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<document id="3CD5D4324292C9F52052F02383F440CB" ID-CLB-Dataset="8744" ID-DOI="10.11646/zootaxa.4838.2.1" ID-GBIF-Dataset="761fcec7-c79d-4e80-bb2b-23cc9e78e0b6" ID-ISSN="1175-5326" ID-Zenodo-Dep="4403783" ID-ZooBank="0AD43D95-7508-466C-980F-CBEF38CA1198" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609352386978" checkinUser="plazi" docAuthor="Gompel, Nicolas" docDate="2020" docId="039187D8FFC0BB17FF1884DDFAEFFBA5" docLanguage="en" docName="zootaxa.4838.2.1.pdf" docOrigin="Zootaxa 4838 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Zarcosia deharvengi Gompel 2020, n. sp." docType="treatment" docUuid="7AC6F8AC-1F44-439A-A651-E48C80B38F53" docUuidSource="ZooBank" docVersion="8" lastPageNumber="157" masterDocId="FFA8FFA0FFC6BB11FF8F844AFF9DFFFF" masterDocTitle="Synopsis of the Asian species in the genus Zarcosia Collado &amp; Alonso-Zarazaga 1996 with new combinations and descriptions of fourteen new species" masterLastPageNumber="178" masterPageNumber="151" pageNumber="157" updateTime="1698850145988" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="14311CB3B589282F01D36203AAAA86CE">Synopsis of the Asian species in the genus Zarcosia Collado &amp; Alonso-Zarazaga 1996 with new combinations and descriptions of fourteen new species</mods:title>
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<mods:name id="1ECBEEFC48D765A3F302C40D94A3F3B7" type="personal">
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<mods:namePart id="89174B4310783030A81A7D295F72414E">Gompel, Nicolas</mods:namePart>
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<mods:title id="DECD13F047B147AB507356B0D734FA87">Zootaxa</mods:title>
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<mods:date id="EDC1E20F76B94A107269A5876AD32C37">2020</mods:date>
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<mods:number id="F0ACA6CFCB4826747790717629406432">2020-08-26</mods:number>
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<mods:number id="AD6EE03F6414CE526947EEA6EFD0FB95">4838</mods:number>
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<mods:classification id="B168B8F5F628D7CC3A68282A3BA45570">journal article</mods:classification>
<mods:identifier id="1761C45C6D5D03D8100C7E04C7B0EC0F" type="CLB-Dataset">8744</mods:identifier>
<mods:identifier id="CF29DD800B6D752B71C397EAFBC479E3" type="DOI">10.11646/zootaxa.4838.2.1</mods:identifier>
<mods:identifier id="363D3ADACD7BF4424BA591D7571A8903" type="GBIF-Dataset">761fcec7-c79d-4e80-bb2b-23cc9e78e0b6</mods:identifier>
<mods:identifier id="5402E568ABED628F96D5ADB38FC6F975" type="ISSN">1175-5326</mods:identifier>
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<treatment id="039187D8FFC0BB17FF1884DDFAEFFBA5" ID-DOI="http://doi.org/10.5281/zenodo.4404249" ID-GBIF-Taxon="177012540" ID-Zenodo-Dep="4404249" LSID="urn:lsid:zoobank.org:act:7AC6F8AC-1F44-439A-A651-E48C80B38F53" httpUri="http://treatment.plazi.org/id/039187D8FFC0BB17FF1884DDFAEFFBA5" lastPageNumber="157" pageId="6" pageNumber="157">
<subSubSection id="C3226545FFC0BB17FF1884DDFE44FF4E" box="[151,473,151,178]" pageId="6" pageNumber="157" type="nomenclature">
<paragraph id="8B8736CEFFC0BB17FF1884DDFE44FF4E" blockId="6.[151,473,151,214]" box="[151,473,151,178]" pageId="6" pageNumber="157">
<heading id="D0CF81A2FFC0BB17FF1884DDFE44FF4E" bold="true" box="[151,473,151,178]" fontSize="11" level="1" pageId="6" pageNumber="157" reason="1">
<emphasis id="B94CEADCFFC0BB17FF1884DDFE44FF4E" bold="true" box="[151,473,151,178]" pageId="6" pageNumber="157">
<taxonomicName id="4C384D4DFFC0BB17FF1884DDFE12FF4D" authority="Gompel, 2020" authorityName="Gompel" authorityYear="2020" box="[151,399,151,178]" class="Insecta" family="Aderidae" genus="Zarcosia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="157" phylum="Arthropoda" rank="species" species="deharvengi" status="sp. nov.">
<emphasis id="B94CEADCFFC0BB17FF1884DDFE12FF4D" bold="true" box="[151,399,151,178]" italics="true" pageId="6" pageNumber="157">Zarcosia deharvengi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27F57A7FFC0BB17FE1884DDFE44FF4E" box="[407,473,151,177]" pageId="6" pageNumber="157" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3226545FFC0BB17FF1884F1FE2DFF29" box="[151,432,187,214]" pageId="6" pageNumber="157" type="description">
<paragraph id="8B8736CEFFC0BB17FF1884F1FE2DFF29" blockId="6.[151,473,151,214]" box="[151,432,187,214]" pageId="6" pageNumber="157">
(
<figureCitation id="13032A4BFFC0BB17FF1084F1FE96FF2A" box="[159,267,187,213]" captionStart="FIGURE 1" captionStartId="3.[151,250,1474,1498]" captionTargetBox="[158,1432,565,1446]" captionTargetId="figure-172@3.[151,1437,561,1451]" captionTargetPageId="3" captionText="FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line." figureDoi="http://doi.org/10.5281/zenodo.4403787" httpUri="https://zenodo.org/record/4403787/files/figure.png" pageId="6" pageNumber="157">Figures 1</figureCitation>
