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<document id="B059D85EC3DBA24141B95979C103E76E" ID-CLB-Dataset="57602" ID-DOI="10.5852/ejt.2023.867.2109" ID-GBIF-Dataset="1d2570da-e150-4d75-94d8-bfc5813062bc" ID-ISSN="2118-9773" ID-Zenodo-Dep="7891021" ID-ZooBank="8DEF295C-A8B1-4A6B-B873-B30949F64E07" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="julia" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="julia" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="julia" checkinTime="1683117081100" checkinUser="felipe" docAuthor="Iniesta, Luiz Felipe Moretti, Bouzan, Rodrigo Salvador &amp; Brescovit, Antonio Domingos" docDate="2023" docId="03F887BA1320B1744D45FAD1FB2E548B" docLanguage="en" docName="ejt-867_1-312.pdf.imf" docOrigin="European Journal of Taxonomy 867 (1)" docSource="http://dx.doi.org/10.5852/ejt.2023.867.2109" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Pseudonannolene alegrensis Silvestri 1897" docType="treatment" docVersion="7" lastPageNumber="22" masterDocId="FFC1FFC21333B1614F56FFECFF9A553E" masterDocTitle="A reassessment of the Neotropical genus Pseudonannolene Silvestri, 1895: cladistic analysis, biogeography, and taxonomic review (Spirostreptida: Pseudonannolenidae)" masterLastPageNumber="312" masterPageNumber="1" pageNumber="20" updateTime="1699515670513" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="69C9D79F2D99FEBF2BBBC912698BFB7E">A reassessment of the Neotropical genus Pseudonannolene Silvestri, 1895: cladistic analysis, biogeography, and taxonomic review (Spirostreptida: Pseudonannolenidae)</mods:title>
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<mods:namePart id="E5E51F625AE551CDEE3016D987DF897C">Iniesta, Luiz Felipe Moretti</mods:namePart>
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<mods:affiliation id="A866D0F3E7B0F711705583D5264407E3">Pós-graduação em Zoologia, Instituto de Biociências, Universidade de São Paulo, São Paulo, Brazil. &amp; Laboratório de Coleções Zoológicas, Instituto Butantan, Avenida Vital Brasil, 1500, 05503 - 090, São Paulo, Brazil.</mods:affiliation>
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<mods:namePart id="F059888F84D9ADD7217661381ABE0A8C">Bouzan, Rodrigo Salvador</mods:namePart>
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<mods:affiliation id="2E16674C25C67DCF3C656AD7A396C524">Pós-graduação em Zoologia, Instituto de Biociências, Universidade de São Paulo, São Paulo, Brazil. &amp; Laboratório de Coleções Zoológicas, Instituto Butantan, Avenida Vital Brasil, 1500, 05503 - 090, São Paulo, Brazil.</mods:affiliation>
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<mods:namePart id="E679CE8B27E67964F73F44F95E5E7519">Brescovit, Antonio Domingos</mods:namePart>
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<mods:affiliation id="9AA6F680E3559A50C6F2E7B06625A216">Laboratório de Coleções Zoológicas, Instituto Butantan, Avenida Vital Brasil, 1500, 05503 - 090, São Paulo, Brazil.</mods:affiliation>
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<taxonomicName id="4C514D2F1320B1724D45FAD1FBBA5069" ID-CoL="4P4M8" authority="Silvestri, 1897" authorityName="Silvestri" authorityYear="1897" box="[531,1056,1341,1368]" class="Diplopoda" family="Pseudonannolenidae" genus="Pseudonannolene" kingdom="Animalia" order="Spirostreptida" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="alegrensis">
<emphasis id="B925EABE1320B1724D45FAD1FCF75066" bold="true" box="[531,877,1341,1368]" italics="true" pageId="19" pageNumber="20">Pseudonannolene alegrensis</emphasis>
Silvestri, 1897
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<subSubSection id="C34B65271320B1724EE1FA8CFBE75045" box="[439,1149,1376,1403]" pageId="19" pageNumber="20" type="description">
<paragraph id="8BEE36AC1320B1724EE1FA8CFBE75045" blockId="19.[439,1149,1341,1403]" box="[439,1149,1376,1403]" pageId="19" pageNumber="20">
<figureCitation id="136A2A291320B1724EE1FA8CFDA75045" box="[439,573,1376,1403]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="158.[189,232,759,785]" captionStartId-1="159.[189,230,1666,1692]" captionTargetBox-0="[189,1384,274,714]" captionTargetBox-1="[189,1398,267,1624]" captionTargetId-0="figure-14@158.[189,1398,262,714]" captionTargetId-1="figure-11@159.[189,1398,267,1624]" captionTargetPageId-0="158" captionTargetPageId-1="159" captionText-0="Fig. 44. Pseudonannolene alegrensis Silvestri, 1897 (MCN 626), in lateral view. A. Anterior region. B. Posterior region. Scale bars: A = 0.5 mm; B = 1 mm." captionText-1="Fig. 45. Pseudonannolene alegrensis Silvestri, 1897, neotype, ♂ (MCN 626). A. First leg-pair. B. Detail of prefemur. C. Second leg-pair. D. Detail of telopodite, in anal view. E. Left gonopod, in anal view. F. Left gonopod, in oral view.Abbreviations: see Material and methods. Scale bars:A, C = 0.5 mm; B, D–F = 0.2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.7891131" figureDoi-1="http://doi.org/10.5281/zenodo.7891137" httpUri-0="https://zenodo.org/record/7891131/files/figure.png" httpUri-1="https://zenodo.org/record/7891137/files/figure.png" pageId="19" pageNumber="20">
Figs 44–
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,
<figureCitation id="136A2A291320B1724D1FFA8CFD4D5045" box="[585,727,1376,1403]" captionStart="Fig" captionStartId="278.[189,232,1928,1954]" captionTargetBox="[199,1388,265,1886]" captionTargetId="figure-14@278.[199,1388,265,1886]" captionTargetPageId="278" captionText="Fig. 175. Stereoscopic images of gnathochilarium and anterior sternum of species of Pseudonannolene Silvestri, 1895. A. P. granulata sp. nov. B. P. alata sp. nov. C. P. curvata sp. nov. D. P. aurea sp. nov. E. P. alegrensis. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.7891435" httpUri="https://zenodo.org/record/7891435/files/figure.png" pageId="19" pageNumber="20">175E, 175B</figureCitation>
,
<figureCitation id="136A2A291320B1724DB5FA8DFC885045" box="[739,786,1377,1403]" captionStart="Fig" captionStartId="283.[189,232,1363,1389]" captionTargetBox="[192,1368,285,1297]" captionTargetId="figure-12@283.[189,1398,265,1328]" captionTargetPageId="283" captionText="Fig. 180. Distribution map of the species Pseudonannolene albiventris Schubart, 1952, P. alegrensis Silvestri, 1897, P. ambuatinga Iniesta &amp; Ferreira, 2013, P. anapophysis Fontanetti, 1996, and P. borelli Silvestri, 1895." figureDoi="http://doi.org/10.5281/zenodo.7891445" httpUri="https://zenodo.org/record/7891445/files/figure.png" pageId="19" pageNumber="20">180</figureCitation>
