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<document id="1352A5A583BBDE73F54E68C86191B9D6" ID-CLB-Dataset="25923" ID-DOI="10.11646/zootaxa.4658.3.5" ID-GBIF-Dataset="d922f25d-1252-4379-8e0f-39124631c5d8" ID-ISSN="1175-5326" ID-Zenodo-Dep="3376104" ID-ZooBank="664EEF6E-ABD3-4DFB-B0B8-12AC082FE0E0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1566575951368" checkinUser="plazi" docAuthor="Groenen, Frans &amp; Wangdi, Karma" docDate="2019" docId="CF458781AD0A2C61FF06FD55FB13AA52" docLanguage="en" docName="zootaxa.4658.3.5.pdf" docOrigin="Zootaxa 4658 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Epiblema albulusana Groenen &amp; Wangdi 2019, sp. nov." docType="treatment" docUuid="656F7FD7-B133-46F9-878F-9E5DB1AC3E24" docUuidSource="ZooBank" docVersion="9" lastPageNumber="534" masterDocId="337CFFF9AD022C69FF91FF8CFFE6AF29" masterDocTitle="The Tortricidae (Lepidoptera) of Bhutan" masterLastPageNumber="540" masterPageNumber="526" pageNumber="534" updateTime="1698752418553" updateUser="ExternalLinkService">
<mods:mods id="796DDEB646CEB9D8192FCBCD0F7FAA58" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="F7FCB420A8F7177D976B1B797E260B09">
<mods:title id="878551518DA983FE884CE40A7D975146">The Tortricidae (Lepidoptera) of Bhutan</mods:title>
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<mods:role id="B16C3EADCCFE5B93B1804C71E53C685D">
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<mods:namePart id="3394392E0D6E3CDF9C96DD698E9F014D">Groenen, Frans</mods:namePart>
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<mods:namePart id="4EFC85246B926C81ACC0E01C96FD9E62">Wangdi, Karma</mods:namePart>
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<mods:title id="BBB0ED2CD0C86D9EB703128B78A8C171">Zootaxa</mods:title>
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<mods:date id="D9CD6ADBC2BDBD19339068E7B042BFE8">2019</mods:date>
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<mods:number id="81C31A18CEE6494AC8AD13DE9FD1AC26">2019-08-23</mods:number>
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<mods:number id="0FE5CAEE1B6691183512AF98919E83AB">4658</mods:number>
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<mods:classification id="AD1AF1F6E270E424DCB8465395D0236F">journal article</mods:classification>
<mods:identifier id="F6B0910D2A6CFC04E7AC6EBD1D6D12EC" type="CLB-Dataset">25923</mods:identifier>
<mods:identifier id="E237127199A186D4601B3D770385C16F" type="DOI">10.11646/zootaxa.4658.3.5</mods:identifier>
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<treatment id="CF458781AD0A2C61FF06FD55FB13AA52" ID-GBIF-Taxon="159244154" LSID="urn:lsid:zoobank.org:act:656F7FD7-B133-46F9-878F-9E5DB1AC3E24" httpUri="http://treatment.plazi.org/id/CF458781AD0A2C61FF06FD55FB13AA52" lastPageNumber="534" pageId="8" pageNumber="534">
<subSubSection id="0FF6651CAD0A2C61FF06FD55FDE0ADDD" box="[151,518,729,756]" pageId="8" pageNumber="534" type="nomenclature">
<paragraph id="47533697AD0A2C61FF06FD55FDE0ADDD" blockId="8.[151,518,729,792]" box="[151,518,729,756]" pageId="8" pageNumber="534">
<heading id="1C1B81FBAD0A2C61FF06FD55FDE0ADDD" bold="true" box="[151,518,729,756]" fontSize="11" level="1" pageId="8" pageNumber="534" reason="1">
<emphasis id="7598EA85AD0A2C61FF06FD55FDE0ADDD" bold="true" box="[151,518,729,756]" pageId="8" pageNumber="534">
<emphasis id="7598EA85AD0A2C61FF06FD55FE47ADDD" bold="true" box="[151,417,729,756]" italics="true" pageId="8" pageNumber="534">
