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<document id="A3D1FBBBC4568DC024643F8AED3F8536" ID-DOI="10.5281/zenodo.198863" ID-GBIF-Dataset="6f286e35-670c-4bb9-920b-604c51cecd9f" ID-ISSN="1175-5326" ID-Zenodo-Dep="198863" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460315829380" checkinUser="plazi" docAuthor="Anker, Arthur" docDate="2010" docId="039F5848FF9AFFE5E5F3F955FECDCD29" docLanguage="en" docName="zt02652p063.pdf" docOrigin="Zootaxa 2652" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Triacanthoneus toro Anker, 2010, n. sp." docType="treatment" docVersion="8" lastPageNumber="54" masterDocId="FFA62030FF98FFE2E564FFD8FFC6CE05" masterDocTitle="A new genus and three new species of alpheid shrimps (Crustacea, Decapoda, Caridea) from the tropical American coasts" masterLastPageNumber="63" masterPageNumber="47" pageNumber="49" updateTime="1698255077161" updateUser="plazi">
<mods:mods id="BA22109891063943FBAD3FDCF270CDD6" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E6841678E08C6D4E53255324188EDB24">
<mods:title id="8B724819AAB832BF52EC2270568B50DF">A new genus and three new species of alpheid shrimps (Crustacea, Decapoda, Caridea) from the tropical American coasts</mods:title>
</mods:titleInfo>
<mods:name id="544D2462D05DAA894A64DE37389079AF" type="personal">
<mods:role id="6BE90E3B4E80616388BEE29E9E360E2F">
<mods:roleTerm id="DBD8815DCAC51EDB252B5E4B57F26B36">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0B986E49D6C3CD1BC5795CEDF1E808E6">Anker, Arthur</mods:namePart>
</mods:name>
<mods:typeOfResource id="5FEFBFA89324A10DCEFFCAF6B9C7F23B">text</mods:typeOfResource>
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<mods:title id="1DB8610803325E4B23185413107A6AD2">Zootaxa</mods:title>
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<mods:part id="90288EDDB35A390EDF7C3F70CA422150">
<mods:date id="62D4F7A507A1F355B328438A9723945E">2010</mods:date>
<mods:detail id="00BAF71738A1EA2A68F74ED7CCC58489" type="volume">
<mods:number id="17E3D80B55FCC1F320996197A29F64DA">2652</mods:number>
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<mods:classification id="35A9515292308C39783031933F5A2FCB">journal article</mods:classification>
<mods:identifier id="B5953838BD02CC26C8A67C7E9A13DD23" type="DOI">10.5281/zenodo.198863</mods:identifier>
<mods:identifier id="36F883111DEC3D961BCC31699D6E694C" type="GBIF-Dataset">6f286e35-670c-4bb9-920b-604c51cecd9f</mods:identifier>
<mods:identifier id="0BA46E256DE1CEA13CAB213FBF56A180" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="5372B27873E66869C072E6FA14D8B5CF" type="Zenodo-Dep">198863</mods:identifier>
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<treatment id="039F5848FF9AFFE5E5F3F955FECDCD29" ID-DOI="http://doi.org/10.5281/zenodo.6206916" ID-GBIF-Taxon="119401710" ID-Zenodo-Dep="6206916" LSID="urn:lsid:plazi:treatment:039F5848FF9AFFE5E5F3F955FECDCD29" httpUri="http://treatment.plazi.org/id/039F5848FF9AFFE5E5F3F955FECDCD29" lastPageId="7" lastPageNumber="54" pageId="2" pageNumber="49">
<subSubSection id="C32CBAD5FF9AFFE0E5F3F955FEBCC8CF" pageId="2" pageNumber="49" type="nomenclature">
<paragraph id="8B89E95EFF9AFFE0E5F3F955FE1EC8A2" blockId="2.[151,472,1677,1738]" box="[151,472,1677,1703]" pageId="2" pageNumber="49">
<heading id="D0C15E32FF9AFFE0E5F3F955FE1EC8A2" bold="true" box="[151,472,1677,1703]" fontSize="11" level="1" pageId="2" pageNumber="49" reason="1">
<emphasis id="B942354CFF9AFFE0E5F3F955FE1EC8A2" bold="true" box="[151,472,1677,1703]" pageId="2" pageNumber="49">
<taxonomicName id="4C3692DDFF9AFFE0E5F3F955FE48C8A2" ID-CoL="85ZFV" box="[151,398,1677,1703]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="49" phylum="Arthropoda" rank="species" species="toro" status="sp. nov.">
<emphasis id="B942354CFF9AFFE0E5F3F955FE48C8A2" bold="true" box="[151,398,1677,1703]" italics="true" pageId="2" pageNumber="49">Triacanthoneus toro</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2718837FF9AFFE0E4F1F955FE1EC8A2" box="[405,472,1677,1703]" pageId="2" pageNumber="49" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B89E95EFF9AFFE0E5F3F968FEBCC8CF" blockId="2.[151,472,1677,1738]" box="[151,378,1712,1738]" pageId="2" pageNumber="49">
(
<figureCitation id="130DF5DBFF9AFFE0E5FBF968FEC9C8CF" box="[159,271,1712,1738]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="3.[151,259,1373,1397]" captionStartId-1="4.[151,259,1821,1845]" captionStartId-2="5.[151,259,1424,1448]" captionStartId-3="6.[151,259,1544,1568]" captionTargetBox-0="[152,1432,538,1343]" captionTargetBox-1="[195,1375,200,1771]" captionTargetBox-2="[157,1425,188,1393]" captionTargetBox-3="[154,1433,226,1501]" captionTargetId-0="figure@3.[151,1436,530,1352]" captionTargetId-1="figure@4.[193,1379,194,1782]" captionTargetId-2="figure@5.[151,1436,174,1418]" captionTargetId-3="figure@6.[151,1436,194,1523]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionText-0="FIGURE 1. Triacanthoneus toro n. gen., n. sp., paratype, ovigerous female (CL 3.4 mm) from Bocas del Toro, Caribbean coast of Panama, USNM 1145376: habitus, lateral." captionText-1="FIGURE 2. Triacanthoneus toro n. gen., n. sp., paratype, ovigerous female (CL 3.4 mm) from Bocas del Toro, Caribbean coast of Panama, USNM 1145376 [A – M]; paratype, ovigerous female (CL 3.4 mm) from the same locality, OUMNH. ZC 2010 - 01 - 005 [N]: A, frontal region, dorsal; B, carapace and frontal appendages, lateral; C, rostrum, lateral; D, carapace, dorsal; E, tooth on mesioventral carina of first segment of antennular peduncle, lateral; F, antennular flagella, dorsal; G, antenna, ventral; H, second pleopod, mesial; I, same, appendix masculina and appendix interna; J, uropod, dorsal; K, same, posterior region of exopod; L, telson, dorsal; M, same, posterior region; N, rostrum, lateral." captionText-2="FIGURE 3. Triacanthoneus toro n. gen., n. sp., paratype, ovigerous female (CL 3.4 mm) from Bocas del Toro, Caribbean coast of Panama, USNM 1145376: A, mandible, mesial; B, same, incisor process, lateral; C, maxillule, lateral; D, maxilla, lateral; E, first maxilliped, lateral; F, second maxilliped, lateral; G, third maxilliped, lateral." captionText-3="FIGURE 4. Triacanthoneus toro n. gen., n. sp., paratype, ovigerous female (CL 3.4 mm) from Bocas del Toro, Caribbean coast of Panama, USNM 1145376: A, right (major) cheliped, lateral; B, same, carpus and chela, mesial; C, same, chela fingers; D, left (minor) cheliped, lateral; E, same, chela, mesial; F, second pereiopod, lateral; G, third pereiopod, lateral; H, same, dactylus; I, fifth pereiopod, lateral." httpUri-0="https://zenodo.org/record/198864/files/figure.png" httpUri-1="https://zenodo.org/record/198865/files/figure.png" httpUri-2="https://zenodo.org/record/198866/files/figure.png" httpUri-3="https://zenodo.org/record/198867/files/figure.png" pageId="2" pageNumber="49">Figs. 1–4</figureCitation>