;
<figureCitation id="13032A4BFFC0BB17FE9784F1FEAEFF2A" box="[280,307,187,213]" captionStart="FIGURE 3" captionStartId="20.[151,250,188,212]" captionTargetId="figure-15@19.[244,1343,179,2016]" captionTargetPageId="19" captionText="FIGURE 3. Habitus of new Zarcosia species. a, Z. lemairei n. sp., ♂ holotype, from Laos. b, Z. lemairei n. sp., ♀ paratype, from Laos. c, Z. ilonae n. sp., ♂ holotype, from Thailand. d, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. e, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). f, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). g, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). h, Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra). i, Z. barclayi n. sp., ♂ holotype, from Malaysia. j, Z. barclayi n. sp., ♀ paratype, from Malaysia. k, Z. schawalleri n. sp., ♂ holotype, from Malaysia. l, Z. schawalleri n. sp., ♀ paratype, from Malaysia. m, Z. bedosae n. sp., ♂ holotype, from Laos. n, Z. sinuata n. sp., ♂ holotype, from Thailand. o, Z. sinuata n. sp., ♀ paratype, from Thailand. p, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. q, Z. dentatifemur (Pic, 1912), ♂ holotype, from Sri Lanka, after remounting (same specimen as in Fig. 2o). r, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. s, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. s, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4453388" httpUri="https://zenodo.org/record/4453388/files/figure.png" pageId="6" pageNumber="157">3f</figureCitation>
;
<figureCitation id="13032A4BFFC0BB17FEB184F1FEC1FF2A" box="[318,348,187,213]" captionStart="FIGURE 4" captionStartId="22.[151,250,186,210]" captionTargetId="figure-15@21.[275,1312,181,1998]" captionTargetPageId="21" captionText="FIGURE 4. Aedeagus of selected Zarcosia species. a, Z. barclayi n. sp., holotype, from Malaysia, lateral and dorsal views. b, Z. bedosae n. sp., holotype, from Laos, lateral and dorsal views. c, Z. bipartita (Pic, 1901) from India, lateral view. d, Z. cephalica (Pic, 1901) from Republic of Indonesia, lateral view. e, Z. deharvengi n. sp., holotype, from Laos, lateral and dorsal views. f, Z. gerstmeieri n. sp., holotype, from Vietnam, lateral and dorsal views. g, Z. ilonae n. sp., holotype, from Thailand, lateral and dorsal views. h, Z. kempi (Blair, 1924), holotype, from India, lateral and dorsal views. i, Z. lemairei n. sp., holotype, from Laos, lateral and dorsal views. j, Z. obscuricornis (Pic, 1927) from Vietnam, lateral view. k, Z. palliditarsis (Pic, 1912) from Sri Lanka, lateral view. l, Z. picina (Fairmaire, 1893), lectotype, from Vietnam, lateral view. m, Z. rubrobasalis (Pic, 1921) (slight uncertainty on the identity of this specimen) from Singapore, lateral view. n, Z. schawalleri n. sp., holotype, from Malaysia, lateral and dorsal views. o, Z. sinuata n. sp., holotype, from Thailand, lateral and dorsal views. p, Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra), lateral and dorsal views. q, Z. troglodytes (Champion, 1916), holotype, from Malaysia, lateral view. r, Z. weigeli n. sp., holotype, from China (Yunnan), lateral and dorsal views. s, Z. spinifemur n. sp., holotype, from Sri Lanka, lateral and dorsal views. t, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka, lateral and dorsal views. u, Z. srilankaensis n. sp., paratype, from Sri Lanka, lateral and dorsal views. v, Z. intermedia n. sp., paratype, from Sri Lanka, lateral and dorsal views. w, Z. brunnea n. sp., paratype, from Sri Lanka, lateral and dorsal views. Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4453392" httpUri="https://zenodo.org/record/4453392/files/figure.png" pageId="6" pageNumber="157">4e</figureCitation>
;
<figureCitation id="13032A4BFFC0BB17FEE784F1FE1DFF29" box="[360,384,187,214]" captionStart="FIGURE 5" captionStartId="25.[151,250,1602,1626]" captionTargetBox="[170,1408,188,1563]" captionTargetId="figure-15@25.[169,1419,181,1578]" captionTargetPageId="25" captionText="FIGURE 5. Morphological details of selected Zarcosia species. a–d, Z. lemairei n. sp., ♂ holotype, from Laos: head in lateral (a), frontal (b) and dorsal (c) views; mesothoracic (left) and metathoracic (right) tarsae (d). e–g, Z. palliditarsis (Pic, 1912), ♂, from Sri Lanka: head in lateral (e), frontal (f) and dorsal (g) views. h, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka: tuft of pubescence at elytral apex. i, Z. weigeli n. sp., ♂ holotype, from China (Yunnan): elytral pubescence on disc. j, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam: elytral pubescence on disc. k–t: male forelegs of Z. ilonae n. sp., holotype, from Thailand (k); Z. deharvengi n. sp., holotype, from Republic of Indonesia (Sumatra) (l); Z. schawalleri n. sp., holotype, from Malaysia (m); Z. sinuata n. sp., holotype, from Thailand (n); Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra) (o); Z. srilankaensis n. sp., paratype, from Sri Lanka (p); Z. intermedia n. sp., paratype, from Sri Lanka (q); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (r); Z. brunnea n. sp., paratype, from Sri Lanka (s); Z. weigeli n. sp., holotype, from China (Yunnan) (t). u–x: male mesothoracic legs of Z. cephalica (Pic, 1901) from Republic of Indonesia (u); Z. gerstmeieri n. sp., holotype, from Vietnam (v); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (w); Z. spinifemur n. sp., holotype, from Sri Lanka (x). Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4403791" httpUri="https://zenodo.org/record/4403791/files/figure.png" pageId="6" pageNumber="157">5l</figureCitation>
;
<figureCitation id="13032A4BFFC0BB17FE0384F1FE3AFF29" box="[396,423,187,214]" captionStart="FIGURE 6" captionStartId="26.[151,250,1631,1655]" captionTargetBox="[158,1431,530,1605]" captionTargetId="figure-126@26.[151,1436,525,1607]" captionTargetPageId="26" captionText="FIGURE 6. Thoraces of selected Zarcosia species in dorsal view. a, Z. lemairei n. sp., ♂ holotype, from Laos (left) and Z. lemairei n. sp., ♀ paratype, from Laos (right). b, Z. ilonae n. sp., ♂ holotype, from Thailand. c, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. d, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). e, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). f, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra) (left) and Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra) (right). g, Z. barclayi n. sp., ♂ holotype, from Malaysia (left) and Z. barclayi n. sp., ♀ paratype, from Malaysia (right). h, Z. schawalleri n. sp., ♂ holotype, from Malaysia (left) and Z. schawalleri n. sp., ♀ paratype, from Malaysia (right). i, Z. bedosae n. sp., ♂ holotype, from Laos. j, Z. sinuata n. sp., ♂ holotype, from Thailand (left) and Z. sinuata n. sp., ♀ paratype, from Thailand (right). k, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka. l, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. m, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. n, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. o, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4403795" httpUri="https://zenodo.org/record/4403795/files/figure.png" pageId="6" pageNumber="157">6e</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3226545FFC0BB17FF188549FA06FEE2" box="[151,1435,259,286]" pageId="6" pageNumber="157" type="etymology">
<paragraph id="8B8736CEFFC0BB17FF188549FA06FEE2" blockId="6.[151,1437,259,1114]" box="[151,1435,259,286]" pageId="6" pageNumber="157">
<emphasis id="B94CEADCFFC0BB17FF188549FE83FEE2" bold="true" box="[151,286,259,285]" pageId="6" pageNumber="157">Etymology.</emphasis>
The species is named after Louis Deharveng in recognition of his efforts to collect material for others.
</paragraph>
</subSubSection>
<subSubSection id="C3226545FFC0BB17FF48856DFAA6FEBE" box="[199,1339,294,322]" pageId="6" pageNumber="157" type="materials_examined">
<paragraph id="8B8736CEFFC0BB17FF48856DFAA6FEBE" blockId="6.[151,1437,259,1114]" box="[199,1339,294,322]" pageId="6" pageNumber="157">
<materialsCitation id="3B503C93FFC0BB17FF48856DFAAAFEBE" ID-GBIF-Occurrence="3020160307" box="[199,1335,294,322]" country="Indonesia" latitude="-0.23333333" location="Ngalau Simarasop" longLatPrecision="1308" longitude="100.46667" municipality="Baso" pageId="6" pageNumber="157" specimenCount="1" stateProvince="Sumatera Barat" typeStatus="holotype">
<emphasis id="B94CEADCFFC0BB17FF48856DFEFBFEBE" bold="true" box="[199,358,295,321]" pageId="6" pageNumber="157">
<typeStatus id="5483886CFFC0BB17FF48856DFE9FFEBE" box="[199,258,295,321]" pageId="6" pageNumber="157">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F32F765EFFC0BB17FEE3856DFDF5FEBE" box="[364,616,295,321]" name="Indonesia" pageId="6" pageNumber="157">Republic of Indonesia</collectingCountry>
:
<collectingRegion id="49FCF82CFFC0BB17FDFD856DFCBFFEBD" box="[626,802,295,322]" country="Indonesia" name="Sumatera Barat" pageId="6" pageNumber="157">Sumatera Barat</collectingRegion>
,
<collectingMunicipality id="6BE3ACB4FFC0BB17FCA18562FCF4FEBD" box="[814,873,296,322]" pageId="6" pageNumber="157">Baso</collectingMunicipality>
,
<location id="8EE76015FFC0BB17FCFB856DFBDFFEBE" LSID="urn:lsid:plazi:treatment:039187D8FFC0BB17FF1884DDFAEFFBA5:8EE76015FFC0BB17FCFB856DFBDFFEBE" box="[884,1090,295,321]" country="Indonesia" latitude="-0.23333333" longLatPrecision="1308" longitude="100.46667" municipality="Baso" name="Ngalau Simarasop" pageId="6" pageNumber="157" stateProvince="Sumatera Barat">Ngalau Simarasop</location>
[
<geoCoordinate id="EE0C5009FFC0BB17FBDF856DFB29FEBE" box="[1104,1204,295,321]" degrees="00" direction="south" minutes="14" orientation="latitude" pageId="6" pageNumber="157" precision="925" value="-0.23333333">00°14’ S</geoCoordinate>