<taxonomicName id="80EC4D14AD0A2C61FF06FD55FE7CADDD" authority="Groenen &amp; Wangdi, 2019" authorityName="Groenen &amp; Wangdi" authorityYear="2019" box="[151,410,729,756]" class="Insecta" family="Tortricidae" genus="Epiblema" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="534" phylum="Arthropoda" rank="species" species="albulusana" status="sp. nov.">Epiblema albulusana</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="6EAB57FEAD0A2C61FE39FD56FDE0ADDD" box="[424,518,730,756]" pageId="8" pageNumber="534" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="0FF6651CAD0A2C61FF06FD71FEFDAC31" box="[151,283,765,792]" pageId="8" pageNumber="534" type="description">
<paragraph id="47533697AD0A2C61FF06FD71FEFDAC31" blockId="8.[151,518,729,792]" box="[151,283,765,792]" pageId="8" pageNumber="534">
<figureCitation id="DFD72A12AD0A2C61FF06FD71FF15AC3E" box="[151,243,765,791]" captionStart="FIGURES 19‒36" captionStartId="10.[151,264,1793,1818]" captionTargetBox="[196,1387,176,1768]" captionTargetId="figure@10.[194,1393,166,1774]" captionTargetPageId="10" captionText="FIGURES 19‒36. Aduts and male genitalia of Cochylini, Archipini, Bactrini, Olethreutini, Eucosmini. 16. Meridemis bathymorpha, male. 17. Meridemis bathymorpha, female. 18. Borneogena trashiyana, female holotype. 19. Isodemis illiberalis, male. 20. Bactra cophinana, female holotype. 21. Lobesia ambigua, male. 22. Penthostola subnigrantis, male, holotype), 23. Metendothenia ordospina, male. 24. M. brunnofasciana, female holotype. 25. Peridaedala nigrifasciana, female holotype. 26. Gibberifera glaciata, male. 27. Crocidosema plebejana, female. 28. Lepteucosma charassuncus, male. 29. Epiblema charadrias, male. 30 E. charadrias, female. 31. E. albulusana, male holotype. 32. Microsarotis bicincta, female. 33. Eupoecilia jakarana, male genitalia, holotype. 34. Eupoecilia jakarana, phallus. 35. E. gedui, male, holotype. 36. E. gedui, phallus." httpUri="https://zenodo.org/record/3376108/files/figure.png" pageId="8" pageNumber="534">Figs. 31</figureCitation>
,
<figureCitation id="DFD72A12AD0A2C61FF6EFD71FEFDAC31" box="[255,283,765,792]" captionStart="FIGURES 37‒55" captionStartId="11.[151,264,1822,1847]" captionTargetBox="[177,1416,170,1798]" captionTargetId="figure@11.[168,1420,166,1802]" captionTargetPageId="11" captionText="FIGURES 37‒55. Male and female genitalia of Tortricini, Archipini, Olethreutini, and Eucosmini. 37. Archips limatus, male. 38. A. limatus, phallus. 39. Chirapsina expleta, male. 40. C. expleta, phallus. 41. Clepsis humana, male. 42. Lumaria phuntschona, male. 43. L. phuntschona, phallus. 44. Meridemis bathymorpha, male. 45. M. bathymorpha, phallus. 46. Isodemis illiberalis, male. 47. I illiberalis, phallus. 48. Lobesia ambigua, male. 49. Penthostola subnigrantis, male holotype. 50. Metendothenia ordospina, male. 51. Gibberifera glaciata, male. 52. Lepteucosma charassuncus, male. 53. Epiblema charasdrias, male. 54. Epiblema albulusana, male holotype. 55. Acleris perfundana, female." httpUri="https://zenodo.org/record/3376110/files/figure.png" pageId="8" pageNumber="534">54</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="0FF6651CAD0A2C61FF06FCCAFB13AA52" pageId="8" pageNumber="534" type="materials_examined">
<paragraph id="47533697AD0A2C61FF06FCCAFDCFACAA" blockId="8.[151,1436,837,1404]" pageId="8" pageNumber="534">
<materialsCitation id="F7843CCAAD0A2C61FF06FCCAFDCFACAA" ID-GBIF-Occurrence="2351015258" collectingDate="2016-08-16" collectionCode="NBCS" country="Bhutan" elevation="2910" latitude="27.544167" location="Jakar" longLatPrecision="20" longitude="90.72278" pageId="8" pageNumber="534" specimenCode="FG3312" specimenCount="1" specimenCount-male="1" stateProvince="Bumthang" typeStatus="holotype">
<emphasis id="7598EA85AD0A2C61FF06FCCAFEE2AC49" bold="true" box="[151,260,838,864]" pageId="8" pageNumber="534">
<typeStatus id="98578835AD0A2C61FF06FCCAFEE2AC49" box="[151,260,838,864]" pageId="8" pageNumber="534" type="holotype">Holotype</typeStatus>
</emphasis>
.