; Supp. file 4:
<figureCitation id="136A2A291320B1724CE3FA8CFBA85045" box="[949,1074,1376,1403]" captionStart="Fig" captionStartId="309.[242,285,365,391]" captionText="Fig. 202. Characters 24–27 and their character states.A. Midbody leg ofPseudonannolene halophila Schubart, 1949 (IBSP 1101). B. Tarsus of midbody leg of P. halophila (IBSP 1101). C. Posterior region of P. alegrensis Silvestri, 1897 (MCN626). D. Posterior region of Epinannolene sp. (ICN). E. Posterior region of P. granulata sp. nov. (MNRJ). F. Posterior region of P. buhrnheimi Schubart, 1960 (IBSP 2397). Scale bars: A = 0.25 mm; B = 0.1 mm; C–F = 0.5 mm." pageId="19" pageNumber="20">Figs 202C</figureCitation>
,
<figureCitation id="136A2A291320B1724B6BFA8DFBE75045" box="[1085,1149,1377,1403]" captionStart="Fig" captionStartId="310.[242,285,883,909]" captionText="Fig. 214. Characters 65–68 and their character states. Gonopods. A. Distal region of gonopods of Phallorthus colombianus Chamberlin, 1952 (FMNH), in anal view. B. Distal region of gonopods of Pseudonannolene buhrnheimi Schubart, 1960 (IBSP 2397), in anal view. C. Left gonopod of P. alegrensis Silvestri, 1897 (MCN 626), in anal view. D. Left gonopod of P. typica Silvestri, 1895 (MCSN), in anal view. E. Distal region of gonopods of P. erikae Iniesta &amp; Ferreira, 2014 (IBSP 3331), in anal view. F. Right gonopod of Holopodostreptus braueri Carl, 1918 (MNRJ), in oral view. Scale bars: A, F = 0.1 mm; B–C = 0.2 mm; D–E = 0.5 mm." pageId="19" pageNumber="20">214C</figureCitation>
</paragraph>
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<paragraph id="8BEE36AC1320B1724FEBFA4DFC055085" blockId="19.[189,927,1441,1467]" box="[189,927,1441,1467]" pageId="19" pageNumber="20">
<treatmentCitationGroup id="AB4111821320B1724FEBFA4DFC055085" box="[189,927,1441,1467]" pageId="19" pageNumber="20">
<treatmentCitation id="0AF010BD1320B1724FEBFA4DFD605085" author="Silvestri F." box="[189,762,1441,1467]" page="19" pageId="19" pageNumber="20" year="1897">
<taxonomicName id="4C514D2F1320B1724FEBFA4DFD605085" ID-CoL="4P4M8" authority="Silvestri, 1897 c: 19" authorityName="Silvestri" authorityPageNumber="19" authorityYear="1897" box="[189,762,1441,1467]" class="Diplopoda" family="Pseudonannolenidae" genus="Pseudonannolene" kingdom="Animalia" order="Spirostreptida" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="alegrensis">
<emphasis id="B925EABE1320B1724FEBFA4DFD965085" box="[189,524,1441,1467]" italics="true" pageId="19" pageNumber="20">Pseudonannolene alegrensis</emphasis>
<bibRefCitation id="EFC04B5D1320B1724D42FA4DFD605085" author="Silvestri F." box="[532,762,1441,1467]" pageId="19" pageNumber="20" pagination="1 - 23" refId="ref108317" refString="Silvestri F. 1897 c. Neue Diplopoden. Abhandlungen und Berichte des Koniglichen Zoologischen und Anthropologisch-Ethnographischen Museums zu Dresden 6 (9): 1 - 23." type="journal article" year="1897">Silvestri, 1897c: 19</bibRefCitation>
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, pl. iii fig. 28.
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<taxonomicName id="4C514D2F1320B1724FEBFA0DFD9550C5" ID-CoL="4P4M8" authorityName="Silvestri" authorityYear="1897" box="[189,527,1505,1531]" class="Diplopoda" family="Pseudonannolenidae" genus="Pseudonannolene" kingdom="Animalia" order="Spirostreptida" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="alegrensis">
<emphasis id="B925EABE1320B1724FEBFA0DFD9550C5" box="[189,527,1505,1531]" italics="true" pageId="19" pageNumber="20">Pseudonannolene alegrensis</emphasis>
</taxonomicName>
–
<treatmentCitation id="0AF010BD1320B1724D62FA0DFCB550C2" author="Brolemann H. W." box="[564,815,1505,1532]" page="56" pageId="19" pageNumber="20" year="1909">
<bibRefCitation id="EFC04B5D1320B1724D62FA0DFCB550C2" author="Brolemann H. W." box="[564,815,1505,1532]" pageId="19" pageNumber="20" refId="ref100208" refString="Brolemann H. W. 1909. Os Myriapodos do Brazil. Catalogos da Fauna Brazileira. Museu Paulista, Sao Paulo, Brasil." type="book" year="1909">Brölemann 1909: 56</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="0AF010BD1320B1724C33FA0DFBC850C5" author="Viggiani V." box="[869,1106,1505,1531]" page="366" pageId="19" pageNumber="20" year="1973">
<bibRefCitation id="EFC04B5D1320B1724C33FA0DFBC850C5" author="Viggiani V." box="[869,1106,1505,1531]" pageId="19" pageNumber="20" pagination="351 - 417" refId="ref108936" refString="Viggiani V. 1973. Le specie descritte da Filippo Silvestri (1873 - 1949). Bollettino del Laboratorio di entomologia agraria &quot; Filippo Silvestri &quot; 30: 351 - 417." type="journal article" year="1973">Viggiani 1973: 366</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="0AF010BD1320B1724BDEFA0DFAD150C5" author="Jeekel C. A. W." box="[1160,1355,1505,1531]" page="88" pageId="19" pageNumber="20" year="2004">
<bibRefCitation id="EFC04B5D1320B1724BDEFA0DFAD150C5" author="Jeekel C. A. W." box="[1160,1355,1505,1531]" pageId="19" pageNumber="20" pagination="43 - 109" refId="ref104347" refString="Jeekel C. A. W. 2004. A bibliographic catalogue of the &quot; Cambaloidea &quot; (Diplopoda, Spirostreptida). Myriapod Memoranda 7: 43 - 109." type="journal article" year="2004">Jeekel 2004: 88</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="0AF010BD1320B1724FBCF9EFFDD15320" author="Iniesta L. F. &amp; Ferreira R. L." box="[234,587,1539,1566]" page="357" pageId="19" pageNumber="20" year="2013">
<bibRefCitation id="EFC04B5D1320B1724FBCF9EFFDD15320" author="Iniesta L. F. &amp; Ferreira R. L." box="[234,587,1539,1566]" pageId="19" pageNumber="20" pagination="357 - 369" refId="ref103738" refString="Iniesta L. F. M &amp; Ferreira R. L. 2013 b. Two new species of Pseudonannolene Silvestri, 1895 from Brazilian limestone caves (Spirostreptida: Pseudonannolenidae): synotopy of a troglophilic and a troglobiotic species. Zootaxa 3702 (4): 357 - 369. https: // doi. org / 10.11646 / zootaxa. 3702.4.3" type="journal article" year="2013">Iniesta &amp; Ferreira 2013b: 357</bibRefCitation>
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.