<geoCoordinate id="EE0C5009FFC0BB17FB35856DFAADFEBE" box="[1210,1328,294,321]" degrees="100" direction="east" minutes="28" orientation="longitude" pageId="6" pageNumber="157" precision="925" value="100.46667">100°28’ E</geoCoordinate>
]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3226545FFC0BB17FF488501FCD4FE9A" box="[199,841,331,357]" pageId="6" pageNumber="157" type="distribution">
<paragraph id="8B8736CEFFC0BB17FF488501FCD4FE9A" blockId="6.[151,1437,259,1114]" box="[199,841,331,357]" pageId="6" pageNumber="157">
<emphasis id="B94CEADCFFC0BB17FF488501FEFCFE9A" bold="true" box="[199,353,331,357]" pageId="6" pageNumber="157">Distribution.</emphasis>
<collectingCountry id="F32F765EFFC0BB17FEE88501FDFEFE9A" box="[359,611,331,357]" name="Indonesia" pageId="6" pageNumber="157">Republic of Indonesia</collectingCountry>
:
<collectingRegion id="49FCF82CFFC0BB17FDE18501FD56FE9A" box="[622,715,331,357]" country="Indonesia" name="Sumatera Utara" pageId="6" pageNumber="157">Sumatra</collectingRegion>
(
<figureCitation id="13032A4BFFC0BB17FD568501FCA5FE9A" box="[729,824,331,357]" captionStart="FIGURE 1" captionStartId="3.[151,250,1474,1498]" captionTargetBox="[158,1432,565,1446]" captionTargetId="figure-172@3.[151,1437,561,1451]" captionTargetPageId="3" captionText="FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line." figureDoi="http://doi.org/10.5281/zenodo.4403787" httpUri="https://zenodo.org/record/4403787/files/figure.png" pageId="6" pageNumber="157">Figure 1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3226545FFC0BB17FF488525FB76FCA1" pageId="6" pageNumber="157" type="description">
<paragraph id="8B8736CEFFC0BB17FF488525FB11FE52" blockId="6.[151,1437,259,1114]" pageId="6" pageNumber="157">
<emphasis id="B94CEADCFFC0BB17FF488525FEC4FE76" bold="true" box="[199,345,367,393]" pageId="6" pageNumber="157">Description.</emphasis>
(
<figureCitation id="13032A4BFFC0BB17FEE78525FE48FE75" box="[360,469,367,394]" captionStart="FIGURE 3" captionStartId="20.[151,250,188,212]" captionTargetId="figure-15@19.[244,1343,179,2016]" captionTargetPageId="19" captionText="FIGURE 3. Habitus of new Zarcosia species. a, Z. lemairei n. sp., ♂ holotype, from Laos. b, Z. lemairei n. sp., ♀ paratype, from Laos. c, Z. ilonae n. sp., ♂ holotype, from Thailand. d, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. e, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). f, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). g, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). h, Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra). i, Z. barclayi n. sp., ♂ holotype, from Malaysia. j, Z. barclayi n. sp., ♀ paratype, from Malaysia. k, Z. schawalleri n. sp., ♂ holotype, from Malaysia. l, Z. schawalleri n. sp., ♀ paratype, from Malaysia. m, Z. bedosae n. sp., ♂ holotype, from Laos. n, Z. sinuata n. sp., ♂ holotype, from Thailand. o, Z. sinuata n. sp., ♀ paratype, from Thailand. p, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. q, Z. dentatifemur (Pic, 1912), ♂ holotype, from Sri Lanka, after remounting (same specimen as in Fig. 2o). r, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. s, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. s, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4453388" httpUri="https://zenodo.org/record/4453388/files/figure.png" pageId="6" pageNumber="157">Figure 3f</figureCitation>
). Female unknown. Body length.
<specimenCount id="9D3EFD47FFC0BB17FCDD853AFCF7FE75" box="[850,874,368,394]" pageId="6" pageNumber="157" type="male">♂</specimenCount>
: 2.43±
<quantity id="4CC09B2BFFC0BB17FC398525FB86FE75" box="[950,1051,367,394]" metricMagnitude="-4" metricUnit="m" metricValue="1.9" pageId="6" pageNumber="157" unit="mm" value="0.19">0.19 mm</quantity>
(n=3). Dark brown, darker anteriorly, gracile, covered with very short, pulverulent pubescence. Appendages orange-brown.
</paragraph>
<paragraph id="8B8736CEFFC0BB17FF4885FDFF57FDE6" blockId="6.[151,1437,259,1114]" pageId="6" pageNumber="157">
<emphasis id="B94CEADCFFC0BB17FF4885FDFE99FE2E" box="[199,260,439,465]" italics="true" pageId="6" pageNumber="157">Head</emphasis>
. Small, dark, barely wider than pronotum. Palpi yellow, last article securiform. Integument, finely and densely punctuated, grainy, looking like velvet. Occiput lined with a fringe of dense and short silver hairs posteriorly.
</paragraph>
<paragraph id="8B8736CEFFC0BB17FF488669FAFFFDC2" blockId="6.[151,1437,259,1114]" box="[199,1378,547,574]" pageId="6" pageNumber="157">
<emphasis id="B94CEADCFFC0BB17FF488669FEACFDC2" box="[199,305,547,573]" italics="true" pageId="6" pageNumber="157">Antennae</emphasis>
. 1.5 the length of elytra, orange-brown, regularly filiform with antenommeres 3–10 cylindrical.
