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<document id="A6934EE9C5D43245AA7EF6933DBA2F8F" ID-CLB-Dataset="3256" ID-DOI="10.5281/zenodo.5733446" ID-GBIF-Dataset="9953df2f-b6bb-408a-980c-146175454d39" ID-ISBN="978-84-16728-19-0" ID-Zenodo-Dep="5733446" IM.illustrations_requiresApprovalFor="plazi" IM.metadata_requiresApprovalFor="plazi" IM.taxonomicNames_requiresApprovalFor="plazi" checkinTime="1600872631645" checkinUser="plazi" docAuthor="Don E. Wilson &amp; Russell A. Mittermeier" docDate="2019" docId="039187AC5C3F2A52F099001F399BF24E" docLanguage="en" docName="hbmw_9_Furipteridae_412.pdf.imf" docOrigin="Handbook of the Mammals of the World Volume 9 Bats, Barcelona: Lynx Edicions" docTitle="Amorphochilus schnablii Peters 1877" docType="treatment" docVersion="11" lastPageNumber="416" masterDocId="FFA8FFD45C3E2A53F03609113F64FFC4" masterDocTitle="Furipteridae" masterLastPageNumber="417" masterPageNumber="412" pageNumber="416" updateTime="1698845888388" updateUser="ExternalLinkService">
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<mods:title id="503B1540D5B6F65EB52CEDF52BB6F271">Furipteridae</mods:title>
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<mods:namePart id="3B8FB298658FB22BC8C880A98045F14E">Don E. Wilson</mods:namePart>
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<mods:namePart id="BF4072C3F9FF2042442CB34BFA13858D">Russell A. Mittermeier</mods:namePart>
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<mods:title id="8190EC0EE21C30BCEEDAA9FFDFED291F">Handbook of the Mammals of the World Volume 9 Bats</mods:title>
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<heading id="D0CF81D65C3F2A52F099001F3FA8F6F8" box="[175,204,2318,2364]" pageId="1" pageNumber="416">1.</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32265315C3F2A52F0E8001F3EDFF6F8" box="[222,443,2318,2364]" pageId="1" pageNumber="416" type="vernacular_names">
<paragraph id="8B8736BA5C3F2A52F0E8001F3EDFF6F8" blockId="1.[172,1188,2318,2443]" box="[222,443,2318,2364]" pageId="1" pageNumber="416">
<heading id="D0CF81D65C3F2A52F0E8001F3EDFF6F8" box="[222,443,2318,2364]" pageId="1" pageNumber="416">
<vernacularName id="053B46945C3F2A52F0E8001F3EDFF6F8" box="[222,443,2318,2364]" pageId="1" pageNumber="416">Smoky Bat</vernacularName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32265315C3F2A52F1DA001F3CDEF6F8" box="[492,954,2318,2364]" pageId="1" pageNumber="416" type="nomenclature">
<paragraph id="8B8736BA5C3F2A52F1DA001F3CDEF6F8" blockId="1.[172,1188,2318,2443]" box="[492,954,2318,2364]" pageId="1" pageNumber="416">
<heading id="D0CF81D65C3F2A52F1DA001F3CDEF6F8" box="[492,954,2318,2364]" pageId="1" pageNumber="416">
<taxonomicName id="4C384D395C3F2A52F1DA001F3CDEF6F8" ID-CoL="66DQS" authorityName="Peters" authorityYear="1877" box="[492,954,2318,2364]" class="Mammalia" family="Furipteridae" genus="Amorphochilus" kingdom="Animalia" order="Chiroptera" pageId="1" pageNumber="416" phylum="Chordata" rank="species" species="schnablii">
<emphasis id="B94CEAA85C3F2A52F1DA001F3CDEF6F8" box="[492,954,2318,2364]" italics="true" pageId="1" pageNumber="416">Amorphochilus schnablii</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32265315C3F2A52F09A005F3D19F64F" pageId="1" pageNumber="416" type="vernacular_names">
<paragraph id="8B8736BA5C3F2A52F09A005F3BC0F6A7" blockId="1.[172,1188,2318,2443]" box="[172,1188,2382,2403]" pageId="1" pageNumber="416">
