treatments-xml/data/03/9F/FD/039FFD65D11FFFDA32DF5831FE5B3CB7.xml

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<mods:title id="998E257C17294DCDDFC736E6A688BAEC">Schistura thavonei, a new species of loach from northwestern Laos (Teleostei: Nemacheilidae)</mods:title>
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<taxonomicName id="4C3637F0D11FFFDC32DF5831FE303D05" authority="Kottelat, 2017" authorityName="Kottelat" authorityYear="2017" box="[254,463,1582,1606]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="396" phylum="Chordata" rank="species" species="thavonei" status="sp. nov.">
<emphasis id="B9429061D11FFFDC32DF5831FE303D05" bold="true" box="[254,463,1582,1606]" italics="true" pageId="1" pageNumber="396">Schistura thavonei</emphasis>
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, 
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<paragraph id="8B894C73D11FFFDC33465851FE053D25" blockId="1.[254,611,1582,1638]" box="[359,506,1614,1638]" pageId="1" pageNumber="396">
(
<figureCitation id="130D50F6D11FFFDC334F5851FE283D25" box="[366,471,1614,1638]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="1.[114,149,865,887]" captionStartId-1="2.[136,171,2064,2086]" captionStartId-2="3.[113,148,1210,1232]" captionTargetBox-0="[113,1451,189,850]" captionTargetBox-1="[276,1334,189,2039]" captionTargetBox-2="[113,1451,189,1195]" captionTargetId-0="figure-586@1.[113,1451,189,850]" captionTargetId-1="figure-7@2.[276,1334,189,2039]" captionTargetId-2="figure-427@3.[113,1451,189,1195]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionText-0="Fig. 1. Schistura thavonei, MHNG 2667.078, holotype, 44.0 mm SL; Laos: Louang Namtha Province: Nam Ma." captionText-1="Fig. 2. Schistura thavonei, paratypes; Laos: Louang Namtha Province: Nam Ma; a, CMK 25944, 44.0 mm SL; b, CMK 26054, 41.8 mm SL; c, CMK 26054, 39.5 mm SL; d, CMK 25944, 44.9 mm SL; e, CMK 26066, 44.7 mm SL." captionText-2="Fig. 3. Schistura thavonei, paratypes; Laos: Louang Namtha Province: Nam Ma; a, CMK 26066, 33.7 mm SL; b, CMK 26054, 34.4 mm SL (right side, reversed); c, CMK 25944, 36.6 mm SL." figureDoi-0="http://doi.org/10.5281/zenodo.5357129" figureDoi-1="http://doi.org/10.5281/zenodo.5357132" figureDoi-2="http://doi.org/10.5281/zenodo.5357134" httpUri-0="https://zenodo.org/record/5357129/files/figure.png" httpUri-1="https://zenodo.org/record/5357132/files/figure.png" httpUri-2="https://zenodo.org/record/5357134/files/figure.png" pageId="1" pageNumber="396">Figs. 1–3</figureCitation>
, 
<figureCitation id="130D50F6D11FFFDC33C55851FE0D3D25" box="[484,498,1614,1638]" captionStart="Fig" captionStartId="5.[112,147,549,571]" captionTargetBox="[113,1451,189,534]" captionTargetId="figure-341@5.[113,1451,189,534]" captionTargetPageId="5" captionText="Fig. 6. Schistura thavonei, CMK 25944, paratype, 50.7 mm SL; Laos: Louang Namtha Province: Nam Ma; immediately after fixation (right side, reversed)." figureDoi="http://doi.org/10.5281/zenodo.5357140" httpUri="https://zenodo.org/record/5357140/files/figure.png" pageId="1" pageNumber="396">6</figureCitation>
)
</paragraph>
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<materialsCitation id="3B5E462ED11FFFDC32505891FE3C3C65" ID-GBIF-Occurrence="3340077305" collectingDate="2015-03-25" collectionCode="MHNG" collectorName="M. Kottelat &amp; T. Phommavong" country="Laos" county="Muang Long District" elevation="638" latitude="21.048056" location="Nam Ma Yen" longLatPrecision="21" longitude="101.02112" municipality="Nam Ma watershed" pageId="1" pageNumber="396" specimenCode="MHNG 2667.078" specimenCount="1" stateProvince="Louangnamtha" typeStatus="holotype">
<emphasis id="B9429061D11FFFDC32505891FF1F3DE5" bold="true" box="[113,224,1678,1702]" pageId="1" pageNumber="396">
<typeStatus id="548DF2D1D11FFFDC32505891FF243DE5" box="[113,219,1678,1702]" pageId="1" pageNumber="396" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCode id="DB90E408D11FFFDC32CB5891FE523DE5" box="[234,429,1678,1702]" collectionCode="MHNG" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="1" pageNumber="396" type="Museum">MHNG 2667.078</specimenCode>
, 44.0 mm SL; 
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: 
<collectingRegion id="49F28291D11FFFDC30BF5890FED33D85" country="Laos" name="Louangnamtha" pageId="1" pageNumber="396">Louang Namtha Province</collectingRegion>
: 
<collectingCounty id="62E834FFD11FFFDC331258B0FDED3D85" box="[307,530,1710,1735]" pageId="1" pageNumber="396">Muang Long District</collectingCounty>
: 
<collectingMunicipality id="6BEDD609D11FFFDC303A58B0FD153D85" box="[539,746,1710,1734]" pageId="1" pageNumber="396">Nam Ma watershed</collectingMunicipality>
: 
<location id="8EE91AA8D11FFFDC325158D0FEF83DA5" LSID="urn:lsid:plazi:treatment:039FFD65D11FFFDA32DF5831FE5B3CB7:8EE91AA8D11FFFDC325158D0FEF83DA5" box="[112,263,1743,1766]" country="Laos" county="Muang Long District" latitude="21.048056" longLatPrecision="21" longitude="101.02112" municipality="Nam Ma watershed" name="Nam Ma Yen" pageId="1" pageNumber="396" stateProvince="Louangnamtha">Nam Ma Yen</location>
, a branch of 
<location id="8EE91AA8D11FFFDC33B858D0FE043DA5" LSID="urn:lsid:plazi:treatment:039FFD65D11FFFDA32DF5831FE5B3CB7:8EE91AA8D11FFFDC33B858D0FE043DA5" box="[409,507,1743,1766]" country="Laos" county="Muang Long District" latitude="21.048056" longLatPrecision="21" longitude="101.02112" municipality="Nam Ma watershed" name="Nam Ma" pageId="1" pageNumber="396" stateProvince="Louangnamtha">Nam Ma</location>
, upstream of 
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, 
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<geoCoordinate id="EE022AB4D11FFFDC330A58F1FE483C45" box="[299,439,1773,1798]" degrees="101" direction="east" minutes="01" orientation="longitude" pageId="1" pageNumber="396" precision="15" seconds="16" value="101.02112">101°01′16″E</geoCoordinate>
, 
<quantity id="4CCEE196D11FFFDC33E558F1FDD53C45" box="[452,554,1774,1798]" metricMagnitude="2" metricUnit="m" metricValue="6.38" pageId="1" pageNumber="396" unit="m" value="638.0">
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; 
<collectorName id="26C329A5D11FFFDC301758F0FD4C3C45" box="[566,691,1774,1798]" pageId="1" pageNumber="396">M. Kottelat</collectorName>
&amp; 
<collectorName id="26C329A5D11FFFDC30F958F0FEF73C65" pageId="1" pageNumber="396">T. Phommavong</collectorName>
, 
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<collectingDate id="EFCC935BD11FFFDC33345911FE403C65" box="[277,447,1806,1830]" pageId="1" pageNumber="396" value="2015-03-25">25 March 2015</collectingDate>
</date>
.
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</paragraph>
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<materialsCitation id="3B5E462ED11FFFDC32505951FD563C25" ID-GBIF-Occurrence="3340077301" accessionNumber="CMK 26066" box="[113,681,1870,1894]" collectionCode="CMK" pageId="1" pageNumber="396" specimenCode="30" specimenCount="1" typeStatus="paratype">
<emphasis id="B9429061D11FFFDC32505951FF0D3C25" bold="true" box="[113,242,1870,1894]" pageId="1" pageNumber="396">
<typeStatus id="548DF2D1D11FFFDC32505951FF123C25" box="[113,237,1870,1894]" pageId="1" pageNumber="396" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectionCode id="ED27D4B6D11FFFDC32DF5951FEC23C25" box="[254,317,1870,1894]" pageId="1" pageNumber="396">CMK</collectionCode>
26066, 
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, 
<quantity id="4CCEE196D11FFFDC33F05951FD863C25" box="[465,633,1870,1894]" metricMagnitude="-2" metricUnit="m" metricValue="3.965" metricValueMax="4.76" metricValueMin="3.17" pageId="1" pageNumber="396" unit="mm" value="39.65" valueMax="47.6" valueMin="31.7">31.7–47.6 mm</quantity>
SL
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; 
<materialsCitation id="3B5E462ED11FFFDC30985951FD383CC5" ID-GBIF-Occurrence="3340077302" accessionNumber="ZRC 56174" collectingDate="2015-03-25" collectionCode="ZRC" collectorName="M. Kottelat &amp; T. Phommavong" country="Laos" county="Muang Long District" elevation="638" latitude="21.048056" location="Nam Ma Yen" longLatPrecision="21" longitude="101.02112" municipality="Nam Ma watershed" pageId="1" pageNumber="396" specimenCode="5, 35.1" specimenCount="1" stateProvince="Louangnamtha" typeStatus="paratype">
<collectionCode id="ED27D4B6D11FFFDC30985951FD113C25" box="[697,750,1870,1894]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="396">ZRC</collectionCode>
56174, 
<specimenCode id="DB90E408D11FFFDC32E95971FF283CC5" box="[200,215,1902,1926]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="396">5</specimenCode>
, 
<specimenCode id="DB90E408D11FFFDC32C95971FEE93CC5" box="[232,278,1902,1926]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="396">35.1</specimenCode>
–43.0 mm SL; same data as holotype
</materialsCitation>
. — 
<materialsCitation id="3B5E462ED11FFFDC32505991FDFC3365" ID-GBIF-Occurrence="3340077306" accessionNumber="CMK 25944" collectingDate="2015-03-21" collectionCode="CMK" collectorName="M. Kottelat &amp; T. Phommavong" country="Laos" county="Muang Long District" elevation="670" latitude="21.081945" location="Nam Ma Oun" longLatPrecision="21" longitude="101.06028" municipality="Nam Ma watershed" pageId="1" pageNumber="396" specimenCode="10" specimenCount="1" stateProvince="Louangnamtha" typeStatus="paratype">
<collectionCode id="ED27D4B6D11FFFDC32505991FF513CE5" box="[113,174,1934,1958]" pageId="1" pageNumber="396">CMK</collectionCode>
25944, 
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, 
<quantity id="4CCEE196D11FFFDC330E5991FE353CE5" box="[303,458,1934,1958]" metricMagnitude="-2" metricUnit="m" metricValue="4.345" metricValueMax="5.07" metricValueMin="3.62" pageId="1" pageNumber="396" unit="mm" value="43.45" valueMax="50.7" valueMin="36.2">36.2–50.7 mm</quantity>
SL; 
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: 
<collectingRegion id="49F28291D11FFFDC30635990FF2B3C85" country="Laos" name="Louangnamtha" pageId="1" pageNumber="396">Louang Namtha Province</collectingRegion>
: 
<collectingCounty id="62E834FFD11FFFDC32C159B0FE353C85" box="[224,458,1966,1991]" pageId="1" pageNumber="396">Muang Long District</collectingCounty>
: 
<collectingMunicipality id="6BEDD609D11FFFDC33F759B0FD503C85" box="[470,687,1966,1990]" pageId="1" pageNumber="396">Nam Ma watershed</collectingMunicipality>
: 
<location id="8EE91AA8D11FFFDC309D59B0FF2E3CA5" LSID="urn:lsid:plazi:treatment:039FFD65D11FFFDA32DF5831FE5B3CB7:8EE91AA8D11FFFDC309D59B0FF2E3CA5" country="Laos" county="Muang Long District" latitude="21.081945" longLatPrecision="21" longitude="101.06028" municipality="Nam Ma watershed" name="Nam Ma Oun" pageId="1" pageNumber="396" stateProvince="Louangnamtha">Nam Ma Oun</location>
, a branch of 
<location id="8EE91AA8D11FFFDC334359D0FE3B3CA5" LSID="urn:lsid:plazi:treatment:039FFD65D11FFFDA32DF5831FE5B3CB7:8EE91AA8D11FFFDC334359D0FE3B3CA5" box="[354,452,1999,2022]" country="Laos" county="Muang Long District" latitude="21.081945" longLatPrecision="21" longitude="101.06028" municipality="Nam Ma watershed" name="Nam Ma" pageId="1" pageNumber="396" stateProvince="Louangnamtha">Nam Ma</location>
, downstream of 
<location id="8EE91AA8D11FFFDC305A59D0FF443345" LSID="urn:lsid:plazi:treatment:039FFD65D11FFFDA32DF5831FE5B3CB7:8EE91AA8D11FFFDC305A59D0FF443345" country="Laos" county="Muang Long District" latitude="21.081945" longLatPrecision="21" longitude="101.06028" municipality="Nam Ma watershed" name="Ban Phang Thong" pageId="1" pageNumber="396" stateProvince="Louangnamtha">Ban Phang Thong</location>