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</paragraph>
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<heading id="D0A681C01320B1724FEBF9AFFDEF5363" bold="true" box="[189,629,1603,1630]" fontSize="11" level="3" pageId="19" pageNumber="20" reason="3">
<emphasis id="B925EABE1320B1724FEBF9AFFDEF5363" bold="true" box="[189,629,1603,1630]" pageId="19" pageNumber="20">
Justification of
<typeStatus id="54EA880E1320B1724E2BF9A8FE455360" box="[381,479,1604,1630]" pageId="19" pageNumber="20" type="neotype">neotype</typeStatus>
designation
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<paragraph id="8BEE36AC1320B1724FEBF99DFD8A5266" blockId="19.[189,1399,1649,1881]" pageId="19" pageNumber="20">
The type material of
<taxonomicName id="4C514D2F1320B1724E83F99EFDEB53B5" ID-CoL="4P4M8" authorityName="Silvestri" authorityYear="1897" box="[469,625,1649,1675]" class="Diplopoda" family="Pseudonannolenidae" genus="Pseudonannolene" kingdom="Animalia" order="Spirostreptida" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="alegrensis">
<emphasis id="B925EABE1320B1724E83F99EFDEB53B5" box="[469,625,1649,1675]" italics="true" pageId="19" pageNumber="20">P. alegrensis</emphasis>
</taxonomicName>
deposited in the Senckenberg Naturhistorische Sammlungen, Dresden,
<collectingCountry id="F346763C1320B1724E67F97FFE045393" box="[305,414,1683,1709]" name="Germany" pageId="19" pageNumber="20">Germany</collectingCountry>
(SMTD), and other types of millipedes described by Silvestri in 1897, were lost during the bombing of Dresden on
<date id="FFEF106C1320B1724D23F959FCA653EE" box="[629,828,1717,1744]" pageId="19" pageNumber="20" value="1944-10-07">October 7, 1944</date>
(
<bibRefCitation id="EFC04B5D1320B1724C19F959FB0C53EE" author="Sierwald P. &amp; Reft A. J." box="[847,1174,1717,1744]" pageId="19" pageNumber="20" pagination="1 - 100" refId="ref107926" refString="Sierwald P. &amp; Reft A. J. 2004. The millipede collections of the world. Fieldiana, N. S. 103 (1532): 1 - 100." type="journal article" year="2004">Sierwald &amp; Reft 2004: 47</bibRefCitation>
). The species was described by
<bibRefCitation id="EFC04B5D1320B1724E0AF934FDB953CC" author="Silvestri F." box="[348,547,1752,1778]" pageId="19" pageNumber="20" pagination="1 - 23" refId="ref108317" refString="Silvestri F. 1897 c. Neue Diplopoden. Abhandlungen und Berichte des Koniglichen Zoologischen und Anthropologisch-Ethnographischen Museums zu Dresden 6 (9): 1 - 23." type="journal article" year="1897">Silvestri (1897c)</bibRefCitation>
based on an adult female collected in Porto Alegre,
<collectingRegion id="4995F84E1320B1724BC1F934FF67522A" country="Brazil" name="Rio Grande do Sul" pageId="19" pageNumber="20">Rio Grande do Sul State</collectingRegion>
,
<collectingCountry id="F346763C1320B1724E51F916FEC8522A" box="[263,338,1786,1812]" name="Brazil" pageId="19" pageNumber="20">Brazil</collectingCountry>
. Since no name-bearing type specimens for
<taxonomicName id="4C514D2F1320B1724C05F917FC7D522A" ID-CoL="4P4M8" authorityName="Silvestri" authorityYear="1897" box="[851,999,1786,1812]" class="Diplopoda" family="Pseudonannolenidae" genus="Pseudonannolene" kingdom="Animalia" order="Spirostreptida" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="alegrensis">
<emphasis id="B925EABE1320B1724C05F917FC7D522A" box="[851,999,1786,1812]" italics="true" pageId="19" pageNumber="20">P. alegrensis</emphasis>
</taxonomicName>
exist and to secure the stability of the nomenclature, we selected topotype material from MCN to designate the
<typeStatus id="54EA880E1320B1724B11F8F0FB3E5208" box="[1095,1188,1820,1846]" pageId="19" pageNumber="20" type="neotype">neotype</typeStatus>
(for more details, see article #75.3.5 of ICZN).
</paragraph>
</subSubSection>
<subSubSection id="C34B65271320B1754FEBF892FDC05478" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20" type="diagnosis">
<paragraph id="8BEE36AC1320B1724FEBF892FEAC52A6" blockId="19.[189,310,1918,1944]" box="[189,310,1918,1944]" pageId="19" pageNumber="20">
<heading id="D0A681C01320B1724FEBF892FEAC52A6" bold="true" box="[189,310,1918,1944]" fontSize="11" level="3" pageId="19" pageNumber="20" reason="3">
<emphasis id="B925EABE1320B1724FEBF892FEAC52A6" bold="true" box="[189,310,1918,1944]" pageId="19" pageNumber="20">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BEE36AC1320B1754FEBF840FDC05478" blockId="19.[189,1399,1964,2025]" lastBlockId="20.[189,1399,265,327]" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20">
Males of
<taxonomicName id="4C514D2F1320B1724E78F841FE5952F8" authorityName="Silvestri" authorityYear="1897" box="[302,451,1964,1990]" class="Diplopoda" family="Pseudonannolenidae" genus="Pseudonannolene" kingdom="Animalia" order="Spirostreptida" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="alegrensis">
<emphasis id="B925EABE1320B1724E78F841FE5952F8" box="[302,451,1964,1990]" italics="true" pageId="19" pageNumber="20">P. alegrensis</emphasis>
</taxonomicName>
resemble those of
<taxonomicName id="4C514D2F1320B1724DF5F841FC8852F8" authorityName="Silvestri" authorityYear="1902" box="[675,786,1965,1990]" class="Diplopoda" family="Pseudonannolenidae" genus="Pseudonannolene" kingdom="Animalia" order="Spirostreptida" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="rocana">
<emphasis id="B925EABE1320B1724DF5F841FC8852F8" box="[675,786,1965,1990]" italics="true" pageId="19" pageNumber="20">P. rocana</emphasis>
</taxonomicName>
by having an elongated telopodite (longer than half of gonocoxa) (
<figureCitation id="136A2A291320B1724E3CF822FE6C52D7" box="[362,502,1998,2025]" captionStart="Fig" captionStartId="159.[189,230,1666,1692]" captionTargetBox="[189,1398,267,1624]" captionTargetId="figure-11@159.[189,1398,267,1624]" captionTargetPageId="159" captionText="Fig. 45. Pseudonannolene alegrensis Silvestri, 1897, neotype, ♂ (MCN 626). A. First leg-pair. B. Detail of prefemur. C. Second leg-pair. D. Detail of telopodite, in anal view. E. Left gonopod, in anal view. F. Left gonopod, in oral view.Abbreviations: see Material and methods. Scale bars:A, C = 0.5 mm; B, D–F = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7891137" httpUri="https://zenodo.org/record/7891137/files/figure.png" pageId="19" pageNumber="20">Fig. 45D–F</figureCitation>
), but differing by the presence of a large prefemoral process on the first leg-pair (
