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<document id="75D5ADB500F2A7579A031C188497298B" ID-DOI="10.11646/zootaxa.2096.1.27" ID-ISSN="1175-5326" ID-Zenodo-Dep="5321990" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629621422523" checkinUser="felipe" docAuthor="Strugnell, Jan, Voight, Janet R., Collins, Patrick C. &amp; Allcock, A. Louise" docDate="2009" docId="03914A02FF959C12FF6692D3FEAEFA1D" docLanguage="en" docName="zt02096p459.pdf" docOrigin="Zootaxa 2096 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2096.1.27" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Benthoctopus rigbyae Vecchione, Allcock, Piatkowski, Strugnell 2009" docType="treatment" docVersion="4" lastPageNumber="449" masterDocId="FFA8327AFF929C15FFF1914AFFD0FFD1" masterDocTitle="Molecular phylogenetic analysis of a known and a new hydrothermal vent octopod: their relationships with the genus Benthoctopus (Cephalopoda: Octopodidae)" masterLastPageNumber="459" masterPageNumber="442" pageNumber="449" updateTime="1699084934570" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="65C0845FB1D20BE4058897041616EE8D" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="F07A865D0855C30ADA810DD2A169F971">
<mods:title id="6A32C41810BB996F5782CBFB6BCFCC39">Molecular phylogenetic analysis of a known and a new hydrothermal vent octopod: their relationships with the genus Benthoctopus (Cephalopoda: Octopodidae)</mods:title>
</mods:titleInfo>
<mods:name id="76283540832511F2E0EAE5C7F8BA4561" type="personal">
<mods:role id="199C1869D5381CEDC702EDC8862D3B8B">
<mods:roleTerm id="860B21407A515814796B43A83E6F0198">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="49815D5CE74813A9B9392265E7AECB25">Strugnell, Jan</mods:namePart>
<mods:affiliation id="E30FF5E2EFB7622A5E727661AF34359C">Department of Zoology, University of Cambridge, Downing Street, Cambridge CB 2 3 EJ, UK School of Biological Sciences, Queen's University Belfast, 97 Lisburn Road, Belfast BT 9 7 BL, UK Department of Zoology, Field Museum of Natural History, 1400 S. Lake Shore Dr, Chicago, IL 60605 - 2496, USA Duke Marine Laboratory, 135 Duke Marine Lab Road, Beaufort, NC 28516, USA The Martin Ryan Marine Science Institute, National University of Ireland, Galway, University Road, Galway, Ireland Corresponding author: jan. strugnell @ gmail. com</mods:affiliation>
</mods:name>
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<mods:role id="CAF9B08F6A3CB70FA6FDA8E453524A24">
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</mods:role>
<mods:namePart id="642EB070F9BDC67AA2DB17DCE29CE487">Voight, Janet R.</mods:namePart>
</mods:name>
<mods:name id="D65982B3574608151298B023879C1274" type="personal">
<mods:role id="773BB9ED4FCC6DF289E1513CB669B8A8">
<mods:roleTerm id="3E307EDBAAB2EB0AA6A8CB6D52C5C403">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C9031ED6CA44ED0CA315BEB976A6FA75">Collins, Patrick C.</mods:namePart>
</mods:name>
<mods:name id="099161EEF813F787DB6E3BC479CE274F" type="personal">
<mods:role id="AAAA7C2F8CB5A1ECC8C7FC216EFE57D7">
<mods:roleTerm id="E1FF7D1D66AC285478D80325AE44431E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9431BE9905A227096EA9BAA0B915A438">Allcock, A. Louise</mods:namePart>
</mods:name>
<mods:typeOfResource id="873AFABC684B4935E305E92DE23D4257">text</mods:typeOfResource>