</paragraph>
<paragraph id="8B8736CEFFC0BB17FF488602FB96FD7A" blockId="6.[151,1437,259,1114]" pageId="6" pageNumber="157">
<emphasis id="B94CEADCFFC0BB17FF488602FEABFD9E" box="[199,310,584,609]" italics="true" pageId="6" pageNumber="157">Pronotum</emphasis>
(
<figureCitation id="13032A4BFFC0BB17FECA860DFE29FD9D" box="[325,436,583,610]" captionStart="FIGURE 6" captionStartId="26.[151,250,1631,1655]" captionTargetBox="[158,1431,530,1605]" captionTargetId="figure-126@26.[151,1436,525,1607]" captionTargetPageId="26" captionText="FIGURE 6. Thoraces of selected Zarcosia species in dorsal view. a, Z. lemairei n. sp., ♂ holotype, from Laos (left) and Z. lemairei n. sp., ♀ paratype, from Laos (right). b, Z. ilonae n. sp., ♂ holotype, from Thailand. c, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. d, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). e, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). f, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra) (left) and Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra) (right). g, Z. barclayi n. sp., ♂ holotype, from Malaysia (left) and Z. barclayi n. sp., ♀ paratype, from Malaysia (right). h, Z. schawalleri n. sp., ♂ holotype, from Malaysia (left) and Z. schawalleri n. sp., ♀ paratype, from Malaysia (right). i, Z. bedosae n. sp., ♂ holotype, from Laos. j, Z. sinuata n. sp., ♂ holotype, from Thailand (left) and Z. sinuata n. sp., ♀ paratype, from Thailand (right). k, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka. l, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. m, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. n, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. o, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4403795" httpUri="https://zenodo.org/record/4403795/files/figure.png" pageId="6" pageNumber="157">Figure 6e</figureCitation>
). Round and convex, as long as wide, sides regularly curved. Base sinuated. Surface mat, rugose, covered with dense, velvety pubescence. Punctation undistinguishable.
</paragraph>
<paragraph id="8B8736CEFFC0BB17FF4886C5FD13FD0E" blockId="6.[151,1437,259,1114]" pageId="6" pageNumber="157">
<emphasis id="B94CEADCFFC0BB17FF4886C5FE90FD56" box="[199,269,655,681]" italics="true" pageId="6" pageNumber="157">Elytra</emphasis>
. Length: 1.72±
<quantity id="4CC09B2BFFC0BB17FE3586C5FD82FD55" box="[442,543,655,682]" metricMagnitude="-4" metricUnit="m" metricValue="1.3" pageId="6" pageNumber="157" unit="mm" value="0.13">0.13 mm</quantity>
(n=3). Punctation fine and shallow, dense (space between points as wide as the points themselves). Disc depressed. Pubescence dense and short, whitish, comprised of very short hairs borne from the points and interspersed pulverulent hairs.
</paragraph>
<paragraph id="8B8736CEFFC0BB17FF4886B6FA97FCE9" blockId="6.[151,1437,259,1114]" box="[199,1290,763,790]" pageId="6" pageNumber="157">
<emphasis id="B94CEADCFFC0BB17FF4886B6FF61FCEA" box="[199,252,764,789]" italics="true" pageId="6" pageNumber="157">Legs</emphasis>
. Particularly elongated. Foretibia thickened in a smooth tooth on their inner side (
<figureCitation id="13032A4BFFC0BB17FB1D86B1FB61FCE9" box="[1170,1276,763,790]" captionStart="FIGURE 5" captionStartId="25.[151,250,1602,1626]" captionTargetBox="[170,1408,188,1563]" captionTargetId="figure-15@25.[169,1419,181,1578]" captionTargetPageId="25" captionText="FIGURE 5. Morphological details of selected Zarcosia species. a–d, Z. lemairei n. sp., ♂ holotype, from Laos: head in lateral (a), frontal (b) and dorsal (c) views; mesothoracic (left) and metathoracic (right) tarsae (d). e–g, Z. palliditarsis (Pic, 1912), ♂, from Sri Lanka: head in lateral (e), frontal (f) and dorsal (g) views. h, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka: tuft of pubescence at elytral apex. i, Z. weigeli n. sp., ♂ holotype, from China (Yunnan): elytral pubescence on disc. j, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam: elytral pubescence on disc. k–t: male forelegs of Z. ilonae n. sp., holotype, from Thailand (k); Z. deharvengi n. sp., holotype, from Republic of Indonesia (Sumatra) (l); Z. schawalleri n. sp., holotype, from Malaysia (m); Z. sinuata n. sp., holotype, from Thailand (n); Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra) (o); Z. srilankaensis n. sp., paratype, from Sri Lanka (p); Z. intermedia n. sp., paratype, from Sri Lanka (q); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (r); Z. brunnea n. sp., paratype, from Sri Lanka (s); Z. weigeli n. sp., holotype, from China (Yunnan) (t). u–x: male mesothoracic legs of Z. cephalica (Pic, 1901) from Republic of Indonesia (u); Z. gerstmeieri n. sp., holotype, from Vietnam (v); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (w); Z. spinifemur n. sp., holotype, from Sri Lanka (x). Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4403791" httpUri="https://zenodo.org/record/4403791/files/figure.png" pageId="6" pageNumber="157">Figure 5l</figureCitation>
).