,
<figureCitation id="130DF5DBFF9AFFE0E478F968FEFEC8CF" box="[284,312,1712,1738]" captionStart="FIGURE 10" captionStartId="14.[151,256,1843,1867]" captionTargetBox="[151,1436,493,1821]" captionTargetId="figure@14.[151,1436,493,1822]" captionTargetPageId="14" captionText="FIGURE 10. Life colouration of three species of Triacanthoneus n. gen.: A – C, Triacanthoneus toro n. sp., paratype, ovigerous female from Bocas del Toro, Caribbean coast of Panama, USNM 1145376; D – F, Triacanthoneus pacificus n. sp., holotype, ovigerous female from Playa Venao, Pacific coast of Panama, USNM 1145374 [D, E], non-type ovigerous female from Bahía Malaga, Pacific coast of Colombia, UVC [F]; G, Triacanthoneus alacranes n. sp., holotype, ovigerous female from Alacranes Reef, Yucatán, Mexico, UNAM-CNCR 26177 [G by J. Duarte-Gutiérrez, all other photographs by author]." httpUri="https://zenodo.org/record/198873/files/figure.png" pageId="2" pageNumber="49">10</figureCitation>
A–C)
</paragraph>
</subSubSection>
<subSubSection id="C32CBAD5FF9AFFE1E5F3F922FC37CED2" lastPageId="3" lastPageNumber="50" pageId="2" pageNumber="49" type="materials_examined">
<paragraph id="8B89E95EFF9AFFE1E5F3F922FC37CED2" blockId="2.[151,1436,1786,2036]" lastBlockId="3.[151,1437,152,476]" lastPageId="3" lastPageNumber="50" pageId="2" pageNumber="49">
<emphasis id="B942354CFF9AFFE0E5F3F922FE8DC911" bold="true" box="[151,331,1786,1812]" pageId="2" pageNumber="49">
<typeStatus id="548D57FCFF9AFFE0E5F3F922FF14C911" box="[151,210,1786,1812]" pageId="2" pageNumber="49">Type</typeStatus>
material.
</emphasis>
<typeStatus id="548D57FCFF9AFFE0E430F922FE00C911" box="[340,454,1786,1812]" pageId="2" pageNumber="49" type="holotype">Holotype</typeStatus>
: ovigerous female (CL
<quantity id="4CCE44BBFF9AFFE0E7BEF922FCFEC911" box="[730,824,1786,1812]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="2" pageNumber="49" unit="mm" value="4.1">4.1 mm</quantity>
),
<collectionCode id="ED27719BFF9AFFE0E629F922FC64C911" LSID="urn:lsid:biocol.org:col:34871" box="[845,930,1786,1812]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="2" pageNumber="49">USNM</collectionCode>
1145375,
<collectingCountry id="F321A9CEFF9AFFE0E179F922FBBBC911" box="[1053,1149,1786,1812]" name="Panama" pageId="2" pageNumber="49">Panama</collectingCountry>
, Caribbean coast, Bocas del
<taxonomicName id="4C3692DDFF9AFFE0E5A6F8F8FF3AC93F" box="[194,252,1824,1850]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="62" phylum="Arthropoda" rank="species" species="toro" status="sp. nov.">Toro</taxonomicName>
,
<collectingCountry id="F321A9CEFF9AFFE0E463F8F8FEF7C93F" box="[263,305,1824,1850]" name="Iceland" pageId="2" pageNumber="49">Isla</collectingCountry>
Colón, Punta Caracol, subtidal sand flat with rubble and sparse seagrass, near mangroves, depth
<quantity id="4CCE44BBFF9AFFE0E5F3F89DFF1BC95A" box="[151,221,1861,1887]" metricMagnitude="0" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="49" unit="m" value="1.5">1.5 m</quantity>
, under rubble in sand, coll. A. Anker,
<date id="FF88CF9EFF9AFFE0E7C4F89DFCFCC95A" box="[672,826,1861,1887]" pageId="2" pageNumber="49" value="2008-04-01">1 April 2008</date>
, fcn 08-022.
<typeStatus id="548D57FCFF9AFFE0E6B1F89DFB8AC95A" box="[981,1100,1861,1887]" pageId="2" pageNumber="49" type="paratype">Paratypes</typeStatus>
: 1 ovigerous female (CL
<quantity id="4CCE44BBFF9AFFE0E01CF89DFF0FC981" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="2" pageNumber="49" unit="mm" value="3.4">3.4 mm</quantity>
),
<collectionCode id="ED27719BFF9AFFE0E5B8F8B2FEF7C981" LSID="urn:lsid:biocol.org:col:34871" box="[220,305,1898,1924]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="2" pageNumber="49">USNM</collectionCode>
1145376,
<collectingCountry id="F321A9CEFF9AFFE0E4CFF8B2FDCDC981" box="[427,523,1898,1924]" name="Panama" pageId="2" pageNumber="49">Panama</collectingCountry>
, Caribbean coast, Bocas del
<taxonomicName id="4C3692DDFF9AFFE0E63CF8B2FC54C981" box="[856,914,1898,1924]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="62" phylum="Arthropoda" rank="species" species="toro" status="sp. nov.">Toro</taxonomicName>
,
<collectingCountry id="F321A9CEFF9AFFE0E6F8F8B2FC00C981" box="[924,966,1898,1924]" name="Iceland" pageId="2" pageNumber="49">Isla</collectingCountry>
Colón, Punta Caracol, subtidal sand flat with rubble and sparse seagrass, near mangroves, depth
<quantity id="4CCE44BBFF9AFFE0E62DF848FC54C9AF" box="[841,914,1936,1962]" metricMagnitude="0" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="49" unit="m" value="1.5">1.5 m</quantity>
, under rubble in sand, coll. A. Anker, J.A. Baeza, I. Marin,
<date id="FF88CF9EFF9AFFE0E438F86DFDC4C9CA" box="[348,514,1973,1999]" pageId="2" pageNumber="49" value="2007-04-28">28 April 2007</date>
, fcn 07-156 (dissected); 1 ovigerous female (CL
<quantity id="4CCE44BBFF9AFFE0E126F86DFB66C9CA" box="[1090,1184,1973,1999]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="2" pageNumber="49" unit="mm" value="3.4">3.4 mm</quantity>
),
<collectionCode id="ED27719BFF9AFFE0E1D0F86DFAE0C9CA" box="[1204,1318,1973,1999]" pageId="2" pageNumber="49">OUMNH</collectionCode>
.ZC 2010- 01-005, same collection data as for previous specimen, fcn 07-157; 1 ovigerous female (CL
<quantity id="4CCE44BBFF9AFFE0E19EF802FA9AC9F1" box="[1274,1372,2010,2036]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="2" pageNumber="49" unit="mm" value="2.5">2.5 mm</quantity>
), UP, Bocas del
<taxonomicName id="4C3692DDFF9BFFE1E475FF40FE8DCEB7" box="[273,331,152,178]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="toro" status="sp. nov.">Toro</taxonomicName>
, east of
<collectingCountry id="F321A9CEFF9BFFE1E4CEFF40FE12CEB7" box="[426,468,152,178]" name="Iceland" pageId="3" pageNumber="50">Isla</collectingCountry>
<collectingCountry id="F321A9CEFF9BFFE1E4BFFF40FDB8CEB7" box="[475,638,152,178]" name="Saint Kitts and Nevis" pageId="3" pageNumber="50">San Cristobal</collectingCountry>
, Laguna Porras, Sta. 9, 9º16’514”N, 82º14’833”W, depth
<quantity id="4CCE44BBFF9BFFE1E047FF40FA97CEB7" box="[1315,1361,152,178]" metricMagnitude="0" metricUnit="m" metricValue="7.0" pageId="3" pageNumber="50" unit="m" value="7.0">7 m</quantity>
, loose coral rubble, coll. S.