<mods:relatedItem id="DEC85E677C8AAB358ECE31DEA8CF9E7B" type="host">
<mods:titleInfo id="454D24878AAA717984A8278DA295C29D">
<mods:title id="F3CF045485D79CEB22A5600827BB7A8D">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="41C7030B951B7647CFD65DE5AEB352FF">
<mods:date id="859007DE798F01E00CEB99BBEC811765">2009</mods:date>
<mods:detail id="D535E730AF88CDB25A536DFF5AF12B9D" type="pubDate">
<mods:number id="A6AC784149E513D399842AD2BB710A95">2009-05-11</mods:number>
</mods:detail>
<mods:detail id="65C0AEEC8FD09B8ADA4A21718A88675E" type="volume">
<mods:number id="D53AA11C5697FBACF858091C1D1EEA5E">2096</mods:number>
</mods:detail>
<mods:detail id="780ED614EA6CE100087CB9A44E1C489C" type="issue">
<mods:number id="4E9C3BB51A28C3D45819D9F138117F41">1</mods:number>
</mods:detail>
<mods:extent id="EB4CA456A37207FBACE4CB3873D951E1" unit="page">
<mods:start id="4108BDE5CDE0DAD885FDCBABBA8DC61C">442</mods:start>
<mods:end id="957109567CE543415AD105D6961F3BA3">459</mods:end>
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<mods:url id="B4B9F7EA3F5E7C5BA16BC96075C71000">https://biotaxa.org/Zootaxa/article/view/zootaxa.2096.1.27</mods:url>
</mods:location>
<mods:classification id="1F10703F64E8ADF562D5A46FE90EAB32">journal article</mods:classification>
<mods:identifier id="EB4851829A03C840B5BBA984ACFEF583" type="DOI">10.11646/zootaxa.2096.1.27</mods:identifier>
<mods:identifier id="5CD93427BCA76B334BDEA510F550B711" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="BA0CB2AF36DC9BC4DB1640B607AD71EB" type="Zenodo-Dep">5321990</mods:identifier>
</mods:mods>
<treatment id="03914A02FF959C12FF6692D3FEAEFA1D" ID-DOI="http://doi.org/10.5281/zenodo.5334612" ID-Zenodo-Dep="5334612" LSID="urn:lsid:plazi:treatment:03914A02FF959C12FF6692D3FEAEFA1D" httpUri="http://treatment.plazi.org/id/03914A02FF959C12FF6692D3FEAEFA1D" lastPageNumber="449" pageId="7" pageNumber="449">
<subSubSection id="C322A89FFF959C12FF6692D3FC26FC63" box="[151,1014,920,947]" pageId="7" pageNumber="449" type="nomenclature">
<paragraph id="8B87FB14FF959C12FF6692D3FC26FC63" blockId="7.[151,1436,920,1484]" box="[151,1014,920,947]" pageId="7" pageNumber="449">
<heading id="D0CF4C78FF959C12FF6692D3FC26FC63" bold="true" box="[151,1014,920,947]" fontSize="11" level="1" pageId="7" pageNumber="449" reason="1">
<taxonomicName id="4C388097FF959C12FF6692D3FC26FC63" authority="Vecchione, Allcock, Piatkowski, Strugnell, 2009" authorityName="Vecchione, Allcock, Piatkowski, Strugnell" authorityYear="2009" box="[151,1014,920,947]" class="Cephalopoda" family="Octopodidae" genus="Benthoctopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Octopoda" pageId="7" pageNumber="449" phylum="Mollusca" rank="species" species="rigbyae">
<emphasis id="B94C2706FF959C12FF6692D3FC26FC63" bold="true" box="[151,1014,920,947]" pageId="7" pageNumber="449">
<emphasis id="B94C2706FF959C12FF6692D3FE4DFC62" bold="true" box="[151,413,921,947]" italics="true" pageId="7" pageNumber="449">Benthoctopus rigbyae</emphasis>