, 
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, 
<quantity id="4CCEE196D11FFFDC33D459F1FDA03345" box="[501,607,2030,2054]" metricMagnitude="2" metricUnit="m" metricValue="6.7" pageId="1" pageNumber="396" unit="m" value="670.0">
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; 
<collectorName id="26C329A5D11FFFDC304F59F0FD103345" box="[622,751,2030,2054]" pageId="1" pageNumber="396">M. Kottelat</collectorName>
&amp; 
<collectorName id="26C329A5D11FFFDC32B15610FEB73365" box="[144,328,2062,2086]" pageId="1" pageNumber="396">T. Phommavong</collectorName>
, 
<date id="FF886AB3D11FFFDC33775611FDFC3365" box="[342,515,2062,2086]" pageId="1" pageNumber="396" value="2015-03-21">
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</materialsCitation>
. — 
<materialsCitation id="3B5E462ED11FFFDC30155611FAE93F05" ID-GBIF-Occurrence="3340077304" accessionNumber="CMK 26054" collectingDate="2015-03-25" collectionCode="CMK" collectorName="M. Kottelat &amp; T. Phommavong" country="Laos" county="Muang Long District" elevation="496" latitude="20.9675" location="Nam Ma" longLatPrecision="21" longitude="100.818886" municipality="Nam Ma watershed" pageId="1" pageNumber="396" specimenCode="24" specimenCount="1" stateProvince="Louangnamtha" typeStatus="paratype">
<collectionCode id="ED27D4B6D11FFFDC30155611FD8E3365" box="[564,625,2062,2086]" pageId="1" pageNumber="396">CMK</collectionCode>
26054, 
<specimenCode id="DB90E408D11FFFDC30EF5611FD143365" box="[718,747,2062,2086]" collectionCode="CMK" pageId="1" pageNumber="396">24</specimenCode>
, 
<quantity id="4CCEE196D11FFFDC310C5DB1FC383885" box="[813,967,942,966]" metricMagnitude="-2" metricUnit="m" metricValue="3.825" metricValueMax="4.34" metricValueMin="3.31" pageId="1" pageNumber="396" unit="mm" value="38.25" valueMax="43.4" valueMin="33.1">33.1–43.4 mm</quantity>
SL; 
<collectingCountry id="F3210CE3D11FFFDC31DD5DB0FBCB3885" box="[1020,1076,943,966]" name="Laos" pageId="1" pageNumber="396">Laos</collectingCountry>
: 
<collectingRegion id="49F28291D11FFFDC361E5DB0FAA93885" box="[1087,1366,942,967]" country="Laos" name="Louangnamtha" pageId="1" pageNumber="396">Louang Namtha Province</collectingRegion>
: 
<collectingCounty id="62E834FFD11FFFDC37415DB0FC3E38A5" pageId="1" pageNumber="396">Muang Long District</collectingCounty>
: 
<collectingMunicipality id="6BEDD609D11FFFDC31EC5DD0FB5A38A5" box="[973,1189,974,998]" pageId="1" pageNumber="396">Nam Ma watershed</collectingMunicipality>
: confluence of 
<location id="8EE91AA8D11FFFDC376D5DD0FA5338A5" LSID="urn:lsid:plazi:treatment:039FFD65D11FFFDA32DF5831FE5B3CB7:8EE91AA8D11FFFDC376D5DD0FA5338A5" box="[1356,1452,975,998]" country="Laos" county="Muang Long District" latitude="20.9675" longLatPrecision="21" longitude="100.818886" municipality="Nam Ma watershed" name="Nam Ma" pageId="1" pageNumber="396" stateProvince="Louangnamtha">Nam Ma</location>
and 
<location id="8EE91AA8D11FFFDC317F5DF0FC293F44" LSID="urn:lsid:plazi:treatment:039FFD65D11FFFDA32DF5831FE5B3CB7:8EE91AA8D11FFFDC317F5DF0FC293F44" box="[862,982,1007,1031]" country="Laos" county="Muang Long District" latitude="20.9675" longLatPrecision="21" longitude="100.818886" municipality="Nam Ma watershed" name="Nam Long" pageId="1" pageNumber="396" stateProvince="Louangnamtha">Nam Long</location>
at 
<location id="8EE91AA8D11FFFDC31DF5DF0FAF93F45" LSID="urn:lsid:plazi:treatment:039FFD65D11FFFDA32DF5831FE5B3CB7:8EE91AA8D11FFFDC31DF5DF0FAF93F45" box="[1022,1286,1006,1031]" country="Laos" county="Muang Long District" latitude="20.9675" longLatPrecision="21" longitude="100.818886" municipality="Nam Ma watershed" name="Ban Louang Pha Kham" pageId="1" pageNumber="396" stateProvince="Louangnamtha">Ban Louang Pha Kham</location>
(
<quantity id="4CCEE196D11FFFDC37395DF1FA983F45" box="[1304,1383,1006,1030]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="396" unit="km" value="0.5">0.5 km</quantity>
north of 
<location id="8EE91AA8D11FFFDC316C5A10FC203F65" LSID="urn:lsid:plazi:treatment:039FFD65D11FFFDA32DF5831FE5B3CB7:8EE91AA8D11FFFDC316C5A10FC203F65" box="[845,991,1038,1063]" country="Laos" county="Muang Long District" latitude="20.9675" longLatPrecision="21" longitude="100.818886" municipality="Nam Ma watershed" name="Muang Long" pageId="1" pageNumber="396" stateProvince="Louangnamtha">Muang Long</location>
), 
<geoCoordinate id="EE022AB4D11FFFDC31D55A11FB8C3F65" box="[1012,1139,1037,1062]" degrees="20" direction="north" minutes="58" orientation="latitude" pageId="1" pageNumber="396" precision="15" seconds="03" value="20.9675">20°58′03″N</geoCoordinate>
<geoCoordinate id="EE022AB4D11FFFDC365C5A11FAF63F65" box="[1149,1289,1037,1062]" degrees="100" direction="east" minutes="49" orientation="longitude" pageId="1" pageNumber="396" precision="15" seconds="08" value="100.818886">100°49′08″E</geoCoordinate>
, 
<quantity id="4CCEE196D11FFFDC37365A11FA803F65" box="[1303,1407,1038,1062]" metricMagnitude="2" metricUnit="m" metricValue="4.96" pageId="1" pageNumber="396" unit="m" value="496.0">
<elevation id="001BAB40D11FFFDC37365A11FA803F65" box="[1303,1407,1038,1062]" metricMagnitude="2" metricUnit="m" metricValue="4.96" pageId="1" pageNumber="396" unit="m" value="496.0">496 masl</elevation>
</quantity>
; 
<collectorName id="26C329A5D11FFFDC37AC5A10FC7C3F05" pageId="1" pageNumber="396">M. Kottelat</collectorName>
&amp; 
<collectorName id="26C329A5D11FFFDC31885A30FBA03F05" box="[937,1119,1070,1094]" pageId="1" pageNumber="396">T. Phommavong</collectorName>
, 
<date id="FF886AB3D11FFFDC364D5A31FAE93F05" box="[1132,1302,1070,1094]" pageId="1" pageNumber="396" value="2015-03-25">
<collectingDate id="EFCC935BD11FFFDC364D5A31FAE93F05" box="[1132,1302,1070,1094]" pageId="1" pageNumber="396" value="2015-03-25">25 March 2015</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C32C1FF8D11FFFDC310C5A71FA9D3D04" pageId="1" pageNumber="396" type="diagnosis">
<paragraph id="8B894C73D11FFFDC310C5A71FA9D3D04" blockId="1.[811,1453,1134,1607]" pageId="1" pageNumber="396">
<emphasis id="B9429061D11FFFDC310C5A71FC5F3FC5" bold="true" box="[813,928,1134,1158]" pageId="1" pageNumber="396">Diagnosis.</emphasis>
<taxonomicName id="4C3637F0D11FFFDC31845A71FB963FC5" authorityName="Kottelat" authorityYear="2017" box="[933,1129,1134,1158]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="396" phylum="Chordata" rank="species" species="thavonei">
<emphasis id="B9429061D11FFFDC31845A71FB963FC5" box="[933,1129,1134,1158]" italics="true" pageId="1" pageNumber="396">Schistura thavonei</emphasis>
</taxonomicName>
is distinguished from the other species of the genus by its unique colour pattern made of two broad dark brown stripes (one middorsal, one midlateral) and between them a pale yellowish-brown stripe (iridescent copper to orange in life) from the upper extremity of the gill opening to the upper half of the base of the caudal fin; overimposed to the midlateral stripe, a row of 12–24 short black bars, located increasingly lower on the flank from head to tail, posterior-most ones restricted to the lower half of the body or forming blotches along the ventral midline of the caudal peduncle. Additional characters useful for identification but not unique to the species are: body very elongate (depth 6.2–7.2 times in SL), with a marked hump behind the head; male without suborbital flap; 8+7 branched caudal-fin rays; 7½ branched dorsal fin rays; 9–10 pectoral-fin rays.
</paragraph>
</subSubSection>
<subSubSection id="C32C1FF8D11FFFD8310C5871FEE13E44" lastPageId="5" lastPageNumber="400" pageId="1" pageNumber="396" type="description">
<paragraph id="8B894C73D11FFFDE310C5871FECC3E1F" blockId="1.[811,1453,1646,2086]" lastBlockId="3.[113,752,1316,1372]" lastPageId="3" lastPageNumber="398" pageId="1" pageNumber="396">
<emphasis id="B9429061D11FFFDC310C5871FC483DC5" bold="true" box="[813,951,1646,1670]" pageId="1" pageNumber="396">Description.</emphasis>
See 
<figureCitation id="130D50F6D11FFFDC31C95871FBB63DC5" box="[1000,1097,1646,1670]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="1.[114,149,865,887]" captionStartId-1="2.[136,171,2064,2086]" captionStartId-2="3.[113,148,1210,1232]" captionTargetBox-0="[113,1451,189,850]" captionTargetBox-1="[276,1334,189,2039]" captionTargetBox-2="[113,1451,189,1195]" captionTargetId-0="figure-586@1.[113,1451,189,850]" captionTargetId-1="figure-7@2.[276,1334,189,2039]" captionTargetId-2="figure-427@3.[113,1451,189,1195]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionText-0="Fig. 1. Schistura thavonei, MHNG 2667.078, holotype, 44.0 mm SL; Laos: Louang Namtha Province: Nam Ma." captionText-1="Fig. 2. Schistura thavonei, paratypes; Laos: Louang Namtha Province: Nam Ma; a, CMK 25944, 44.0 mm SL; b, CMK 26054, 41.8 mm SL; c, CMK 26054, 39.5 mm SL; d, CMK 25944, 44.9 mm SL; e, CMK 26066, 44.7 mm SL." captionText-2="Fig. 3. Schistura thavonei, paratypes; Laos: Louang Namtha Province: Nam Ma; a, CMK 26066, 33.7 mm SL; b, CMK 26054, 34.4 mm SL (right side, reversed); c, CMK 25944, 36.6 mm SL." figureDoi-0="http://doi.org/10.5281/zenodo.5357129" figureDoi-1="http://doi.org/10.5281/zenodo.5357132" figureDoi-2="http://doi.org/10.5281/zenodo.5357134" httpUri-0="https://zenodo.org/record/5357129/files/figure.png" httpUri-1="https://zenodo.org/record/5357132/files/figure.png" httpUri-2="https://zenodo.org/record/5357134/files/figure.png" pageId="1" pageNumber="396">Figs. 1–3</figureCitation>
for general appearance and 
<tableCitation id="C6B479C8D11FFFDC374E5871FCC53DE5" captionStart="Table 1" captionStartId="6.[136,191,189,211]" captionText="Table 1. Morphometric data of type material of Schistura thavonei (n=11). Range and mean include holotype data." pageId="1" pageNumber="396">Table 1</tableCitation>
for morphometric data of 
<typeStatus id="548DF2D1D11FFFDC36695891FB5B3DE5" box="[1096,1188,1678,1702]" pageId="1" pageNumber="396" type="holotype">holotype</typeStatus>
and 
<specimenCount id="9D3087FAD11FFFDC36F45891FAA33DE4" box="[1237,1372,1678,1703]" pageId="1" pageNumber="396" type="generic" typeStatus="paratypes">10 paratypes</specimenCount>
. A very elongate nemacheilid with body depth increasing up to about middle of pre-dorsal area of trunk, then decreasing slowly to caudal-fin base. Dorsal profile with hump immediately behind head. Head slightly depressed; body from slightly compressed anteriorly to compressed posteriorly. Interorbital area slightly arched, more so in smallest specimens. In lateral view, eye slightly below or flushed with dorsal profile of head. Cheeks not swollen. Snout pointed. Caudal peduncle 1.5–1.8 times longer than deep, tapering posteriorly. In some of largest specimens, a low dorsal crest on posterior 
<superScript id="7C43E13BD11FFFDC310C59F2FCCA3CB8" attach="right" box="[813,821,2029,2043]" fontSize="6" pageId="1" pageNumber="396">1</superScript>
/ 
<subScript id="17B24E36D11FFFDC311D59E5FCB3334B" attach="left" box="[828,844,2042,2056]" fontSize="6" pageId="1" pageNumber="396">10</subScript>
of post-dorsal area, and a low ventral crest on posterior half of caudal peduncle. Dorsal crest separated from upper margin of caudal fin by a small concavity. Largest recorded size 
<quantity id="4CCEE196D11DFFDE32825B5BFEFB3E18" box="[163,260,1348,1372]" metricMagnitude="-2" metricUnit="m" metricValue="5.07" pageId="3" pageNumber="398" unit="mm" value="50.7">50.7 mm</quantity>
SL.