<collectingCountry id="F321A9CEFF9BFFE1E4E8FF65FE68CED2" box="[396,430,189,215]" name="South Georgia and the South Sandwich Islands" pageId="3" pageNumber="50">De</collectingCountry>
Grave &amp; M. Salazar,
<date id="FF88CF9EFF9BFFE1E7CBFF65FC98CED2" box="[687,862,189,215]" pageId="3" pageNumber="50" value="2005-08-02">2 August 2005</date>
, fcn 08-254.
</paragraph>
</subSubSection>
<subSubSection id="C32CBAD5FF9BFFE5E5A2FF3AFC12CFD9" lastPageId="7" lastPageNumber="54" pageId="3" pageNumber="50" type="description">
<paragraph id="8B89E95EFF9BFFE1E5A2FF3AFD12CFD9" blockId="3.[151,1437,152,476]" pageId="3" pageNumber="50">
<emphasis id="B942354CFF9BFFE1E5A2FF3AFE99CEF9" bold="true" box="[198,351,226,252]" pageId="3" pageNumber="50">Description.</emphasis>
Small-sized alpheid shrimp. Carapace with one strong sharp mediodorsal tooth situated slightly posterior to carapace mid-length and two strong sharp dorsolateral teeth situated ventral and anterior to mediodorsal tooth, in post-hepatic area (
<figureCitation id="130DF5DBFF9BFFE1E7F3FEF5FD2CCF42" box="[663,746,301,327]" captionStart="FIGURE 1" captionStartId="3.[151,259,1373,1397]" captionTargetBox="[152,1432,538,1343]" captionTargetId="figure@3.[151,1436,530,1352]" captionTargetPageId="3" captionText="FIGURE 1. Triacanthoneus toro n. gen., n. sp., paratype, ovigerous female (CL 3.4 mm) from Bocas del Toro, Caribbean coast of Panama, USNM 1145376: habitus, lateral." httpUri="https://zenodo.org/record/198864/files/figure.png" pageId="3" pageNumber="50">Figs. 1</figureCitation>
,
<figureCitation id="130DF5DBFF9BFFE1E79DFEF5FCCFCF42" box="[761,777,301,327]" captionStart="FIGURE 2" captionStartId="4.[151,259,1821,1845]" captionTargetBox="[195,1375,200,1771]" captionTargetId="figure@4.[193,1379,194,1782]" captionTargetPageId="4" captionText="FIGURE 2. Triacanthoneus toro n. gen., n. sp., paratype, ovigerous female (CL 3.4 mm) from Bocas del Toro, Caribbean coast of Panama, USNM 1145376 [A – M]; paratype, ovigerous female (CL 3.4 mm) from the same locality, OUMNH. ZC 2010 - 01 - 005 [N]: A, frontal region, dorsal; B, carapace and frontal appendages, lateral; C, rostrum, lateral; D, carapace, dorsal; E, tooth on mesioventral carina of first segment of antennular peduncle, lateral; F, antennular flagella, dorsal; G, antenna, ventral; H, second pleopod, mesial; I, same, appendix masculina and appendix interna; J, uropod, dorsal; K, same, posterior region of exopod; L, telson, dorsal; M, same, posterior region; N, rostrum, lateral." httpUri="https://zenodo.org/record/198865/files/figure.png" pageId="3" pageNumber="50">2</figureCitation>
B, D). Rostrum straight, more than 1.5 times as long as broad at base; lateral margins slightly concave; tip overreaching distal margin of first segment of antennular peduncle; ventral margin with distinct, anteriorly directed subapical dent; rostral carina well developed, reaching posteriorly to mediodorsal tooth; orbital teeth absent (
<figureCitation id="130DF5DBFF9BFFE1E6D9FE45FBCCCFB2" box="[957,1034,413,439]" captionStart="FIGURE 2" captionStartId="4.[151,259,1821,1845]" captionTargetBox="[195,1375,200,1771]" captionTargetId="figure@4.[193,1379,194,1782]" captionTargetPageId="4" captionText="FIGURE 2. Triacanthoneus toro n. gen., n. sp., paratype, ovigerous female (CL 3.4 mm) from Bocas del Toro, Caribbean coast of Panama, USNM 1145376 [A – M]; paratype, ovigerous female (CL 3.4 mm) from the same locality, OUMNH. ZC 2010 - 01 - 005 [N]: A, frontal region, dorsal; B, carapace and frontal appendages, lateral; C, rostrum, lateral; D, carapace, dorsal; E, tooth on mesioventral carina of first segment of antennular peduncle, lateral; F, antennular flagella, dorsal; G, antenna, ventral; H, second pleopod, mesial; I, same, appendix masculina and appendix interna; J, uropod, dorsal; K, same, posterior region of exopod; L, telson, dorsal; M, same, posterior region; N, rostrum, lateral." httpUri="https://zenodo.org/record/198865/files/figure.png" pageId="3" pageNumber="50">Fig. 2</figureCitation>
A–C). Pterygostomial angle not protruding anteriorly, broadly rounded (
<figureCitation id="130DF5DBFF9BFFE1E70DFE1AFD77CFD9" box="[617,689,450,476]" captionStart="FIGURE 2" captionStartId="4.[151,259,1821,1845]" captionTargetBox="[195,1375,200,1771]" captionTargetId="figure@4.[193,1379,194,1782]" captionTargetPageId="4" captionText="FIGURE 2. Triacanthoneus toro n. gen., n. sp., paratype, ovigerous female (CL 3.4 mm) from Bocas del Toro, Caribbean coast of Panama, USNM 1145376 [A – M]; paratype, ovigerous female (CL 3.4 mm) from the same locality, OUMNH. ZC 2010 - 01 - 005 [N]: A, frontal region, dorsal; B, carapace and frontal appendages, lateral; C, rostrum, lateral; D, carapace, dorsal; E, tooth on mesioventral carina of first segment of antennular peduncle, lateral; F, antennular flagella, dorsal; G, antenna, ventral; H, second pleopod, mesial; I, same, appendix masculina and appendix interna; J, uropod, dorsal; K, same, posterior region of exopod; L, telson, dorsal; M, same, posterior region; N, rostrum, lateral." httpUri="https://zenodo.org/record/198865/files/figure.png" pageId="3" pageNumber="50">Fig. 2</figureCitation>
B).
