issue">
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<mods:classification id="0D558CF847C08CF8CD09A24205105AD1">journal article</mods:classification>
<mods:identifier id="80134FE1A2091EB54AD5E7CCD7C8070B" type="CLB-Dataset">46589</mods:identifier>
<mods:identifier id="4D5B1BEAF36EB0201614FE36D7752FF8" type="DOI">10.11646/zootaxa.3755.4.1</mods:identifier>
<mods:identifier id="4C4523FF98E5F42674E80CCAFBFD14E3" type="GBIF-Dataset">5106994e-9a64-477c-8340-340b42733531</mods:identifier>
<mods:identifier id="D127633624C282A8298BAB0F59F523EC" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="85679B5C601C99E1C8412A464D06DAC8" type="Zenodo-Dep">251910</mods:identifier>
<mods:identifier id="806D23248E6A29D0B796B14D76B99CB3" type="ZooBank">220B94FD-F265-40B5-BEE8-53DB0194AB35</mods:identifier>
</mods:mods>
<treatment id="03AD7C3FD47F4337B6D8FE27FD4B255E" ID-DOI="http://doi.org/10.5281/zenodo.6140819" ID-GBIF-Taxon="119412772" ID-Zenodo-Dep="6140819" LSID="urn:lsid:plazi:treatment:03AD7C3FD47F4337B6D8FE27FD4B255E" httpUri="http://treatment.plazi.org/id/03AD7C3FD47F4337B6D8FE27FD4B255E" lastPageNumber="322" pageId="21" pageNumber="322">
<subSubSection id="C31E9EA2D47F4337B6D8FE27FEA3278E" pageId="21" pageNumber="322" type="nomenclature">
<paragraph id="8BBBCD29D47F4337B6D8FE27FD84239F" blockId="21.[151,583,404,463]" box="[151,583,404,430]" pageId="21" pageNumber="322">
<heading id="D0F37A45D47F4337B6D8FE27FD84239F" bold="true" box="[151,583,404,430]" fontSize="11" level="1" pageId="21" pageNumber="322" reason="1">
<emphasis id="B970113BD47F4337B6D8FE27FD84239F" bold="true" box="[151,583,404,430]" pageId="21" pageNumber="322">
<taxonomicName id="4C04B6AAD47F4337B6D8FE27FE3E239F" box="[151,509,404,430]" class="Insecta" family="Ichneumonidae" genus="Netelia" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="322" phylum="Arthropoda" rank="species" species="longipad" status="sp. nov." subGenus="Bessobates">
<emphasis id="B970113BD47F4337B6D8FE27FF2E239F" bold="true" box="[151,237,404,430]" italics="true" pageId="21" pageNumber="322">Netelia</emphasis>
(
<emphasis id="B970113BD47F4337B6B1FE27FE41239F" bold="true" box="[254,386,404,430]" italics="true" pageId="21" pageNumber="322">Bessobates</emphasis>
)
<emphasis id="B970113BD47F4337B7DCFE27FE3E239F" bold="true" box="[403,509,404,430]" italics="true" pageId="21" pageNumber="322">longipad</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A243AC40D47F4337B44BFE27FD84239F" box="[516,583,404,430]" pageId="21" pageNumber="322" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BBBCD29D47F4337B6D8FE05FEB823FE" blockId="21.[151,583,404,463]" box="[151,379,438,463]" pageId="21" pageNumber="322">
(
<figureCitation id="133FD1ACD47F4337B6D0FE05FEE323FE" box="[159,288,438,463]" captionStart="FIGURES 53 – 57" captionStartId="22.[151,264,1958,1981]" captionTargetBox="[240,1346,193,1906]" captionTargetId="figure@22.[240,1346,193,1938]" captionTargetPageId="22" captionText="FIGURES 53 – 57. Male of Netelia (Bessobates) longipad. 53, Head in dorsal view; 54, head in frontal view; 55, mesosoma in dorsal view; 56, fore wing; 57, hind wing." httpUri="https://zenodo.org/record/251922/files/figure.png" pageId="21" pageNumber="322">Figs 53–57</figureCitation>
,
<figureCitation id="133FD1ACD47F4337B764FE05FEB023FE" box="[299,371,438,463]" captionStart="FIGURES 63 – 68" captionStartId="25.[151,264,1405,1428]" captionTargetBox="[170,1425,195,1385]" captionTargetId="figure@25.[163,1428,193,1390]" captionTargetPageId="25" captionText="FIGURES 63 – 68. Male subgenital plate and genitalia of Netelia (Bessobates) spp. 63 – 65, Netelia (Bessobates) longipad; 66 – 68, N. (B.) nanaki; 63, 66, subgenital plate; 64, 67, paramere in mesal view; 65, 68, aedeagus in lateral view." httpUri="https://zenodo.org/record/251924/files/figure.png" pageId="21" pageNumber="322">63–65</figureCitation>
)
</paragraph>