<bibRefCitation id="EFA986E5FF959C12FE5492D2FC26FC63" author="Vecchione, Allcock, Piatkowski, Strugnell" box="[421,1014,920,946]" firstAuthor="Vecchione" pageId="7" pageNumber="449" pagination="13 - 28" refId="ref11818" refString="Vecchione, M., Allcock, L., Piatkowski, U. &amp; Strugnell, J. (2009) Benthoctopus rigbyae n. sp., a new species of cephalopod (Octopoda; Incirrata) from near the Antarctic Peninsula. Malacologia, 51, 13 - 28." type="journal article" year="2009">Vecchione, Allcock, Piatkowski, Strugnell, 2009</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C322A89FFF959C12FF6692F6FEAEFA1D" pageId="7" pageNumber="449" type="materials_examined">
<paragraph id="8B87FB14FF959C12FF6692F6FC6FFB9B" blockId="7.[151,1436,920,1484]" pageId="7" pageNumber="449">
<emphasis id="B94C2706FF959C12FF6692F6FE69FC04" box="[151,441,956,981]" italics="true" pageId="7" pageNumber="449">Type material examined:</emphasis>
<taxonomicName id="4C388097FF959C12FE3092F6FAFEFC07" authority="Vecchione, Allcock, Piatkowski, Strugnell, 2009: 18" authorityName="Vecchione, Allcock, Piatkowski, Strugnell" authorityPageNumber="18" authorityYear="2009" box="[449,1326,956,982]" class="Cephalopoda" family="Octopodidae" genus="Benthoctopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Octopoda" pageId="7" pageNumber="449" phylum="Mollusca" rank="species" species="rigbyae">
<emphasis id="B94C2706FF959C12FE3092F6FD11FC04" box="[449,705,956,981]" italics="true" pageId="7" pageNumber="449">Benthoctopus rigbyae</emphasis>
<bibRefCitation id="EFA986E5FF959C12FD3992F6FAFEFC07" author="Vecchione, Allcock, Piatkowski, Strugnell" box="[712,1326,956,982]" firstAuthor="Vecchione" page="18" pageId="7" pageNumber="449" pagination="13 - 28" refId="ref11818" refString="Vecchione, M., Allcock, L., Piatkowski, U. &amp; Strugnell, J. (2009) Benthoctopus rigbyae n. sp., a new species of cephalopod (Octopoda; Incirrata) from near the Antarctic Peninsula. Malacologia, 51, 13 - 28." type="journal article" year="2009">
Vecchione, Allcock, Piatkowski, Strugnell, 2009:
<specimenCount id="9D3E309DFF959C12FAE092F6FAFEFC07" box="[1297,1326,956,982]" pageId="7" pageNumber="449" type="generic" typeStatus="holotype">18</specimenCount>
</bibRefCitation>
</taxonomicName>
holotype
<collectionCode id="ED2963D1FF959C12FF6692A8FF3CFC2D" box="[151,236,994,1020]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="7" pageNumber="449" type="Museum">USNM</collectionCode>
1117765,
<emphasis id="B94C2706FF959C12FE9692A8FE5AFC2A" box="[359,394,994,1019]" italics="true" pageId="7" pageNumber="449">
<collectionCode id="ED2963D1FF959C12FE9692A8FE5AFC2A" box="[359,394,994,1019]" country="Italy" name="Collection of Leptospira Strains" pageId="7" pageNumber="449">RV</collectionCode>
</emphasis>
Polarstern,
<geoCoordinate id="EE0C9DD3FF959C12FDE992A8FDABFC2D" box="[536,635,994,1020]" degrees="61" direction="south" minutes="14" orientation="latitude" pageId="7" pageNumber="449" precision="925" value="-61.233334">61° 14'S</geoCoordinate>
<geoCoordinate id="EE0C9DD3FF959C12FD7592A8FCC0FC2D" box="[644,784,994,1020]" degrees="56" direction="west" minutes="25.8" orientation="longitude" pageId="7" pageNumber="449" precision="92" value="-56.43">56° 25.8' W</geoCoordinate>
(Antarctic Peninsula), Stn 42/022,
<date id="FF86DDD4FF959C12FB4092A8FA47FC2D" box="[1201,1431,994,1020]" pageId="7" pageNumber="449" value="1996-11-21">21 November 1996</date>
, 394-
<quantity id="4CC056F1FF959C12FF3D9543FECDFBF2" box="[204,285,1033,1059]" metricMagnitude="2" metricUnit="m" metricValue="4.12" pageId="7" pageNumber="449" unit="m" value="412.0">412 m</quantity>
, male.