</paragraph>
<paragraph id="8B8736CEFFC0BB17FF488755FB76FCA1" blockId="6.[151,1437,259,1114]" pageId="6" pageNumber="157">
<emphasis id="B94CEADCFFC0BB17FF488755FEA9FCC6" box="[199,308,799,825]" italics="true" pageId="6" pageNumber="157">Aedeagus</emphasis>
(
<figureCitation id="13032A4BFFC0BB17FEC98755FE24FCC6" box="[326,441,799,825]" captionStart="FIGURE 4" captionStartId="22.[151,250,186,210]" captionTargetId="figure-15@21.[275,1312,181,1998]" captionTargetPageId="21" captionText="FIGURE 4. Aedeagus of selected Zarcosia species. a, Z. barclayi n. sp., holotype, from Malaysia, lateral and dorsal views. b, Z. bedosae n. sp., holotype, from Laos, lateral and dorsal views. c, Z. bipartita (Pic, 1901) from India, lateral view. d, Z. cephalica (Pic, 1901) from Republic of Indonesia, lateral view. e, Z. deharvengi n. sp., holotype, from Laos, lateral and dorsal views. f, Z. gerstmeieri n. sp., holotype, from Vietnam, lateral and dorsal views. g, Z. ilonae n. sp., holotype, from Thailand, lateral and dorsal views. h, Z. kempi (Blair, 1924), holotype, from India, lateral and dorsal views. i, Z. lemairei n. sp., holotype, from Laos, lateral and dorsal views. j, Z. obscuricornis (Pic, 1927) from Vietnam, lateral view. k, Z. palliditarsis (Pic, 1912) from Sri Lanka, lateral view. l, Z. picina (Fairmaire, 1893), lectotype, from Vietnam, lateral view. m, Z. rubrobasalis (Pic, 1921) (slight uncertainty on the identity of this specimen) from Singapore, lateral view. n, Z. schawalleri n. sp., holotype, from Malaysia, lateral and dorsal views. o, Z. sinuata n. sp., holotype, from Thailand, lateral and dorsal views. p, Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra), lateral and dorsal views. q, Z. troglodytes (Champion, 1916), holotype, from Malaysia, lateral view. r, Z. weigeli n. sp., holotype, from China (Yunnan), lateral and dorsal views. s, Z. spinifemur n. sp., holotype, from Sri Lanka, lateral and dorsal views. t, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka, lateral and dorsal views. u, Z. srilankaensis n. sp., paratype, from Sri Lanka, lateral and dorsal views. v, Z. intermedia n. sp., paratype, from Sri Lanka, lateral and dorsal views. w, Z. brunnea n. sp., paratype, from Sri Lanka, lateral and dorsal views. Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4453392" httpUri="https://zenodo.org/record/4453392/files/figure.png" pageId="6" pageNumber="157">Figure 4e</figureCitation>
). Sides weakly and progressively convergent in dorsal view, apex rounded. Long, extremely bent in lateral view, apex with a brief downward deflection. Appendages visible near apex.
</paragraph>
</subSubSection>
<subSubSection id="C3226545FFC0BB17FF48872DFAEFFBA5" pageId="6" pageNumber="157" type="materials_examined">
<paragraph id="8B8736CEFFC0BB17FF48872DFD16FBC9" blockId="6.[151,1437,259,1114]" pageId="6" pageNumber="157">
<emphasis id="B94CEADCFFC0BB17FF48872DFEEBFC7E" bold="true" box="[199,374,871,897]" pageId="6" pageNumber="157">Type material.</emphasis>
<materialsCitation id="3B503C93FFC0BB17FEF1872DFB18FC5A" ID-GBIF-Occurrence="3020160352" collectingDate="2000-07-10" collectionCode="MNHN, A" collectorName="L. Deharveng &amp; Bedos" country="Indonesia" elevation="740" latitude="-0.23333333" location="Ngalau Simarasop" longLatPrecision="1308" longitude="100.46667" municipality="Baso" pageId="6" pageNumber="157" specimenCount="1" specimenCount-male="1" stateProvince="Sumatera Utara" typeStatus="holotype">
<typeStatus id="5483886CFFC0BB17FEF1872DFE7BFC7E" box="[382,486,871,897]" pageId="6" pageNumber="157" type="holotype">Holotype</typeStatus>
(
<specimenCount id="9D3EFD47FFC0BB17FE77872DFD92FC7E" box="[504,527,871,897]" pageId="6" pageNumber="157" type="male">♂</specimenCount>
,
<collectionCode id="ED29AE0BFFC0BB17FD938722FDEAFC7D" box="[540,631,872,898]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="6" pageNumber="157">MNHN</collectionCode>
).
<emphasis id="B94CEADCFFC0BB17FD05872DFC0FFC7E" bold="true" box="[650,914,871,897]" pageId="6" pageNumber="157">
<collectingCountry id="F32F765EFFC0BB17FD05872DFC0FFC7E" box="[650,914,871,897]" name="Indonesia" pageId="6" pageNumber="157">Republic of Indonesia</collectingCountry>
</emphasis>
:
<collectingRegion id="49FCF82CFFC0BB17FC2E872DFB9CFC7E" box="[929,1025,871,897]" country="Indonesia" name="Sumatera Utara" pageId="6" pageNumber="157">Sumatra</collectingRegion>
,
<collectingRegion id="49FCF82CFFC0BB17FB828722FB30FC7E" box="[1037,1197,871,898]" country="Indonesia" name="Sumatera Barat" pageId="6" pageNumber="157">West Sumatra</collectingRegion>
,
<collectingMunicipality id="6BE3ACB4FFC0BB17FB368722FB69FC7D" box="[1209,1268,872,898]" pageId="6" pageNumber="157">Baso</collectingMunicipality>
,
<location id="8EE76015FFC0BB17FA8F872DFF4BFC59" LSID="urn:lsid:plazi:treatment:039187D8FFC0BB17FF1884DDFAEFFBA5:8EE76015FFC0BB17FA8F872DFF4BFC59" country="Indonesia" latitude="-0.23333333" longLatPrecision="1308" longitude="100.46667" municipality="Baso" name="Ngalau Simarasop" pageId="6" pageNumber="157" stateProvince="Sumatera Utara">Ngalau Simarasop</location>
, (
<quantity id="4CC09B2BFFC0BB17FF6387C1FEA4FC59" box="[236,313,907,934]" metricMagnitude="2" metricUnit="m" metricValue="7.4" pageId="6" pageNumber="157" unit="m" value="740.0">
<elevation id="0015D1FDFFC0BB17FF6387C1FEA4FC59" box="[236,313,907,934]" metricMagnitude="2" metricUnit="m" metricValue="7.4" pageId="6" pageNumber="157" unit="m" value="740.0">740 m</elevation>
</quantity>
) cave,
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<geoCoordinate id="EE0C5009FFC0BB17FE7387C1FDEDFC59" box="[508,624,907,934]" degrees="100" direction="east" minutes="28" orientation="longitude" pageId="6" pageNumber="157" precision="925" value="100.46667">100°28’ E</geoCoordinate>
(
<collectorName id="26CD5318FFC0BB17FD0D87C1FCBDFC5A" box="[642,800,907,933]" pageId="6" pageNumber="157">L. Deharveng</collectorName>
&amp;
<collectionCode id="ED29AE0BFFC0BB17FCC887C1FCC3FC5A" box="[839,862,907,933]" pageId="6" pageNumber="157">A</collectionCode>
.