</paragraph>
<caption id="DF491CFBD11CFFDF32A9560FFBD53301" ID-DOI="http://doi.org/10.5281/zenodo.5357132" ID-Zenodo-Dep="5357132" httpUri="https://zenodo.org/record/5357132/files/figure.png" pageId="2" pageNumber="397" startId="2.[136,171,2064,2086]" targetBox="[276,1334,189,2039]" targetPageId="2">
<paragraph id="8B894C73D11CFFDF32A9560FFBD53301" blockId="2.[136,1474,2064,2115]" pageId="2" pageNumber="397">
Fig. 2. 
<taxonomicName id="4C3637F0D11CFFDF32EF560FFE7B3366" authorityName="Kottelat" authorityYear="2017" box="[206,388,2064,2085]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="397" phylum="Chordata" rank="species" species="thavonei">
<emphasis id="B9429061D11CFFDF32EF560FFE7B3366" box="[206,388,2064,2085]" italics="true" pageId="2" pageNumber="397">Schistura thavonei</emphasis>
</taxonomicName>
, paratypes; Laos: Louang Namtha Province: Nam Ma; a, CMK 25944, 44.0 mm SL; b, CMK 26054, 41.8 mm SL; c, CMK 26054, 39.5 mm SL; d, CMK 25944, 44.9 mm SL; e, CMK 26066, 44.7 mm SL.
</paragraph>
</caption>
<caption id="DF491CFBD11DFFDE32505AA5FD6F3FAF" ID-DOI="http://doi.org/10.5281/zenodo.5357134" ID-Zenodo-Dep="5357134" httpUri="https://zenodo.org/record/5357134/files/figure.png" pageId="3" pageNumber="398" startId="3.[113,148,1210,1232]" targetBox="[113,1451,189,1195]" targetPageId="3">
<paragraph id="8B894C73D11DFFDE32505AA5FD6F3FAF" blockId="3.[113,1451,1210,1261]" pageId="3" pageNumber="398">
Fig. 3. 
<taxonomicName id="4C3637F0D11DFFDE32965AA5FE933F8C" authorityName="Kottelat" authorityYear="2017" box="[183,364,1210,1231]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="398" phylum="Chordata" rank="species" species="thavonei">
<emphasis id="B9429061D11DFFDE32965AA5FE933F8C" box="[183,364,1210,1231]" italics="true" pageId="3" pageNumber="398">Schistura thavonei</emphasis>
</taxonomicName>
, paratypes; Laos: Louang Namtha Province: Nam Ma; a, CMK 26066, 33.7 mm SL; b, CMK 26054, 34.4 mm SL (right side, reversed); c, CMK 25944, 36.6 mm SL.
</paragraph>
</caption>
<paragraph id="8B894C73D11DFFDE32505B9BFD5C3CD8" blockId="3.[113,754,1411,1948]" pageId="3" pageNumber="398">Dorsal fin with 4 unbranched and 7½ (10*) or 8½ (1) branched rays; distal margin straight; second branched ray longest. Pectoral fin with 1 unbranched and 8 (9*) or 9 (2) branched rays (including small last ray, usually unbranched); rounded but with angled tip; reaching about halfway of distance to pelvic-fin base. Axillary pectoral lobe present. Pelvic fin with 1 unbranched and 6 (6) or 7 (5*) branched rays (including small last ray, usually unbranched); reaching to or at least ⅔ of distance to anus (or about halfway to anal-fin base); triangular; origin from slightly in front of dorsal-fin origin to below base of last unbranched dorsal-fin ray; axillary lobe present, entirely free. Anus situated about 2½–3 times eye diameter in front of anal fin. Anal fin with 3 unbranched and 5½ branched rays; distal margin straight to slightly convex. Caudal fin with 8+7 (10*) or 8+6 (1) branched rays; dorsal and ventral procurrent rays cannot be counted; forked, lobes rounded, lower lobe slightly longer than upper one.</paragraph>
<paragraph id="8B894C73D11DFFDE325059DBFACE3EDF" blockId="3.[113,752,1988,2108]" lastBlockId="3.[813,1451,1316,1436]" pageId="3" pageNumber="398">
On pectoral and pelvic fins, thick unculiferous pads (sensu 
<bibRefCitation id="EFA73182D11DFFDE325059FBFEB73CBF" author="Conway KW &amp; Lujan NK &amp; Lundberg JG &amp; Mayden RL &amp; Siegel DS" box="[113,328,2020,2044]" pageId="3" pageNumber="398" pagination="1127 - 1149" refId="ref4898" refString="Conway KW, Lujan NK, Lundberg JG, Mayden RL &amp; Siegel DS (2012) Microanatomy of the paired-fin pads of ostariophysan fishes (Teleostei: Ostariophysi). Journal of Morphology, 273: 1127 - 1149." type="journal article" year="2012">Conway et al., 2012</bibRefCitation>
) along anterior edge of unbranched ray, and on dorsal side on whole width of membrane between unbranched and first branched rays, and along posterior margin of rays on membranes between remaining rays; on ventral side on whole width of membrane between unbranched and first branched ray, and along anterior margin of rays on membranes between remaining rays.
</paragraph>
<paragraph id="8B894C73D11DFFDE310C5BDBFB293DF8" blockId="3.[812,1452,1476,1724]" pageId="3" pageNumber="398">Body entirely covered by scales, except belly in front of pelvic fins. Scales embedded, increasingly less densely-set from level of dorsal fin towards head. Lateral line incomplete, reaching between vertical of pelvic-fin origin to vertical of end of anal-fin base, often with interruption near posterior extremity, with 44–91 pores. Cephalic lateral line system with 6 supraorbital, 4 + 10–12 infraorbital, 9 preoperculomandibular and 3 supratemporal pores.</paragraph>
<paragraph id="8B894C73D11DFFD9310C58FBFEB539A7" blockId="3.[811,1453,1764,2108]" lastBlockId="4.[136,776,652,740]" lastPageId="4" lastPageNumber="399" pageId="3" pageNumber="398">
Anterior nare pierced in front side of a pointed flap-like tube. Posterior nare adjacent to anterior one. Mouth arched, gape about 2.2–2.9 times wider than long (
<figureCitation id="130D50F6D11DFFDE36F1593BFAEA3C7F" box="[1232,1301,1828,1852]" captionStart="Fig" captionStartId="4.[136,171,575,597]" captionTargetBox="[136,775,189,560]" captionTargetId="figure-7@4.[136,775,189,560]" captionTargetPageId="4" captionText="Fig. 4. Schistura thavonei, CMK 25944, 44 mm SL; mouth." figureDoi="http://doi.org/10.5281/zenodo.5357136" httpUri="https://zenodo.org/record/5357136/files/figure.png" pageId="3" pageNumber="398">Fig. 4</figureCitation>
). Lips fleshy. Upper lip without or (rarely) with a small median notch, with many shallow weakly-marked furrows, edge almost smooth. Processus dentiformis present. Lower lip with narrow median interruption; median part with 2–3 feebly marked sulci, lateral part smooth. Tip of lower jaw not exposed. No median notch or concavity in lower jaw. Inner rostral barbel reaching base of outer one; outer one reaching vertical of posterior nare. Maxillary barbel reaching at most vertical of posterior margin of eye. Intestine with a weak bend behind stomach (
<figureCitation id="130D50F6D11AFFD932D35CB3FECA3987" box="[242,309,684,708]" captionStart="Fig" captionStartId="4.[835,870,885,907]" captionTargetBox="[1017,1293,190,869]" captionTargetId="figure-20@4.[1013,1297,186,872]" captionTargetPageId="4" captionText="Fig. 5. Schistura thavonei, CMK 25944, 42.4 mm SL; digestive tract. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5357138" httpUri="https://zenodo.org/record/5357138/files/figure.png" pageId="4" pageNumber="399">Fig. 5</figureCitation>
). Air bladder without posterior chamber in abdominal cavity.
</paragraph>
<caption id="DF491CFBD11AFFD932A95C20FD293916" ID-DOI="http://doi.org/10.5281/zenodo.5357136" ID-Zenodo-Dep="5357136" box="[136,726,575,597]" httpUri="https://zenodo.org/record/5357136/files/figure.png" pageId="4" pageNumber="399" startId="4.[136,171,575,597]" targetBox="[136,775,189,560]" targetPageId="4">
<paragraph id="8B894C73D11AFFD932A95C20FD293916" blockId="4.[136,726,575,597]" box="[136,726,575,597]" pageId="4" pageNumber="399">
Fig. 4. 
<taxonomicName id="4C3637F0D11AFFD932EE5C20FE363917" authority=", CMK" authorityName="CMK" box="[207,457,575,596]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="399" phylum="Chordata" rank="species" species="thavonei">
<emphasis id="B9429061D11AFFD932EE5C20FE7A3917" box="[207,389,575,596]" italics="true" pageId="4" pageNumber="399">Schistura thavonei</emphasis>
, CMK
</taxonomicName>
25944, 44 mm SL; mouth.
</paragraph>
</caption>
<paragraph id="8B894C73D11AFFD932A95D13FED738E7" blockId="4.[135,775,780,932]" pageId="4" pageNumber="399">
<emphasis id="B9429061D11AFFD932A95D13FE943867" bold="true" box="[136,363,780,804]" pageId="4" pageNumber="399">Sexual dimorphism.</emphasis>
None observed. None of the characters usually associated with sexual dimorphism in 
<taxonomicName id="4C3637F0D11AFFD930465D33FCF93807" box="[615,774,812,836]" class="Actinopterygii" family="Nemacheilidae" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="399" phylum="Chordata" rank="family">Nemacheilidae</taxonomicName>
was observed (no suborbital flap, groove or slit; no tubercles; no modified pectoral fins in males). Ovigerous females deeper bodied.
</paragraph>
<caption id="DF491CFBD11AFFD931625D6AFBE138E4" ID-DOI="http://doi.org/10.5281/zenodo.5357138" ID-Zenodo-Dep="5357138" httpUri="https://zenodo.org/record/5357138/files/figure.png" pageId="4" pageNumber="399" startId="4.[835,870,885,907]" targetBox="[1017,1293,190,869]" targetPageId="4">
<paragraph id="8B894C73D11AFFD931625D6AFBE138E4" blockId="4.[835,1474,885,935]" pageId="4" pageNumber="399">
Fig. 5. 
<taxonomicName id="4C3637F0D11AFFD931AF5D6AFB7538C9" authority=", CMK" authorityName="CMK" box="[910,1162,885,906]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="399" phylum="Chordata" rank="species" species="thavonei">
<emphasis id="B9429061D11AFFD931AF5D6AFBBB38C9" box="[910,1092,885,906]" italics="true" pageId="4" pageNumber="399">Schistura thavonei</emphasis>
, CMK
</taxonomicName>
25944, 42.4 mm SL; digestive tract. Scale bar 1 mm.
</paragraph>
</caption>
<paragraph id="8B894C73D11AFFD932A95DD3FE323E87" blockId="4.[136,777,972,1476]" pageId="4" pageNumber="399">
<emphasis id="B9429061D11AFFD932A95DD3FEF538A7" bold="true" box="[136,266,972,996]" pageId="4" pageNumber="399">Coloration.</emphasis>
After one month in formalin. Head and body background colour pale yellowish brown; except otherwise stated, markings dark brown to black. Dorsal half of head plain dark brown. Barbels orange or black, lips sometimes orange. Body with broad middorsal stripe from occiput almost to caudal-fin base, leaving a whitish patch about equal to eye diameter between stripe and fin base. A broad midlateral stripe from head almost to caudal-fin base; stripe made of surface pigments anteriorly, and both deep and surface pigments posteriorly. Interspace between middorsal and midlateral stripe appearing as a yellowish brown stripe from upper extremity of gill opening to upper half of caudal-fin base, more contrasted and orange posteriorly. Along anal-fin base and ventral edge of caudal peduncle: a narrow band of greyish pigments, in a few specimens in contact with midlateral stripe.