</paragraph>
<caption id="DF49B9D6FF9BFFE1E5F3FA85FCEFCB90" httpUri="https://zenodo.org/record/198864/files/figure.png" pageId="3" pageNumber="50" targetBox="[152,1432,538,1343]" targetPageId="3">
<paragraph id="8B89E95EFF9BFFE1E5F3FA85FCEFCB90" blockId="3.[151,1436,1373,1429]" pageId="3" pageNumber="50">
<emphasis id="B942354CFF9BFFE1E5F3FA85FEE4CB70" bold="true" box="[151,290,1373,1397]" pageId="3" pageNumber="50">FIGURE 1.</emphasis>
<taxonomicName id="4C3692DDFF9BFFE1E44DFA86FDD7CB70" box="[297,529,1374,1397]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="50" phylum="Arthropoda" rank="species" species="toro" status="gen. nov.">
<emphasis id="B942354CFF9BFFE1E44DFA86FDD7CB70" box="[297,529,1374,1397]" italics="true" pageId="3" pageNumber="50">Triacanthoneus toro</emphasis>
</taxonomicName>
<emphasis id="B942354CFF9BFFE1E77DFA85FD7DCB70" bold="true" box="[537,699,1373,1397]" pageId="3" pageNumber="50">
<taxonomicNameLabel id="A2718837FF9BFFE1E77DFA85FDADCB70" box="[537,619,1373,1397]" pageId="3" pageNumber="50" rank="genus">n. gen.</taxonomicNameLabel>
, n. sp.
</emphasis>
, paratype, ovigerous female (CL 3.4 mm) from Bocas del
<taxonomicName id="4C3692DDFF9BFFE1E004FA86FA5ECB70" box="[1376,1432,1374,1397]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="toro" status="sp. nov.">Toro</taxonomicName>
, Caribbean coast of Panama, USNM 1145376: habitus, lateral.
</paragraph>
</caption>
<paragraph id="8B89E95EFF9BFFE1E5A1FA1BFD86C89D" blockId="3.[151,1437,1475,2024]" pageId="3" pageNumber="50">
Pleura of first to fourth abdominal somites rounded posteroventrally; fifth pleuron with posteroventral angle produced in small tooth (
<figureCitation id="130DF5DBFF9BFFE1E775FA31FD9EC806" box="[529,600,1513,1539]" captionStart="FIGURE 1" captionStartId="3.[151,259,1373,1397]" captionTargetBox="[152,1432,538,1343]" captionTargetId="figure@3.[151,1436,530,1352]" captionTargetPageId="3" captionText="FIGURE 1. Triacanthoneus toro n. gen., n. sp., paratype, ovigerous female (CL 3.4 mm) from Bocas del Toro, Caribbean coast of Panama, USNM 1145376: habitus, lateral." httpUri="https://zenodo.org/record/198864/files/figure.png" pageId="3" pageNumber="50">Fig. 1</figureCitation>
). Telson narrow, at least 3.5 times as long as wide in proximal third, tapering distally; dorsal surface with two pairs of strong spiniform setae situated at some distance from lateral margins, at about 0.5 and 0.8 telson length, respectively; posterior margin straight, each posterolateral angle with two pairs of spiniform setae, mesial being 0.6 times as long as lateral, and two plumose setae between mesial spiniform setae (
<figureCitation id="130DF5DBFF9BFFE1E4CAF9A6FE32C89D" box="[430,500,1662,1688]" captionStart="FIGURE 2" captionStartId="4.[151,259,1821,1845]" captionTargetBox="[195,1375,200,1771]" captionTargetId="figure@4.[193,1379,194,1782]" captionTargetPageId="4" captionText="FIGURE 2. Triacanthoneus toro n. gen., n. sp., paratype, ovigerous female (CL 3.4 mm) from Bocas del Toro, Caribbean coast of Panama, USNM 1145376 [A – M]; paratype, ovigerous female (CL 3.4 mm) from the same locality, OUMNH. ZC 2010 - 01 - 005 [N]: A, frontal region, dorsal; B, carapace and frontal appendages, lateral; C, rostrum, lateral; D, carapace, dorsal; E, tooth on mesioventral carina of first segment of antennular peduncle, lateral; F, antennular flagella, dorsal; G, antenna, ventral; H, second pleopod, mesial; I, same, appendix masculina and appendix interna; J, uropod, dorsal; K, same, posterior region of exopod; L, telson, dorsal; M, same, posterior region; N, rostrum, lateral." httpUri="https://zenodo.org/record/198865/files/figure.png" pageId="3" pageNumber="50">Fig. 2</figureCitation>
L, M).
</paragraph>
<paragraph id="8B89E95EFF9BFFE1E5A1F97BFB0BC8E6" blockId="3.[151,1437,1475,2024]" pageId="3" pageNumber="50">
Eyes partly exposed in dorsal and lateral views; cornea somewhat reduced, well pigmented, occupying anterolateral portion of eyestalk, anteromesial margin with small blunt process (
<figureCitation id="130DF5DBFF9BFFE1E15BF911FB43C8E6" box="[1087,1157,1737,1763]" captionStart="FIGURE 2" captionStartId="4.[151,259,1821,1845]" captionTargetBox="[195,1375,200,1771]" captionTargetId="figure@4.[193,1379,194,1782]" captionTargetPageId="4" captionText="FIGURE 2. Triacanthoneus toro n. gen., n. sp., paratype, ovigerous female (CL 3.4 mm) from Bocas del Toro, Caribbean coast of Panama, USNM 1145376 [A – M]; paratype, ovigerous female (CL 3.4 mm) from the same locality, OUMNH. ZC 2010 - 01 - 005 [N]: A, frontal region, dorsal; B, carapace and frontal appendages, lateral; C, rostrum, lateral; D, carapace, dorsal; E, tooth on mesioventral carina of first segment of antennular peduncle, lateral; F, antennular flagella, dorsal; G, antenna, ventral; H, second pleopod, mesial; I, same, appendix masculina and appendix interna; J, uropod, dorsal; K, same, posterior region of exopod; L, telson, dorsal; M, same, posterior region; N, rostrum, lateral." httpUri="https://zenodo.org/record/198865/files/figure.png" pageId="3" pageNumber="50">Fig. 2</figureCitation>
A, B).
</paragraph>
<paragraph id="8B89E95EFF9BFFE1E5A1F936FC08C9ED" blockId="3.[151,1437,1475,2024]" pageId="3" pageNumber="50">
Antennular peduncle stout, with second segment distinctly longer than wide; first segment with distally acute stylocerite reaching to about 0.4 length of second segment; ventromesial carina of first segment with anteriorly directed tooth; lateral flagellum biramous, fused portion composed of at least three segments; accessory ramus composed of four or so partly fused segments with groups of aesthetascs (
<figureCitation id="130DF5DBFF9BFFE1E18CF886FAF6C97D" box="[1256,1328,1886,1912]" captionStart="FIGURE 2" captionStartId="4.[151,259,1821,1845]" captionTargetBox="[195,1375,200,1771]" captionTargetId="figure@4.[193,1379,194,1782]" captionTargetPageId="4" captionText="FIGURE 2. Triacanthoneus toro n. gen., n. sp., paratype, ovigerous female (CL 3.4 mm) from Bocas del Toro, Caribbean coast of Panama, USNM 1145376 [A – M]; paratype, ovigerous female (CL 3.4 mm) from the same locality, OUMNH. ZC 2010 - 01 - 005 [N]: A, frontal region, dorsal; B, carapace and frontal appendages, lateral; C, rostrum, lateral; D, carapace, dorsal; E, tooth on mesioventral carina of first segment of antennular peduncle, lateral; F, antennular flagella, dorsal; G, antenna, ventral; H, second pleopod, mesial; I, same, appendix masculina and appendix interna; J, uropod, dorsal; K, same, posterior region of exopod; L, telson, dorsal; M, same, posterior region; N, rostrum, lateral." httpUri="https://zenodo.org/record/198865/files/figure.png" pageId="3" pageNumber="50">Fig. 2</figureCitation>
A, B, F). Antenna with stout basicerite bearing sharp distoventral tooth; scaphocerite reaching to end of antennular peduncle, ovate; blade broad, with anterior margin convex, reaching slightly beyond strong distolateral tooth; carpocerite barely reaching 0.5 length of scaphocerite (
<figureCitation id="130DF5DBFF9BFFE1E67FF816FCA7C9ED" box="[795,865,1998,2024]" captionStart="FIGURE 2" captionStartId="4.[151,259,1821,1845]" captionTargetBox="[195,1375,200,1771]" captionTargetId="figure@4.[193,1379,194,1782]" captionTargetPageId="4" captionText="FIGURE 2. Triacanthoneus toro n. gen., n. sp., paratype, ovigerous female (CL 3.4 mm) from Bocas del Toro, Caribbean coast of Panama, USNM 1145376 [A – M]; paratype, ovigerous female (CL 3.4 mm) from the same locality, OUMNH. ZC 2010 - 01 - 005 [N]: A, frontal region, dorsal; B, carapace and frontal appendages, lateral; C, rostrum, lateral; D, carapace, dorsal; E, tooth on mesioventral carina of first segment of antennular peduncle, lateral; F, antennular flagella, dorsal; G, antenna, ventral; H, second pleopod, mesial; I, same, appendix masculina and appendix interna; J, uropod, dorsal; K, same, posterior region of exopod; L, telson, dorsal; M, same, posterior region; N, rostrum, lateral." httpUri="https://zenodo.org/record/198865/files/figure.png" pageId="3" pageNumber="50">Fig. 2</figureCitation>
A, B, G).