<figureCitation id="136A2A291327B1754FACFEE5FE10541A" box="[250,394,265,292]" captionStart="Fig" captionStartId="159.[189,230,1666,1692]" captionTargetBox="[189,1398,267,1624]" captionTargetId="figure-11@159.[189,1398,267,1624]" captionTargetPageId="159" captionText="Fig. 45. Pseudonannolene alegrensis Silvestri, 1897, neotype, ♂ (MCN 626). A. First leg-pair. B. Detail of prefemur. C. Second leg-pair. D. Detail of telopodite, in anal view. E. Left gonopod, in anal view. F. Left gonopod, in oral view.Abbreviations: see Material and methods. Scale bars:A, C = 0.5 mm; B, D–F = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7891137" httpUri="https://zenodo.org/record/7891137/files/figure.png" pageId="20" pageNumber="21">Fig. 45A–B</figureCitation>
); subtriangular solenomere (
<figureCitation id="136A2A291327B1754D81FEE5FCDE541A" box="[727,836,265,292]" captionStart="Fig" captionStartId="159.[189,230,1666,1692]" captionTargetBox="[189,1398,267,1624]" captionTargetId="figure-11@159.[189,1398,267,1624]" captionTargetPageId="159" captionText="Fig. 45. Pseudonannolene alegrensis Silvestri, 1897, neotype, ♂ (MCN 626). A. First leg-pair. B. Detail of prefemur. C. Second leg-pair. D. Detail of telopodite, in anal view. E. Left gonopod, in anal view. F. Left gonopod, in oral view.Abbreviations: see Material and methods. Scale bars:A, C = 0.5 mm; B, D–F = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7891137" httpUri="https://zenodo.org/record/7891137/files/figure.png" pageId="20" pageNumber="21">Fig. 45D</figureCitation>
); and by having projections bearing setae on the paraproct (Supp. file 4:
<figureCitation id="136A2A291327B1754E85FEC0FDD75478" box="[467,589,300,326]" captionStart="Fig" captionStartId="309.[242,285,365,391]" captionText="Fig. 202. Characters 24–27 and their character states.A. Midbody leg ofPseudonannolene halophila Schubart, 1949 (IBSP 1101). B. Tarsus of midbody leg of P. halophila (IBSP 1101). C. Posterior region of P. alegrensis Silvestri, 1897 (MCN626). D. Posterior region of Epinannolene sp. (ICN). E. Posterior region of P. granulata sp. nov. (MNRJ). F. Posterior region of P. buhrnheimi Schubart, 1960 (IBSP 2397). Scale bars: A = 0.25 mm; B = 0.1 mm; C–F = 0.5 mm." pageId="20" pageNumber="21">Fig. 202C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34B65271327B1754FEBFE81FC60548A" box="[189,1018,365,437]" pageId="20" pageNumber="21" type="etymology">
<paragraph id="8BEE36AC1327B1754FEBFE81FEDE54B9" blockId="20.[189,324,365,391]" box="[189,324,365,391]" pageId="20" pageNumber="21">
<heading id="D0A681C01327B1754FEBFE81FEDE54B9" bold="true" box="[189,324,365,391]" fontSize="11" level="3" pageId="20" pageNumber="21" reason="3">
<emphasis id="B925EABE1327B1754FEBFE81FEDE54B9" bold="true" box="[189,324,365,391]" pageId="20" pageNumber="21">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BEE36AC1327B1754FEBFE76FC60548A" blockId="20.[189,1018,410,437]" box="[189,1018,410,437]" pageId="20" pageNumber="21">
Adjective referring to the
<typeStatus id="54EA880E1327B1754EB8FE77FDBB548B" box="[494,545,411,437]" pageId="20" pageNumber="21">type</typeStatus>
locality of the species (
<bibRefCitation id="EFC04B5D1327B1754C60FE76FC77548A" author="Silvestri F." box="[822,1005,410,436]" pageId="20" pageNumber="21" pagination="1 - 23" refId="ref108317" refString="Silvestri F. 1897 c. Neue Diplopoden. Abhandlungen und Berichte des Koniglichen Zoologischen und Anthropologisch-Ethnographischen Museums zu Dresden 6 (9): 1 - 23." type="journal article" year="1897">Silvestri 1897c</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34B65271327B1754FEBFE36FD945630" pageId="20" pageNumber="21" type="materials_examined">
<paragraph id="8BEE36AC1327B1754FEBFE36FE3054CA" blockId="20.[189,426,474,500]" box="[189,426,474,500]" pageId="20" pageNumber="21">
<heading id="D0A681C01327B1754FEBFE36FE3054CA" bold="true" box="[189,426,474,500]" fontSize="11" level="3" pageId="20" pageNumber="21" reason="3">
<emphasis id="B925EABE1327B1754FEBFE36FE3054CA" bold="true" box="[189,426,474,500]" pageId="20" pageNumber="21">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BEE36AC1327B1754FBCFDE4FCB95758" blockId="20.[189,1398,520,615]" pageId="20" pageNumber="21">
<materialsCitation id="3B393CF11327B1754FBCFDE4FCB95758" ID-GBIF-Occurrence="4101309735" collectingDate="2010-07-17" collectionCode="MCN" collectorName="M. Poiret" country="Brazil" elevation="124" latitude="-30.11695" location="Lomba do Pinheiro" longLatPrecision="1" longitude="-51.101997" municipality="Porto Alegre" pageId="20" pageNumber="21" specimenCode="MCN 626" specimenCount="1" specimenCount-male="1" stateProvince="Rio Grande do Sul" typeStatus="neotype">
<emphasis id="B925EABE1327B1754FBCFDE4FECB571C" bold="true" box="[234,337,520,546]" pageId="20" pageNumber="21">
<typeStatus id="54EA880E1327B1754FBCFDE4FECB571C" box="[234,337,520,546]" pageId="20" pageNumber="21" type="neotype">Neotype</typeStatus>
</emphasis>
(here designated)
<collectingCountry id="F346763C1327B1754FEBFDC6FEBC577A" box="[189,294,554,580]" name="Brazil" pageId="20" pageNumber="21">BRAZIL</collectingCountry>
•
<specimenCount id="9D57FD251327B1754E1FFDC6FEE7577B" box="[329,381,554,581]" pageId="20" pageNumber="21" type="male">1 ♂</specimenCount>
;
<collectingRegion id="4995F84E1327B1754ED8FDC6FDE3577A" box="[398,633,554,580]" country="Brazil" name="Rio Grande do Sul" pageId="20" pageNumber="21">Rio Grande do Sul</collectingRegion>
,
<collectingMunicipality id="6B8AACD61327B1754DDAFDC7FCB0577A" box="[652,810,554,581]" pageId="20" pageNumber="21">Porto Alegre</collectingMunicipality>
,
<location id="8E8E60771327B1754C6CFDC6FBB0577A" LSID="urn:lsid:plazi:treatment:03F887BA1320B1744D45FAD1FB2E548B:8E8E60771327B1754C6CFDC6FBB0577A" box="[826,1066,554,580]" country="Brazil" latitude="-30.11695" longLatPrecision="1" longitude="-51.101997" municipality="Porto Alegre" name="Lomba do Pinheiro" pageId="20" pageNumber="21" stateProvince="Rio Grande do Sul">Lomba do Pinheiro</location>
; [
<geoCoordinate id="EE65506B1327B1754B14FDC6FB54577A" box="[1090,1230,554,580]" degrees="30.116949" direction="south" orientation="latitude" pageId="20" pageNumber="21" precision="1" value="-30.11695">-30.116949</geoCoordinate>
,
<geoCoordinate id="EE65506B1327B1754B88FDC5FAF1577A" box="[1246,1387,553,580]" degrees="51.101999" direction="west" orientation="longitude" pageId="20" pageNumber="21" precision="1" value="-51.101997">-51.101999</geoCoordinate>
];
<elevation id="007CD19F1327B1754FEBFDA0FEDD5758" box="[189,327,588,615]" metricMagnitude="2" metricUnit="m" metricValue="1.24" pageId="20" pageNumber="21" unit="m" value="124.0">
<quantity id="4CA99B491327B1754FEBFDA0FE9D5759" box="[189,263,588,615]" metricMagnitude="2" metricUnit="m" metricValue="1.24" pageId="20" pageNumber="21" unit="m" value="124.0">124 m</quantity>
a.s.l.