</paragraph>
<paragraph id="8B894C73D11AFFD932A95BF3FE0D3CE7" blockId="4.[135,777,1516,1956]" pageId="4" pageNumber="399">Flank with 12–24 dark bars, upper tip at or slightly above upper edge of midlateral stripe, starting higher in predorsal area, sometimes even in contact with middorsal stripe; upper extremity of bars decreasingly lower towards back, last ones restricted to lower half of caudal peduncle. Lower extremity of bars about at level of pectoral-fin base anteriorly, decreasingly lower towards back, reaching ventral midline at anal-fin base and on caudal peduncle; last bars continuous with contralaterals, sometimes fused to form one or two elongated blotches along ventral midline, always leaving a whitish patch between last bar and fin base. Bars in front of dorsal fin narrow, narrower than or as wide as interspaces. Bars behind dorsal fin becoming increasingly darker and wider, narrower than interspaces.</paragraph>
<paragraph id="8B894C73D11AFFD932A959D3FBE33ED4" blockId="4.[136,775,1996,2117]" lastBlockId="4.[834,1476,991,1431]" pageId="4" pageNumber="399">Black pattern at caudal-fin base: all stripes and bars on caudal peduncle ending shortly before caudal-fin base, leaving a vertical band devoid of any pigment. On upper half: a patch of black pigments on basal part of upper 4–5 principal rays and on last 0–2 procurrent rays (on rays only, not on fleshy base of fin). In some largest specimens, patches extending backwards up to about ¼ of length of rays. Anterior edge usually more posterior than that of upper patch on lower half of fin. On lower half: two patches of black pigments on fleshy base of fin and on proximal-most part of principal rays; first patch on upper 2–3 rays, second on remaining rays. Patches made of superficial and deep pigments, both patches may be differently dark and large. Patches usually fused or adjacent, with lower one weaker or sometimes missing, fused patches often appearing as a slanted mark, with lower patch slightly more posterior than upper. In some largest specimens, patches extending backwards up to about ¼ of length of rays.</paragraph>
<paragraph id="8B894C73D11AFFD931625BA0FBF33C34" blockId="4.[834,1476,1471,1911]" pageId="4" pageNumber="399">Dorsal fin hyaline with a small black spot at base of simple rays and first branched ray and an elongated blotch at base of branched rays 2–4, both low and sometimes fused; middle part of last unbranched ray blackish, anterior edge orange; pigments on rays in area of first branching point, beyond only along edges of rays. Caudal fin hyaline; beside black pattern at base described above, an orange patch along dorsal and ventral edges, extending from black blotch backwards; black pigments along edges of rays on their entire length. Anal fin hyaline; black pigments along edges of rays in area of first branching point; tip of last unbranched rays orangish. Pectoral and pelvic fins hyaline, membranes between anterior rays pale orange; greyish pigments along edges of rays on their entire length.</paragraph>
<paragraph id="8B894C73D11AFFD831625980FD143984" blockId="4.[834,1474,1951,2103]" lastBlockId="5.[113,751,655,711]" lastPageId="5" lastPageNumber="400" pageId="4" pageNumber="399">
In specimens less than about 
<quantity id="4CCEE196D11AFFD9365E5980FB353CF5" box="[1151,1226,1951,1975]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="4" pageNumber="399" unit="mm" value="40.0">40 mm</quantity>
SL (
<figureCitation id="130D50F6D11AFFD937215980FABE3CF4" box="[1280,1345,1951,1975]" captionStart="Fig" captionStartId="3.[113,148,1210,1232]" captionTargetBox="[113,1451,189,1195]" captionTargetId="figure-427@3.[113,1451,189,1195]" captionTargetPageId="3" captionText="Fig. 3. Schistura thavonei, paratypes; Laos: Louang Namtha Province: Nam Ma; a, CMK 26066, 33.7 mm SL; b, CMK 26054, 34.4 mm SL (right side, reversed); c, CMK 25944, 36.6 mm SL." figureDoi="http://doi.org/10.5281/zenodo.5357134" httpUri="https://zenodo.org/record/5357134/files/figure.png" pageId="4" pageNumber="399">Fig. 3</figureCitation>
), middorsal stripe present but faint; midlateral stripe and pale area between stripes absent or poorly distinct, appearing first in postdorsal part of body. Bars less regular, especially in anterior half of flank where most reach middorsal stripe; some bars interrupted at level of lateral line. Pattern at caudal-fin base and on fins as in adults. Smaller individuals not available.
</paragraph>
<caption id="DF491CFBD11BFFD832515C3AFEBC391B" ID-DOI="http://doi.org/10.5281/zenodo.5357140" ID-Zenodo-Dep="5357140" httpUri="https://zenodo.org/record/5357140/files/figure.png" pageId="5" pageNumber="400" startId="5.[112,147,549,571]" targetBox="[113,1451,189,534]" targetPageId="5">
<paragraph id="8B894C73D11BFFD832515C3AFEBC391B" blockId="5.[112,1451,549,600]" pageId="5" pageNumber="400">
Fig. 6. 
<taxonomicName id="4C3637F0D11BFFD8329A5C3AFE483979" authority=", CMK" authorityName="CMK" box="[187,439,549,571]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="400" phylum="Chordata" rank="species" species="thavonei">
<emphasis id="B9429061D11BFFD8329A5C3AFE8D3979" box="[187,370,549,570]" italics="true" pageId="5" pageNumber="400">Schistura thavonei</emphasis>
, CMK
</taxonomicName>
25944, paratype, 50.7 mm SL; Laos: Louang Namtha Province: Nam Ma; immediately after fixation (right side, reversed).
</paragraph>
</caption>
<paragraph id="8B894C73D11BFFD832505CEFFDAD38E4" blockId="5.[113,752,751,935]" pageId="5" pageNumber="400">
In life, from photographs taken immediately after capture and fixation (
<figureCitation id="130D50F6D11BFFD833235D10FEB93864" box="[258,326,783,807]" captionStart="Fig" captionStartId="5.[112,147,549,571]" captionTargetBox="[113,1451,189,534]" captionTargetId="figure-341@5.[113,1451,189,534]" captionTargetPageId="5" captionText="Fig. 6. Schistura thavonei, CMK 25944, paratype, 50.7 mm SL; Laos: Louang Namtha Province: Nam Ma; immediately after fixation (right side, reversed)." figureDoi="http://doi.org/10.5281/zenodo.5357140" httpUri="https://zenodo.org/record/5357140/files/figure.png" pageId="5" pageNumber="400">Fig. 6</figureCitation>
): body yellowish brown, stripes darker brown, bars blackish brown. Pale stripe iridescent brown, bright yellow at extremity of caudal peduncle. Tip of snout, barbels and lips reddish orange. Dorsal, caudal and anal fins yellowish, pectoral and pelvic fins orangish.
</paragraph>
<paragraph id="8B894C73D11BFFD832505DD0FEE13E44" blockId="5.[113,752,975,1287]" pageId="5" pageNumber="400">
<emphasis id="B9429061D11BFFD832505DD0FEC538A4" bold="true" box="[113,314,975,999]" pageId="5" pageNumber="400">Notes on biology.</emphasis>
A dissected female (CMK 25066, 
<quantity id="4CCEE196D11BFFD830E05DD0FF643F44" metricMagnitude="-2" metricUnit="m" metricValue="4.66" pageId="5" pageNumber="400" unit="mm" value="46.6">46.6 mm</quantity>
SL) had unripe ovaries with white, irregular, not mature, ova about 1.0 mm diameter. The stomach of a 
<quantity id="4CCEE196D11BFFD832535A30FF2C3F04" box="[114,211,1071,1095]" metricMagnitude="-2" metricUnit="m" metricValue="4.24" pageId="5" pageNumber="400" unit="mm" value="42.4">42.4 mm</quantity>
SL specimen was filled with insect larvae about 
<quantity id="4CCEE196D11BFFD832505A50FF303F24" box="[113,207,1103,1127]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="5" pageNumber="400" unit="mm" value="3.5" valueMax="5.0" valueMin="2.0">2–5 mm</quantity>
long. 
<taxonomicName id="4C3637F0D11BFFD8333D5A50FE133F24" box="[284,492,1103,1127]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="400" phylum="Chordata" rank="species" species="thavonei">
<emphasis id="B9429061D11BFFD8333D5A50FE133F24" box="[284,492,1103,1127]" italics="true" pageId="5" pageNumber="400">Schistura thavonei</emphasis>
</taxonomicName>
was observed in clear water [as expected for a benthic fish with bright coloration and contrasted pattern]. At all sites, 
<taxonomicName id="4C3637F0D11BFFD8302B5A8FFD7A3FE4" box="[522,645,1167,1191]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="400" phylum="Chordata" rank="species" species="thavonei">
<emphasis id="B9429061D11BFFD8302B5A8FFD7A3FE4" box="[522,645,1167,1191]" italics="true" pageId="5" pageNumber="400">S. thavonei</emphasis>
</taxonomicName>
has been collected in stretches of streams with riffles (in the sheltered parts with somewhat quieter current), over gravel to stone bottom (
<figureCitation id="130D50F6D11BFFD832EC5AF0FEEE3E44" box="[205,273,1263,1287]" captionStart="Fig" captionStartId="5.[813,847,1069,1091]" captionTargetBox="[813,1452,655,1053]" captionTargetId="figure-365@5.[813,1452,655,1053]" captionTargetPageId="5" captionText="Fig.7. Nam Ma Yen, Nam Ma watershed,Louang Namtha Province, Laos; type locality of Schistura thavonei; 25 March 2015." figureDoi="http://doi.org/10.5281/zenodo.5357142" httpUri="https://zenodo.org/record/5357142/files/figure.png" pageId="5" pageNumber="400">Fig. 7</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF491CFBD11BFFD8310C5A32FA983F1C" ID-DOI="http://doi.org/10.5281/zenodo.5357142" ID-Zenodo-Dep="5357142" httpUri="https://zenodo.org/record/5357142/files/figure.png" pageId="5" pageNumber="400" startId="5.[813,847,1069,1091]" targetBox="[813,1452,655,1053]" targetPageId="5">
<paragraph id="8B894C73D11BFFD8310C5A32FA983F1C" blockId="5.[812,1452,1069,1120]" pageId="5" pageNumber="400">
Fig. 7. Nam Ma Yen, Nam Ma watershed, Louang Namtha Province, Laos; type locality of 
<taxonomicName id="4C3637F0D11BFFD836295A55FB433F1C" box="[1032,1212,1098,1119]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="400" phylum="Chordata" rank="species" species="thavonei">
<emphasis id="B9429061D11BFFD836295A55FB433F1C" box="[1032,1212,1098,1119]" italics="true" pageId="5" pageNumber="400">Schistura thavonei</emphasis>
</taxonomicName>
; 25 March 2015.
</paragraph>
</caption>
<subSubSection id="C32C1FF8D11BFFD832505B30FD2B3E84" pageId="5" pageNumber="400" type="distribution">
<paragraph id="8B894C73D11BFFD832505B30FD2B3E84" blockId="5.[112,752,1327,1479]" pageId="5" pageNumber="400">
<emphasis id="B9429061D11BFFD832505B30FEFB3E04" bold="true" box="[113,260,1327,1351]" pageId="5" pageNumber="400">Distribution.</emphasis>
<taxonomicName id="4C3637F0D11BFFD8332F5B30FE253E04" box="[270,474,1327,1351]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="400" phylum="Chordata" rank="species" species="thavonei">
<emphasis id="B9429061D11BFFD8332F5B30FE253E04" box="[270,474,1327,1351]" italics="true" pageId="5" pageNumber="400">Schistura thavonei</emphasis>
</taxonomicName>
is presently known only from the watershed of the Nam Ma in 
<collectingRegion id="49F28291D11BFFD830185B4FFF2C3EC4" country="Laos" name="Louangnamtha" pageId="5" pageNumber="400">Louang Namtha Province</collectingRegion>
, a tributary of the Mekong, in northeastern 
<collectingCountry id="F3210CE3D11BFFD8309D5B6FFD0F3EC4" box="[700,752,1392,1415]" name="Laos" pageId="5" pageNumber="400">Laos</collectingCountry>
(not to be confused with the Nam Ma in Houa Phan Province, which flows to 
<collectingCountry id="F3210CE3D11BFFD833385BB0FE8A3E84" box="[281,373,1455,1479]" name="Vietnam" pageId="5" pageNumber="400">Vietnam</collectingCountry>
and enters the Gulf of 
<collectingCountry id="F3210CE3D11BFFD830565BB0FD363E84" box="[631,713,1455,1479]" name="Vietnam" pageId="5" pageNumber="400">Tonkin</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="C32C1FF8D11BFFD832505BF0FBFE3F8C" pageId="5" pageNumber="400" type="etymology">
<paragraph id="8B894C73D11BFFD832505BF0FBFE3F8C" blockId="5.[113,752,1519,1575]" lastBlockId="5.[811,1450,1175,1231]" pageId="5" pageNumber="400">
<emphasis id="B9429061D11BFFD832505BF0FEFF3D44" bold="true" box="[113,256,1519,1543]" pageId="5" pageNumber="400">Etymology.</emphasis>
The species is named for Mr. Thavone Phommavong, in appreciation for his help and companionship during several, and sometimes difficult, fish surveys in 
<collectingCountry id="F3210CE3D11BFFD8374E5A88FA593FED" box="[1391,1446,1175,1198]" name="Laos" pageId="5" pageNumber="400">Laos</collectingCountry>
. A noun in genitive.