<heading id="D0CF81D65C3F2A52F09A005F3BC0F6A7" box="[172,1188,2382,2403]" pageId="1" pageNumber="416">
<emphasis id="B94CEAA85C3F2A52F09A005F3F93F6A7" bold="true" box="[172,247,2382,2403]" pageId="1" pageNumber="416">French:</emphasis>
<vernacularName id="053B46945C3F2A52F137005F3EC0F6A7" box="[257,420,2382,2403]" pageId="1" pageNumber="416">Furie de Schnabl</vernacularName>
/
<emphasis id="B94CEAA85C3F2A52F18C005F3D70F6A7" bold="true" box="[442,532,2382,2403]" pageId="1" pageNumber="416">German:</emphasis>
<vernacularName id="053B46945C3F2A52F228005F3C32F6A7" box="[542,854,2382,2403]" pageId="1" pageNumber="416">Felsenklsten-Stummeldaumen</vernacularName>
/
<emphasis id="B94CEAA85C3F2A52F35A005F3CA3F6A7" bold="true" box="[876,967,2382,2403]" pageId="1" pageNumber="416">Spanish:</emphasis>
<vernacularName id="053B46945C3F2A52F3E4005F3B24F6A7" box="[978,1088,2382,2403]" pageId="1" pageNumber="416">Murciélago</vernacularName>
ahumado
</heading>
</paragraph>
<paragraph id="8B8736BA5C3F2A52F09A00673D19F64F" blockId="1.[172,1188,2318,2443]" box="[172,637,2422,2443]" pageId="1" pageNumber="416">
<heading id="D0CF81D65C3F2A52F09A00673D19F64F" box="[172,637,2422,2443]" pageId="1" pageNumber="416">
<emphasis id="B94CEAA85C3F2A52F09A00673EC7F64F" bold="true" box="[172,419,2422,2443]" pageId="1" pageNumber="416">Other common names:</emphasis>
<vernacularName id="053B46945C3F2A52F19B00673D19F64F" box="[429,637,2422,2443]" pageId="1" pageNumber="416">Schnabl's Smoky Bat</vernacularName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32265315C3F2A52F32400D13C38F5C4" pageId="1" pageNumber="416" type="reference_group">
<paragraph id="8B8736BA5C3F2A52F32400D13C38F5C4" blockId="1.[786,1377,2496,2915]" pageId="1" pageNumber="416">
<emphasis id="B94CEAA85C3F2A52F32400D13CC9F61D" bold="true" box="[786,941,2496,2521]" pageId="1" pageNumber="416">Taxonomy.</emphasis>
<taxonomicName id="4C384D395C3F2A52F38B00D13C3CF5C4" ID-CoL="66DQS" authority="Peters, 1877" authorityName="Peters" authorityYear="1877" class="Mammalia" family="Furipteridae" genus="Amorphochilus" kingdom="Animalia" order="Chiroptera" pageId="1" pageNumber="416" phylum="Chordata" rank="species" species="schnablii">Amorphochilus schnablii Peters, 1877</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C32265315C3F2A52F34600F63B19F559" pageId="1" pageNumber="416" type="materials_examined">
<paragraph id="8B8736BA5C3F2A52F34600F63B64F5B2" blockId="1.[786,1377,2496,2915]" pageId="1" pageNumber="416">
<materialsCitation id="3B503CE75C3F2A52F34600F63B64F5B2" ID-GBIF-Occurrence="3747872302" pageId="1" pageNumber="416">
“Tumbez un noérdlichen
<collectingCountry id="F32F762A5C3F2A52F4D200F63A42F5C4" box="[1252,1318,2535,2560]" name="Peru" pageId="1" pageNumber="416">Peru</collectingCountry>
, an der Grenze von
<collectingCountry id="F32F762A5C3F2A52F3C3031B3B0AF5E3" box="[1013,1134,2570,2599]" name="Ecuador" pageId="1" pageNumber="416">Ecuador</collectingCountry>
.” Restricted by A. Cabrera in 1958 to
<collectingRegion id="49FCF8585C3F2A52F41D03203BFFF58A" box="[1067,1179,2609,2638]" country="Peru" name="Tumbes" pageId="1" pageNumber="416">Tumbes</collectingRegion>
, departamento
<collectingRegion id="49FCF8585C3F2A52F30E03483CCCF5B2" box="[824,936,2649,2678]" country="Peru" name="Tumbes" pageId="1" pageNumber="416">Tumbes</collectingRegion>
,
<collectingCountry id="F32F762A5C3F2A52F38C03483C98F5B2" box="[954,1020,2649,2678]" name="Peru" pageId="1" pageNumber="416">Peru</collectingCountry>
.