<paragraph id="8BBBCD29D47F4337B6D8FE4CFC3820FA" blockId="21.[151,1436,510,1903]" pageId="21" pageNumber="322">
♂. Head (
<figureCitation id="133FD1ACD47F4337B74CFE4DFE412026" box="[259,386,510,535]" captionStart="FIGURES 53 – 57" captionStartId="22.[151,264,1958,1981]" captionTargetBox="[240,1346,193,1906]" captionTargetId="figure@22.[240,1346,193,1938]" captionTargetPageId="22" captionText="FIGURES 53 – 57. Male of Netelia (Bessobates) longipad. 53, Head in dorsal view; 54, head in frontal view; 55, mesosoma in dorsal view; 56, fore wing; 57, hind wing." httpUri="https://zenodo.org/record/251922/files/figure.png" pageId="21" pageNumber="322">Figs 53, 54</figureCitation>
) 1.7–1.8 times as wide as long; vertex in dorsal view evenly rounded; occiput and interocellar area impunctate and polished; vertex and gena weakly and finely punctate; geno-orbital index 2.0–2.2; frons impunctate and finely coriaceous, with a weak median longitudinal groove; face about 1.4 times as wide as long, rather more strongly punctate than vertex; clypeus 2.1–2.3 times as wide as long, rather more sparsely punctate than face, with apical margin truncate and slightly concave medially; antenna with 50–57 flagellomeres, 1st flagellomere 1.5–1.6 times as long as 2nd, 2nd 2.5–2.6 times as long as wide.
</paragraph>
<paragraph id="8BBBCD29D47F4337B688FD64FC272106" blockId="21.[151,1436,510,1903]" pageId="21" pageNumber="322">
Mesosoma (
<figureCitation id="133FD1ACD47F4337B71EFD65FE6520DE" box="[337,422,726,751]" captionStart="FIGURES 53 – 57" captionStartId="22.[151,264,1958,1981]" captionTargetBox="[240,1346,193,1906]" captionTargetId="figure@22.[240,1346,193,1938]" captionTargetPageId="22" captionText="FIGURES 53 – 57. Male of Netelia (Bessobates) longipad. 53, Head in dorsal view; 54, head in frontal view; 55, mesosoma in dorsal view; 56, fore wing; 57, hind wing." httpUri="https://zenodo.org/record/251922/files/figure.png" pageId="21" pageNumber="322">Fig. 55</figureCitation>
): mesoscutum 1.2–1.3 times as long as wide; scutellum 1.4–1.6 times as long as distance between lateral carinae at its base, with lateral carina fading out before basal 1/3 of scutellum; propodeum transstriate on about basal 0.7, with sublateral crest present and weakly carinate.
</paragraph>
<paragraph id="8BBBCD29D47F4337B688FCF0FCEA214E" blockId="21.[151,1436,510,1903]" pageId="21" pageNumber="322">Fore tibial spur 0.4–0.5 times as long as 1st tarsomere, which is 8.5–9.8 times as long as wide; fore tarsal claw with 21–26 pectines; hind tarsal claw with 16–20 pectines.</paragraph>
<paragraph id="8BBBCD29D47F4337B688FC38FC5921F6" blockId="21.[151,1436,510,1903]" pageId="21" pageNumber="322">
Fore wing (
<figureCitation id="133FD1ACD47F4337B703FC39FE5C2192" box="[332,415,906,931]" captionStart="FIGURES 53 – 57" captionStartId="22.[151,264,1958,1981]" captionTargetBox="[240,1346,193,1906]" captionTargetId="figure@22.[240,1346,193,1938]" captionTargetPageId="22" captionText="FIGURES 53 – 57. Male of Netelia (Bessobates) longipad. 53, Head in dorsal view; 54, head in frontal view; 55, mesosoma in dorsal view; 56, fore wing; 57, hind wing." httpUri="https://zenodo.org/record/251922/files/figure.png" pageId="21" pageNumber="322">Fig. 56</figureCitation>
) with areolet pointed or petiolate above; cu-a opposite Rs&amp;M, almost straight, inclivous; hind wing (
<figureCitation id="133FD1ACD47F4337B758FC1DFEAB21F6" box="[279,360,942,967]" captionStart="FIGURES 53 – 57" captionStartId="22.[151,264,1958,1981]" captionTargetBox="[240,1346,193,1906]" captionTargetId="figure@22.[240,1346,193,1938]" captionTargetPageId="22" captionText="FIGURES 53 – 57. Male of Netelia (Bessobates) longipad. 53, Head in dorsal view; 54, head in frontal view; 55, mesosoma in dorsal view; 56, fore wing; 57, hind wing." httpUri="https://zenodo.org/record/251922/files/figure.png" pageId="21" pageNumber="322">Fig. 57</figureCitation>
) with 7–11 distal hamuli; nervellar index 0.7–0.9.