</paragraph>
</subSubSection>
<subSubSection id="C32C1FF8D11BFFDA310C5AE8FE5B3CB7" lastPageId="7" lastPageNumber="402" pageId="5" pageNumber="400" type="discussion">
<paragraph id="8B894C73D11BFFDA310C5AE8FDF23FF0" blockId="5.[811,1453,1271,1583]" lastBlockId="7.[112,754,667,1203]" lastPageId="7" lastPageNumber="402" pageId="5" pageNumber="400">
<emphasis id="B9429061D11BFFD8310C5AE8FC643E4C" bold="true" box="[813,923,1271,1295]" pageId="5" pageNumber="400">Remarks.</emphasis>
The striking colour pattern of 
<taxonomicName id="4C3637F0D11BFFD836F45AE8FAB63E4C" box="[1237,1353,1271,1295]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="400" phylum="Chordata" rank="species" species="thavonei">
<emphasis id="B9429061D11BFFD836F45AE8FAB63E4C" box="[1237,1353,1271,1295]" italics="true" pageId="5" pageNumber="400">S. thavonei</emphasis>
</taxonomicName>
is shared with no other 
<taxonomicName id="4C3637F0D11BFFD831E35B08FB993E6C" box="[962,1126,1303,1327]" class="Actinopterygii" family="Nemacheilidae" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="400" phylum="Chordata" rank="family">Nemacheilidae</taxonomicName>
. Some elements, however, are shared with 
<taxonomicName id="4C3637F0D11BFFD831955B28FBF93E0C" box="[948,1030,1335,1359]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="400" phylum="Chordata" rank="species" species="bella">
<emphasis id="B9429061D11BFFD831955B28FBF93E0C" box="[948,1030,1335,1359]" italics="true" pageId="5" pageNumber="400">S. bella</emphasis>
</taxonomicName>
(
<figureCitation id="130D50F6D11BFFD836365B28FBA63E0C" box="[1047,1113,1335,1359]" captionStart="Fig" captionStartId="5.[113,148,2093,2115]" captionTargetBox="[113,1451,1656,2077]" captionTargetId="figure-388@5.[113,1451,1656,2077]" captionTargetPageId="5" captionText="Fig. 8. Schistura bella, CMK 26052, 38.4 mm SL; Laos: Louang Namtha Province: Nam Ma." figureDoi="http://doi.org/10.5281/zenodo.5357144" httpUri="https://zenodo.org/record/5357144/files/figure.png" pageId="5" pageNumber="400">Fig. 8</figureCitation>
), like the yellow (iridescent in life) stripe between the upper extremity of the gill opening and the upper half of caudal-fin base, and bars more or less restricted to part of body below the pale stripe. However, in 
<taxonomicName id="4C3637F0D11BFFD8316A5BA8FC613E8C" box="[843,926,1463,1487]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="400" phylum="Chordata" rank="species" species="bella">
<emphasis id="B9429061D11BFFD8316A5BA8FC613E8C" box="[843,926,1463,1487]" italics="true" pageId="5" pageNumber="400">S. bella</emphasis>
</taxonomicName>
the bars are very variable, in some individuals regular and wider than interspaces, in others they are narrow and interconnected, forming a marmorated or variegated pattern; the middorsal stripe, when present, is very irregular, but there is usually a succession of blotches; the black mark at the base of the caudal fin differs (compare 
<figureCitation id="130D50F6D119FFDA307C5CA4FD3D3990" box="[605,706,699,723]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="1.[114,149,865,887]" captionStartId-1="2.[136,171,2064,2086]" captionStartId-2="3.[113,148,1210,1232]" captionTargetBox-0="[113,1451,189,850]" captionTargetBox-1="[276,1334,189,2039]" captionTargetBox-2="[113,1451,189,1195]" captionTargetId-0="figure-586@1.[113,1451,189,850]" captionTargetId-1="figure-7@2.[276,1334,189,2039]" captionTargetId-2="figure-427@3.[113,1451,189,1195]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionText-0="Fig. 1. Schistura thavonei, MHNG 2667.078, holotype, 44.0 mm SL; Laos: Louang Namtha Province: Nam Ma." captionText-1="Fig. 2. Schistura thavonei, paratypes; Laos: Louang Namtha Province: Nam Ma; a, CMK 25944, 44.0 mm SL; b, CMK 26054, 41.8 mm SL; c, CMK 26054, 39.5 mm SL; d, CMK 25944, 44.9 mm SL; e, CMK 26066, 44.7 mm SL." captionText-2="Fig. 3. Schistura thavonei, paratypes; Laos: Louang Namtha Province: Nam Ma; a, CMK 26066, 33.7 mm SL; b, CMK 26054, 34.4 mm SL (right side, reversed); c, CMK 25944, 36.6 mm SL." figureDoi-0="http://doi.org/10.5281/zenodo.5357129" figureDoi-1="http://doi.org/10.5281/zenodo.5357132" figureDoi-2="http://doi.org/10.5281/zenodo.5357134" httpUri-0="https://zenodo.org/record/5357129/files/figure.png" httpUri-1="https://zenodo.org/record/5357132/files/figure.png" httpUri-2="https://zenodo.org/record/5357134/files/figure.png" pageId="7" pageNumber="402">Figs. 1–3</figureCitation>
and 
<figureCitation id="130D50F6D119FFDA32505CC4FF7F39B0" box="[113,128,731,755]" captionStart="Fig" captionStartId="5.[113,148,2093,2115]" captionTargetBox="[113,1451,1656,2077]" captionTargetId="figure-388@5.[113,1451,1656,2077]" captionTargetPageId="5" captionText="Fig. 8. Schistura bella, CMK 26052, 38.4 mm SL; Laos: Louang Namtha Province: Nam Ma." figureDoi="http://doi.org/10.5281/zenodo.5357144" httpUri="https://zenodo.org/record/5357144/files/figure.png" pageId="7" pageNumber="402">8</figureCitation>
). The contrasted colour pattern at the base of the caudalfin is species-specific in many 
<taxonomicName id="4C3637F0D119FFDA33F35CE4FD893850" box="[466,630,763,787]" class="Actinopterygii" family="Nemacheilidae" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="402" phylum="Chordata" rank="family">Nemacheilidae</taxonomicName>
and likely has a species-recognition or advertising function. 
<taxonomicName id="4C3637F0D119FFDA30AC5D04FF323810" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="402" phylum="Chordata" rank="species" species="thavonei">
<emphasis id="B9429061D119FFDA30AC5D04FF323810" italics="true" pageId="7" pageNumber="402">Schistura thavonei</emphasis>
</taxonomicName>
has usually 7½ branched dorsal-fin rays (vs 8–9½ in 
<taxonomicName id="4C3637F0D119FFDA32AC5D44FF233830" box="[141,220,859,883]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="402" phylum="Chordata" rank="species" species="bella">
<emphasis id="B9429061D119FFDA32AC5D44FF233830" box="[141,220,859,883]" italics="true" pageId="7" pageNumber="402">S. bella</emphasis>
</taxonomicName>
), 8+7 branched caudal-fin rays (vs 9+8), the male does not have a suborbital flap (vs present) and there is no sexual dimorphism in the shape of the pectoral fins (vs the shape of the pectoral-fin rays is sexually dimorphic). The two species were collected together at one site (CMK 26052), in the Nam Ma proper. 
<taxonomicName id="4C3637F0D119FFDA33685DE4FE173F50" box="[329,488,1019,1043]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="402" phylum="Chordata" rank="species" species="bella">
<emphasis id="B9429061D119FFDA33685DE4FE173F50" box="[329,488,1019,1043]" italics="true" pageId="7" pageNumber="402">Schistura bella</emphasis>
</taxonomicName>
was in somewhat deeper and faster water than 
<taxonomicName id="4C3637F0D119FFDA33745A04FE343F70" box="[341,459,1051,1075]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="402" phylum="Chordata" rank="species" species="thavonei">
<emphasis id="B9429061D119FFDA33745A04FE343F70" box="[341,459,1051,1075]" italics="true" pageId="7" pageNumber="402">S. thavonei</emphasis>
</taxonomicName>
. In the field, they were first confused in the net because of the similar iridescent pale stripe. The relationships of 
<taxonomicName id="4C3637F0D119FFDA33815A44FE0D3F30" box="[416,498,1115,1139]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="402" phylum="Chordata" rank="species" species="bella">
<emphasis id="B9429061D119FFDA33815A44FE0D3F30" box="[416,498,1115,1139]" italics="true" pageId="7" pageNumber="402">S. bella</emphasis>
</taxonomicName>
, however, seems to be with a group of species commonly placed in 
<taxonomicName id="4C3637F0D119FFDA306C5A64FD0F3FD0" box="[589,752,1147,1171]" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="402" phylum="Chordata" rank="genus">
<emphasis id="B9429061D119FFDA306C5A64FD0F3FD0" box="[589,752,1147,1171]" italics="true" pageId="7" pageNumber="402">Physoschistura</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C3637F0D119FFDA32815A84FE9E3FF0" authorityName="Bohlen &amp; Slechtova" authorityYear="2011" box="[160,353,1179,1203]" class="Actinopterygii" family="Nemacheilidae" genus="Pteronemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="402" phylum="Chordata" rank="genus">
<emphasis id="B9429061D119FFDA32815A84FE9E3FF0" box="[160,353,1179,1203]" italics="true" pageId="7" pageNumber="402">Pteronemacheilus</emphasis>
</taxonomicName>
(Kottelat, ms.).
</paragraph>
<caption id="DF491CFBD11BFFD832505632FBF13301" ID-DOI="http://doi.org/10.5281/zenodo.5357144" ID-Zenodo-Dep="5357144" box="[113,1038,2092,2115]" httpUri="https://zenodo.org/record/5357144/files/figure.png" pageId="5" pageNumber="400" startId="5.[113,148,2093,2115]" targetBox="[113,1451,1656,2077]" targetPageId="5">
<paragraph id="8B894C73D11BFFD832505632FBF13301" blockId="5.[113,1038,2092,2115]" box="[113,1038,2092,2115]" pageId="5" pageNumber="400">
Fig. 8. 
<taxonomicName id="4C3637F0D11BFFD832985632FE6F3301" authority=", CMK" authorityName="CMK" box="[185,400,2093,2114]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="400" phylum="Chordata" rank="species" species="bella">
<emphasis id="B9429061D11BFFD832985632FEB43301" box="[185,331,2093,2114]" italics="true" pageId="5" pageNumber="400">Schistura bella</emphasis>
, CMK
</taxonomicName>
26052, 38.4 mm SL; Laos: Louang Namtha Province: Nam Ma.
</paragraph>
</caption>
<caption id="DF491CFBD118FFDB32A95EA2FB0B3B90" box="[136,1268,189,211]" pageId="6" pageNumber="401" startId="6.[136,191,189,211]">
<paragraph id="8B894C73D118FFDB32A95EA2FB0B3B90" blockId="6.[136,1268,189,211]" box="[136,1268,189,211]" pageId="6" pageNumber="401">
Table 1. Morphometric data of type material of 
<taxonomicName id="4C3637F0D118FFDB30425EA2FCE63B91" box="[611,793,189,210]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="401" phylum="Chordata" rank="species" species="thavonei">
<emphasis id="B9429061D118FFDB30425EA2FCE63B91" box="[611,793,189,210]" italics="true" pageId="6" pageNumber="401">Schistura thavonei</emphasis>
</taxonomicName>
(n=11). Range and mean include holotype data.
</paragraph>
</caption>
<paragraph id="8B894C73D118FFDB30DE5F1AFAB53E3C" pageId="6" pageNumber="401">
<table id="F936BED3D118002232B65F1AFAA33E3C" box="[151,1372,261,1407]" gridcols="4" gridrows="35" pageId="6" pageNumber="401">
<tr id="35064E31D118002232B65F1AFAA33A59" box="[151,1372,261,282]" gridrow="0" pageId="6" pageNumber="401" rowspan-0="1">
<th id="76D7274DD118002230DE5F1AFCA33A59" box="[767,860,261,282]" gridcol="1" gridrow="0" pageId="6" pageNumber="401">
<emphasis id="B9429061D118FFDB30DE5F1AFCA33A59" bold="true" box="[767,860,261,282]" pageId="6" pageNumber="401">Holotype</emphasis>
</th>
<th id="76D7274DD118002231DB5F1AFB8D3A59" box="[1018,1138,261,282]" gridcol="2" gridrow="0" pageId="6" pageNumber="401">
<emphasis id="B9429061D118FFDB36345F1AFBA93A59" bold="true" box="[1045,1110,261,282]" pageId="6" pageNumber="401">Range</emphasis>
</th>
<th id="76D7274DD118002237005F1AFAA33A59" box="[1313,1372,261,282]" gridcol="3" gridrow="0" pageId="6" pageNumber="401">