<emphasis id="B94C2706FF959C12FE849543FE09FBF3" box="[373,473,1033,1058]" italics="true" pageId="7" pageNumber="449">Octopus</emphasis>
<taxonomicName id="4C388097FF959C12FE139543FD23FBF2" authority="Hoyle, 1885: 220" authorityName="Hoyle" authorityPageNumber="220" authorityYear="1885" box="[482,755,1033,1059]" class="Cephalopoda" family="Octopodidae" genus="Benthoctopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Octopoda" pageId="7" pageNumber="449" phylum="Mollusca" rank="species" species="levis">
<emphasis id="B94C2706FF959C12FE139543FDC9FBF3" box="[482,537,1033,1058]" italics="true" pageId="7" pageNumber="449">levis</emphasis>
<bibRefCitation id="EFA986E5FF959C12FDD09543FD23FBF2" author="Hoyle" box="[545,755,1033,1059]" firstAuthor="Hoyle" page="220" pageId="7" pageNumber="449" pagination="222 - 236" refId="ref10924" refString="Hoyle, W. E. (1885) Diagnoses of new species of Cephalopoda collected during the cruise of H. M. S. ' Challenger' - Part I. The Octopoda. Annals and Magazine of Natural History, Series 5, 15, 222 - 236." type="journal article" year="1885">
Hoyle, 1885:
<specimenCount id="9D3E309DFF959C12FD369543FD23FBF2" box="[711,755,1033,1059]" pageId="7" pageNumber="449" type="generic" typeStatus="holotype">220</specimenCount>
</bibRefCitation>
</taxonomicName>
holotype
<collectionCode id="ED2963D1FF959C12FC9F9543FC17FBF2" box="[878,967,1033,1059]" pageId="7" pageNumber="449">BMNH</collectionCode>
1889.4.24.43,
<emphasis id="B94C2706FF959C12FB8F9543FB72FBF3" box="[1150,1186,1033,1058]" italics="true" pageId="7" pageNumber="449">
<collectionCode id="ED2963D1FF959C12FB8F9543FB72FBF3" box="[1150,1186,1033,1058]" country="Italy" name="Collection of Leptospira Strains" pageId="7" pageNumber="449">RV</collectionCode>
</emphasis>
Challenger,
<geoCoordinate id="EE0C9DD3FF959C12FACF9543FA4CFBF2" box="[1342,1436,1032,1059]" degrees="52" direction="south" minutes="59" orientation="latitude" pageId="7" pageNumber="449" precision="925" value="-52.983334">52°59'S</geoCoordinate>
<geoCoordinate id="EE0C9DD3FF959C12FF66957AFF24FB9B" box="[151,244,1071,1098]" degrees="73" direction="east" minutes="33" orientation="longitude" pageId="7" pageNumber="449" precision="925" value="73.55">73°33'E</geoCoordinate>
(
<collectingCountry id="F32FBB84FF959C12FEF2957AFE70FB9B" box="[259,416,1072,1098]" name="Heard Island and McDonald Islands" pageId="7" pageNumber="449">Heard Island</collectingCountry>
), Stn 151,
<date id="FF86DDD4FF959C12FDEC957AFD31FB9B" box="[541,737,1072,1098]" pageId="7" pageNumber="449" value="1874-02-07">7 February 1874</date>
, 75 fathoms, male.
</paragraph>
<paragraph id="8B87FB14FF959C12FF37951DFBDCFB35" blockId="7.[151,1436,920,1484]" pageId="7" pageNumber="449">
<emphasis id="B94C2706FF959C12FF37951DFC85FBA1" box="[198,853,1111,1136]" italics="true" pageId="7" pageNumber="449">
Additional material examined:
<taxonomicName id="4C388097FF959C12FDB4951DFC9AFBA1" authorityName="Vecchione, Allcock, Piatkowski, Strugnell" authorityYear="2009" box="[581,842,1111,1136]" class="Cephalopoda" family="Octopodidae" genus="Benthoctopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Octopoda" pageId="7" pageNumber="449" phylum="Mollusca" rank="species" species="rigbyae">Benthoctopus rigbyae</taxonomicName>
:
</emphasis>
*
<materialsCitation id="3B50F149FF959C12FC9C951CFBB8FBA1" box="[877,1128,1110,1136]" collectionCode="NMSZ" pageId="7" pageNumber="449" specimenCode="NMSZ 2002037.032" specimenCount="1">
<specimenCode id="DB9E536FFF959C12FC9C951CFBB8FBA1" box="[877,1128,1110,1136]" collectionCode="NMSZ" pageId="7" pageNumber="449">NMSZ 2002037.032</specimenCode>
</materialsCitation>
,