</paragraph>
<paragraph id="8BBBCD29D47F4337B688FC60FF3426F2" blockId="21.[151,1436,510,1903]" pageId="21" pageNumber="322">
Metasoma with 1st tergum 2.4–2.8 times as long as apical width and 1.4–1.5 times as long as 2nd; subgenital plate (
<figureCitation id="133FD1ACD47F4337B691FC45FEF2263E" box="[222,305,1014,1039]" captionStart="FIGURES 63 – 68" captionStartId="25.[151,264,1405,1428]" captionTargetBox="[170,1425,195,1385]" captionTargetId="figure@25.[163,1428,193,1390]" captionTargetPageId="25" captionText="FIGURES 63 – 68. Male subgenital plate and genitalia of Netelia (Bessobates) spp. 63 – 65, Netelia (Bessobates) longipad; 66 – 68, N. (B.) nanaki; 63, 66, subgenital plate; 64, 67, paramere in mesal view; 65, 68, aedeagus in lateral view." httpUri="https://zenodo.org/record/251924/files/figure.png" pageId="21" pageNumber="322">Fig. 63</figureCitation>
) short with apical margin weakly convex; paramere (
<figureCitation id="133FD1ACD47F4337B5DDFC45FC26263E" box="[914,997,1014,1039]" captionStart="FIGURES 63 – 68" captionStartId="25.[151,264,1405,1428]" captionTargetBox="[170,1425,195,1385]" captionTargetId="figure@25.[163,1428,193,1390]" captionTargetPageId="25" captionText="FIGURES 63 – 68. Male subgenital plate and genitalia of Netelia (Bessobates) spp. 63 – 65, Netelia (Bessobates) longipad; 66 – 68, N. (B.) nanaki; 63, 66, subgenital plate; 64, 67, paramere in mesal view; 65, 68, aedeagus in lateral view." httpUri="https://zenodo.org/record/251924/files/figure.png" pageId="21" pageNumber="322">Fig. 64</figureCitation>
) with subapical portions of dorsal and ventral margins slightly emarginate; hairs on lamina volsellaris small; digitus truncate with corners rounded and ventro-apical corner without teeth; membrane of median surface without papillae; pad long, extending anteriorly and reaching apical 2/3 of paramere; subapical sclerotized plate not produced to form an overhang; penis valve (
<figureCitation id="133FD1ACD47F4337B6D0FB35FF3326AE" box="[159,240,1158,1183]" captionStart="FIGURES 63 – 68" captionStartId="25.[151,264,1405,1428]" captionTargetBox="[170,1425,195,1385]" captionTargetId="figure@25.[163,1428,193,1390]" captionTargetPageId="25" captionText="FIGURES 63 – 68. Male subgenital plate and genitalia of Netelia (Bessobates) spp. 63 – 65, Netelia (Bessobates) longipad; 66 – 68, N. (B.) nanaki; 63, 66, subgenital plate; 64, 67, paramere in mesal view; 65, 68, aedeagus in lateral view." httpUri="https://zenodo.org/record/251924/files/figure.png" pageId="21" pageNumber="322">Fig. 65</figureCitation>
) slightly broadened toward apex and slightly curved ventrally; basal apodeme of aedeagus strongly curved dorsally.