</paragraph>
<caption id="DF49B9D6FF9CFFE6E5F3F8C5FAB4C9D0" httpUri="https://zenodo.org/record/198865/files/figure.png" pageId="4" pageNumber="51" targetBox="[195,1375,200,1771]" targetPageId="4">
<paragraph id="8B89E95EFF9CFFE6E5F3F8C5FAB4C9D0" blockId="4.[151,1437,1821,2005]" pageId="4" pageNumber="51">
<emphasis id="B942354CFF9CFFE6E5F3F8C5FEE7C930" bold="true" box="[151,289,1821,1845]" pageId="4" pageNumber="51">FIGURE 2.</emphasis>
<taxonomicName id="4C3692DDFF9CFFE6E44DF8C6FDD7C930" box="[297,529,1822,1845]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="51" phylum="Arthropoda" rank="species" species="toro" status="gen. nov.">
<emphasis id="B942354CFF9CFFE6E44DF8C6FDD7C930" box="[297,529,1822,1845]" italics="true" pageId="4" pageNumber="51">Triacanthoneus toro</emphasis>
</taxonomicName>
<emphasis id="B942354CFF9CFFE6E77DF8C5FD7DC930" bold="true" box="[537,699,1821,1845]" pageId="4" pageNumber="51">
<taxonomicNameLabel id="A2718837FF9CFFE6E77DF8C5FDADC930" box="[537,619,1821,1845]" pageId="4" pageNumber="51" rank="genus">n. gen.</taxonomicNameLabel>
, n. sp.
</emphasis>
, paratype, ovigerous female (CL 3.4 mm) from Bocas del
<taxonomicName id="4C3692DDFF9CFFE6E004F8C6FA5EC930" box="[1376,1432,1822,1845]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="62" phylum="Arthropoda" rank="species" species="toro" status="sp. nov.">Toro</taxonomicName>
, Caribbean coast of Panama, USNM 1145376 [A–M]; paratype, ovigerous female (CL 3.4 mm) from the same locality, OUMNH.ZC 2010-01-005 [N]: A, frontal region, dorsal; B, carapace and frontal appendages, lateral; C, rostrum, lateral; D, carapace, dorsal; E, tooth on mesioventral carina of first segment of antennular peduncle, lateral; F, antennular flagella, dorsal; G, antenna, ventral; H, second pleopod, mesial; I, same, appendix masculina and appendix interna; J, uropod, dorsal; K, same, posterior region of exopod; L, telson, dorsal; M, same, posterior region; N, rostrum, lateral.
</paragraph>
</caption>
<caption id="DF49B9D6FF9DFFE7E5F3FA48FB26CBE2" httpUri="https://zenodo.org/record/198866/files/figure.png" pageId="5" pageNumber="52" targetBox="[157,1425,188,1393]" targetPageId="5">
<paragraph id="8B89E95EFF9DFFE7E5F3FA48FB26CBE2" blockId="5.[151,1436,1424,1511]" pageId="5" pageNumber="52">
<emphasis id="B942354CFF9DFFE7E5F3FA48FEE7CBAD" bold="true" box="[151,289,1424,1448]" pageId="5" pageNumber="52">FIGURE 3.</emphasis>
<taxonomicName id="4C3692DDFF9DFFE7E44DFA49FDD7CBAD" box="[297,529,1425,1448]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="52" phylum="Arthropoda" rank="species" species="toro" status="gen. nov.">
<emphasis id="B942354CFF9DFFE7E44DFA49FDD7CBAD" box="[297,529,1425,1448]" italics="true" pageId="5" pageNumber="52">Triacanthoneus toro</emphasis>
</taxonomicName>
<emphasis id="B942354CFF9DFFE7E77DFA48FD7DCBAD" bold="true" box="[537,699,1424,1448]" pageId="5" pageNumber="52">
<taxonomicNameLabel id="A2718837FF9DFFE7E77DFA48FDADCBAD" box="[537,619,1424,1448]" pageId="5" pageNumber="52" rank="genus">n. gen.</taxonomicNameLabel>
, n. sp.
</emphasis>
, paratype, ovigerous female (CL 3.4 mm) from Bocas del
<taxonomicName id="4C3692DDFF9DFFE7E004FA48FA5ECBA2" box="[1376,1432,1424,1447]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="62" phylum="Arthropoda" rank="species" species="toro" status="sp. nov.">Toro</taxonomicName>
, Caribbean coast of Panama, USNM 1145376: A, mandible, mesial; B, same, incisor process, lateral; C, maxillule, lateral; D, maxilla, lateral; E, first maxilliped, lateral; F, second maxilliped, lateral; G, third maxilliped, lateral.
</paragraph>
</caption>
<paragraph id="8B89E95EFF9DFFE7E5A1F9CEFD67C8A5" blockId="5.[151,1437,1558,2032]" pageId="5" pageNumber="52">
Mouthparts typical for genus, as illustrated (
<figureCitation id="130DF5DBFF9DFFE7E781F9CEFCEBC835" box="[741,813,1558,1584]" captionStart="FIGURE 3" captionStartId="5.[151,259,1424,1448]" captionTargetBox="[157,1425,188,1393]" captionTargetId="figure@5.[151,1436,174,1418]" captionTargetPageId="5" captionText="FIGURE 3. Triacanthoneus toro n. gen., n. sp., paratype, ovigerous female (CL 3.4 mm) from Bocas del Toro, Caribbean coast of Panama, USNM 1145376: A, mandible, mesial; B, same, incisor process, lateral; C, maxillule, lateral; D, maxilla, lateral; E, first maxilliped, lateral; F, second maxilliped, lateral; G, third maxilliped, lateral." httpUri="https://zenodo.org/record/198866/files/figure.png" pageId="5" pageNumber="52">Fig. 3</figureCitation>
A–F). Third maxilliped with coxa bearing rounded lateral plate above mastigobranch; antepenultimate segment elongate, flattened; penultimate segment elongate, slender; ultimate segment tapering distally, ending in corneous tip, without robust setiform setae on apex; arthrobranch well developed (
<figureCitation id="130DF5DBFF9DFFE7E758F95EFDBCC8A5" box="[572,634,1670,1696]" captionStart="FIGURE 3" captionStartId="5.[151,259,1424,1448]" captionTargetBox="[157,1425,188,1393]" captionTargetId="figure@5.[151,1436,174,1418]" captionTargetPageId="5" captionText="FIGURE 3. Triacanthoneus toro n. gen., n. sp., paratype, ovigerous female (CL 3.4 mm) from Bocas del Toro, Caribbean coast of Panama, USNM 1145376: A, mandible, mesial; B, same, incisor process, lateral; C, maxillule, lateral; D, maxilla, lateral; E, first maxilliped, lateral; F, second maxilliped, lateral; G, third maxilliped, lateral." httpUri="https://zenodo.org/record/198866/files/figure.png" pageId="5" pageNumber="52">Fig 3</figureCitation>
G).