<emphasis id="B9429061D118FFDB37005F1AFAA33A59" bold="true" box="[1313,1372,261,282]" pageId="6" pageNumber="401">Mean</emphasis>
</th>
</tr>
<tr id="35064E31D118002232B65F24FAA33A12" box="[151,1372,315,337]" gridrow="1" pageId="6" pageNumber="401" rowspan-3="1">
<th id="76D7274DD118002232B65F24FE223A12" box="[151,477,315,337]" gridcol="0" gridrow="1" pageId="6" pageNumber="401">Standard length (mm)</th>
<td id="76D7274DD118002230DE5F24FCA33A12" box="[767,860,315,337]" gridcol="1" gridrow="1" pageId="6" pageNumber="401">44.0</td>
<td id="76D7274DD118002231DB5F24FB8D3A12" box="[1018,1138,315,337]" gridcol="2" gridrow="1" pageId="6" pageNumber="401">41.0–47.6</td>
</tr>
<tr id="35064E31D118002232B65F44FAA33A32" box="[151,1372,347,369]" gridrow="2" pageId="6" pageNumber="401" rowspan-3="1">
<th id="76D7274DD118002232B65F44FE223A32" box="[151,477,347,369]" gridcol="0" gridrow="2" pageId="6" pageNumber="401">Total length (mm)</th>
<td id="76D7274DD118002230DE5F44FCA33A32" box="[767,860,347,369]" gridcol="1" gridrow="2" pageId="6" pageNumber="401">51.7</td>
<td id="76D7274DD118002231DB5F44FB8D3A32" box="[1018,1138,347,369]" gridcol="2" gridrow="2" pageId="6" pageNumber="401">48.5–55.9</td>
</tr>
<tr id="35064E31D118002232B65F9DFAA33ADB" box="[151,1372,386,408]" gridrow="3" pageId="6" pageNumber="401">
<th id="76D7274DD118002232B65F9DFAA33ADB" box="[151,1372,386,408]" colspan="4" colspanRight="3" gridcol="0" gridrow="3" pageId="6" pageNumber="401">
<emphasis id="B9429061D118FFDB32B65F9DFE353AD4" bold="true" box="[151,458,386,408]" pageId="6" pageNumber="401">In percent of standard length</emphasis>
</th>
</tr>
<tr id="35064E31D118002232B65FBDFAA33AFB" box="[151,1372,418,440]" gridrow="4" pageId="6" pageNumber="401">
<th id="76D7274DD118002232B65FBDFE223AFB" box="[151,477,418,440]" gridcol="0" gridrow="4" pageId="6" pageNumber="401">Total length</th>
<td id="76D7274DD118002230DE5FBDFCA33AFB" box="[767,860,418,440]" gridcol="1" gridrow="4" pageId="6" pageNumber="401">117.4</td>
<td id="76D7274DD118002231DB5FBDFB8D3AFB" box="[1018,1138,418,440]" gridcol="2" gridrow="4" pageId="6" pageNumber="401">115.3–119.1</td>
<td id="76D7274DD118002237005FBDFAA33AFB" box="[1313,1372,418,440]" gridcol="3" gridrow="4" pageId="6" pageNumber="401">117.6</td>
</tr>
<tr id="35064E31D118002232B65FDEFAA33A94" box="[151,1372,449,471]" gridrow="5" pageId="6" pageNumber="401">
<th id="76D7274DD118002232B65FDEFE223A94" box="[151,477,449,471]" gridcol="0" gridrow="5" pageId="6" pageNumber="401">Head length (dorsal)</th>
<td id="76D7274DD118002230DE5FDEFCA33A94" box="[767,860,449,471]" gridcol="1" gridrow="5" pageId="6" pageNumber="401">18.0</td>
<td id="76D7274DD118002231DB5FDEFB8D3A94" box="[1018,1138,449,471]" gridcol="2" gridrow="5" pageId="6" pageNumber="401">17.8–20.4</td>
<td id="76D7274DD118002237005FDEFAA33A94" box="[1313,1372,449,471]" gridcol="3" gridrow="5" pageId="6" pageNumber="401">18.5</td>
</tr>
<tr id="35064E31D118002232B65FFEFAA33AB4" box="[151,1372,481,503]" gridrow="6" pageId="6" pageNumber="401">
<th id="76D7274DD118002232B65FFEFE223AB4" box="[151,477,481,503]" gridcol="0" gridrow="6" pageId="6" pageNumber="401">Head length (lateral)</th>
<td id="76D7274DD118002230DE5FFEFCA33AB4" box="[767,860,481,503]" gridcol="1" gridrow="6" pageId="6" pageNumber="401">19.6</td>
<td id="76D7274DD118002231DB5FFEFB8D3AB4" box="[1018,1138,481,503]" gridcol="2" gridrow="6" pageId="6" pageNumber="401">19.6–21.5</td>
<td id="76D7274DD118002237005FFEFAA33AB4" box="[1313,1372,481,503]" gridcol="3" gridrow="6" pageId="6" pageNumber="401">20.6</td>
</tr>
<tr id="35064E31D118002232B65C1EFAA33954" box="[151,1372,513,535]" gridrow="7" pageId="6" pageNumber="401">
<th id="76D7274DD118002232B65C1EFE223954" box="[151,477,513,535]" gridcol="0" gridrow="7" pageId="6" pageNumber="401">Predorsal length</th>
<td id="76D7274DD118002230DE5C1EFCA33954" box="[767,860,513,535]" gridcol="1" gridrow="7" pageId="6" pageNumber="401">54.3</td>
<td id="76D7274DD118002231DB5C1EFB8D3954" box="[1018,1138,513,535]" gridcol="2" gridrow="7" pageId="6" pageNumber="401">52.2–56.0</td>
<td id="76D7274DD118002237005C1EFAA33954" box="[1313,1372,513,535]" gridcol="3" gridrow="7" pageId="6" pageNumber="401">54.4</td>
</tr>
<tr id="35064E31D118002232B65C3EFAA33974" box="[151,1372,545,567]" gridrow="8" pageId="6" pageNumber="401">
<th id="76D7274DD118002232B65C3EFE223974" box="[151,477,545,567]" gridcol="0" gridrow="8" pageId="6" pageNumber="401">Prepelvic length</th>
<td id="76D7274DD118002230DE5C3EFCA33974" box="[767,860,545,567]" gridcol="1" gridrow="8" pageId="6" pageNumber="401">49.5</td>
<td id="76D7274DD118002231DB5C3EFB8D3974" box="[1018,1138,545,567]" gridcol="2" gridrow="8" pageId="6" pageNumber="401">48.8–52.0</td>
<td id="76D7274DD118002237005C3EFAA33974" box="[1313,1372,545,567]" gridcol="3" gridrow="8" pageId="6" pageNumber="401">50.6</td>
</tr>
<tr id="35064E31D118002232B65C5FFAA33915" box="[151,1372,576,598]" gridrow="9" pageId="6" pageNumber="401">
<th id="76D7274DD118002232B65C5FFE223915" box="[151,477,576,598]" gridcol="0" gridrow="9" pageId="6" pageNumber="401">Pre-anus length</th>
<td id="76D7274DD118002230DE5C5FFCA33915" box="[767,860,576,598]" gridcol="1" gridrow="9" pageId="6" pageNumber="401">66.7</td>
<td id="76D7274DD118002231DB5C5FFB8D3915" box="[1018,1138,576,598]" gridcol="2" gridrow="9" pageId="6" pageNumber="401">66.2–69.1</td>
<td id="76D7274DD118002237005C5FFAA33915" box="[1313,1372,576,598]" gridcol="3" gridrow="9" pageId="6" pageNumber="401">67.8</td>
</tr>
<tr id="35064E31D118002232B65C7FFAA33935" box="[151,1372,608,630]" gridrow="10" pageId="6" pageNumber="401">
<th id="76D7274DD118002232B65C7FFE223935" box="[151,477,608,630]" gridcol="0" gridrow="10" pageId="6" pageNumber="401">Pre-anal length</th>
<td id="76D7274DD118002230DE5C7FFCA33935" box="[767,860,608,630]" gridcol="1" gridrow="10" pageId="6" pageNumber="401">75.9</td>
<td id="76D7274DD118002231DB5C7FFB8D3935" box="[1018,1138,608,630]" gridcol="2" gridrow="10" pageId="6" pageNumber="401">74.7–78.7</td>
<td id="76D7274DD118002237005C7FFAA33935" box="[1313,1372,608,630]" gridcol="3" gridrow="10" pageId="6" pageNumber="401">77.2</td>
</tr>
<tr id="35064E31D118002232B65C9FFAA339D5" box="[151,1372,640,662]" gridrow="11" pageId="6" pageNumber="401">
<th id="76D7274DD118002232B65C9FFE2239D5" box="[151,477,640,662]" gridcol="0" gridrow="11" pageId="6" pageNumber="401">Head depth</th>
<td id="76D7274DD118002230DE5C9FFCA339D5" box="[767,860,640,662]" gridcol="1" gridrow="11" pageId="6" pageNumber="401">9.7</td>
<td id="76D7274DD118002231DB5C9FFB8D39D5" box="[1018,1138,640,662]" gridcol="2" gridrow="11" pageId="6" pageNumber="401">9.0–10.2</td>
<td id="76D7274DD118002237005C9FFAA339D5" box="[1313,1372,640,662]" gridcol="3" gridrow="11" pageId="6" pageNumber="401">9.4</td>
</tr>
<tr id="35064E31D118002232B65CBFFAA339F5" box="[151,1372,672,694]" gridrow="12" pageId="6" pageNumber="401">
<th id="76D7274DD118002232B65CBFFE2239F5" box="[151,477,672,694]" gridcol="0" gridrow="12" pageId="6" pageNumber="401">Body depth at dorsal-fin origin</th>
<td id="76D7274DD118002230DE5CBFFCA339F5" box="[767,860,672,694]" gridcol="1" gridrow="12" pageId="6" pageNumber="401">14.9</td>
<td id="76D7274DD118002231DB5CBFFB8D39F5" box="[1018,1138,672,694]" gridcol="2" gridrow="12" pageId="6" pageNumber="401">13.9–16.2</td>
<td id="76D7274DD118002237005CBFFAA339F5" box="[1313,1372,672,694]" gridcol="3" gridrow="12" pageId="6" pageNumber="401">14.9</td>
</tr>
<tr id="35064E31D118002232B65CA0FAA33996" box="[151,1372,703,725]" gridrow="13" pageId="6" pageNumber="401">
<th id="76D7274DD118002232B65CA0FE223996" box="[151,477,703,725]" gridcol="0" gridrow="13" pageId="6" pageNumber="401">Depth of caudal peduncle</th>
<td id="76D7274DD118002230DE5CA0FCA33996" box="[767,860,703,725]" gridcol="1" gridrow="13" pageId="6" pageNumber="401">10.7</td>
<td id="76D7274DD118002231DB5CA0FB8D3996" box="[1018,1138,703,725]" gridcol="2" gridrow="13" pageId="6" pageNumber="401">9.3–11.3</td>
<td id="76D7274DD118002237005CA0FAA33996" box="[1313,1372,703,725]" gridcol="3" gridrow="13" pageId="6" pageNumber="401">10.0</td>
</tr>
<tr id="35064E31D118002232B65CC0FAA339B6" box="[151,1372,735,757]" gridrow="14" pageId="6" pageNumber="401">
<th id="76D7274DD118002232B65CC0FE2239B6" box="[151,477,735,757]" gridcol="0" gridrow="14" pageId="6" pageNumber="401">Length of caudal peduncle</th>
<td id="76D7274DD118002230DE5CC0FCA339B6" box="[767,860,735,757]" gridcol="1" gridrow="14" pageId="6" pageNumber="401">18.0</td>
<td id="76D7274DD118002231DB5CC0FB8D39B6" box="[1018,1138,735,757]" gridcol="2" gridrow="14" pageId="6" pageNumber="401">14.8–18.0</td>
<td id="76D7274DD118002237005CC0FAA339B6" box="[1313,1372,735,757]" gridcol="3" gridrow="14" pageId="6" pageNumber="401">16.5</td>
</tr>
<tr id="35064E31D118002232B65CE0FAA33856" box="[151,1372,767,789]" gridrow="15" pageId="6" pageNumber="401">
<th id="76D7274DD118002232B65CE0FE223856" box="[151,477,767,789]" gridcol="0" gridrow="15" pageId="6" pageNumber="401">Head width</th>
<td id="76D7274DD118002230DE5CE0FCA33856" box="[767,860,767,789]" gridcol="1" gridrow="15" pageId="6" pageNumber="401">11.5</td>
<td id="76D7274DD118002231DB5CE0FB8D3856" box="[1018,1138,767,789]" gridcol="2" gridrow="15" pageId="6" pageNumber="401">11.4–12.7</td>
<td id="76D7274DD118002237005CE0FAA33856" box="[1313,1372,767,789]" gridcol="3" gridrow="15" pageId="6" pageNumber="401">12.0</td>
</tr>
<tr id="35064E31D118002232B65D00FAA33876" box="[151,1372,799,821]" gridrow="16" pageId="6" pageNumber="401">
<th id="76D7274DD118002232B65D00FE223876" box="[151,477,799,821]" gridcol="0" gridrow="16" pageId="6" pageNumber="401">Body width at dorsal-fin origin</th>
<td id="76D7274DD118002230DE5D00FCA33876" box="[767,860,799,821]" gridcol="1" gridrow="16" pageId="6" pageNumber="401">12.4</td>
<td id="76D7274DD118002231DB5D00FB8D3876" box="[1018,1138,799,821]" gridcol="2" gridrow="16" pageId="6" pageNumber="401">10.7–12.9</td>
<td id="76D7274DD118002237005D00FAA33876" box="[1313,1372,799,821]" gridcol="3" gridrow="16" pageId="6" pageNumber="401">11.8</td>
</tr>
<tr id="35064E31D118002232B65D20FAA33816" box="[151,1372,831,853]" gridrow="17" pageId="6" pageNumber="401">
<th id="76D7274DD118002232B65D20FE223816" box="[151,477,831,853]" gridcol="0" gridrow="17" pageId="6" pageNumber="401">Snout length</th>
<td id="76D7274DD118002230DE5D20FCA33816" box="[767,860,831,853]" gridcol="1" gridrow="17" pageId="6" pageNumber="401">7.2</td>
<td id="76D7274DD118002231DB5D20FB8D3816" box="[1018,1138,831,853]" gridcol="2" gridrow="17" pageId="6" pageNumber="401">7.2–8.7</td>
<td id="76D7274DD118002237005D20FAA33816" box="[1313,1372,831,853]" gridcol="3" gridrow="17" pageId="6" pageNumber="401">8.1</td>
</tr>
<tr id="35064E31D118002232B65D41FAA33837" box="[151,1372,862,884]" gridrow="18" pageId="6" pageNumber="401">
<th id="76D7274DD118002232B65D41FE223837" box="[151,477,862,884]" gridcol="0" gridrow="18" pageId="6" pageNumber="401">Eye diameter</th>
