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<document ID-DOI="http://doi.org/10.5281/zenodo.5575685" ID-GBIF-Dataset="887f4c2c-1812-4aa5-b994-58388f6a45c5" ID-GBIF-Taxon="131977836" ID-Zenodo-Dep="5575685" checkinTime="1497949082728" checkinUser="jeremy" docAuthor="Deeleman-Reinhold, Christa" docDate="2001" docId="03B3FE72C97FFFA664F289AEDE5EF8F8" docLanguage="en" docName="Deeleman-Reinhold_2001_LIOCRANIDAE_mat_cite_checked_IV.imf" docOrigin="Forest Spiders of South East Asia With a revision of the sac and ground spiders- Family Liocranidae, Leiden, Netherlands: Brill Leiden; Boston; Köln" docStyle="DocumentStyle{}" docTitle="Jacaena Thorell" docType="treatment" docVersion="21" lastPageNumber="465" masterDocId="FF8A860AC93EFFE765528965DD50FFD6" masterDocTitle="Forest Spiders of South East Asia With a revision of the sac and ground spiders (Araneae: Clubionidae, Corinnidae, Liocranidae, Gnaphosidae, Prodidomidae and Trochanteriidae)." masterLastPageNumber="505" masterPageNumber="400" pageId="65" pageNumber="465" updateTime="1634566706600" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Forest Spiders of South East Asia With a revision of the sac and ground spiders (Araneae: Clubionidae, Corinnidae, Liocranidae, Gnaphosidae, Prodidomidae and Trochanteriidae).</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Deeleman-Reinhold, Christa</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:originInfo>
<mods:dateIssued>2001</mods:dateIssued>
<mods:publisher>Brill Leiden; Boston; Köln</mods:publisher>
<mods:place>
<mods:placeTerm>Leiden, Netherlands</mods:placeTerm>
</mods:place>
</mods:originInfo>
<mods:titleInfo>
<mods:title>Forest Spiders of South East Asia With a revision of the sac and ground spiders- Family Liocranidae</mods:title>
</mods:titleInfo>
<mods:part>
<mods:extent unit="page">
<mods:start>400</mods:start>
<mods:end>505</mods:end>
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<mods:classification>book chapter</mods:classification>
<mods:identifier type="DOI">10.5281/zenodo.814704</mods:identifier>
<mods:identifier type="GBIF-Dataset">887f4c2c-1812-4aa5-b994-58388f6a45c5</mods:identifier>
<mods:identifier type="Zenodo-Dep">814704</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.5575685" ID-GBIF-Taxon="131977836" ID-Zenodo-Dep="5575685" LSID="urn:lsid:plazi:treatment:03B3FE72C97FFFA664F289AEDE5EF8F8" httpUri="http://treatment.plazi.org/id/03B3FE72C97FFFA664F289AEDE5EF8F8" lastPageNumber="465" pageId="65" pageNumber="465">
<subSubSection box="[416,730,203,231]" pageId="65" pageNumber="465" type="nomenclature">
<paragraph blockId="65.[416,731,201,231]" box="[416,730,203,231]" pageId="65" pageNumber="465">
<heading box="[416,730,203,231]" centered="true" fontSize="14" level="1" pageId="65" pageNumber="465" reason="1">
Genus
<taxonomicName authority="Thorell" authorityName="Thorell" box="[509,622,203,231]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="65" pageNumber="465" phylum="Arthropoda" rank="genus">
<emphasis box="[509,622,203,231]" italics="true" pageId="65" pageNumber="465">Jacaena</emphasis>
</taxonomicName>
Thorell
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="65" pageNumber="465" type="reference_group">
<paragraph blockId="65.[44,1102,266,328]" pageId="65" pageNumber="465">
<taxonomicName authority="Thorell, 1897: 230" authorityName="Thorell" authorityPageNumber="230" authorityYear="1897" box="[44,369,269,293]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="65" pageNumber="465" phylum="Arthropoda" rank="genus">
<emphasis box="[44,139,269,293]" italics="true" pageId="65" pageNumber="465">Jacaena</emphasis>
Thorell, 1897: 230
</taxonomicName>
. Type species:
<taxonomicName authority="Thorell, 1897" authorityName="Thorell" authorityYear="1897" box="[546,843,269,293]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="65" pageNumber="465" phylum="Arthropoda" rank="species" species="distincta">
<emphasis box="[546,672,269,293]" italics="true" pageId="65" pageNumber="465">J. distincta</emphasis>
Thorell, 1897
</taxonomicName>
, no figures, ♀. Burma, Tenasserim.