</paragraph>
<paragraph id="8B89E95EFF9DFFE7E5A1F973FB2CC9F5" blockId="5.[151,1437,1558,2032]" pageId="5" pageNumber="52">
Chelipeds very unequal in size and asymmetrical in shape, carried folded (
<figureCitation id="130DF5DBFF9DFFE7E12DF973FB54C8C0" box="[1097,1170,1707,1733]" captionStart="FIGURE 4" captionStartId="6.[151,259,1544,1568]" captionTargetBox="[154,1433,226,1501]" captionTargetId="figure@6.[151,1436,194,1523]" captionTargetPageId="6" captionText="FIGURE 4. Triacanthoneus toro n. gen., n. sp., paratype, ovigerous female (CL 3.4 mm) from Bocas del Toro, Caribbean coast of Panama, USNM 1145376: A, right (major) cheliped, lateral; B, same, carpus and chela, mesial; C, same, chela fingers; D, left (minor) cheliped, lateral; E, same, chela, mesial; F, second pereiopod, lateral; G, third pereiopod, lateral; H, same, dactylus; I, fifth pereiopod, lateral." httpUri="https://zenodo.org/record/198867/files/figure.png" pageId="5" pageNumber="52">Fig. 4</figureCitation>
A, D). Major cheliped enlarged, elongate, slender; ischium rather long, slender, ventrolateral surface with three stout spiniform setae; merus much longer than ischium, slender, smooth, ventrally flattened; carpus distinctly shorter than merus, subcylindrical, distally somewhat broadening, smooth; chela subcylindrical, smooth; fingers longer than palm, compressed, slightly twisted, cutting edges evenly serrated, with at least 18 subtriangular teeth (
<figureCitation id="130DF5DBFF9DFFE7E00FF898FF60C985" captionStart="FIGURE 4" captionStartId="6.[151,259,1544,1568]" captionTargetBox="[154,1433,226,1501]" captionTargetId="figure@6.[151,1436,194,1523]" captionTargetPageId="6" captionText="FIGURE 4. Triacanthoneus toro n. gen., n. sp., paratype, ovigerous female (CL 3.4 mm) from Bocas del Toro, Caribbean coast of Panama, USNM 1145376: A, right (major) cheliped, lateral; B, same, carpus and chela, mesial; C, same, chela fingers; D, left (minor) cheliped, lateral; E, same, chela, mesial; F, second pereiopod, lateral; G, third pereiopod, lateral; H, same, dactylus; I, fifth pereiopod, lateral." httpUri="https://zenodo.org/record/198867/files/figure.png" pageId="5" pageNumber="52">Fig. 4</figureCitation>
A–C). Minor cheliped not particularly enlarged, slender; ischium long, ventrolateral surface with three stout spiniform setae; merus slightly longer than ischium, somewhat compressed, smooth; carpus slightly shorter than merus, subcylindrical, slightly widening distally, smooth; chela simple, smooth, with fingers slightly longer than palm, finger cutting edges straight, except for a few small distal teeth (
<figureCitation id="130DF5DBFF9DFFE7E13AF80EFB63C9F5" box="[1118,1189,2006,2032]" captionStart="FIGURE 4" captionStartId="6.[151,259,1544,1568]" captionTargetBox="[154,1433,226,1501]" captionTargetId="figure@6.[151,1436,194,1523]" captionTargetPageId="6" captionText="FIGURE 4. Triacanthoneus toro n. gen., n. sp., paratype, ovigerous female (CL 3.4 mm) from Bocas del Toro, Caribbean coast of Panama, USNM 1145376: A, right (major) cheliped, lateral; B, same, carpus and chela, mesial; C, same, chela fingers; D, left (minor) cheliped, lateral; E, same, chela, mesial; F, second pereiopod, lateral; G, third pereiopod, lateral; H, same, dactylus; I, fifth pereiopod, lateral." httpUri="https://zenodo.org/record/198867/files/figure.png" pageId="5" pageNumber="52">Fig. 4</figureCitation>
D, E).
</paragraph>
<caption id="DF49B9D6FF9EFFE4E5F3F9D0FCFEC885" httpUri="https://zenodo.org/record/198867/files/figure.png" pageId="6" pageNumber="53" targetBox="[154,1433,226,1501]" targetPageId="6">
<paragraph id="8B89E95EFF9EFFE4E5F3F9D0FCFEC885" blockId="6.[151,1437,1544,1664]" pageId="6" pageNumber="53">
<emphasis id="B942354CFF9EFFE4E5F3F9D0FEE7C825" bold="true" box="[151,289,1544,1568]" pageId="6" pageNumber="53">FIGURE 4.</emphasis>
<taxonomicName id="4C3692DDFF9EFFE4E44DF9D1FDD7C825" box="[297,529,1545,1568]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="53" phylum="Arthropoda" rank="species" species="toro" status="gen. nov.">
<emphasis id="B942354CFF9EFFE4E44DF9D1FDD7C825" box="[297,529,1545,1568]" italics="true" pageId="6" pageNumber="53">Triacanthoneus toro</emphasis>
</taxonomicName>
<emphasis id="B942354CFF9EFFE4E77DF9D0FD7DC825" bold="true" box="[537,699,1544,1568]" pageId="6" pageNumber="53">
<taxonomicNameLabel id="A2718837FF9EFFE4E77DF9D0FDADC825" box="[537,619,1544,1568]" pageId="6" pageNumber="53" rank="genus">n. gen.</taxonomicNameLabel>
, n. sp.
</emphasis>
, paratype, ovigerous female (CL 3.4 mm) from Bocas del
<taxonomicName id="4C3692DDFF9EFFE4E004F9D1FA5EC825" box="[1376,1432,1545,1568]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="62" phylum="Arthropoda" rank="species" species="toro" status="sp. nov.">Toro</taxonomicName>
, Caribbean coast of Panama, USNM 1145376: A, right (major) cheliped, lateral; B, same, carpus and chela, mesial; C, same, chela fingers; D, left (minor) cheliped, lateral; E, same, chela, mesial; F, second pereiopod, lateral; G, third pereiopod, lateral; H, same, dactylus; I, fifth pereiopod, lateral.