<collectorName id="26CD5318FFC0BB17FCE387C1FC2FFC5A" box="[876,946,907,933]" pageId="6" pageNumber="157">Bedos</collectorName>
leg.)
<date id="FF86100EFFC0BB17FC7787C1FB18FC5A" box="[1016,1157,907,933]" pageId="6" pageNumber="157" value="2000-07-10">
<collectingDate id="EFC2E9E6FFC0BB17FC7787C1FB18FC5A" box="[1016,1157,907,933]" pageId="6" pageNumber="157" value="2000-07-10">10.VII.2000</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="3B503C93FFC0BB17FB1D87C6FF72FBEE" ID-GBIF-Occurrence="3020160365" collectingDate="1984-07-26" collectionCode="NGPC" country="Indonesia" latitude="5.45" location="Jungle Quarry Cave" longLatPrecision="1305" longitude="95.23333" municipality="Besar Regency" pageId="6" pageNumber="157" specimenCount="2" specimenCount-male="1" stateProvince="Sumatera Utara" typeStatus="paratype">
<typeStatus id="5483886CFFC0BB17FB1D87C6FA9EFC59" box="[1170,1283,908,934]" pageId="6" pageNumber="157" type="paratype">Paratypes</typeStatus>
.
<emphasis id="B94CEADCFFC0BB17FA9F87C1FE91FC36" bold="true" pageId="6" pageNumber="157">
<collectingCountry id="F32F765EFFC0BB17FA9F87C1FE91FC36" name="Indonesia" pageId="6" pageNumber="157">Republic of Indonesia</collectingCountry>
</emphasis>
:
<collectingRegion id="49FCF82CFFC0BB17FE9387E5FEE1FC36" box="[284,380,943,969]" country="Indonesia" name="Sumatera Utara" pageId="6" pageNumber="157">Sumatra</collectingRegion>
,
<collectingRegion id="49FCF82CFFC0BB17FE0887E5FE58FC36" box="[391,453,943,969]" country="Indonesia" name="Aceh" pageId="6" pageNumber="157">Aceh</collectingRegion>
,
<collectingRegion id="49FCF82CFFC0BB17FE5F87E5FD96FC36" box="[464,523,943,969]" country="Indonesia" name="Aceh" pageId="6" pageNumber="157">Aceh</collectingRegion>
<collectingMunicipality id="6BE3ACB4FFC0BB17FD9C87FAFD23FC35" box="[531,702,944,970]" pageId="6" pageNumber="157">Besar Regency</collectingMunicipality>
,
<collectorName id="26CD5318FFC0BB17FD4587E5FCE3FC35" box="[714,894,943,970]" pageId="6" pageNumber="157">Lhok’Nga, P. T.</collectorName>
Semen Andalas,
<geoCoordinate id="EE0C5009FFC0BB17FBCB87E5FB36FC35" box="[1092,1195,943,970]" degrees="05" direction="north" minutes="27" orientation="latitude" pageId="6" pageNumber="157" precision="925" value="5.45">05°27’ N</geoCoordinate>
<geoCoordinate id="EE0C5009FFC0BB17FB3C87E5FA84FC35" box="[1203,1305,943,970]" degrees="95" direction="east" minutes="14" orientation="longitude" pageId="6" pageNumber="157" precision="925" value="95.23333">95°14’ E</geoCoordinate>
,
<collectorName id="26CD5318FFC0BB17FAA987E5FF7AFC12" pageId="6" pageNumber="157">Limestone Quarry</collectorName>
,
<location id="8EE76015FFC0BB17FF7C8799FE4AFC12" LSID="urn:lsid:plazi:treatment:039187D8FFC0BB17FF1884DDFAEFFBA5:8EE76015FFC0BB17FF7C8799FE4AFC12" box="[243,471,979,1005]" country="Indonesia" latitude="5.45" longLatPrecision="1305" longitude="95.23333" municipality="Besar Regency" name="Jungle Quarry Cave" pageId="6" pageNumber="157" stateProvince="Sumatera Utara">Jungle Quarry Cave</location>
, 2nd cavern, twilight zone, on sand near guano (
<collectorName id="26CD5318FFC0BB17FC718799FBC4FC11" box="[1022,1113,979,1006]" pageId="6" pageNumber="157">I. Davis</collectorName>
)
<date id="FF86100EFFC0BB17FBE78799FB6FFC12" box="[1128,1266,979,1005]" pageId="6" pageNumber="157" value="1984-07-26">
<collectingDate id="EFC2E9E6FFC0BB17FBE78799FB6FFC12" box="[1128,1266,979,1005]" pageId="6" pageNumber="157" value="1984-07-26">26.VII.1984</collectingDate>
</date>
(
<specimenCount id="9D3EFD47FFC0BB17FA8B8799FAB0FC11" box="[1284,1325,979,1006]" pageId="6" pageNumber="157" type="male">1 ♂</specimenCount>