</paragraph>
</subSubSection>
<subSubSection pageId="65" pageNumber="465" type="discussion">
<paragraph blockId="65.[44,1106,358,1360]" pageId="65" pageNumber="465">
This genus was described by Thorell (date of issue
<date box="[658,837,361,385]" pageId="65" pageNumber="465" value="1897-01-02">2 january 1897</date>
) in the
<taxonomicName box="[931,1072,361,385]" class="Arachnida" family="Clubionidae" kingdom="Animalia" order="Araneae" pageId="65" pageNumber="465" phylum="Arthropoda" rank="family">Clubionidae</taxonomicName>
in the subfamily Thamphilinae (=Sphingiinae). This subfamily was composed of the genera
<emphasis box="[46,183,431,455]" italics="true" pageId="65" pageNumber="465">Thamphilus</emphasis>
,
<taxonomicName box="[203,317,431,455]" class="Arachnida" family="Liocranidae" genus="Sphingius" kingdom="Animalia" order="Araneae" pageId="65" pageNumber="465" phylum="Arthropoda" rank="genus">
<emphasis box="[203,317,431,455]" italics="true" pageId="65" pageNumber="465">Sphingius</emphasis>
</taxonomicName>
and
<emphasis box="[386,641,431,455]" italics="true" pageId="65" pageNumber="465">
<taxonomicName box="[386,486,431,455]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="65" pageNumber="465" phylum="Arthropoda" rank="genus">Jacaena</taxonomicName>
. Thamphilus
</emphasis>
Thorell 1895 was soon synonymised with
<taxonomicName box="[104,217,465,489]" class="Arachnida" family="Liocranidae" genus="Sphingius" kingdom="Animalia" order="Araneae" pageId="65" pageNumber="465" phylum="Arthropoda" rank="genus">
<emphasis box="[104,217,465,489]" italics="true" pageId="65" pageNumber="465">Sphingius</emphasis>
</taxonomicName>
(Simon, 1897,
<collectionCode box="[402,492,465,489]" pageId="65" pageNumber="465">HNAII</collectionCode>
: 158). Apparently, Simon did not see any specimens of
<emphasis box="[76,178,500,524]" italics="true" pageId="65" pageNumber="465">
<taxonomicName box="[76,174,500,524]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="65" pageNumber="465" phylum="Arthropoda" rank="genus">Jacaena</taxonomicName>
,
</emphasis>
but he did somehow receive Thorells data before the first part of HNAII went to press. Consequently, he mentioned the genus only in a footnote (1897, HNA
<collectionCode box="[1017,1037,535,559]" pageId="65" pageNumber="465">II</collectionCode>
: 157, date of issue
<date box="[217,427,570,594]" pageId="65" pageNumber="465" value="1897-02-15">15 february 1897</date>
), where he placed it close to
<emphasis box="[802,922,570,594]" italics="true" pageId="65" pageNumber="465">
<taxonomicName box="[802,918,570,594]" class="Arachnida" family="Liocranidae" genus="Sphingius" kingdom="Animalia" order="Araneae" pageId="65" pageNumber="465" phylum="Arthropoda" rank="genus">Sphingius</taxonomicName>
.