</materialsCitation>
</paragraph>
<paragraph id="8B8736BA5C3F2A52F32503913B19F559" blockId="1.[786,1377,2496,2915]" box="[787,1149,2688,2717]" pageId="1" pageNumber="416">This species is monotypic.</paragraph>
</subSubSection>
<subSubSection id="C32265315C3F2A52F32503B93A63F4D0" pageId="1" pageNumber="416" type="distribution">
<caption id="DF4766325C3F2A52F32503B93A63F4D0" ID-DOI="http://doi.org/10.5281/zenodo.5733448" ID-Zenodo-Dep="5733448" httpUri="https://zenodo.org/record/5733448/files/figure.png" inLine="true" pageId="1" pageNumber="416" targetBox="[171,763,2496,2911]" targetPageId="1">
<paragraph id="8B8736BA5C3F2A52F32503B93A63F4D0" blockId="1.[786,1377,2496,2915]" pageId="1" pageNumber="416">
<emphasis id="B94CEAA85C3F2A52F32503B93CA7F501" bold="true" box="[787,963,2728,2757]" pageId="1" pageNumber="416">Distribution.</emphasis>
Known only from a narrow strip Wofthe Andes in C &amp; S
<collectingCountry id="F32F762A5C3F2A52F48403DE3A47F528" box="[1202,1315,2767,2796]" name="Ecuador" pageId="1" pageNumber="416">Ecuador</collectingCountry>
(including Puna I),
<collectingCountry id="F32F762A5C3F2A52F43203E63B22F4D0" box="[1028,1094,2807,2836]" name="Peru" pageId="1" pageNumber="416">Peru</collectingCountry>
, and N
<collectingCountry id="F32F762A5C3F2A52F48103E63A67F4D0" box="[1207,1283,2807,2836]" name="Chile" pageId="1" pageNumber="416">Chile</collectingCountry>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="C32265315C3F2A52F322020F3D01F249" pageId="1" pageNumber="416" type="description">
<paragraph id="8B8736BA5C3F2A52F322020F3D01F249" blockId="1.[786,1377,2496,2915]" lastBlockId="1.[173,1378,2926,3469]" pageId="1" pageNumber="416">
<emphasis id="B94CEAA85C3F2A52F322020F3B56F4FF" bold="true" box="[788,1074,2846,2875]" pageId="1" pageNumber="416">Descriptive notes.</emphasis>
Head-body 39-47 mm, tail 27-34 mm, ear 12-15 mm, hindfoot 6-10 mm, forearm 34-38 mm; weight 3-10 g. Tail extends nearly fourfifths the length of uropatagium andis entirely enclosed by that membrane. Height of braincase, including auditory bullae, is equal to distance from frontal angle to most posterior point of occipital region. Muzzleis relatively simple and well-furred; no fleshystructures occur under chin; palateis short, not extending much beyond last molar; and mesopterygoid fossa is longer than wide. The Smoky Bat has long, dark smokygrayfur, often with brownish tinge; hair tips are blackish; front and top of head have pale buffy suffusion; underparts are nearly the same color as the upperparts; ears are light brown; and wing and tail membranes arelittle darker than ears. Snout is prominently rimmedand blunt; lips are conspicuouslyfleshy; and chin excrescences and lowerlips havefleshy wart-like structures. On average, third metacarpal is
<quantity id="4CC09B5F5C3F2A52F10704313EEAF2F9" box="[305,398,3360,3389]" metricMagnitude="-2" metricUnit="m" metricValue="3.4" pageId="1" pageNumber="416" unit="mm" value="34.0">34 mm</quantity>
, tibia is 15-5 mm, total skull length is
<quantity id="4CC09B5F5C3F2A52F3FA04313B48F2F9" box="[972,1068,3360,3389]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="416" unit="mm" value="12.0">12 mm</quantity>
, braincase breadthis
<quantity id="4CC09B5F5C3F2A52F09904593E66F2A1" box="[175,258,3400,3429]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="1" pageNumber="416" unit="mm" value="6.0">6 mm</quantity>
, and uppertooth rowlength (with canine) is
<quantity id="4CC09B5F5C3F2A52F39504593C9CF2A1" box="[931,1016,3400,3429]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="416" unit="mm" value="5.0">5 mm</quantity>
. Dental formula is I 2/3, Cl1/1,P2/3.M 3/3 (x2) =36.