</elevation>
;
<date id="FFEF106C1327B1754E03FDA0FE725758" box="[341,488,588,614]" pageId="20" pageNumber="21" value="2010-07-17">
<collectingDate id="EFABE9841327B1754E03FDA0FE725758" box="[341,488,588,614]" pageId="20" pageNumber="21" value="2010-07-17">17 Jul. 2010</collectingDate>
</date>
;
<collectorName id="26A4537A1327B1754EA2FDA1FDF95758" box="[500,611,588,615]" pageId="20" pageNumber="21">M. Poiret</collectorName>
leg.;
<specimenCode id="DBF79ED71327B1754DF3FDA0FC855758" box="[677,799,588,614]" pageId="20" pageNumber="21">MCN 626</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BEE36AC1327B1754FBCFD60FCA25799" blockId="20.[189,1399,652,782]" box="[234,824,652,681]" pageId="20" pageNumber="21">
<emphasis id="B925EABE1327B1754FBCFD60FE3C5798" bold="true" box="[234,422,652,678]" pageId="20" pageNumber="21">Other material</emphasis>
(total:
<specimenCount id="9D57FD251327B1754EADFD61FDA75797" box="[507,573,653,681]" pageId="20" pageNumber="21" type="male">
3
<emphasis id="B925EABE1327B1754D47FD61FDA75797" bold="true" box="[529,573,653,681]" pageId="20" pageNumber="21">♂♂</emphasis>
</specimenCount>
,
<specimenCount id="9D57FD251327B1754D1DFD60FD175797" box="[587,653,652,681]" pageId="20" pageNumber="21" type="female">
6
<emphasis id="B925EABE1327B1754D37FD61FD175797" bold="true" box="[609,653,653,681]" pageId="20" pageNumber="21">♀♀</emphasis>
</specimenCount>
,
<specimenCount id="9D57FD251327B1754DCAFD61FCAA5799" box="[668,816,652,679]" pageId="20" pageNumber="21" type="immature">4 immatures</specimenCount>
)
</paragraph>
<paragraph id="8BEE36AC1327B1754FEBFD43FD945630" blockId="20.[189,1399,652,782]" pageId="20" pageNumber="21">
<materialsCitation id="3B393CF11327B1754FEBFD43FF7357D5" ID-GBIF-Occurrence="4101309906" collectingDate="2010-07-17" collectionCode="MCN" collectorName="M. Poiret" country="Brazil" elevation="124" latitude="-30.11695" location="Rio Grande do Sul" longLatPrecision="1" longitude="-51.101997" municipality="Porto Alegre" pageId="20" pageNumber="21" specimenCode="MCN 626" specimenCount="11" specimenCount-female="5" specimenCount-immature="4" specimenCount-male="2" stateProvince="Rio Grande do Sul" typeStatus="neotype">
<collectingCountry id="F346763C1327B1754FEBFD43FEBC57F7" box="[189,294,687,713]" name="Brazil" pageId="20" pageNumber="21">BRAZIL</collectingCountry>
–
<emphasis id="B925EABE1327B1754E15FD42FDB757F6" bold="true" box="[323,557,686,713]" pageId="20" pageNumber="21">
<collectingRegion id="4995F84E1327B1754E15FD42FDB757F6" box="[323,557,686,713]" country="Brazil" name="Rio Grande do Sul" pageId="20" pageNumber="21">Rio Grande do Sul</collectingRegion>
</emphasis>
•
<specimenCount id="9D57FD251327B1754D13FD43FD1357F7" box="[581,649,687,713]" pageId="20" pageNumber="21" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="9D57FD251327B1754DC0FD42FD4357F7" box="[662,729,686,713]" pageId="20" pageNumber="21" type="female">5 ♀♀</specimenCount>
,
<specimenCount id="9D57FD251327B1754DB0FD43FCE657F7" box="[742,892,687,713]" pageId="20" pageNumber="21" type="immature">4 immatures</specimenCount>
; same collection data as for
<typeStatus id="54EA880E1327B1754B91FD43FAB257F7" box="[1223,1320,687,713]" pageId="20" pageNumber="21" type="neotype">neotype</typeStatus>
;
<specimenCode id="DBF79ED71327B1754A62FD43FF7357D5" pageId="20" pageNumber="21">MCN 626</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B393CF11327B1754E53FD3DFD905630" ID-GBIF-Occurrence="4101310132" collectingDate="2003-12-18" collectionCode="R" collectorName="Ott" country="Brazil" elevation="17" latitude="-30.292875" location="Fazenda Boa Vista" longLatPrecision="1" longitude="-51.316044" municipality="Barra do Ribeiro" pageId="20" pageNumber="21" specimenCode="MCN 521" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Rio Grande do Sul">
<specimenCount id="9D57FD251327B1754E53FD3DFEAF57D5" box="[261,309,721,747]" pageId="20" pageNumber="21" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D57FD251327B1754E15FD3DFEE857D5" box="[323,370,721,747]" pageId="20" pageNumber="21" type="female">1 ♀</specimenCount>
;
<collectingMunicipality id="6B8AACD61327B1754ED4FD3DFDD757D5" box="[386,589,721,747]" pageId="20" pageNumber="21">Barra do Ribeiro</collectingMunicipality>
,
<location id="8E8E60771327B1754D0CFD3DFCA757D5" LSID="urn:lsid:plazi:treatment:03F887BA1320B1744D45FAD1FB2E548B:8E8E60771327B1754D0CFD3DFCA757D5" box="[602,829,721,747]" country="Brazil" latitude="-30.292875" longLatPrecision="1" longitude="-51.316044" municipality="Barra do Ribeiro" name="Fazenda Boa Vista" pageId="20" pageNumber="21" stateProvince="Rio Grande do Sul">Fazenda Boa Vista</location>
; [
<geoCoordinate id="EE65506B1327B1754C07FD3DFC4757D2" box="[849,989,721,748]" degrees="30.292875" direction="south" orientation="latitude" pageId="20" pageNumber="21" precision="1" value="-30.292875">-30.292875</geoCoordinate>
,
<geoCoordinate id="EE65506B1327B1754CBDFD3CFBED57D5" box="[1003,1143,720,747]" degrees="51.316045" direction="west" orientation="longitude" pageId="20" pageNumber="21" precision="1" value="-51.316044">-51.316045</geoCoordinate>
];
<elevation id="007CD19F1327B1754BDAFD3DFA9157D5" box="[1164,1291,721,747]" metricMagnitude="1" metricUnit="m" metricValue="1.7" pageId="20" pageNumber="21" unit="m" value="17.0">
<quantity id="4CA99B491327B1754BDAFD3DFB5357D5" box="[1164,1225,721,747]" metricMagnitude="1" metricUnit="m" metricValue="1.7" pageId="20" pageNumber="21" unit="m" value="17.0">17 m</quantity>
a.s.l.