</paragraph>
<paragraph id="8BBBCD29D47F4337B688FB7DFD592746" blockId="21.[151,1436,510,1903]" pageId="21" pageNumber="322">Head yellow; occiput and interocellar area light brown; antenna reddish-brown. Mesosoma yellow; mesoscutum, scutellum, postscutellum, irregular markings on mesopleuron and mesosternum light brown. Fore and middle legs brownish-yellow; hind leg light brown except tarsus whitish-yellow. Wings hyaline; basal 2/3 tinged with yellow and distal 1/3 tinged with gray; vein C and pterostigma of fore wing brownish-yellow, other veins blackish-brown. Metasoma light brown.</paragraph>
<paragraph id="8BBBCD29D47F4337B688FA31FDA227AB" blockId="21.[151,1436,510,1903]" box="[199,609,1410,1435]" pageId="21" pageNumber="322">Length of fore wing: 13.2–17.0 mm.</paragraph>
<paragraph id="8BBBCD29D47F4337B688FA14FEA3278E" blockId="21.[151,1436,510,1903]" box="[199,352,1446,1471]" pageId="21" pageNumber="322">♀. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C31E9EA2D47F4337B688FA79FD6D2442" pageId="21" pageNumber="322" type="materials_examined">
<paragraph id="8BBBCD29D47F4337B688FA79FC2527D2" blockId="21.[151,1436,510,1903]" box="[199,998,1482,1507]" pageId="21" pageNumber="322">
<typeStatus id="54BF738BD47F4337B688FA79FEF727D2" box="[199,308,1482,1507]" pageId="21" pageNumber="322" type="holotype">Holotype</typeStatus>
: ♂, Odaigahara, Nara Pref.,
<date id="FFBAEBE9D47F4337B43EFA78FCEB27D2" box="[625,808,1482,1507]" pageId="21" pageNumber="322" value="1983-07-13" valueMax="1983-07-14" valueMin="1983-07-13">13–14. vii. 1983</date>
(S. Ito),
<collectionCode id="ED1555ECD47F4337B5C3FA78FC2127D2" box="[908,994,1483,1507]" httpUri="http://grbio.org/cool/h0ih-38y8" name="National Institute for Agro-Environmental Sciences" pageId="21" pageNumber="322">NIAES</collectionCode>
.
</paragraph>
<paragraph id="8BBBCD29D47F4337B688FA5CFD6D2442" blockId="21.[151,1436,510,1903]" pageId="21" pageNumber="322">
<typeStatus id="54BF738BD47F4337B688FA5CFEFB2436" box="[199,312,1519,1543]" pageId="21" pageNumber="322" type="paratype">Paratypes</typeStatus>
: [Honshu] Nara Pref.:
<specimenCount id="9D0206A0D47F4337B405FA5CFDAE2436" box="[586,621,1519,1543]" pageId="21" pageNumber="322" type="generic">3♂</specimenCount>
(
<date id="FFBAEBE9D47F4337B4CEFA5CFC842436" box="[641,839,1518,1543]" pageId="21" pageNumber="322" value="1983-07-13" valueMax="1983-07-14" valueMin="1983-07-13">13–14. vii. 1983</date>
, S. Ito) &amp;
<specimenCount id="9D0206A0D47F4337B59FFA5CFC302436" box="[976,1011,1519,1543]" pageId="21" pageNumber="322" type="generic">1♂</specimenCount>
(
<date id="FFBAEBE9D47F4337B247FA5DFB582436" box="[1032,1179,1518,1543]" pageId="21" pageNumber="322" value="1984-06-25">25. vi. 1984</date>
, K. Konishi at light), Odaigahara,
<collectionCode id="ED1555ECD47F4337B763F9A0FE41241A" box="[300,386,1555,1579]" httpUri="http://grbio.org/cool/h0ih-38y8" name="National Institute for Agro-Environmental Sciences" pageId="21" pageNumber="322">NIAES</collectionCode>
.
