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<mods:title id="730F77B0D4BF805F7EE556668EA96BC0">An annotated checklist of the Vespidae (Hymenoptera: Vespoidea) of Bhutan with new records</mods:title>
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<mods:namePart id="515CA4C3F396B92062F7E766BBA4D378">Nidup, Tshering</mods:namePart>
<mods:affiliation id="93F454CEDD65C8C47DDE1DCE33248483">Sherubtse College, Royal University of Bhutan, Kanglung, Bhutan.</mods:affiliation>
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<mods:namePart id="55918FFC770DA446B3C311E73B8A2857">Klein, Wim</mods:namePart>
<mods:affiliation id="C099BACD72213EDBF1933879476ED032">Naturalis Biodiversity Center, 2300 RA Leiden, Netherlands.</mods:affiliation>
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<mods:affiliation id="CB5E0596A6D351862B5BC2AEB8485A70">College of Natural Resources, Royal University of Bhutan, Lobesa, Bhutan.</mods:affiliation>
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<mods:namePart id="27177F10DFD8E7F609DB34ADB186E84D">Daglio, Anthony</mods:namePart>
<mods:affiliation id="AE3EDFFF59D43AD0A72DD07E3D7B21D3">Independent Researcher. anthonydaglio 88 @ yahoo. com</mods:affiliation>
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<paragraph id="4275C55FFFE24032FF796BD6FD94C690" blockId="22.[151,616,151,178]" box="[151,616,151,178]" pageId="22" pageNumber="323">
<heading id="193D7233FFE24032FF796BD6FD94C690" bold="true" box="[151,616,151,178]" fontSize="11" level="1" pageId="22" pageNumber="323" reason="1">
<emphasis id="70BE194DFFE24032FF796BD6FD94C690" bold="true" box="[151,616,151,178]" pageId="22" pageNumber="323">
61.
<taxonomicName id="85CABEDCFFE24032FF296BD6FD94C690" authority="van der Vecht, 1959" authorityName="van der Vecht" authorityYear="1959" box="[199,616,151,178]" class="Insecta" family="Vespidae" genus="Vespa" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="323" phylum="Arthropoda" rank="species" species="fumida">
<emphasis id="70BE194DFFE24032FF296BD6FE97C693" bold="true" box="[199,363,151,178]" italics="true" pageId="22" pageNumber="323">Vespa fumida</emphasis>
<bibRefCitation id="265BB8AEFFE24032FE9C6BD6FD94C690" author="van der Vecht, J." box="[370,616,151,177]" pageId="22" pageNumber="323" pagination="205 - 232" refId="ref22637" refString="van der Vecht, J. (1959) Notes on Oriental Vespinae, including some species from China and Japan (Hymenoptera, Vespidae). Zoologische Mededelingen, XXXVI (13), 205 - 232." type="journal article" year="1959">van der Vecht, 1959</bibRefCitation>
</taxonomicName>
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<taxonomicName id="85CABEDCFFE24032FF796B9DFDB6C6D5" ID-CoL="7G3C5" authority="du Buysson, 1905" authorityName="du Buysson" authorityYear="1905" box="[151,586,219,244]" class="Insecta" family="Vespidae" genus="Vespa" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="323" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis id="70BE194DFFE24032FF796B9DFEC7C6D5" box="[151,315,220,244]" italics="true" pageId="22" pageNumber="323">Vespa variabilis</emphasis>
du Buysson, 1905 (1904)
</taxonomicName>
: 522.