<specimenCount id="51EAFD1EAD0A2C61FE83FCCAFECFAC49" box="[274,297,838,864]" pageId="8" pageNumber="534" type="male">♂</specimenCount>
,
<collectingCountry id="3FFB7607AD0A2C61FEA4FCC9FE6FAC76" box="[309,393,837,863]" name="Bhutan" pageId="8" pageNumber="534">Bhutan</collectingCountry>
,
<collectingRegion id="8528F875AD0A2C61FE02FCC9FDEDAC76" box="[403,523,837,863]" country="Bhutan" name="Bumthang" pageId="8" pageNumber="534">Bumthang</collectingRegion>
,
<quantity id="80149B72AD0A2C61FD87FCC9FDA9AC76" box="[534,591,837,863]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="534" unit="km" value="1.0">1 km</quantity>
NE
<location id="4233604CAD0A2C61FD13FCC9FD59AC76" LSID="urn:lsid:plazi:treatment:CF458781AD0A2C61FF06FD55FB13AA52:4233604CAD0A2C61FD13FCC9FD59AC76" box="[642,703,837,863]" country="Bhutan" latitude="27.544167" longLatPrecision="20" longitude="90.72278" name="Jakar" pageId="8" pageNumber="534" stateProvince="Bumthang">Jakar</location>
,
<geoCoordinate id="22D85050AD0A2C61FD58FCC9FCBEAC76" box="[713,856,837,863]" degrees="27" direction="north" minutes="32" orientation="latitude" pageId="8" pageNumber="534" precision="15" seconds="39" value="27.544167">27º32´39´´N</geoCoordinate>
,
<geoCoordinate id="22D85050AD0A2C61FCF5FCC9FC17AC76" box="[868,1009,837,863]" degrees="90" direction="east" minutes="43" orientation="longitude" pageId="8" pageNumber="534" precision="15" seconds="22" value="90.72278">90º43´22´´E</geoCoordinate>
,
<quantity id="80149B72AD0A2C61FC6DFCC9FBB4AC49" box="[1020,1106,837,864]" metricMagnitude="3" metricUnit="m" metricValue="2.91" pageId="8" pageNumber="534" unit="m" value="2910.0">
<elevation id="CCC1D1A4AD0A2C61FC6DFCC9FBB4AC49" box="[1020,1106,837,864]" metricMagnitude="3" metricUnit="m" metricValue="2.91" pageId="8" pageNumber="534" unit="m" value="2910.0">2910 m</elevation>
</quantity>
,
<date id="33521057AD0A2C61FBCFFCC9FB0EAC76" box="[1118,1256,837,863]" pageId="8" pageNumber="534" value="2016-08-16">
<collectingDate id="2316E9BFAD0A2C61FBCFFCC9FB0EAC76" box="[1118,1256,837,863]" pageId="8" pageNumber="534" value="2016-08-16">16.viii.2016</collectingDate>
</date>
, sta. 24, C. Gie- lis, genitalia slide
<specimenCode id="174A9EECAD0A2C61FEF2FCE5FE59ACAA" box="[355,447,873,899]" collectionCode="FG" country="Germany" name="Palaontologische Hauptsammlung der Bergakadmie" pageId="8" pageNumber="534">FG3312</specimenCode>
(
<collectionCode id="21FDAE52AD0A2C61FE5FFCE5FDFBACAA" box="[462,541,873,899]" pageId="8" pageNumber="534">NBCS</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="47533697AD0A2C61FF56FC01FB12ACC6" blockId="8.[151,1436,837,1404]" pageId="8" pageNumber="534">
<emphasis id="7598EA85AD0A2C61FF56FC01FEDCAC81" bold="true" box="[199,314,909,936]" pageId="8" pageNumber="534">Diagnosis</emphasis>
. This species is described from a single, worn specimen with a pattern similar to species of