<materialsCitation id="3B50F149FF959C12FB8D951CFEA3FB35" collectingDate="2002-03-16" collectionCode="RV" collectorName="Antarctic Peninsula" county="Stn" determinerName="AL Allcock." latitude="-61.166668" location="Polarstern" longLatPrecision="1027" longitude="-54.566666" municipality="Antarctic Peninsula" pageId="7" pageNumber="449" specimenCount="1">
<collectionCode id="ED2963D1FF959C12FB8D951CFB75FBA1" box="[1148,1189,1110,1136]" country="Italy" name="Collection of Leptospira Strains" pageId="7" pageNumber="449">RV</collectionCode>
<emphasis id="B94C2706FF959C12FB5E951DFAFCFBA1" box="[1199,1324,1111,1136]" italics="true" pageId="7" pageNumber="449">
<location id="8EE7ADCFFF959C12FB5E951DFAFCFBA1" LSID="urn:lsid:plazi:treatment:03914A02FF959C12FF6692D3FEAEFA1D:8EE7ADCFFF959C12FB5E951DFAFCFBA1" box="[1199,1324,1111,1136]" county="Stn" latitude="-61.166668" longLatPrecision="1027" longitude="-54.566666" municipality="Antarctic Peninsula" name="Polarstern" pageId="7" pageNumber="449">Polarstern</location>
</emphasis>
,
<geoCoordinate id="EE0C9DD3FF959C12FAB1951CFA4CFBA1" box="[1344,1436,1110,1136]" degrees="61" direction="south" minutes="10" orientation="latitude" pageId="7" pageNumber="449" precision="925" value="-61.166668">61°10'S</geoCoordinate>
<geoCoordinate id="EE0C9DD3FF959C12FF669537FED1FB46" box="[151,257,1148,1175]" degrees="54" direction="west" minutes="34" orientation="longitude" pageId="7" pageNumber="449" precision="925" value="-54.566666">54°34'W</geoCoordinate>
(
<collectingMunicipality id="6BE3616EFF959C12FEE29537FDDDFB46" box="[275,525,1149,1175]" pageId="7" pageNumber="449">Antarctic Peninsula</collectingMunicipality>
),
<collectingCounty id="62E68398FF959C12FDD59537FD9CFB46" box="[548,588,1149,1175]" pageId="7" pageNumber="449">Stn</collectingCounty>
61/048-1,
<date id="FF86DDD4FF959C12FD249537FC44FB46" box="[725,916,1149,1175]" pageId="7" pageNumber="449" value="2002-03-16">
<collectingDate id="EFC2243CFF959C12FD249537FC44FB46" box="[725,916,1149,1175]" pageId="7" pageNumber="449" value="2002-03-16">
16 March
<quantity id="4CC056F1FF959C12FCA59537FC44FB46" box="[852,916,1149,1175]" metricMagnitude="3" metricUnit="m" metricValue="2.002343" pageId="7" pageNumber="449" unit="m" value="2002.343">2002</quantity>
</collectingDate>
</date>
, 343 m, male, leg. &amp; det.
<determinerName id="7E5B17D0FF959C12FB2A9537FAA5FB46" box="[1243,1397,1149,1175]" pageId="7" pageNumber="449">AL Allcock.</determinerName>
10 additional specimens from the
<collectorName id="26CD9EC2FF959C12FDEB95EEFCDFFB6F" box="[538,783,1188,1214]" pageId="7" pageNumber="449">Antarctic Peninsula</collectorName>
detailed in
<bibRefCitation id="EFA986E5FF959C12FC5795EEFB1BFB6F" author="Vecchione" box="[934,1227,1188,1214]" etAl="et al." firstAuthor="Vecchione" pageId="7" pageNumber="449" pagination="13 - 28" refId="ref11818" refString="Vecchione, M., Allcock, L., Piatkowski, U. &amp; Strugnell, J. (2009) Benthoctopus rigbyae n. sp., a new species of cephalopod (Octopoda; Incirrata) from near the Antarctic Peninsula. Malacologia, 51, 13 - 28." type="journal article" year="2009">
Vecchione
<emphasis id="B94C2706FF959C12FBC195EEFBBFFB6C" box="[1072,1135,1188,1213]" italics="true" pageId="7" pageNumber="449">et al.</emphasis>
(2009)
</bibRefCitation>
preserved under catalogue numbers
</materialsCitation>
<materialsCitation id="3B50F149FF959C12FE8B9580FDA1FB35" box="[378,625,1226,1252]" collectionCode="NMSZ" pageId="7" pageNumber="449" specimenCode="NMSZ 2002037.030" specimenCount="1">
<specimenCode id="DB9E536FFF959C12FE8B9580FDA1FB35" box="[378,625,1226,1252]" collectionCode="NMSZ" pageId="7" pageNumber="449">NMSZ 2002037.030</specimenCode>
</materialsCitation>
-
<materialsCitation id="3B50F149FF959C12FD889580FD04FB35" box="[633,724,1226,1252]" collectionCode="NMSZ" isTailEnumeration="true" pageId="7" pageNumber="449" specimenCode="031" specimenCount="1">