<typeStatus id="9D717BFDFFE24032FD676B9DFD12C6D5" box="[649,750,220,244]" pageId="22" pageNumber="323" type="lectotype">Lectotype</typeStatus>
female (designated by
<treatmentCitation id="C36BE34EFFE24032FC316B9DFB19C6D5" author="van der Vecht, J." box="[991,1253,219,244]" page="228" pageId="22" pageNumber="323" year="1959">
<bibRefCitation id="265BB8AEFFE24032FC316B9DFB19C6D5" author="van der Vecht, J." box="[991,1253,219,244]" pageId="22" pageNumber="323" pagination="205 - 232" refId="ref22637" refString="van der Vecht, J. (1959) Notes on Oriental Vespinae, including some species from China and Japan (Hymenoptera, Vespidae). Zoologische Mededelingen, XXXVI (13), 205 - 232." type="journal article" year="1959">van der Vecht, 1959: 228</bibRefCitation>
</treatmentCitation>
);
<collectingCountry id="3ADD85CFFFE24032FB166B9DFAC4C6D5" box="[1272,1336,220,244]" name="China" pageId="22" pageNumber="323">China</collectingCountry>
:
<location id="47159384FFE24032FAAD6B9DFA60C6D5" LSID="urn:lsid:plazi:treatment:CA637449FFE24032FF796BD6FB17C7E0:47159384FFE24032FAAD6B9DFA60C6D5" box="[1347,1436,220,244]" country="China" name="Mou-pin" pageId="22" pageNumber="323">Mou-pin</location>
(PARIS). Nec
<taxonomicName id="85CABEDCFFE24032FEB76BBAFD5FC733" ID-CoL="7G3C5" authority="Fabricius, 1781" authorityName="Fabricius" authorityYear="1781" box="[345,675,250,274]" class="Insecta" family="Vespidae" genus="Vespa" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="323" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis id="70BE194DFFE24032FEB76BBAFE00C733" box="[345,508,250,274]" italics="true" pageId="22" pageNumber="323">Vespa variabilis</emphasis>
Fabricius, 1781
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.
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<treatmentCitationGroup id="62DAE271FFE24032FF796A5BFB04C710" box="[151,1272,281,305]" pageId="22" pageNumber="323">
<treatmentCitation id="C36BE34EFFE24032FF796A5BFD71C710" author="van der Vecht, J." box="[151,653,281,305]" page="228" pageId="22" pageNumber="323" year="1959">
<taxonomicName id="85CABEDCFFE24032FF796A5BFD71C710" ID-CoL="5LR76" authority="van der Vecht, 1959: 228" authorityName="van der Vecht" authorityPageNumber="228" authorityYear="1959" box="[151,653,281,305]" class="Insecta" family="Vespidae" genus="Vespa" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="323" phylum="Arthropoda" rank="subSpecies" species="variabilis" subSpecies="fumida">
<emphasis id="70BE194DFFE24032FF796A5BFE7AC710" box="[151,390,281,305]" italics="true" pageId="22" pageNumber="323">Vespa variabilis fumida</emphasis>
<bibRefCitation id="265BB8AEFFE24032FE636A58FD71C710" author="van der Vecht, J." box="[397,653,281,305]" pageId="22" pageNumber="323" pagination="205 - 232" refId="ref22637" refString="van der Vecht, J. (1959) Notes on Oriental Vespinae, including some species from China and Japan (Hymenoptera, Vespidae). Zoologische Mededelingen, XXXVI (13), 205 - 232." type="journal article" year="1959">van der Vecht, 1959: 228</bibRefCitation>
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.
<typeStatus id="9D717BFDFFE24032FD796A58FD09C710" box="[663,757,281,305]" pageId="22" pageNumber="323" type="holotype">Holotype</typeStatus>
female;
<collectingCountry id="3ADD85CFFFE24032FCA06A58FC7AC710" box="[846,902,281,305]" name="India" pageId="22" pageNumber="323">India</collectingCountry>
:
<location id="47159384FFE24032FC7D6A58FBCBC710" LSID="urn:lsid:plazi:treatment:CA637449FFE24032FF796BD6FB17C7E0:47159384FFE24032FC7D6A58FBCBC710" box="[915,1079,281,305]" country="India" municipality="Padong'" name="British Bootan'" pageId="22" pageNumber="323">British Bootan</location>
,
<collectingMunicipality id="A2115F25FFE24032FBAF6A58FB6FC710" box="[1089,1171,281,305]" pageId="22" pageNumber="323">Padong</collectingMunicipality>
(PARIS).