</elevation>
;
<date id="FFEF106C1327B1754A4CFD3DFF615633" pageId="20" pageNumber="21" value="2003-12-18">
<collectingDate id="EFABE9841327B1754A4CFD3DFF615633" pageId="20" pageNumber="21" value="2003-12-18">18 Dec. 2003</collectingDate>
</date>
; R. Ott leg.;
<specimenCode id="DBF79ED71327B1754EC7FD1FFD905630" box="[401,522,755,782]" pageId="20" pageNumber="21">MCN 521</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34B65271327B1744FEBFCDFFAD65478" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21" type="description">
<paragraph id="8BEE36AC1327B1754FEBFCDFFE095673" blockId="20.[189,403,819,845]" box="[189,403,819,845]" pageId="20" pageNumber="21">
<heading id="D0A681C01327B1754FEBFCDFFE095673" bold="true" box="[189,403,819,845]" fontSize="11" level="3" pageId="20" pageNumber="21" reason="3">
<emphasis id="B925EABE1327B1754FEBFCDFFE095673" bold="true" box="[189,403,819,845]" pageId="20" pageNumber="21">Descriptive notes</emphasis>
</heading>
</paragraph>
<paragraph id="8BEE36AC1327B1754FEBFC8DFEA656FE" blockId="20.[189,1399,865,960]" pageId="20" pageNumber="21">
<smallCapsWord id="8D08A0701327B1754FEBFC8DFEF45647" baselines="884,884" box="[189,366,865,891]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="20" pageNumber="21">MEASUREMENTS</smallCapsWord>
. 58–60 body rings (1–2 apodous + telson). Males: body length
<quantity id="4CA99B491327B1754B0DFC8DFB695645" box="[1115,1267,865,892]" metricMagnitude="-2" metricUnit="m" metricValue="3.875" metricValueMax="4.25" metricValueMin="3.5" pageId="20" pageNumber="21" unit="mm" value="38.75" valueMax="42.5" valueMin="35.0">35–42.5 mm</quantity>
; maximum midbody diameter
<quantity id="4CA99B491327B1754EFDFC6FFDDE56A3" box="[427,580,899,925]" metricMagnitude="-3" metricUnit="m" metricValue="1.85" metricValueMax="1.9" metricValueMin="1.8" pageId="20" pageNumber="21" unit="mm" value="1.85" valueMax="1.9" valueMin="1.8">1.8–1.9 mm</quantity>
. Females: body length
<quantity id="4CA99B491327B1754C22FC6FFC6456A3" box="[884,1022,899,925]" metricMagnitude="-2" metricUnit="m" metricValue="3.85" metricValueMax="4.1" metricValueMin="3.6" pageId="20" pageNumber="21" unit="mm" value="38.5" valueMax="41.0" valueMin="36.0">36–41 mm</quantity>
; maximum midbody diameter
<quantity id="4CA99B491327B1754FEBFC49FEAD56FE" box="[189,311,933,960]" metricMagnitude="-3" metricUnit="m" metricValue="2.55" metricValueMax="3.1" metricValueMin="2.0" pageId="20" pageNumber="21" unit="mm" value="2.55" valueMax="3.1" valueMin="2.0">2–3.1 mm</quantity>
.
</paragraph>
<paragraph id="8BEE36AC1327B1754FEBFC09FCF5511F" blockId="20.[188,1398,997,1058]" pageId="20" pageNumber="21">
<smallCapsWord id="8D08A0701327B1754FEBFC09FE9356C0" baselines="1016,1017" box="[189,265,997,1023]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Color" pageId="20" pageNumber="21">COLOR</smallCapsWord>
. Body color greyish; prozonites darker; metazonites with a medial dark grey band and a posterior lighter one; head, collum, antennae and legs lighter brown.
</paragraph>
<paragraph id="8BEE36AC1327B1754FEBFBA4FC165199" blockId="20.[189,1399,1096,1191]" pageId="20" pageNumber="21">
<smallCapsWord id="8D08A0701327B1754FEBFBA4FF66515E" baselines="1115,1115" box="[189,252,1096,1122]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Head" pageId="20" pageNumber="21">HEAD</smallCapsWord>
. Antennae short (
<figureCitation id="136A2A291327B1754E86FBA4FDA4515C" box="[464,574,1096,1122]" captionStart="Fig" captionStartId="158.[189,232,759,785]" captionTargetBox="[189,1384,274,714]" captionTargetId="figure-14@158.[189,1398,262,714]" captionTargetPageId="158" captionText="Fig. 44. Pseudonannolene alegrensis Silvestri, 1897 (MCN 626), in lateral view. A. Anterior region. B. Posterior region. Scale bars: A = 0.5 mm; B = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7891131" httpUri="https://zenodo.org/record/7891131/files/figure.png" pageId="20" pageNumber="21">Fig. 44A</figureCitation>
), just reaching back to end of ring 4 when extended dorsally; relative antennomere lengths 1&lt;2&lt;3&gt;4&lt;5≈6&gt;7. Mandibular cardo with ventral margin swollen. Ommatidial cluster well-developed, elliptical; ca 20 ommatidia in 5 rows.
</paragraph>
<paragraph id="8BEE36AC1327B1754FEBFB21FC975073" blockId="20.[189,1399,1228,1357]" pageId="20" pageNumber="21">
<smallCapsWord id="8D08A0701327B1754FEBFB21FF6451DB" baselines="1248,1248" box="[189,254,1229,1255]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Body" pageId="20" pageNumber="21">BODY</smallCapsWord>
<smallCapsWord id="8D08A0701327B1754E5DFB3EFED351DB" baselines="1248" box="[267,329,1234,1253]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="rings" pageId="20" pageNumber="21">RINGS</smallCapsWord>
. Collum with lateral lobes broadly rounded, with ca 6 striae (
<figureCitation id="136A2A291327B1754B0FFB20FB5751D9" box="[1113,1229,1228,1255]" captionStart="Fig" captionStartId="158.[189,232,759,785]" captionTargetBox="[189,1384,274,714]" captionTargetId="figure-14@158.[189,1398,262,714]" captionTargetPageId="158" captionText="Fig. 44. Pseudonannolene alegrensis Silvestri, 1897 (MCN 626), in lateral view. A. Anterior region. B. Posterior region. Scale bars: A = 0.5 mm; B = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7891131" httpUri="https://zenodo.org/record/7891131/files/figure.png" pageId="20" pageNumber="21">Fig. 44A</figureCitation>
). Very faintly constricted between prozonite and metazonite; prozonites smooth; metazonites laterally with transverse striae below ozopore. Anterior sternum subrectangular, without transverse striae (
<figureCitation id="136A2A291327B1754B2EFAFDFB6B5012" box="[1144,1265,1297,1324]" captionStart="Fig" captionStartId="278.[189,232,1928,1954]" captionTargetBox="[199,1388,265,1886]" captionTargetId="figure-14@278.[199,1388,265,1886]" captionTargetPageId="278" captionText="Fig. 175. Stereoscopic images of gnathochilarium and anterior sternum of species of Pseudonannolene Silvestri, 1895. A. P. granulata sp. nov. B. P. alata sp. nov. C. P. curvata sp. nov. D. P. aurea sp. nov. E. P. alegrensis. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.7891435" httpUri="https://zenodo.org/record/7891435/files/figure.png" pageId="20" pageNumber="21">Fig. 175E</figureCitation>
). Paraproct with rounded projections bearing setae (
<figureCitation id="136A2A291327B1754DC2FADFFC9A5073" box="[660,768,1331,1357]" captionStart="Fig" captionStartId="158.[189,232,759,785]" captionTargetBox="[189,1384,274,714]" captionTargetId="figure-14@158.[189,1398,262,714]" captionTargetPageId="158" captionText="Fig. 44. Pseudonannolene alegrensis Silvestri, 1897 (MCN 626), in lateral view. A. Anterior region. B. Posterior region. Scale bars: A = 0.5 mm; B = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7891131" httpUri="https://zenodo.org/record/7891131/files/figure.png" pageId="20" pageNumber="21">Fig. 44B</figureCitation>
).