<taxonomicName id="80EC4D14AD0A2C61FABEFC01FA7AAC8E" authorityName="Hubner" authorityYear="1825" box="[1327,1436,909,935]" class="Insecta" family="Tortricidae" genus="Notocelia" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="534" phylum="Arthropoda" rank="genus">
<emphasis id="7598EA85AD0A2C61FABEFC01FA7AAC8E" box="[1327,1436,909,935]" italics="true" pageId="8" pageNumber="534">Notocelia</emphasis>
</taxonomicName>
with a black basal fascia and a triangle spot before the dorsum. It differs from species of
<taxonomicName id="80EC4D14AD0A2C61FBFFFC3DFB3DACE2" authorityName="Hubner" authorityYear="1825" box="[1134,1243,945,971]" class="Insecta" family="Tortricidae" genus="Notocelia" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="534" phylum="Arthropoda" rank="genus">
<emphasis id="7598EA85AD0A2C61FBFFFC3DFB3DACE2" box="[1134,1243,945,971]" italics="true" pageId="8" pageNumber="534">Notocelia</emphasis>
</taxonomicName>
in the absence of non-deciduous cornuti in the terminal part of the vesical, and the presence of a clasper on the valva.
</paragraph>
<paragraph id="47533697AD0A2C61FF56FC75FCD1AA1A" blockId="8.[151,1436,837,1404]" pageId="8" pageNumber="534">
<emphasis id="7598EA85AD0A2C61FF56FC75FEB7AB3D" bold="true" box="[199,337,1017,1044]" pageId="8" pageNumber="534">Description</emphasis>
. Male. Head (
<figureCitation id="DFD72A12AD0A2C61FE60FC75FDA3AB3A" box="[497,581,1017,1043]" captionStart="FIGURES 19‒36" captionStartId="10.[151,264,1793,1818]" captionTargetBox="[196,1387,176,1768]" captionTargetId="figure@10.[194,1393,166,1774]" captionTargetPageId="10" captionText="FIGURES 19‒36. Aduts and male genitalia of Cochylini, Archipini, Bactrini, Olethreutini, Eucosmini. 16. Meridemis bathymorpha, male. 17. Meridemis bathymorpha, female. 18. Borneogena trashiyana, female holotype. 19. Isodemis illiberalis, male. 20. Bactra cophinana, female holotype. 21. Lobesia ambigua, male. 22. Penthostola subnigrantis, male, holotype), 23. Metendothenia ordospina, male. 24. M. brunnofasciana, female holotype. 25. Peridaedala nigrifasciana, female holotype. 26. Gibberifera glaciata, male. 27. Crocidosema plebejana, female. 28. Lepteucosma charassuncus, male. 29. Epiblema charadrias, male. 30 E. charadrias, female. 31. E. albulusana, male holotype. 32. Microsarotis bicincta, female. 33. Eupoecilia jakarana, male genitalia, holotype. 34. Eupoecilia jakarana, phallus. 35. E. gedui, male, holotype. 36. E. gedui, phallus." httpUri="https://zenodo.org/record/3376108/files/figure.png" pageId="8" pageNumber="534">Fig. 31</figureCitation>
): White, labial palpus white, labial palpus ca. 2 times diameter of compound eye. Thorax: Nota and tegulae white. Forewing length
<quantity id="80149B72AD0A2C61FD6AFB91FCD9AB11" box="[763,831,1053,1080]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="8" pageNumber="534" unit="mm" value="8.0">8 mm</quantity>
; forewings elongate, subtriangular; costal fold to ⅓ of costa; costa slightly curved, apex slightly protruding, termen and dorsum straight; ground colour white overlaid with grey scales, basal fascia and pretornal spot indicated by black scales, a sinuate black mark before termen. Fringe white mixed with black. Hindwings shiny white, tips of scales fuscous. Fringe white. Abdomen: Genitalia (