</paragraph>
</caption>
<paragraph id="8B89E95EFF9EFFE5E5A1F976FCBBCEF9" blockId="6.[151,1437,1710,2035]" lastBlockId="7.[151,1437,152,812]" lastPageId="7" lastPageNumber="54" pageId="6" pageNumber="53">
Second pereiopod slender; ischium about as long as merus, with three spiniform setae on ventrolateral surface; carpus five-segmented, first segment longest, slightly longer than sum of four other segments, ratio of carpal segments (from proximal to distal) more or less equal to: 5.5: 1: 1: 1.5: 1.5; chela simple, longer than most-distal carpal article (
<figureCitation id="130DF5DBFF9EFFE4E4B0F8C6FDD8C93D" box="[468,542,1822,1848]" captionStart="FIGURE 4" captionStartId="6.[151,259,1544,1568]" captionTargetBox="[154,1433,226,1501]" captionTargetId="figure@6.[151,1436,194,1523]" captionTargetPageId="6" captionText="FIGURE 4. Triacanthoneus toro n. gen., n. sp., paratype, ovigerous female (CL 3.4 mm) from Bocas del Toro, Caribbean coast of Panama, USNM 1145376: A, right (major) cheliped, lateral; B, same, carpus and chela, mesial; C, same, chela fingers; D, left (minor) cheliped, lateral; E, same, chela, mesial; F, second pereiopod, lateral; G, third pereiopod, lateral; H, same, dactylus; I, fifth pereiopod, lateral." httpUri="https://zenodo.org/record/198867/files/figure.png" pageId="6" pageNumber="53">Fig. 4</figureCitation>
F). Third pereiopod very slender, compressed; ischium elongate, with two spaced spiniform setae on ventrolateral surface; merus elongate, almost 10 times as long as wide, smooth; carpus longer than merus, with slender stiff distoventral seta; propodus with one slender spiniform seta on ventral margin and one stouter distal spiniform seta adjacent to dactylus; dactylus almost 0.5 propodus length, simple, conical, very slender, gradually curving distally, furnished with setae on dorsal margin (
<figureCitation id="130DF5DBFF9EFFE4E068F86BFA94C9C8" box="[1292,1362,1971,1997]" captionStart="FIGURE 4" captionStartId="6.[151,259,1544,1568]" captionTargetBox="[154,1433,226,1501]" captionTargetId="figure@6.[151,1436,194,1523]" captionTargetPageId="6" captionText="FIGURE 4. Triacanthoneus toro n. gen., n. sp., paratype, ovigerous female (CL 3.4 mm) from Bocas del Toro, Caribbean coast of Panama, USNM 1145376: A, right (major) cheliped, lateral; B, same, carpus and chela, mesial; C, same, chela fingers; D, left (minor) cheliped, lateral; E, same, chela, mesial; F, second pereiopod, lateral; G, third pereiopod, lateral; H, same, dactylus; I, fifth pereiopod, lateral." httpUri="https://zenodo.org/record/198867/files/figure.png" pageId="6" pageNumber="53">Fig. 4</figureCitation>
G, H). Fourth pereiopod almost identical to third pereiopod. Fifth pereiopod longer and more slender than third or fourth pereiopods; ischium unarmed; carpus distinctly longer than merus, propodus with two small spiniform setae on ventral margin and at least six rows of slender serrulate setae on distolateral surface, most-distal row with longest setae reaching well beyond dactylar base (
<figureCitation id="130DF5DBFF9FFFE5E679FF3AFCA3CEF9" box="[797,869,226,252]" captionStart="FIGURE 4" captionStartId="6.[151,259,1544,1568]" captionTargetBox="[154,1433,226,1501]" captionTargetId="figure@6.[151,1436,194,1523]" captionTargetPageId="6" captionText="FIGURE 4. Triacanthoneus toro n. gen., n. sp., paratype, ovigerous female (CL 3.4 mm) from Bocas del Toro, Caribbean coast of Panama, USNM 1145376: A, right (major) cheliped, lateral; B, same, carpus and chela, mesial; C, same, chela fingers; D, left (minor) cheliped, lateral; E, same, chela, mesial; F, second pereiopod, lateral; G, third pereiopod, lateral; H, same, dactylus; I, fifth pereiopod, lateral." httpUri="https://zenodo.org/record/198867/files/figure.png" pageId="7" pageNumber="54">
Fig.
<date id="FF88CF9EFF9FFFE5E630FF3AFCA3CEF9" box="[852,869,226,252]" pageId="7" pageNumber="54">4</date>
</figureCitation>
I).
</paragraph>
<paragraph id="8B89E95EFF9FFFE5E5A1FED0FC7DCF97" blockId="7.[151,1437,152,812]" pageId="7" pageNumber="54">
Second pleopod of all four (ovigerous) specimens with appendix interna and appendix masculina, latter shorter than appendix interna, with stiff apical and subapical setae (
<figureCitation id="130DF5DBFF9FFFE5E6B8FEF5FBEFCF42" box="[988,1065,301,327]" captionStart="FIGURE 2" captionStartId="4.[151,259,1821,1845]" captionTargetBox="[195,1375,200,1771]" captionTargetId="figure@4.[193,1379,194,1782]" captionTargetPageId="4" captionText="FIGURE 2. Triacanthoneus toro n. gen., n. sp., paratype, ovigerous female (CL 3.4 mm) from Bocas del Toro, Caribbean coast of Panama, USNM 1145376 [A – M]; paratype, ovigerous female (CL 3.4 mm) from the same locality, OUMNH. ZC 2010 - 01 - 005 [N]: A, frontal region, dorsal; B, carapace and frontal appendages, lateral; C, rostrum, lateral; D, carapace, dorsal; E, tooth on mesioventral carina of first segment of antennular peduncle, lateral; F, antennular flagella, dorsal; G, antenna, ventral; H, second pleopod, mesial; I, same, appendix masculina and appendix interna; J, uropod, dorsal; K, same, posterior region of exopod; L, telson, dorsal; M, same, posterior region; N, rostrum, lateral." httpUri="https://zenodo.org/record/198865/files/figure.png" pageId="7" pageNumber="54">
Fig.
<date id="FF88CF9EFF9FFFE5E173FEF5FBEFCF42" box="[1047,1065,301,327]" pageId="7" pageNumber="54">2</date>
</figureCitation>
I). Uropod with lateral lobe of protopod ending in small subacute tooth; exopod and endopod slender; exopod with sinuous diaeresis and blunt distolateral tooth adjacent to robust spiniform seta (
<figureCitation id="130DF5DBFF9FFFE5E651FEA0FCBACF97" box="[821,892,376,402]" captionStart="FIGURE 2" captionStartId="4.[151,259,1821,1845]" captionTargetBox="[195,1375,200,1771]" captionTargetId="figure@4.[193,1379,194,1782]" captionTargetPageId="4" captionText="FIGURE 2. Triacanthoneus toro n. gen., n. sp., paratype, ovigerous female (CL 3.4 mm) from Bocas del Toro, Caribbean coast of Panama, USNM 1145376 [A – M]; paratype, ovigerous female (CL 3.4 mm) from the same locality, OUMNH. ZC 2010 - 01 - 005 [N]: A, frontal region, dorsal; B, carapace and frontal appendages, lateral; C, rostrum, lateral; D, carapace, dorsal; E, tooth on mesioventral carina of first segment of antennular peduncle, lateral; F, antennular flagella, dorsal; G, antenna, ventral; H, second pleopod, mesial; I, same, appendix masculina and appendix interna; J, uropod, dorsal; K, same, posterior region of exopod; L, telson, dorsal; M, same, posterior region; N, rostrum, lateral." httpUri="https://zenodo.org/record/198865/files/figure.png" pageId="7" pageNumber="54">Fig. 2</figureCitation>
J, K).