</paragraph>
<paragraph id="8BEE36AC1327B1754FEBFA98FD4F50CA" blockId="20.[189,1398,1395,1524]" pageId="20" pageNumber="21">
<smallCapsWord id="8D08A0701327B1754FEBFA98FF6050B2" baselines="1415,1415" box="[189,250,1396,1422]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="First" pageId="20" pageNumber="21">FIRST</smallCapsWord>
<smallCapsWord id="8D08A0701327B1754E53FA95FEFE50B2" baselines="1415,1415,1415" box="[261,356,1396,1422]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="leg-pair" pageId="20" pageNumber="21">LEG-PAIR</smallCapsWord>
<smallCapsWord id="8D08A0701327B1754E39FA95FE1350B2" baselines="1415" box="[367,393,1401,1420]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="of" pageId="20" pageNumber="21">OF</smallCapsWord>
<smallCapsWord id="8D08A0701327B1754EC2FA95FE4050B2" baselines="1415" box="[404,474,1401,1420]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="males" pageId="20" pageNumber="21">MALES</smallCapsWord>
. Coxae (
<emphasis id="B925EABE1327B1754D1DFA98FDFD50B0" bold="true" box="[587,615,1396,1422]" italics="true" pageId="20" pageNumber="21">cx</emphasis>
) elongated (as long as the sum of remaining podomere lengths), subtriangular, with the base arched, densely setose (
<figureCitation id="136A2A291327B1754C6FFA79FC30508E" box="[825,938,1429,1456]" captionStart="Fig" captionStartId="159.[189,230,1666,1692]" captionTargetBox="[189,1398,267,1624]" captionTargetId="figure-11@159.[189,1398,267,1624]" captionTargetPageId="159" captionText="Fig. 45. Pseudonannolene alegrensis Silvestri, 1897, neotype, ♂ (MCN 626). A. First leg-pair. B. Detail of prefemur. C. Second leg-pair. D. Detail of telopodite, in anal view. E. Left gonopod, in anal view. F. Left gonopod, in oral view.Abbreviations: see Material and methods. Scale bars:A, C = 0.5 mm; B, D–F = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7891137" httpUri="https://zenodo.org/record/7891137/files/figure.png" pageId="20" pageNumber="21">Fig. 45A</figureCitation>
); prefemoral process (
<emphasis id="B925EABE1327B1754BEBFA79FB7B5091" bold="true" box="[1213,1249,1429,1455]" italics="true" pageId="20" pageNumber="21">prf</emphasis>
) as wide as half width of prefemur, subcylindrical, densely setose up to its median region (
<figureCitation id="136A2A291327B1754B2FFA5BFB7D50EC" box="[1145,1255,1463,1490]" captionStart="Fig" captionStartId="159.[189,230,1666,1692]" captionTargetBox="[189,1398,267,1624]" captionTargetId="figure-11@159.[189,1398,267,1624]" captionTargetPageId="159" captionText="Fig. 45. Pseudonannolene alegrensis Silvestri, 1897, neotype, ♂ (MCN 626). A. First leg-pair. B. Detail of prefemur. C. Second leg-pair. D. Detail of telopodite, in anal view. E. Left gonopod, in anal view. F. Left gonopod, in oral view.Abbreviations: see Material and methods. Scale bars:A, C = 0.5 mm; B, D–F = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7891137" httpUri="https://zenodo.org/record/7891137/files/figure.png" pageId="20" pageNumber="21">Fig. 45B</figureCitation>
); remaining podomeres with setae along the mesal region.
</paragraph>
<paragraph id="8BEE36AC1327B1754FEBF9F6FE4D5346" blockId="20.[188,1398,1562,1657]" pageId="20" pageNumber="21">
<smallCapsWord id="8D08A0701327B1754FEBF9F6FE8E530D" baselines="1581,1582" box="[189,276,1562,1588]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Second" pageId="20" pageNumber="21">SECOND</smallCapsWord>
<smallCapsWord id="8D08A0701327B1754E4AF9CCFEE1530D" baselines="1582,1582,1582" box="[284,379,1563,1589]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="leg-pair" pageId="20" pageNumber="21">LEG-PAIR</smallCapsWord>
<smallCapsWord id="8D08A0701327B1754ED4F9CCFE06530D" baselines="1582" box="[386,412,1568,1587]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="of" pageId="20" pageNumber="21">OF</smallCapsWord>
<smallCapsWord id="8D08A0701327B1754EF5F9CCFE73530D" baselines="1582" box="[419,489,1568,1587]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="males" pageId="20" pageNumber="21">MALES</smallCapsWord>
. Coxa (
<emphasis id="B925EABE1327B1754D10F9F6FDF8530A" bold="true" box="[582,610,1562,1588]" italics="true" pageId="20" pageNumber="21">cx</emphasis>
) large and rounded; penis (
<emphasis id="B925EABE1327B1754CF6F9F6FC25530A" bold="true" box="[928,959,1562,1588]" italics="true" pageId="20" pageNumber="21">pn</emphasis>
) located at proximal region, rounded, not extended basally (
<figureCitation id="136A2A291327B1754E9EF9D0FDA25369" box="[456,568,1596,1623]" captionStart="Fig" captionStartId="159.[189,230,1666,1692]" captionTargetBox="[189,1398,267,1624]" captionTargetId="figure-11@159.[189,1398,267,1624]" captionTargetPageId="159" captionText="Fig. 45. Pseudonannolene alegrensis Silvestri, 1897, neotype, ♂ (MCN 626). A. First leg-pair. B. Detail of prefemur. C. Second leg-pair. D. Detail of telopodite, in anal view. E. Left gonopod, in anal view. F. Left gonopod, in oral view.Abbreviations: see Material and methods. Scale bars:A, C = 0.5 mm; B, D–F = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7891137" httpUri="https://zenodo.org/record/7891137/files/figure.png" pageId="20" pageNumber="21">Fig. 45C</figureCitation>
); prefemur compressed dorsoventrally; remaining podomeres setose, with long setae mesally.
</paragraph>
<paragraph id="8BEE36AC1327B1754FEBF973FAC152B8" blockId="20.[189,1399,1695,1926]" pageId="20" pageNumber="21">
<smallCapsWord id="8D08A0701327B1754FEBF973FEAE5389" baselines="1714,1714" box="[189,308,1695,1721]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Gonopods" pageId="20" pageNumber="21">GONOPODS</smallCapsWord>
. Gonocoxa (
<emphasis id="B925EABE1327B1754E84F973FE665387" bold="true" box="[466,508,1695,1721]" italics="true" pageId="20" pageNumber="21">gcx</emphasis>
) elongated, but less than twice the length of the telopodite, with the base arched; slightly flattened antero-posteriorly (
<figureCitation id="136A2A291327B1754DB7F92DFCEF53E2" box="[737,885,1729,1756]" captionStart="Fig" captionStartId="159.[189,230,1666,1692]" captionTargetBox="[189,1398,267,1624]" captionTargetId="figure-11@159.[189,1398,267,1624]" captionTargetPageId="159" captionText="Fig. 45. Pseudonannolene alegrensis Silvestri, 1897, neotype, ♂ (MCN 626). A. First leg-pair. B. Detail of prefemur. C. Second leg-pair. D. Detail of telopodite, in anal view. E. Left gonopod, in anal view. F. Left gonopod, in oral view.Abbreviations: see Material and methods. Scale bars:A, C = 0.5 mm; B, D–F = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7891137" httpUri="https://zenodo.org/record/7891137/files/figure.png" pageId="20" pageNumber="21">Fig. 45D–F</figureCitation>
); with rows of papillae mesally. Seminal groove (
<emphasis id="B925EABE1327B1754E49F90FFEA353C3" bold="true" box="[287,313,1763,1789]" italics="true" pageId="20" pageNumber="21">sg</emphasis>
) curved; arising medially on mesal cavity and terminating apically on the seminal apophysis (