<figureCitation id="DFD72A12AD0A2C61FAD2FB05FA72AB8D" box="[1347,1428,1161,1188]" captionStart="FIGURES 37‒55" captionStartId="11.[151,264,1822,1847]" captionTargetBox="[177,1416,170,1798]" captionTargetId="figure@11.[168,1420,166,1802]" captionTargetPageId="11" captionText="FIGURES 37‒55. Male and female genitalia of Tortricini, Archipini, Olethreutini, and Eucosmini. 37. Archips limatus, male. 38. A. limatus, phallus. 39. Chirapsina expleta, male. 40. C. expleta, phallus. 41. Clepsis humana, male. 42. Lumaria phuntschona, male. 43. L. phuntschona, phallus. 44. Meridemis bathymorpha, male. 45. M. bathymorpha, phallus. 46. Isodemis illiberalis, male. 47. I illiberalis, phallus. 48. Lobesia ambigua, male. 49. Penthostola subnigrantis, male holotype. 50. Metendothenia ordospina, male. 51. Gibberifera glaciata, male. 52. Lepteucosma charassuncus, male. 53. Epiblema charasdrias, male. 54. Epiblema albulusana, male holotype. 55. Acleris perfundana, female." httpUri="https://zenodo.org/record/3376110/files/figure.png" pageId="8" pageNumber="534">Fig. 54</figureCitation>
) with tegumen high, uncus rudimentary, haired; socii club-shaped, slender at base, with long setae; gnathos a slender rim; valva with a wide excavation with a triangular clasper at distal edge of excavation, sacculus slender, setose, valve neck curved, short, cucullus ovate, setose, with a large ventral lobe, dorsal lobe slenderer and longer than ventral lobe. Phallus a simple broad tube. Female unknown.
</paragraph>
<paragraph id="47533697AD0A2C61FF56FAB1FE26AA7E" blockId="8.[151,1436,837,1404]" box="[199,448,1341,1368]" pageId="8" pageNumber="534">
<emphasis id="7598EA85AD0A2C61FF56FAB1FEBFAA71" bold="true" box="[199,345,1341,1368]" pageId="8" pageNumber="534">Distribution</emphasis>
.
<collectingCountry id="3FFB7607AD0A2C61FEF6FAB1FE5AAA7E" box="[359,444,1341,1367]" name="Bhutan" pageId="8" pageNumber="534">Bhutan</collectingCountry>
.
</paragraph>
<paragraph id="47533697AD0A2C61FF56FAEEFB13AA52" blockId="8.[151,1436,837,1404]" box="[199,1269,1377,1404]" pageId="8" pageNumber="534">
<emphasis id="7598EA85AD0A2C61FF56FAEEFEAEAA55" bold="true" box="[199,328,1378,1404]" pageId="8" pageNumber="534">Etymology</emphasis>
. The specific epithet
<taxonomicName id="80EC4D14AD0A2C61FDABFAEDFD50AA52" authorityName="Groenen &amp; Wangdi" authorityYear="2019" box="[570,694,1377,1403]" class="Insecta" family="Tortricidae" genus="Epiblema" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="534" phylum="Arthropoda" rank="species" species="albulusana">
<emphasis id="7598EA85AD0A2C61FDABFAEDFD50AA52" box="[570,694,1377,1403]" italics="true" pageId="8" pageNumber="534">albulusana</emphasis>
</taxonomicName>
refers to the mostly white forewing of this species.
</paragraph>
</subSubSection>
</treatment>
</document>
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