<td id="76D7274DD118002230DE5D41FCA33837" box="[767,860,862,884]" gridcol="1" gridrow="18" pageId="6" pageNumber="401">3.1</td>
<td id="76D7274DD118002231DB5D41FB8D3837" box="[1018,1138,862,884]" gridcol="2" gridrow="18" pageId="6" pageNumber="401">2.6–3.4</td>
<td id="76D7274DD118002237005D41FAA33837" box="[1313,1372,862,884]" gridcol="3" gridrow="18" pageId="6" pageNumber="401">2.9</td>
</tr>
<tr id="35064E31D118002232B65D61FAA338D7" box="[151,1372,894,916]" gridrow="19" pageId="6" pageNumber="401">
<th id="76D7274DD118002232B65D61FE2238D7" box="[151,477,894,916]" gridcol="0" gridrow="19" pageId="6" pageNumber="401">Interorbital width</th>
<td id="76D7274DD118002230DE5D61FCA338D7" box="[767,860,894,916]" gridcol="1" gridrow="19" pageId="6" pageNumber="401">5.2</td>
<td id="76D7274DD118002231DB5D61FB8D38D7" box="[1018,1138,894,916]" gridcol="2" gridrow="19" pageId="6" pageNumber="401">4.8–6.1</td>
<td id="76D7274DD118002237005D61FAA338D7" box="[1313,1372,894,916]" gridcol="3" gridrow="19" pageId="6" pageNumber="401">5.4</td>
</tr>
<tr id="35064E31D118002232B65D81FAA338F7" box="[151,1372,926,948]" gridrow="20" pageId="6" pageNumber="401">
<th id="76D7274DD118002232B65D81FE2238F7" box="[151,477,926,948]" gridcol="0" gridrow="20" pageId="6" pageNumber="401">Length of dorsal fin</th>
<td id="76D7274DD118002230DE5D81FCA338F7" box="[767,860,926,948]" gridcol="1" gridrow="20" pageId="6" pageNumber="401">16.2</td>
<td id="76D7274DD118002231DB5D81FB8D38F7" box="[1018,1138,926,948]" gridcol="2" gridrow="20" pageId="6" pageNumber="401">14.9–16.6</td>
<td id="76D7274DD118002237005D81FAA338F7" box="[1313,1372,926,948]" gridcol="3" gridrow="20" pageId="6" pageNumber="401">15.9</td>
</tr>
<tr id="35064E31D118002232B65DA2FAA33897" box="[151,1372,957,980]" gridrow="21" pageId="6" pageNumber="401">
<th id="76D7274DD118002232B65DA2FE223897" box="[151,477,957,980]" gridcol="0" gridrow="21" pageId="6" pageNumber="401">Length of upper caudal-fin lobe</th>
<td id="76D7274DD118002230DE5DA2FCA33897" box="[767,860,957,980]" gridcol="1" gridrow="21" pageId="6" pageNumber="401">16.0</td>
<td id="76D7274DD118002231DB5DA2FB8D3897" box="[1018,1138,957,980]" gridcol="2" gridrow="21" pageId="6" pageNumber="401">16.0–18.5</td>
<td id="76D7274DD118002237005DA2FAA33897" box="[1313,1372,957,980]" gridcol="3" gridrow="21" pageId="6" pageNumber="401">17.2</td>
</tr>
<tr id="35064E31D118002232B65DC2FAA338B0" box="[151,1372,989,1011]" gridrow="22" pageId="6" pageNumber="401">
<th id="76D7274DD118002232B65DC2FE2238B0" box="[151,477,989,1011]" gridcol="0" gridrow="22" pageId="6" pageNumber="401">Length of median caudal-fin rays</th>
<td id="76D7274DD118002230DE5DC2FCA338B0" box="[767,860,989,1011]" gridcol="1" gridrow="22" pageId="6" pageNumber="401">11.8</td>
<td id="76D7274DD118002231DB5DC2FB8D38B0" box="[1018,1138,989,1011]" gridcol="2" gridrow="22" pageId="6" pageNumber="401">11.8–15.8</td>
<td id="76D7274DD118002237005DC2FAA338B0" box="[1313,1372,989,1011]" gridcol="3" gridrow="22" pageId="6" pageNumber="401">13.3</td>
</tr>
<tr id="35064E31D118002232B65DE2FAA33F50" box="[151,1372,1021,1043]" gridrow="23" pageId="6" pageNumber="401">
<th id="76D7274DD118002232B65DE2FE223F50" box="[151,477,1021,1043]" gridcol="0" gridrow="23" pageId="6" pageNumber="401">Length of lower caudal-fin lobe</th>
<td id="76D7274DD118002230DE5DE2FCA33F50" box="[767,860,1021,1043]" gridcol="1" gridrow="23" pageId="6" pageNumber="401">18.8</td>
<td id="76D7274DD118002231DB5DE2FB8D3F50" box="[1018,1138,1021,1043]" gridcol="2" gridrow="23" pageId="6" pageNumber="401">17.2–19.7</td>
<td id="76D7274DD118002237005DE2FAA33F50" box="[1313,1372,1021,1043]" gridcol="3" gridrow="23" pageId="6" pageNumber="401">18.3</td>
</tr>
<tr id="35064E31D118002232B65A02FAA33F70" box="[151,1372,1053,1075]" gridrow="24" pageId="6" pageNumber="401">
<th id="76D7274DD118002232B65A02FE223F70" box="[151,477,1053,1075]" gridcol="0" gridrow="24" pageId="6" pageNumber="401">Length of anal fin</th>
<td id="76D7274DD118002230DE5A02FCA33F70" box="[767,860,1053,1075]" gridcol="1" gridrow="24" pageId="6" pageNumber="401">14.7</td>
<td id="76D7274DD118002231DB5A02FB8D3F70" box="[1018,1138,1053,1075]" gridcol="2" gridrow="24" pageId="6" pageNumber="401">13.5–15.7</td>
<td id="76D7274DD118002237005A02FAA33F70" box="[1313,1372,1053,1075]" gridcol="3" gridrow="24" pageId="6" pageNumber="401">14.8</td>
</tr>
<tr id="35064E31D118002232B65A23FAA33F10" box="[151,1372,1084,1107]" gridrow="25" pageId="6" pageNumber="401">
<th id="76D7274DD118002232B65A23FE223F10" box="[151,477,1084,1107]" gridcol="0" gridrow="25" pageId="6" pageNumber="401">Length of pelvic fin</th>
<td id="76D7274DD118002230DE5A23FCA33F10" box="[767,860,1084,1107]" gridcol="1" gridrow="25" pageId="6" pageNumber="401">15.1</td>
<td id="76D7274DD118002231DB5A23FB8D3F10" box="[1018,1138,1084,1107]" gridcol="2" gridrow="25" pageId="6" pageNumber="401">14.2–16.9</td>
<td id="76D7274DD118002237005A23FAA33F10" box="[1313,1372,1084,1107]" gridcol="3" gridrow="25" pageId="6" pageNumber="401">15.0</td>
</tr>
<tr id="35064E31D118002232B65A43FAA33F31" box="[151,1372,1116,1138]" gridrow="26" pageId="6" pageNumber="401">
<th id="76D7274DD118002232B65A43FE223F31" box="[151,477,1116,1138]" gridcol="0" gridrow="26" pageId="6" pageNumber="401">Length of pectoral fin</th>
<td id="76D7274DD118002230DE5A43FCA33F31" box="[767,860,1116,1138]" gridcol="1" gridrow="26" pageId="6" pageNumber="401">16.6</td>
<td id="76D7274DD118002231DB5A43FB8D3F31" box="[1018,1138,1116,1138]" gridcol="2" gridrow="26" pageId="6" pageNumber="401">15.8–17.4</td>
<td id="76D7274DD118002237005A43FAA33F31" box="[1313,1372,1116,1138]" gridcol="3" gridrow="26" pageId="6" pageNumber="401">16.5</td>
</tr>
<tr id="35064E31D118002232B65A9CFAA33FDA" box="[151,1372,1155,1177]" gridrow="27" pageId="6" pageNumber="401">
<th id="76D7274DD118002232B65A9CFAA33FDA" box="[151,1372,1155,1177]" colspan="4" colspanRight="3" gridcol="0" gridrow="27" pageId="6" pageNumber="401">
<emphasis id="B9429061D118FFDB32B65A9CFE183FDB" bold="true" box="[151,487,1155,1177]" pageId="6" pageNumber="401">In percent of dorsal head length</emphasis>
</th>
</tr>
<tr id="35064E31D118002232B65ABCFAA33FFA" box="[151,1372,1187,1209]" gridrow="28" pageId="6" pageNumber="401">
<th id="76D7274DD118002232B65ABCFE223FFA" box="[151,477,1187,1209]" gridcol="0" gridrow="28" pageId="6" pageNumber="401">Snout length</th>
<td id="76D7274DD118002230DE5ABCFCA33FFA" box="[767,860,1187,1209]" gridcol="1" gridrow="28" pageId="6" pageNumber="401">40</td>
<td id="76D7274DD118002231DB5ABCFB8D3FFA" box="[1018,1138,1187,1209]" gridcol="2" gridrow="28" pageId="6" pageNumber="401">39–48</td>
<td id="76D7274DD118002237005ABCFAA33FFA" box="[1313,1372,1187,1209]" gridcol="3" gridrow="28" pageId="6" pageNumber="401">44</td>
</tr>
<tr id="35064E31D118002232B65ADCFAA33F9A" box="[151,1372,1219,1241]" gridrow="29" pageId="6" pageNumber="401">
<th id="76D7274DD118002232B65ADCFE223F9A" box="[151,477,1219,1241]" gridcol="0" gridrow="29" pageId="6" pageNumber="401">Eye diameter</th>
<td id="76D7274DD118002230DE5ADCFCA33F9A" box="[767,860,1219,1241]" gridcol="1" gridrow="29" pageId="6" pageNumber="401">17</td>
<td id="76D7274DD118002231DB5ADCFB8D3F9A" box="[1018,1138,1219,1241]" gridcol="2" gridrow="29" pageId="6" pageNumber="401">14–19</td>
<td id="76D7274DD118002237005ADCFAA33F9A" box="[1313,1372,1219,1241]" gridcol="3" gridrow="29" pageId="6" pageNumber="401">16</td>
</tr>
<tr id="35064E31D118002232B65AFCFAA33FBA" box="[151,1372,1251,1273]" gridrow="30" pageId="6" pageNumber="401">
<th id="76D7274DD118002232B65AFCFE223FBA" box="[151,477,1251,1273]" gridcol="0" gridrow="30" pageId="6" pageNumber="401">Interorbital width</th>
<td id="76D7274DD118002230DE5AFCFCA33FBA" box="[767,860,1251,1273]" gridcol="1" gridrow="30" pageId="6" pageNumber="401">29</td>
<td id="76D7274DD118002231DB5AFCFB8D3FBA" box="[1018,1138,1251,1273]" gridcol="2" gridrow="30" pageId="6" pageNumber="401">26–33</td>
<td id="76D7274DD118002237005AFCFAA33FBA" box="[1313,1372,1251,1273]" gridcol="3" gridrow="30" pageId="6" pageNumber="401">29</td>
</tr>
<tr id="35064E31D118002232B65B16FAA33E5C" box="[151,1372,1289,1311]" gridrow="31" pageId="6" pageNumber="401">
<th id="76D7274DD118002232B65B16FAA33E5C" box="[151,1372,1289,1311]" colspan="4" colspanRight="3" gridcol="0" gridrow="31" pageId="6" pageNumber="401">
<emphasis id="B9429061D118FFDB32B65B15FE153E5C" bold="true" box="[151,490,1289,1311]" pageId="6" pageNumber="401">In percent of lateral head length</emphasis>
</th>
</tr>
<tr id="35064E31D118002232B65B36FAA33E7C" box="[151,1372,1321,1343]" gridrow="32" pageId="6" pageNumber="401">
<th id="76D7274DD118002232B65B36FE223E7C" box="[151,477,1321,1343]" gridcol="0" gridrow="32" pageId="6" pageNumber="401">Snout length</th>
<td id="76D7274DD118002230DE5B36FCA33E7C" box="[767,860,1321,1343]" gridcol="1" gridrow="32" pageId="6" pageNumber="401">37</td>
<td id="76D7274DD118002231DB5B36FB8D3E7C" box="[1018,1138,1321,1343]" gridcol="2" gridrow="32" pageId="6" pageNumber="401">35–43</td>
<td id="76D7274DD118002237005B36FAA33E7C" box="[1313,1372,1321,1343]" gridcol="3" gridrow="32" pageId="6" pageNumber="401">39</td>
</tr>
<tr id="35064E31D118002232B65B56FAA33E1C" box="[151,1372,1353,1375]" gridrow="33" pageId="6" pageNumber="401">
<th id="76D7274DD118002232B65B56FE223E1C" box="[151,477,1353,1375]" gridcol="0" gridrow="33" pageId="6" pageNumber="401">Eye diameter</th>
<td id="76D7274DD118002230DE5B56FCA33E1C" box="[767,860,1353,1375]" gridcol="1" gridrow="33" pageId="6" pageNumber="401">16</td>
<td id="76D7274DD118002231DB5B56FB8D3E1C" box="[1018,1138,1353,1375]" gridcol="2" gridrow="33" pageId="6" pageNumber="401">12–16</td>
<td id="76D7274DD118002237005B56FAA33E1C" box="[1313,1372,1353,1375]" gridcol="3" gridrow="33" pageId="6" pageNumber="401">14</td>
</tr>
<tr id="35064E31D118002232B65B76FAA33E3C" box="[151,1372,1385,1407]" gridrow="34" pageId="6" pageNumber="401">
<th id="76D7274DD118002232B65B76FE223E3C" box="[151,477,1385,1407]" gridcol="0" gridrow="34" pageId="6" pageNumber="401">Interorbital width</th>
<td id="76D7274DD118002230DE5B76FCA33E3C" box="[767,860,1385,1407]" gridcol="1" gridrow="34" pageId="6" pageNumber="401">27</td>
<td id="76D7274DD118002231DB5B76FB8D3E3C" box="[1018,1138,1385,1407]" gridcol="2" gridrow="34" pageId="6" pageNumber="401">23–29</td>
<td id="76D7274DD118002237005B76FAA33E3C" box="[1313,1372,1385,1407]" gridcol="3" gridrow="34" pageId="6" pageNumber="401">26</td>
</tr>
</table>
</paragraph>
<caption id="DF491CFBD118FFDB32A9592BFA913C09" ID-DOI="http://doi.org/10.5281/zenodo.5357146" ID-Zenodo-Dep="5357146" box="[136,1390,1844,1867]" httpUri="https://zenodo.org/record/5357146/files/figure.png" pageId="6" pageNumber="401" startId="6.[136,171,1844,1866]" targetBox="[135,1474,1486,1829]" targetPageId="6">
<paragraph id="8B894C73D118FFDB32A9592BFA913C09" blockId="6.[136,1390,1844,1867]" box="[136,1390,1844,1867]" pageId="6" pageNumber="401">
Fig. 9. 