</paragraph>
<paragraph id="8B89E95EFF9FFFE5E5A1FE45FC9ECFB2" blockId="7.[151,1437,152,812]" box="[197,856,413,439]" pageId="7" pageNumber="54">Gill-exopod formula as described for genus (see above).</paragraph>
<paragraph id="8B89E95EFF9FFFE5E5A1FE1AFC12CFD9" blockId="7.[151,1437,152,812]" box="[197,980,450,476]" pageId="7" pageNumber="54">
Colour in life semitransparent-white, gonads yellow (
<figureCitation id="130DF5DBFF9FFFE5E653FE1AFC4DCFD9" box="[823,907,450,476]" captionStart="FIGURE 10" captionStartId="14.[151,256,1843,1867]" captionTargetBox="[151,1436,493,1821]" captionTargetId="figure@14.[151,1436,493,1822]" captionTargetPageId="14" captionText="FIGURE 10. Life colouration of three species of Triacanthoneus n. gen.: A – C, Triacanthoneus toro n. sp., paratype, ovigerous female from Bocas del Toro, Caribbean coast of Panama, USNM 1145376; D – F, Triacanthoneus pacificus n. sp., holotype, ovigerous female from Playa Venao, Pacific coast of Panama, USNM 1145374 [D, E], non-type ovigerous female from Bahía Malaga, Pacific coast of Colombia, UVC [F]; G, Triacanthoneus alacranes n. sp., holotype, ovigerous female from Alacranes Reef, Yucatán, Mexico, UNAM-CNCR 26177 [G by J. Duarte-Gutiérrez, all other photographs by author]." httpUri="https://zenodo.org/record/198873/files/figure.png" pageId="7" pageNumber="54">Fig. 10</figureCitation>
A–C).
</paragraph>
</subSubSection>
<subSubSection id="C32CBAD5FF9FFFE5E5A1FE3FFE40CC92" pageId="7" pageNumber="54" type="etymology">
<paragraph id="8B89E95EFF9FFFE5E5A1FE3FFA5BCC22" blockId="7.[151,1437,152,812]" pageId="7" pageNumber="54">
<emphasis id="B942354CFF9FFFE5E5A1FE3FFE90CC04" bold="true" box="[197,342,487,513]" pageId="7" pageNumber="54">Etymology.</emphasis>
Specific name derived from the third word of the
<typeStatus id="548D57FCFF9FFFE5E6A5FE30FC33CC07" box="[961,1013,488,514]" pageId="7" pageNumber="54">type</typeStatus>
locality, Bocas del
<taxonomicName id="4C3692DDFF9FFFE5E183FE30FAE2CC07" box="[1255,1316,488,514]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="toro" status="sp. nov.">Toro</taxonomicName>
;
<taxonomicName id="4C3692DDFF9FFFE5E055FE30FAA5CC04" box="[1329,1379,488,513]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="54" phylum="Arthropoda" rank="species" species="toro">
<emphasis id="B942354CFF9FFFE5E055FE30FAA5CC04" box="[1329,1379,488,513]" italics="true" pageId="7" pageNumber="54">toro</emphasis>
</taxonomicName>
also means bull in Spanish, alluding to the two horn-like projections on the carapace; used as a noun in opposition.
</paragraph>
<paragraph id="8B89E95EFF9FFFE5E5A1FDEAFE40CC92" blockId="7.[151,1437,152,812]" pageId="7" pageNumber="54">
<emphasis id="B942354CFF9FFFE5E5A1FDEAFEE9CC49" bold="true" box="[197,303,562,588]" pageId="7" pageNumber="54">Habitat.</emphasis>
Sand flats with abundant rubble and seagrass, at some distance from fringing mangroves, at depth range of
<quantity id="4CCE44BBFF9FFFE5E422FD80FE6FCC77" box="[326,425,600,626]" metricMagnitude="0" metricUnit="m" metricValue="4.25" metricValueMax="7.0" metricValueMin="1.5" pageId="7" pageNumber="54" unit="m" value="4.25" valueMax="7.0" valueMin="1.5">1.5–7 m</quantity>
; most specimens were found on sand deep under large pieces of rubble, one was found among loose rubble.
</paragraph>
</subSubSection>
<subSubSection id="C32CBAD5FF9FFFE5E5A1FD7AFCBFCCB9" box="[197,889,674,700]" pageId="7" pageNumber="54" type="materials_examined">
<paragraph id="8B89E95EFF9FFFE5E5A1FD7AFCBFCCB9" blockId="7.[151,1437,152,812]" box="[197,889,674,700]" pageId="7" pageNumber="54">
<emphasis id="B942354CFF9FFFE5E5A1FD7AFEACCCB9" bold="true" box="[197,362,674,700]" pageId="7" pageNumber="54">
<typeStatus id="548D57FCFF9FFFE5E5A1FD7AFEC4CCB9" box="[197,258,674,700]" pageId="7" pageNumber="54">Type</typeStatus>
locality.
</emphasis>
Bocas del
<taxonomicName id="4C3692DDFF9FFFE5E488FD7AFDE0CCB9" box="[492,550,674,700]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="toro" status="sp. nov.">Toro</taxonomicName>
, Caribbean coast of
<collectingCountry id="F321A9CEFF9FFFE5E670FD7AFCB3CCB9" box="[788,885,674,700]" name="Panama" pageId="7" pageNumber="54">Panama</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C32CBAD5FF9FFFE5E5A2FD1FFAD6CCE7" box="[198,1296,711,738]" pageId="7" pageNumber="54" type="distribution">
<paragraph id="8B89E95EFF9FFFE5E5A2FD1FFAD6CCE7" blockId="7.[151,1437,152,812]" box="[198,1296,711,738]" pageId="7" pageNumber="54">
<emphasis id="B942354CFF9FFFE5E5A2FD1FFEA1CCE4" bold="true" box="[198,359,711,737]" pageId="7" pageNumber="54">Distribution.</emphasis>
Presently known only from the
<typeStatus id="548D57FCFF9FFFE5E784FD10FCD4CCE7" box="[736,786,712,738]" pageId="7" pageNumber="54">type</typeStatus>
locality on the Caribbean coast of
<collectingCountry id="F321A9CEFF9FFFE5E1CFFD10FACACCE7" box="[1195,1292,712,738]" name="Panama" pageId="7" pageNumber="54">Panama</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C32CBAD5FF9FFFE5E5A2FD35FECDCD29" pageId="7" pageNumber="54" type="discussion">
<paragraph id="8B89E95EFF9FFFE5E5A2FD35FECDCD29" blockId="7.[151,1437,152,812]" pageId="7" pageNumber="54">
<emphasis id="B942354CFF9FFFE5E5A2FD35FEFECD02" bold="true" box="[198,312,749,775]" pageId="7" pageNumber="54">Remarks</emphasis>
. One of the
<typeStatus id="548D57FCFF9FFFE5E4AFFD35FDFDCD02" box="[459,571,749,775]" pageId="7" pageNumber="54" type="paratype">paratypes</typeStatus>
has a somewhat aberrant rostrum, with two instead of one subapical dent (
<figureCitation id="130DF5DBFF9FFFE5E5FBFCCAFF20CD29" box="[159,230,786,812]" captionStart="FIGURE 2" captionStartId="4.[151,259,1821,1845]" captionTargetBox="[195,1375,200,1771]" captionTargetId="figure@4.[193,1379,194,1782]" captionTargetPageId="4" captionText="FIGURE 2. Triacanthoneus toro n. gen., n. sp., paratype, ovigerous female (CL 3.4 mm) from Bocas del Toro, Caribbean coast of Panama, USNM 1145376 [A – M]; paratype, ovigerous female (CL 3.4 mm) from the same locality, OUMNH. ZC 2010 - 01 - 005 [N]: A, frontal region, dorsal; B, carapace and frontal appendages, lateral; C, rostrum, lateral; D, carapace, dorsal; E, tooth on mesioventral carina of first segment of antennular peduncle, lateral; F, antennular flagella, dorsal; G, antenna, ventral; H, second pleopod, mesial; I, same, appendix masculina and appendix interna; J, uropod, dorsal; K, same, posterior region of exopod; L, telson, dorsal; M, same, posterior region; N, rostrum, lateral." httpUri="https://zenodo.org/record/198865/files/figure.png" pageId="7" pageNumber="54">Fig. 2</figureCitation>
N).
</paragraph>
</subSubSection>
</treatment>
</document>
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