<emphasis id="B925EABE1327B1754F91F8E9FF7B5221" bold="true" box="[199,225,1797,1823]" italics="true" pageId="20" pageNumber="21">sa</emphasis>
). Shoulder (
<emphasis id="B925EABE1327B1754E2CF8E9FE0C521E" bold="true" box="[378,406,1797,1824]" italics="true" pageId="20" pageNumber="21">sh</emphasis>
) inconspicuous. Telopodite (
<emphasis id="B925EABE1327B1754DA5F8E9FC905221" bold="true" box="[755,778,1797,1823]" italics="true" pageId="20" pageNumber="21">tp</emphasis>
) elongated, trunk stout (
<figureCitation id="136A2A291327B1754B60F8E9FB3C521E" box="[1078,1190,1797,1824]" captionStart="Fig" captionStartId="159.[189,230,1666,1692]" captionTargetBox="[189,1398,267,1624]" captionTargetId="figure-11@159.[189,1398,267,1624]" captionTargetPageId="159" captionText="Fig. 45. Pseudonannolene alegrensis Silvestri, 1897, neotype, ♂ (MCN 626). A. First leg-pair. B. Detail of prefemur. C. Second leg-pair. D. Detail of telopodite, in anal view. E. Left gonopod, in anal view. F. Left gonopod, in oral view.Abbreviations: see Material and methods. Scale bars:A, C = 0.5 mm; B, D–F = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7891137" httpUri="https://zenodo.org/record/7891137/files/figure.png" pageId="20" pageNumber="21">Fig. 45D</figureCitation>
); solenomere (
<emphasis id="B925EABE1327B1754A0FF8E9FAF7521E" bold="true" box="[1369,1389,1797,1824]" italics="true" pageId="20" pageNumber="21">sl</emphasis>
) with apicomesal process (
<emphasis id="B925EABE1327B1754EBBF8C4FDBB527C" bold="true" box="[493,545,1832,1858]" italics="true" pageId="20" pageNumber="21">amp</emphasis>
) subtriangular; ectal process absent;
<emphasis id="B925EABE1327B1754C82F8C4FC74527C" bold="true" box="[980,1006,1832,1858]" italics="true" pageId="20" pageNumber="21">sa</emphasis>
located at medial portion, visible apically. Internal branch (
<emphasis id="B925EABE1327B1754EBAF8A5FD99525A" bold="true" box="[492,515,1865,1892]" italics="true" pageId="20" pageNumber="21">ib</emphasis>
) subtriangular, narrow, curved ectad at midlength, surrounding the base of
<emphasis id="B925EABE1327B1754FE8F880FF4E52B8" bold="true" box="[190,212,1900,1926]" italics="true" pageId="20" pageNumber="21">tp</emphasis>
;
<emphasis id="B925EABE1327B1754FB5F887FF6052B8" bold="true" box="[227,250,1899,1926]" italics="true" pageId="20" pageNumber="21">ib</emphasis>
with setae along its entire margin not exceeding apically the seminal region of
<emphasis id="B925EABE1327B1754BCBF887FB2B52B8" bold="true" box="[1181,1201,1899,1926]" italics="true" pageId="20" pageNumber="21">sl</emphasis>
(
<figureCitation id="136A2A291327B1754B97F887FAD552B8" box="[1217,1359,1899,1926]" captionStart="Fig" captionStartId="159.[189,230,1666,1692]" captionTargetBox="[189,1398,267,1624]" captionTargetId="figure-11@159.[189,1398,267,1624]" captionTargetPageId="159" captionText="Fig. 45. Pseudonannolene alegrensis Silvestri, 1897, neotype, ♂ (MCN 626). A. First leg-pair. B. Detail of prefemur. C. Second leg-pair. D. Detail of telopodite, in anal view. E. Left gonopod, in anal view. F. Left gonopod, in oral view.Abbreviations: see Material and methods. Scale bars:A, C = 0.5 mm; B, D–F = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7891137" httpUri="https://zenodo.org/record/7891137/files/figure.png" pageId="20" pageNumber="21">Fig. 45D–F</figureCitation>
).
</paragraph>
<paragraph id="8BEE36AC1326B1744FEBFEE5FAD65478" blockId="21.[188,1398,265,326]" pageId="21" pageNumber="22">
<smallCapsWord id="8D08A0701326B1744FEBFEE5FE8E541C" baselines="284,285" box="[189,276,265,291]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Vulvae" pageId="21" pageNumber="22">VULVAE</smallCapsWord>
. As typical for the genus. Bursa subtriangular, glabrous (
<figureCitation id="136A2A291326B1744CF2FEE5FB87541D" box="[932,1053,265,291]" captionStart="Fig" captionStartId="280.[189,232,1787,1813]" captionTargetBox="[250,1337,265,1739]" captionTargetId="figure-14@280.[250,1337,265,1739]" captionTargetPageId="280" captionText="Fig. 177. Stereoscopic images of left vulva of species of Pseudonannolene Silvestri, 1895, in oral view. A. P. albiventris Schubart, 1952. B. P. alegrensis Silvestri, 1897. C. P. ambuatinga Iniesta &amp; Ferreira, 2013. D. P. anapophysis Fontanetti, 1996. E. P. buhrnheimi Schubart, 1960. F. P. caatinga Iniesta &amp; Ferreira, 2014. G. P. callipyge Brölemann, 1902. H. P. erikae Iniesta &amp; Ferreira, 2014. I. P. fontanettiae Iniesta &amp; Ferreira, 2014. J. P. halophila Schubart, 1949. K. P. imbirensis Fontanetti, 1996. L. P. inops Brölemann, 1929. M. P. leucocephalus Schubart, 1944. N. P. longicornis (Porat, 1888). O. P. lundi Iniesta &amp; Ferreira, 2015. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.7891439" httpUri="https://zenodo.org/record/7891439/files/figure.png" pageId="21" pageNumber="22">Fig. 177B</figureCitation>
); internal valve subtriangular, with mesal region rounded; operculum narrow, curved medially; external valve wide, subtriangular.
</paragraph>
</subSubSection>
<subSubSection id="C34B65271326B1744FEBFE80FB2E548B" box="[189,1204,364,437]" pageId="21" pageNumber="22" type="distribution">
<paragraph id="8BEE36AC1326B1744FEBFE80FECC54B8" blockId="21.[189,342,364,390]" box="[189,342,364,390]" pageId="21" pageNumber="22">
<heading id="D0A681C01326B1744FEBFE80FECC54B8" bold="true" box="[189,342,364,390]" fontSize="11" level="3" pageId="21" pageNumber="22" reason="3">
<emphasis id="B925EABE1326B1744FEBFE80FECC54B8" bold="true" box="[189,342,364,390]" pageId="21" pageNumber="22">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BEE36AC1326B1744FEBFE77FB2E548B" blockId="21.[189,1204,410,437]" box="[189,1204,410,437]" pageId="21" pageNumber="22">
Known only from the
<typeStatus id="54EA880E1326B1744E94FE77FE6F548B" box="[450,501,411,437]" pageId="21" pageNumber="22">type</typeStatus>
locality Porto Alegre,
<collectingRegion id="4995F84E1326B1744DA9FE76FC47548A" box="[767,989,410,436]" country="Brazil" name="Rio Grande do Sul" pageId="21" pageNumber="22">Rio Grande do Sul</collectingRegion>
,
<collectingCountry id="F346763C1326B1744CBEFE76FBAA548A" box="[1000,1072,410,436]" name="Brazil" pageId="21" pageNumber="22">Brazil</collectingCountry>
(
<figureCitation id="136A2A291326B1744B16FE76FB3D548B" box="[1088,1191,410,437]" captionStart="Fig" captionStartId="283.[189,232,1363,1389]" captionTargetBox="[192,1368,285,1297]" captionTargetId="figure-12@283.[189,1398,265,1328]" captionTargetPageId="283" captionText="Fig. 180. Distribution map of the species Pseudonannolene albiventris Schubart, 1952, P. alegrensis Silvestri, 1897, P. ambuatinga Iniesta &amp; Ferreira, 2013, P. anapophysis Fontanetti, 1996, and P. borelli Silvestri, 1895." figureDoi="http://doi.org/10.5281/zenodo.7891445" httpUri="https://zenodo.org/record/7891445/files/figure.png" pageId="21" pageNumber="22">Fig. 180</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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