<taxonomicName id="4C3637F0D118FFDB32EE592BFE6E3C0A" baseAuthorityName="Zhu" baseAuthorityYear="1982" box="[207,401,1844,1866]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="401" phylum="Chordata" rank="species" species="longa">
<emphasis id="B9429061D118FFDB32EE592BFED53C0A" box="[207,298,1844,1865]" italics="true" pageId="6" pageNumber="401">Schistura</emphasis>
aff. 
<emphasis id="B9429061D118FFDB337D592BFE6E3C0A" box="[348,401,1844,1865]" italics="true" pageId="6" pageNumber="401">longa</emphasis>
</taxonomicName>
, 78.0 mm SL; China: Yunnan: “Fugong” (?) (photographed at Yunnan University, Kunming, 1986).
</paragraph>
</caption>
<caption id="DF491CFBD119FFDA32505C51FBA53927" ID-DOI="http://doi.org/10.5281/zenodo.5357148" ID-Zenodo-Dep="5357148" box="[113,1114,590,612]" httpUri="https://zenodo.org/record/5357148/files/figure.png" pageId="7" pageNumber="402" startId="7.[113,148,590,612]" targetBox="[113,1451,189,575]" targetPageId="7">
<paragraph id="8B894C73D119FFDA32505C51FBA53927" blockId="7.[113,1114,590,612]" box="[113,1114,590,612]" pageId="7" pageNumber="402">
Fig. 10. 
<taxonomicName id="4C3637F0D119FFDA32E45C51FE273920" authority=", KIZ" authorityName="KIZ" box="[197,472,590,612]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="402" phylum="Chordata" rank="species" species="thavonei">
<emphasis id="B9429061D119FFDA32E45C51FEDF3920" box="[197,288,590,611]" italics="true" pageId="7" pageNumber="402">Schistura</emphasis>
aff. 
<emphasis id="B9429061D119FFDA33735C51FE5C3920" box="[338,419,590,611]" italics="true" pageId="7" pageNumber="402">thavonei</emphasis>
, KIZ
</taxonomicName>
uncat., 39.9 mm SL; China: Yunnan: Mekong drainage: Yangbi.
</paragraph>
</caption>
<paragraph id="8B894C73D119FFDA32505AC4FE553D30" blockId="7.[113,753,1243,1651]" pageId="7" pageNumber="402">
There are no clues to the affinities of 
<taxonomicName id="4C3637F0D119FFDA30245AC4FD833FB0" box="[517,636,1243,1267]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="402" phylum="Chordata" rank="species" species="thavonei">
<emphasis id="B9429061D119FFDA30245AC4FD833FB0" box="[517,636,1243,1267]" italics="true" pageId="7" pageNumber="402">S. thavonei</emphasis>
</taxonomicName>
. There are a few species of 
<taxonomicName id="4C3637F0D119FFDA33095AE4FE723E50" box="[296,397,1275,1299]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="402" phylum="Chordata" rank="genus">
<emphasis id="B9429061D119FFDA33095AE4FE723E50" box="[296,397,1275,1299]" italics="true" pageId="7" pageNumber="402">Schistura</emphasis>
</taxonomicName>
with a similarly elongated body, of which apparently only one is named, 
<taxonomicName id="4C3637F0D119FFDA300E5B04FD763E70" baseAuthorityName="Zhu" baseAuthorityYear="1982" box="[559,649,1307,1331]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="402" phylum="Chordata" rank="species" species="longa">
<emphasis id="B9429061D119FFDA300E5B04FD763E70" box="[559,649,1307,1331]" italics="true" pageId="7" pageNumber="402">S. longa</emphasis>
</taxonomicName>
from the upper-middle Salween in 
<collectingRegion id="49F28291D119FFDA33AE5B23FE1B3E10" box="[399,484,1340,1363]" country="China" name="Yunnan" pageId="7" pageNumber="402">Yunnan</collectingRegion>
(
<bibRefCitation id="EFA73182D119FFDA33D75B24FD5B3E10" author="Zhu S-Q" box="[502,676,1339,1363]" pageId="7" pageNumber="402" pagination="104 - 111" refId="ref5841" refString="Zhu S-Q (1982) Five new species of fishes of the genus Nemachilus from Yunnan Province, China. Acta Zootaxonomica Sinica, 7 (1): 104 - 111. [In Chinese, English summary]." type="journal article" year="1982">Zhu, 1982: 105</bibRefCitation>
, 
<bibRefCitation id="EFA73182D119FFDA30925B24FF703E30" author="Zhu S-Q" pageId="7" pageNumber="402" refId="ref5882" refString="Zhu S-Q (1989) The loaches of the subfamily Nemacheilinae in China (Cypriniformes: Cobitidae). Jiangsu Science and Technology Publishing House, Nanjing, 150 pp. [In Chinese, English summary]." type="book" year="1989">1989: 51</bibRefCitation>
). Indeed, 
<taxonomicName id="4C3637F0D119FFDA32DA5B44FE8C3E30" box="[251,371,1371,1395]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="402" phylum="Chordata" rank="species" species="thavonei">
<emphasis id="B9429061D119FFDA32DA5B44FE8C3E30" box="[251,371,1371,1395]" italics="true" pageId="7" pageNumber="402">S. thavonei</emphasis>
</taxonomicName>
was figured and identified as 
<emphasis id="B9429061D119FFDA30975B44FD353E31" box="[694,714,1371,1394]" italics="true" pageId="7" pageNumber="402">S.</emphasis>
sp. 
<taxonomicName id="4C3637F0D119FFDA32535B64FF2D3ED0" baseAuthorityName="Zhu" baseAuthorityYear="1982" box="[114,210,1403,1427]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="402" phylum="Chordata" rank="species" species="longa">
cf. 
<emphasis id="B9429061D119FFDA32B45B64FF2D3ED0" box="[149,210,1403,1427]" italics="true" pageId="7" pageNumber="402">longa</emphasis>
</taxonomicName>
by 
<bibRefCitation id="EFA73182D119FFDA32DD5B64FE193ED0" author="Rainboth WJ &amp; Vidthayanon C &amp; Mai DY" box="[252,486,1403,1427]" pageId="7" pageNumber="402" pagination="1 - 173" refId="ref5793" refString="Rainboth WJ, Vidthayanon C &amp; Mai DY (2012) Fishes of the Greater Mekong ecosystem with species list and photographic atlas. Miscellaneous Publications, Museum of Zoology, University of Michigan, 201: i - vi, 1 - 173, 121 pls." type="journal article" year="2012">Rainboth et al. (2012</bibRefCitation>
: pl. 36). 
<taxonomicName id="4C3637F0D119FFDA30665B64FD0F3ED0" baseAuthorityName="Zhu" baseAuthorityYear="1982" box="[583,752,1403,1427]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="402" phylum="Chordata" rank="species" species="longa">
<emphasis id="B9429061D119FFDA30665B64FD0F3ED0" box="[583,752,1403,1427]" italics="true" pageId="7" pageNumber="402">Schistura longa</emphasis>
</taxonomicName>
does not have stripes on the body, the bars on the posterior half of the body are fewer, broader and extend from dorsal midline almost to ventral midline. The figure of 
<taxonomicName id="4C3637F0D119FFDA30565BC4FD303EB0" baseAuthorityName="Zhu" baseAuthorityYear="1982" box="[631,719,1499,1523]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="402" phylum="Chordata" rank="species" species="longa">
<emphasis id="B9429061D119FFDA30565BC4FD303EB0" box="[631,719,1499,1523]" italics="true" pageId="7" pageNumber="402">S. longa</emphasis>
</taxonomicName>
by Yang (in 
<bibRefCitation id="EFA73182D119FFDA32FD5BE4FE243D50" author="Chu X-L &amp; Chen Y-R" box="[220,475,1531,1555]" pageId="7" pageNumber="402" refId="ref4863" refString="Chu X-L &amp; Chen Y-R (1990) (eds.) The fishes of Yunnan, China. Part II. Science Press, Beijing, 313 pp. [In Chinese]." type="book" year="1990">Chu &amp; Chen, 1990: 43</bibRefCitation>
) shows the black pattern at the base of the caudal fin made of a vertical bar; it is possibly misidentified and might be the apparently unnamed species figured here (
<figureCitation id="130D50F6D119FFDA337B5844FE613D30" box="[346,414,1627,1651]" captionStart="Fig" captionStartId="6.[136,171,1844,1866]" captionTargetBox="[135,1474,1486,1829]" captionTargetId="figure-240@6.[135,1474,1486,1829]" captionTargetPageId="6" captionText="Fig. 9. Schistura aff. longa, 78.0 mm SL; China: Yunnan: “Fugong” (?) (photographed at Yunnan University, Kunming, 1986)." figureDoi="http://doi.org/10.5281/zenodo.5357146" httpUri="https://zenodo.org/record/5357146/files/figure.png" pageId="7" pageNumber="402">Fig. 9</figureCitation>
).
</paragraph>
<paragraph id="8B894C73D119FFDA32505884FE0E3C37" blockId="7.[113,753,1691,1908]" pageId="7" pageNumber="402">
A number of other elongate nemacheilid species exist, unnamed, including one from the Nam Ngiep in 
<collectingCountry id="F3210CE3D119FFDA30AA58A3FD403D90" box="[651,703,1724,1747]" name="Laos" pageId="7" pageNumber="402">Laos</collectingCountry>
that will be described elsewhere, and a species I collected in Yangbi (Yunnan) in 1986 that has some similarities with 
<taxonomicName id="4C3637F0D119FFDA32505904FF143C70" box="[113,235,1819,1843]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="402" phylum="Chordata" rank="species" species="thavonei">
<emphasis id="B9429061D119FFDA32505904FF143C70" box="[113,235,1819,1843]" italics="true" pageId="7" pageNumber="402">S. thavonei</emphasis>
</taxonomicName>
(
<figureCitation id="130D50F6D119FFDA32DC5904FEAF3C70" box="[253,336,1819,1843]" captionStart="Fig" captionStartId="7.[113,148,590,612]" captionTargetBox="[113,1451,189,575]" captionTargetId="figure-765@7.[113,1451,189,575]" captionTargetPageId="7" captionText="Fig. 10. Schistura aff. thavonei, KIZ uncat., 39.9 mm SL; China: Yunnan: Mekong drainage: Yangbi." figureDoi="http://doi.org/10.5281/zenodo.5357148" httpUri="https://zenodo.org/record/5357148/files/figure.png" pageId="7" pageNumber="402">Fig. 10</figureCitation>
). Unfortunately, I do not have access to this material. Future research is likely to show that 
<taxonomicName id="4C3637F0D119FFDA30FD5924FF323C30" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="402" phylum="Chordata" rank="species" species="thavonei">
<emphasis id="B9429061D119FFDA30FD5924FF323C30" italics="true" pageId="7" pageNumber="402">S. thavonei</emphasis>
</taxonomicName>
represent a distinct genus.
</paragraph>
<paragraph id="8B894C73D119FFDA32505984FE5B3CB7" blockId="7.[113,753,1947,2036]" pageId="7" pageNumber="402">
<emphasis id="B9429061D119FFDA32505984FE9A3CF0" bold="true" box="[113,357,1947,1971]" pageId="7" pageNumber="402">Comparison material.</emphasis>
<taxonomicName id="4C3637F0D119FFDA33485984FDB33CF0" authority=": CMK" authorityName="CMK" box="[361,588,1947,1971]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="402" phylum="Chordata" rank="species" species="bella">
<emphasis id="B9429061D119FFDA33485984FDFB3CF0" box="[361,516,1947,1971]" italics="true" pageId="7" pageNumber="402">Schistura bella</emphasis>
: 
<materialsCitation id="3B5E462ED119FFDA30315984FDB33CF0" ID-GBIF-Occurrence="3340077303" accessionNumber="CMK 26052" box="[528,588,1947,1971]" country="Laos" county="Muang Long District" location="Nam Long" municipality="Nam Ma" pageId="7" pageNumber="402" specimenCount="1" stateProvince="Louangnamtha">CMK</materialsCitation>
</taxonomicName>
26052, 7; 
<collectingCountry id="F3210CE3D119FFDA30975983FD123CF0" box="[694,749,1948,1971]" name="Laos" pageId="7" pageNumber="402">Laos</collectingCountry>
: 
<collectingRegion id="49F28291D119FFDA325059A3FE793C90" box="[113,390,1979,2004]" country="Laos" name="Louangnamtha" pageId="7" pageNumber="402">Louang Namtha Province</collectingRegion>
: 
<collectingCounty id="62E834FFD119FFDA33AE59A3FD8C3C90" box="[399,627,1979,2004]" pageId="7" pageNumber="402">Muang Long District</collectingCounty>
: confluence of 
<collectingMunicipality id="6BEDD609D119FFDA32B159C3FF0E3CB0" box="[144,241,2012,2035]" pageId="7" pageNumber="402">Nam Ma</collectingMunicipality>
and 
<location id="8EE91AA8D119FFDA330959C3FE5E3CB7" LSID="urn:lsid:plazi:treatment:039FFD65D11FFFDA32DF5831FE5B3CB7:8EE91AA8D119FFDA330959C3FE5E3CB7" box="[296,417,2012,2036]" country="Laos" county="Muang Long District" municipality="Nam Ma" name="Nam Long" pageId="7" pageNumber="402" stateProvince="Louangnamtha">Nam Long</location>
.
</paragraph>
</subSubSection>
</treatment>
</document>