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</paragraph>
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<paragraph id="4275C55FFFE24032FF796A1EFB17C7E0" blockId="22.[151,1437,351,450]" pageId="22" pageNumber="323">
<emphasis id="70BE194DFFE24032FF796A1EFED5C758" bold="true" box="[151,297,351,377]" pageId="22" pageNumber="323">Distribution</emphasis>
:
<collectingCountry id="3ADD85CFFFE24032FED76A1EFE72C758" box="[313,398,351,377]" name="Bhutan" pageId="22" pageNumber="323">Bhutan</collectingCountry>
:
<collectingRegion id="800E0BBDFFE24032FE776A21FE0EC75B" box="[409,498,352,378]" country="Bhutan" name="Mongar" pageId="22" pageNumber="323">Mongar</collectingRegion>
(Dorji, Gyeltshen,
<emphasis id="70BE194DFFE24032FD226A21FCF9C758" box="[716,773,351,377]" italics="true" pageId="22" pageNumber="323">et al.</emphasis>
2017; Dorji, Klein,
<emphasis id="70BE194DFFE24032FC046A21FBDEC758" box="[1002,1058,351,377]" italics="true" pageId="22" pageNumber="323">et al.</emphasis>
2017a).
<emphasis id="70BE194DFFE24032FB686A1EFB06C758" box="[1158,1274,351,377]" italics="true" pageId="22" pageNumber="323">Elsewhere</emphasis>
:
<collectingCountry id="3ADD85CFFFE24032FAE76A1EFAB3C758" box="[1289,1359,351,377]" name="Nepal" pageId="22" pageNumber="323">Nepal</collectingCountry>
;
<collectingCountry id="3ADD85CFFFE24032FAB56A1EFA64C758" box="[1371,1432,351,377]" name="India" pageId="22" pageNumber="323">India</collectingCountry>
;
<collectingCountry id="3ADD85CFFFE24032FF796AC5FEF4C7BF" box="[151,264,388,414]" name="Myanmar" pageId="22" pageNumber="323">Myanmar</collectingCountry>
;
<collectingCountry id="3ADD85CFFFE24032FEFD6AC2FEAAC7BC" box="[275,342,387,413]" name="China" pageId="22" pageNumber="323">China</collectingCountry>
(
<bibRefCitation id="265BB8AEFFE24032FE8B6AC2FE0BC7BC" author="Archer, M. E." box="[357,503,387,413]" pageId="22" pageNumber="323" pagination="1 - 41" refId="ref15519" refString="Archer, M. E. (1989) A key to the world species of the Vespinae (Hymenoptera). Research Monographs of the College of Ripon &amp; York St. John, 2, 1 - 41." type="journal article" year="1989">Archer 1989</bibRefCitation>
,
<bibRefCitation id="265BB8AEFFE24032FDEC6AC2FDC2C7BC" author="Archer, M. E." box="[514,574,387,413]" pageId="22" pageNumber="323" refId="ref15717" refString="Archer, M. E. (2012) Vespine Wasps of the World: Behaviour, Ecology &amp; Taxonomy of the Vespinae. Monograph Series 4. SIRI Scientific Press, Manchester, 352 pp." type="book" year="2012">2012</bibRefCitation>
;
<bibRefCitation id="265BB8AEFFE24032FDA76AC2FC8EC7BC" author="Carpenter, J. M. &amp; Kojima, J. I." box="[585,882,387,413]" pageId="22" pageNumber="323" pagination="51 - 92" refId="ref16036" refString="Carpenter, J. M. &amp; Kojima, J. I. (1997) Checklist of the species in the subfamily Vespinae (Insecta: Hymenoptera: Vespidae). Natural History Bulletin of Ibaraki University, 1, 51 - 92." type="journal article" year="1997">Carpenter &amp; Kojima 1997</bibRefCitation>
; Dorji, Gyeltshen,