<collectionCode id="ED1555ECD47F4337B7DCF9A0FE1C241A" box="[403,479,1555,1579]" httpUri="http://grbio.org/cool/vy23-a9cj" name="Museum of Nature and Human Activities" pageId="21" pageNumber="322">Hyogo</collectionCode>
Pref.:
<specimenCount id="9D0206A0D47F4337B477F9A0FD9F241A" box="[568,604,1555,1579]" pageId="21" pageNumber="322" type="generic">1♂</specimenCount>
, Kobe,
<date id="FFBAEBE9D47F4337B48FF9A1FC8A241A" box="[704,841,1554,1579]" pageId="21" pageNumber="322" value="1959-05-26">26. v. 1959</date>
(T. Naito),
<collectionCode id="ED1555ECD47F4337B590F9A0FBFB241A" box="[991,1080,1555,1579]" pageId="21" pageNumber="322">MNHA</collectionCode>
. [Shikoku]
<collectingCountry id="F3138DB9D47F4337B288F9A1FAC8241A" box="[1223,1291,1554,1579]" name="India" pageId="21" pageNumber="322">Kochi</collectingCountry>
Pref.:
<specimenCount id="9D0206A0D47F4337B32CF9A0FA5B241A" box="[1379,1432,1555,1579]" pageId="21" pageNumber="322" type="generic">22♂</specimenCount>
, Teragawa, Hongawa,
<date id="FFBAEBE9D47F4337B7DEF984FDDB247E" box="[401,536,1590,1615]" pageId="21" pageNumber="322" value="1995-07-01">1. vii. 1995</date>
(K. Konishi, at light),
<collectionCode id="ED1555ECD47F4337B567F984FCBD247E" box="[808,894,1591,1615]" httpUri="http://grbio.org/cool/h0ih-38y8" name="National Institute for Agro-Environmental Sciences" pageId="21" pageNumber="322">NIAES</collectionCode>
. [Kyushu] Oita Pref.:
<specimenCount id="9D0206A0D47F4337B2CCF984FB64247E" box="[1155,1191,1591,1615]" pageId="21" pageNumber="322" type="generic">1♂</specimenCount>
, Mt. Kurodake, Mts. Kuju,
<date id="FFBAEBE9D47F4337B692F9E8FE962442" box="[221,341,1626,1651]" pageId="21" pageNumber="322" value="1985-06-08">8. vi. 1985</date>
(K. Konishi, at light),
<collectionCode id="ED1555ECD47F4337B41BF9E8FD692442" box="[596,682,1627,1651]" httpUri="http://grbio.org/cool/h0ih-38y8" name="National Institute for Agro-Environmental Sciences" pageId="21" pageNumber="322">NIAES</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="C31E9EA2D47F4337B688F9CDFD3324A6" box="[199,752,1662,1687]" pageId="21" pageNumber="322" type="distribution">
<paragraph id="8BBBCD29D47F4337B688F9CDFD3324A6" blockId="21.[151,1436,510,1903]" box="[199,752,1662,1687]" pageId="21" pageNumber="322">
<emphasis id="B970113BD47F4337B688F9CDFEA324A6" bold="true" box="[199,352,1662,1687]" pageId="21" pageNumber="322">Distribution.</emphasis>
<collectingCountry id="F3138DB9D47F4337B728F9CCFE6424A7" box="[359,423,1663,1686]" name="Japan" pageId="21" pageNumber="322">Japan</collectingCountry>
(Honshu, Shikoku, Kyushu).
</paragraph>
</subSubSection>
<subSubSection id="C31E9EA2D47F4337B688F911FC36248B" box="[199,1013,1698,1723]" pageId="21" pageNumber="322" type="etymology">
<paragraph id="8BBBCD29D47F4337B688F911FC36248B" blockId="21.[151,1436,510,1903]" box="[199,1013,1698,1723]" pageId="21" pageNumber="322">
<emphasis id="B970113BD47F4337B688F911FE8D248A" bold="true" box="[199,334,1698,1723]" pageId="21" pageNumber="322">Etymology.</emphasis>
The specific name is based on the long pad of the paramere.
</paragraph>
</subSubSection>
<subSubSection id="C31E9EA2D47F4337B688F975FD4B255E" pageId="21" pageNumber="322" type="discussion">
<paragraph id="8BBBCD29D47F4337B688F975FD4B255E" blockId="21.[151,1436,510,1903]" pageId="21" pageNumber="322">
<emphasis id="B970113BD47F4337B688F975FEF824EE" bold="true" box="[199,315,1734,1759]" pageId="21" pageNumber="322">Remarks.</emphasis>
This new species can easily be distinguished from other species of
<emphasis id="B970113BD47F4337B22BF975FB2224EE" box="[1124,1249,1734,1759]" italics="true" pageId="21" pageNumber="322">Bessobates</emphasis>
by the strongly produced and elongate pad. This species can not be assigned to any species group, because of the following combination of characters: vertex evenly rounded in dorsal view; lateral scutellar carina fading out before middle of scutellum; sublateral crest of propodeum present; digitus truncate, without teeth on ventro-apical corner; pad membranous; penis valve moderately broad.
</paragraph>
</subSubSection>
</treatment>
</document>
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