</paragraph>
</subSubSection>
<subSubSection id="C32265315C3F2A52F59E016C3952F6A4" pageId="1" pageNumber="416" type="biology_ecology">
<paragraph id="8B8736BA5C3F2A52F59E016C3952F6A4" blockId="1.[1443,2657,2173,3469]" pageId="1" pageNumber="416">
<emphasis id="B94CEAA85C3F2A52F59E016C397DF75E" bold="true" box="[1448,1561,2173,2202]" pageId="1" pageNumber="416">Habitat.</emphasis>
South American west coast deserts from sea level up to elevations of
<quantity id="4CC09B5F5C3F2A52F9DB016C3532F75E" box="[2541,2646,2173,2202]" metricMagnitude="3" metricUnit="m" metricValue="2.68" pageId="1" pageNumber="416" unit="m" value="2680.0">2680 m</quantity>
. Two Smoky Bats were found on Hacienda Limon,
<collectingRegion id="49FCF8585C3F2A52F8BC01B43646F706" box="[2186,2338,2213,2242]" country="Peru" name="Cajamarca" pageId="1" pageNumber="416">Cajamarca</collectingRegion>
,
<collectingCountry id="F32F762A5C3F2A52F90001B4361EF706" box="[2358,2426,2213,2242]" name="Peru" pageId="1" pageNumber="416">Peru</collectingCountry>
, in a small arid valley above the Rio Maranon in the Amazon Basin. It is found in arid regions and cultivated areas; it is a cave-dweller and known to roost in culverts, caves, sugar mills, irrigation tunnels, and abandoned urban-rural buildings near crop areas (banana, rice; etc.).
</paragraph>
</subSubSection>
<subSubSection id="C32265315C3F2A52F59F007B36E0F66A" pageId="1" pageNumber="416" type="food_feeding">
<paragraph id="8B8736BA5C3F2A52F59F007B36E0F66A" blockId="1.[1443,2657,2173,3469]" pageId="1" pageNumber="416">
<emphasis id="B94CEAA85C3F2A52F59F007B39D5F643" bold="true" box="[1449,1713,2410,2439]" pageId="1" pageNumber="416">Food and Feeding.</emphasis>
Smoky Bats eat moths and butterflies (Lepidoptera). Adult moth scales were found in stomachs offive specimens from central
<collectingCountry id="F32F762A5C3F2A52F93D008036E4F66A" box="[2315,2432,2449,2478]" name="Ecuador" pageId="1" pageNumber="416">Ecuador</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C32265315C3F2A52F59F00A8372CF4FD" pageId="1" pageNumber="416" type="breeding">
<paragraph id="8B8736BA5C3F2A52F59F00A8372CF4FD" blockId="1.[1443,2657,2173,3469]" pageId="1" pageNumber="416">
<emphasis id="B94CEAA85C3F2A52F59F00A83952F612" bold="true" box="[1449,1590,2489,2518]" pageId="1" pageNumber="416">Breeding.</emphasis>
In central
<collectingCountry id="F32F762A5C3F2A52F6EB00A8383DF612" box="[1757,1881,2489,2518]" name="Ecuador" pageId="1" pageNumber="416">Ecuador</collectingCountry>
in middle November, a large colony included males that did not show testes in the scrotal position, and several females were pregnant (eight of ten) with single fetuses but not lactating. This high percentage of pregnant females suggested a relatively synchronized seasonal breeding period, undoubtedly related to the rainy season (January-May); similarly, a breeding colony was found in the rainy months (December-March) in southern
<collectingCountry id="F32F762A5C3F2A52F9D20393354FF55F" box="[2532,2603,2690,2715]" name="Peru" pageId="1" pageNumber="416">Peru</collectingCountry>
. In a single reproductively inactive male, the small, paired testes were attached to the posterior body wall and inferior pole of the kidney by a suspensoryligament. The Smoky Bat has an ampullary gland. One specimen appeared to contain a tiny baculum, capping the urethra at tip ofglans.
</paragraph>
</subSubSection>
<subSubSection id="C32265315C3F2A52F59E02523639F46B" pageId="1" pageNumber="416" type="activity">
<paragraph id="8B8736BA5C3F2A52F59E02523639F46B" blockId="1.[1443,2657,2173,3469]" pageId="1" pageNumber="416">
<emphasis id="B94CEAA85C3F2A52F59E025239F6F4A4" bold="true" box="[1448,1682,2883,2912]" pageId="1" pageNumber="416">Activity patterns.</emphasis>
Echolocation calls of the SmokyBat are the FMtype, with an average duration of4-4 milliseconds, important for foraging in dense vegetation and taking small insects while flying. Maximum averagefrequencyat 75-8 kHz.