<emphasis id="70BE194DFFE24032FBA96AC5FB7CC7BC" box="[1095,1152,387,413]" italics="true" pageId="22" pageNumber="323">et al.</emphasis>
2017; Dorji, Klein,
<emphasis id="70BE194DFFE24032FA8A6AC5FA61C7BC" box="[1380,1437,387,413]" italics="true" pageId="22" pageNumber="323">et al.</emphasis>
2017a;
<bibRefCitation id="265BB8AEFFE24032FF046AE6FE45C7E0" author="Gawas, S. M. &amp; Kumar, P. G. &amp; Pannure, A. &amp; Gupta, A. &amp; Carpenter, J. M." box="[234,441,423,449]" pageId="22" pageNumber="323" pagination="1 - 87" refId="ref16788" refString="Gawas, S. M., Kumar, P. G., Pannure, A., Gupta, A. &amp; Carpenter, J. M. (2020) An annotated distributional checklist of Vespidae (Hymenoptera: Vespoidea) of India. Zootaxa, 4784 (1), 1 - 87. https: // doi. org / 10.11646 / zootaxa. 4784.1.1" type="journal article" year="2020">
Gawas
<emphasis id="70BE194DFFE24032FED06AE9FE8AC7E0" box="[318,374,423,449]" italics="true" pageId="22" pageNumber="323">et al.</emphasis>
2020
</bibRefCitation>
;
<bibRefCitation id="265BB8AEFFE24032FE2D6AE6FD72C7E0" author="Ghosh, D. &amp; Kumar, P. G. &amp; Tripathy, B. &amp; Chandra, K." box="[451,654,423,449]" pageId="22" pageNumber="323" pagination="36 - 43" refId="ref16932" refString="Ghosh, D., Kumar, P. G., Tripathy, B. &amp; Chandra, K. (2020) An updated checklist of Vespidae (Insecta: Hymenoptera) from Arunachal Pradesh, India with new distributional records from the State. Halteres, 11, 36 - 43. https: // doi. org / 10.5281 / zenodo. 4042968" type="journal article" year="2020">
Ghosh
<emphasis id="70BE194DFFE24032FDFD6AE9FDB0C7E0" box="[531,588,423,449]" italics="true" pageId="22" pageNumber="323">et al.</emphasis>
2020
</bibRefCitation>
;
<bibRefCitation id="265BB8AEFFE24032FD776AE9FC38C7E0" author="Srinivasan, G. &amp; Kumar, P. G." box="[665,964,423,450]" pageId="22" pageNumber="323" pagination="1313 - 1322" refId="ref21953" refString="Srinivasan, G. &amp; Kumar, P. G. (2010) New records of potter wasps (Hymenoptera: Vespidae: Eumeninae) from Arunachal Pradesh, India: five genera and ten species. Journal of Threatened Taxa, 2 (12), 1313 - 1322. https: // doi. org / 10.11609 / JoTT. o 2468.1313 - 22." type="journal article" year="2010">Kumar &amp; Srinivasan 2010</bibRefCitation>
;
<bibRefCitation id="265BB8AEFFE24032FC216AE6FB23C7E0" author="Smith-Pardo, A. H. &amp; Carpenter, J. M. &amp; Kimsey, L." box="[975,1247,423,449]" pageId="22" pageNumber="323" pagination="1 - 17" refId="ref21749" refString="Smith-Pardo, A. H., Carpenter, J. M. &amp; Kimsey, L. (2020) The Diversity of Hornets in the Genus Vespa (Hymenoptera: Vespidae; Vespinae), Their Importance and Interceptions in the United States. Insect Systematics and Diversity, 4 (3), 1 - 17. https: // doi. org / 10.1093 / isd / ixaa 006" type="journal article" year="2020">
Smith-Pardo
<emphasis id="70BE194DFFE24032FB8D6AE9FB60C7E0" box="[1123,1180,423,449]" italics="true" pageId="22" pageNumber="323">et al.</emphasis>
2020
</bibRefCitation>
).
</paragraph>
</subSubSection>
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