</emphasis>
Simon ranged
<taxonomicName box="[46,159,605,629]" class="Arachnida" family="Liocranidae" genus="Sphingius" kingdom="Animalia" order="Araneae" pageId="65" pageNumber="465" phylum="Arthropoda" rank="genus">
<emphasis box="[46,159,605,629]" italics="true" pageId="65" pageNumber="465">Sphingius</emphasis>
</taxonomicName>
erroneously in the Molycriae, a tribe he wrongly believed to belong to the subfamily
<taxonomicName box="[133,258,640,664]" class="Arachnida" family="Gnaphosidae" kingdom="Animalia" order="Araneae" pageId="65" pageNumber="465" phylum="Arthropoda" rank="subFamily" subFamily="Micariinae">Micariinae</taxonomicName>
and which originally included
<taxonomicName box="[645,750,640,664]" class="Arachnida" family="Prodidomidae" genus="Molycria" kingdom="Animalia" order="Araneae" pageId="65" pageNumber="465" phylum="Arthropoda" rank="genus">
<emphasis box="[645,750,640,664]" italics="true" pageId="65" pageNumber="465">Molycria</emphasis>
</taxonomicName>
and
<emphasis box="[817,930,640,664]" italics="true" pageId="65" pageNumber="465">
<taxonomicName box="[817,926,640,664]" class="Arachnida" family="Prodidomidae" genus="Myandra" kingdom="Animalia" order="Araneae" pageId="65" pageNumber="465" phylum="Arthropoda" rank="genus">Myandra</taxonomicName>
.
</emphasis>
Bonnet 1959, (2[5]:4395) recognised the genus
<emphasis box="[452,594,674,698]" italics="true" pageId="65" pageNumber="465">Thamphilus,</emphasis>
but subsequent cataloguers Roewer, Brignoli and Platnick followed Simon in synonymising
<emphasis box="[650,786,709,733]" italics="true" pageId="65" pageNumber="465">Thamphilus</emphasis>
with
<emphasis box="[854,975,709,733]" italics="true" pageId="65" pageNumber="465">
<taxonomicName box="[854,971,709,733]" class="Arachnida" family="Liocranidae" genus="Sphingius" kingdom="Animalia" order="Araneae" pageId="65" pageNumber="465" phylum="Arthropoda" rank="genus">Sphingius</taxonomicName>
.
</emphasis>
Today, the
<taxonomicName box="[46,171,744,768]" class="Arachnida" family="Gnaphosidae" kingdom="Animalia" order="Araneae" pageId="65" pageNumber="465" phylum="Arthropoda" rank="subFamily" subFamily="Micariinae">Micariinae</taxonomicName>
are mainly defined by the apomorphic iridescent scales on the abdomen, which alienate them from all groups treated here. It is now clear that for
<taxonomicName box="[833,946,779,803]" class="Arachnida" family="Liocranidae" genus="Sphingius" kingdom="Animalia" order="Araneae" pageId="65" pageNumber="465" phylum="Arthropoda" rank="genus">
<emphasis box="[833,946,779,803]" italics="true" pageId="65" pageNumber="465">Sphingius</emphasis>
</taxonomicName>
and
<taxonomicName box="[1007,1102,779,803]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="65" pageNumber="465" phylum="Arthropoda" rank="genus">
<emphasis box="[1007,1102,779,803]" italics="true" pageId="65" pageNumber="465">Jacaena</emphasis>
</taxonomicName>
there is no close relationship at all with either
<taxonomicName box="[616,721,813,837]" class="Arachnida" family="Prodidomidae" genus="Myandra" kingdom="Animalia" order="Araneae" pageId="65" pageNumber="465" phylum="Arthropoda" rank="genus">
<emphasis box="[616,721,813,837]" italics="true" pageId="65" pageNumber="465">Myandra</emphasis>
</taxonomicName>
or
<emphasis box="[769,881,813,837]" italics="true" pageId="65" pageNumber="465">
<taxonomicName box="[769,878,813,837]" class="Arachnida" family="Prodidomidae" genus="Molycria" kingdom="Animalia" order="Araneae" pageId="65" pageNumber="465" phylum="Arthropoda" rank="genus">Molycria</taxonomicName>
.