<specimenCode id="DB9E536FFF959C12FD889580FD73FB35" box="[633,675,1226,1252]" collectionCode="NMSZ" pageId="7" pageNumber="449">031</specimenCode>
and
</materialsCitation>
<materialsCitation id="3B50F149FF959C12FD2D9580FC04FB35" box="[732,980,1226,1252]" collectionCode="NMSZ" pageId="7" pageNumber="449" specimenCode="NMSZ 2002037.033" specimenCount="1">
<specimenCode id="DB9E536FFF959C12FD2D9580FC04FB35" box="[732,980,1226,1252]" collectionCode="NMSZ" pageId="7" pageNumber="449">NMSZ 2002037.033</specimenCode>
</materialsCitation>
-
<materialsCitation id="3B50F149FF959C12FC2A9580FBD8FB35" box="[987,1032,1226,1252]" collectionCode="NMSZ" isTailEnumeration="true" pageId="7" pageNumber="449" specimenCode="034" specimenCount="1">
<specimenCode id="DB9E536FFF959C12FC2A9580FBD8FB35" box="[987,1032,1226,1252]" collectionCode="NMSZ" pageId="7" pageNumber="449">034</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8B87FB14FF959C12FF3795BBFD65FAAE" blockId="7.[151,1436,920,1484]" pageId="7" pageNumber="449">
<emphasis id="B94C2706FF959C12FF3795BBFEEDFADB" box="[198,317,1265,1290]" italics="true" pageId="7" pageNumber="449">Diagnosis</emphasis>
. Arms approximately 3–4 times
<collectionCode id="ED2963D1FF959C12FD4D95BBFD3CFADA" box="[700,748,1265,1291]" country="France" name="Musee de Lectoure" pageId="7" pageNumber="449">ML</collectionCode>
. Arm formula variable. Funnel organ W-shaped. Gills with 5–8 lamellae per inner demibranch and 6–8 lamellae per outer demibranch. LLI 6–16 (mean 10). Maximum spermatophore length 102, maximum SpLI 110, maximum number of spermatophores 28. Mature males without enlarged suckers. Integument smooth.
</paragraph>
<paragraph id="8B87FB14FF959C12FF3794C6FEAEFA1D" blockId="7.[151,1436,920,1484]" pageId="7" pageNumber="449">
<emphasis id="B94C2706FF959C12FF3794C6FEE5FA74" box="[198,309,1420,1445]" italics="true" pageId="7" pageNumber="449">Remarks.</emphasis>
This species differs from
<taxonomicName id="4C388097FF959C12FD9294C6FCEFFA74" authorityName="Hoyle" authorityYear="1885" box="[611,831,1420,1445]" class="Cephalopoda" family="Octopodidae" genus="Benthoctopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Octopoda" pageId="7" pageNumber="449" phylum="Mollusca" rank="species" species="levis">
<emphasis id="B94C2706FF959C12FD9294C6FCEFFA74" box="[611,831,1420,1445]" italics="true" pageId="7" pageNumber="449">Benthoctopus levis</emphasis>
</taxonomicName>
by the funnel organ (
<collectionCode id="ED2963D1FF959C12FBC694C6FBB3FA77" box="[1079,1123,1420,1446]" pageId="7" pageNumber="449">VV</collectionCode>
in
<taxonomicName id="4C388097FF959C12FB7694C6FB0DFA74" authorityName="Hoyle" authorityYear="1885" box="[1159,1245,1420,1445]" class="Cephalopoda" family="Octopodidae" genus="Benthoctopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Octopoda" pageId="7" pageNumber="449" phylum="Mollusca" rank="species" species="levis">
<emphasis id="B94C2706FF959C12FB7694C6FB0DFA74" box="[1159,1245,1420,1445]" italics="true" pageId="7" pageNumber="449">B. levis</emphasis>
</taxonomicName>
) and arm length (shorter in
<taxonomicName id="4C388097FF959C12FEE694F9FEBDFA1D" authorityName="Hoyle" authorityYear="1885" box="[279,365,1459,1484]" class="Cephalopoda" family="Octopodidae" genus="Benthoctopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Octopoda" pageId="7" pageNumber="449" phylum="Mollusca" rank="species" species="levis">
<emphasis id="B94C2706FF959C12FEE694F9FEBDFA1D" box="[279,365,1459,1484]" italics="true" pageId="7" pageNumber="449">B. levis</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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