</paragraph>
</subSubSection>
<subSubSection id="C32265315C3F2A52F59C02A43625F389" pageId="1" pageNumber="416" type="biology_ecology">
<paragraph id="8B8736BA5C3F2A52F59C02A43625F389" blockId="1.[1443,2657,2173,3469]" pageId="1" pageNumber="416">
<emphasis id="B94CEAA85C3F2A52F59C02A4370EF412" bold="true" box="[1450,2154,2997,3030]" pageId="1" pageNumber="416">Movements, Home range and Social organization.</emphasis>
In
<collectingRegion id="49FCF8585C3F2A52F89602A43667F412" box="[2208,2307,2997,3030]" country="Ecuador" name="Guayas" pageId="1" pageNumber="416">Guayas</collectingRegion>
,
<collectingCountry id="F32F762A5C3F2A52F92002A436EEF412" box="[2326,2442,2997,3030]" name="Ecuador" pageId="1" pageNumber="416">Ecuador</collectingCountry>
, a large monospecific colony(c.300 individuals) of Smoky Bats included males and females but not juveniles. This bat species has been collected along with Long-snouted Bat (
<taxonomicName id="4C384D395C3F2A52F9D00519392AF389" authorityName="Thomas" authorityYear="1928" class="Mammalia" family="Phyllostomidae" genus="Platalina" kingdom="Animalia" order="Chiroptera" pageId="1" pageNumber="416" phylum="Chordata" rank="species" species="genovensium">Platalina genovensium</taxonomicName>
) and Small Big-eared Brown Bat (
<taxonomicName id="4C384D395C3F2A52F807052137C2F389" authorityName="Gervais" authorityYear="1856" box="[2097,2214,3120,3149]" class="Mammalia" family="Vespertilionidae" genus="Histiotus" kingdom="Animalia" order="Chiroptera" pageId="1" pageNumber="416" phylum="Chordata" rank="genus">Histiotus</taxonomicName>
montanus).
</paragraph>
</subSubSection>
<subSubSection id="C32265315C3F2A52F59C05463892F2D7" pageId="1" pageNumber="416" type="conservation">
<paragraph id="8B8736BA5C3F2A52F59C05463892F2D7" blockId="1.[1443,2657,2173,3469]" pageId="1" pageNumber="416">
<emphasis id="B94CEAA85C3F2A52F59C05463869F3B0" bold="true" box="[1450,1805,3159,3188]" pageId="1" pageNumber="416">Status and Conservation.</emphasis>
Classified as Vulnerable on The IUCNRed List. The Smoky Bat is considered an endangered species in
<collectingCountry id="F32F762A5C3F2A52F82B056F37F7F35F" box="[2077,2195,3198,3227]" name="Ecuador" pageId="1" pageNumber="416">Ecuador</collectingCountry>
because known populations are small, restricted to areas smaller than
<quantity id="4CC09B5F5C3F2A52F78805B73770F307" box="[1982,2068,3238,3267]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="416" unit="km" value="20.0">20 km</quantity>
?, and known fromless than five localities. In
<collectingCountry id="F32F762A5C3F2A52F5E505C23969F328" box="[1491,1549,3283,3308]" name="Peru" pageId="1" pageNumber="416">Peru</collectingCountry>
, it is considered a vulnerable species. In
<collectingCountry id="F32F762A5C3F2A52F87B05C237FDF328" box="[2125,2201,3283,3308]" name="Chile" pageId="1" pageNumber="416">Chile</collectingCountry>
, it has not been evaluated, but it is thought to be beneficial to agroforestry.
</paragraph>
</subSubSection>
<subSubSection id="C32265315C3F2A52F59D0434399BF24E" pageId="1" pageNumber="416" type="bibRefCitation_list">
<paragraph id="8B8736BA5C3F2A52F59D0434399BF24E" blockId="1.[1443,2657,2173,3469]" pageId="1" pageNumber="416">
<emphasis id="B94CEAA85C3F2A52F59D04343927F2FE" bold="true" box="[1451,1603,3365,3386]" pageId="1" pageNumber="416">Bibliography.</emphasis>
Abuja (1999), Aragon &amp; Aguirre (2014), Cabrera (1958), Gardner (2008h), Ibanez (1986), Krutzsch (2000), Miller (1907), Morgan &amp; Czaplewski (1999), Nowak (1994), Rodriguez-San Pedro et al. (2016), Tirira et al. (2012), Ugarte-Nunez (2014).
</paragraph>
</subSubSection>
</treatment>
</document>