<typeStatus id="5483886CFFC0BB17FABA879EFA05FC11" box="[1333,1432,980,1006]" pageId="6" pageNumber="157" type="paratype">paratype</typeStatus>
,
<collectionCode id="ED29AE0BFFC0BB17FF1887BDFF7AFBEE" box="[151,231,1015,1041]" pageId="6" pageNumber="157">NGPC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B503C93FFC0BB17FF7387BDFD1AFBC9" ID-GBIF-Occurrence="3020160450" collectingDate="1984-07-26" collectionCode="BMNH" collectorName="C. Covill" country="Indonesia" county="Penen" latitude="3.3258333" location="Kota Tinggi" longLatPrecision="21" longitude="98.6325" municipality="Rampah Cave" pageId="6" pageNumber="157" specimenCount="1" specimenCount-male="1" stateProvince="Sumatera Utara" typeStatus="paratype">
<collectingRegion id="49FCF82CFFC0BB17FF7387BDFEC1FBEE" box="[252,348,1015,1041]" country="Indonesia" name="Sumatera Utara" pageId="6" pageNumber="157">Sumatra</collectingRegion>
,
<collectingRegion id="49FCF82CFFC0BB17FEE587BDFD89FBEE" box="[362,532,1015,1041]" country="Indonesia" name="Sumatera Utara" pageId="6" pageNumber="157">North Sumatra</collectingRegion>
,
<collectingCounty id="62E64E42FFC0BB17FDAD87B2FDF4FBED" box="[546,617,1016,1042]" pageId="6" pageNumber="157">Penen</collectingCounty>
,
<collectingMunicipality id="6BE3ACB4FFC0BB17FDF987BDFC84FBEE" box="[630,793,1015,1041]" pageId="6" pageNumber="157">Rampah Cave</collectingMunicipality>
,
<quantity id="4CC09B2BFFC0BB17FCA987BDFCFFFBEE" box="[806,866,1015,1041]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="157" unit="km" value="1.0">1 km</quantity>
E. of
<location id="8EE76015FFC0BB17FC2387B2FBA5FBEE" LSID="urn:lsid:plazi:treatment:039187D8FFC0BB17FF1884DDFAEFFBA5:8EE76015FFC0BB17FC2387B2FBA5FBEE" box="[940,1080,1015,1042]" country="Indonesia" county="Penen" latitude="3.3258333" longLatPrecision="21" longitude="98.6325" municipality="Rampah Cave" name="Kota Tinggi" pageId="6" pageNumber="157" stateProvince="Sumatera Utara">Kota Tinggi</location>
,
<geoCoordinate id="EE0C5009FFC0BB17FBCA87BDFB4AFBED" box="[1093,1239,1015,1042]" degrees="03" direction="north" minutes="19" orientation="latitude" pageId="6" pageNumber="157" precision="15" seconds="33" value="3.3258333">03°19’33” N</geoCoordinate>
<geoCoordinate id="EE0C5009FFC0BB17FB6E87BDFAEDFBED" box="[1249,1392,1015,1042]" degrees="98" direction="east" minutes="37" orientation="longitude" pageId="6" pageNumber="157" precision="15" seconds="57" value="98.6325">98°37’57” E</geoCoordinate>
(
<collectorName id="26CD5318FFC0BB17FA0D87BDFF42FBC9" pageId="6" pageNumber="157">C. Covill</collectorName>
)
<date id="FF86100EFFC0BB17FF618051FEEAFBCA" box="[238,375,1051,1077]" pageId="6" pageNumber="157" value="1984-07-26">
<collectingDate id="EFC2E9E6FFC0BB17FF618051FEEAFBCA" box="[238,375,1051,1077]" pageId="6" pageNumber="157" value="1984-07-26">26.VII.1984</collectingDate>
</date>
(
<specimenCount id="9D3EFD47FFC0BB17FE078051FE2CFBC9" box="[392,433,1051,1078]" pageId="6" pageNumber="157" type="male">1 ♂</specimenCount>
<typeStatus id="5483886CFFC0BB17FE378056FD86FBC9" box="[440,539,1052,1078]" pageId="6" pageNumber="157" type="paratype">paratype</typeStatus>
,
<collectionCode id="ED29AE0BFFC0BB17FDA98056FDE2FBC9" box="[550,639,1052,1078]" pageId="6" pageNumber="157">BMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B8736CEFFC0BB17FF488075FAEFFBA5" blockId="6.[151,1437,259,1114]" box="[199,1394,1087,1114]" pageId="6" pageNumber="157">
<emphasis id="B94CEADCFFC0BB17FF488075FF62FBA6" bold="true" box="[199,255,1087,1113]" pageId="6" pageNumber="157">Note</emphasis>
. The extremely thin and elongated appendages of species likely represent adaptations to life in caves.
</paragraph>
</subSubSection>
</treatment>
</document>
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