</emphasis>
Simons reassessment of the taxonomic position of both these genera was very much beside the mark. The genus
<taxonomicName box="[122,221,883,907]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="65" pageNumber="465" phylum="Arthropoda" rank="genus">
<emphasis box="[122,221,883,907]" italics="true" pageId="65" pageNumber="465">Jacaena</emphasis>
</taxonomicName>
: “genre....différant surtout des autres (Molycriae) par ses tibias antérieurs armés de deux séries d'épines”. Roewer (1954) followed Simon; Reiskind (1969: 166) moved
<taxonomicName box="[134,248,953,977]" class="Arachnida" family="Liocranidae" genus="Sphingius" kingdom="Animalia" order="Araneae" pageId="65" pageNumber="465" phylum="Arthropoda" rank="genus">
<emphasis box="[134,248,953,977]" italics="true" pageId="65" pageNumber="465">Sphingius</emphasis>
</taxonomicName>
to the
<taxonomicName box="[338,467,953,977]" class="Arachnida" family="Corinnidae" kingdom="Animalia" order="Araneae" pageId="65" pageNumber="465" phylum="Arthropoda" rank="subFamily" subFamily="Corinninae">Corinninae</taxonomicName>
and moved the bulk of
<taxonomicName box="[762,888,953,977]" class="Arachnida" family="Gnaphosidae" kingdom="Animalia" order="Araneae" pageId="65" pageNumber="465" phylum="Arthropoda" rank="subFamily" subFamily="Micariinae">Micariinae</taxonomicName>
as a whole to the
<taxonomicName box="[46,200,988,1012]" class="Arachnida" family="Gnaphosidae" kingdom="Animalia" order="Araneae" pageId="65" pageNumber="465" phylum="Arthropoda" rank="family">Gnaphosidae</taxonomicName>
, and in this latter family
<taxonomicName box="[501,597,988,1012]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="65" pageNumber="465" phylum="Arthropoda" rank="genus">
<emphasis box="[501,597,988,1012]" italics="true" pageId="65" pageNumber="465">Jacaena</emphasis>
</taxonomicName>
remains in Platnick 1989,
<date box="[927,1100,988,1012]" pageId="65" pageNumber="465" value="1991" valueMax="1998">1991 and 1998</date>
. During my stay in Genova
<date box="[402,587,1022,1046]" pageId="65" pageNumber="465" value="1986" valueMax="1989">1986 and 1989</date>
, I found there was no specimen labelled
<taxonomicName box="[44,248,1057,1081]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="65" pageNumber="465" phylum="Arthropoda" rank="species" species="distincta">
<emphasis box="[44,248,1057,1081]" italics="true" pageId="65" pageNumber="465">Jacaena distincta</emphasis>
</taxonomicName>
in existence. However there was one female labelled
<emphasis box="[884,1103,1057,1081]" italics="true" pageId="65" pageNumber="465">Cadicus distinctus,</emphasis>
agreeing fully with the description of
<emphasis box="[514,653,1092,1116]" italics="true" pageId="65" pageNumber="465">
<taxonomicName box="[514,649,1092,1116]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="65" pageNumber="465" phylum="Arthropoda" rank="species" species="distincta">J. distincta</taxonomicName>
,
</emphasis>
including the unusual pattern of the upper- and underside of the abdomen, and which was taken at the same locality Tenasserim: Mooleyit. No species by the generic name of
<emphasis box="[767,860,1161,1185]" italics="true" pageId="65" pageNumber="465">Cadicus</emphasis>
has ever been published. I consider this case as an inadvertent omission by Thorell to adjust the name on the label to that in the description and I recognise the specimen as the type of
<emphasis italics="true" pageId="65" pageNumber="465">
<taxonomicName box="[1001,1099,1231,1255]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="65" pageNumber="465" phylum="Arthropoda" rank="genus">Jacaena</taxonomicName>
.
<taxonomicName box="[45,250,1266,1290]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="65" pageNumber="465" phylum="Arthropoda" rank="species" species="distincta">Jacaena distincta</taxonomicName>
</emphasis>
is closely related to
<emphasis box="[495,608,1266,1290]" italics="true" pageId="65" pageNumber="465">
<taxonomicName box="[495,604,1266,1290]" class="Arachnida" family="Liocranidae" genus="Sesieutes" kingdom="Animalia" order="Araneae" pageId="65" pageNumber="465" phylum="Arthropoda" rank="genus">Sesieutes</taxonomicName>
.
</emphasis>
The principal distinction between the two is in the extraordinary structure of the vulva, which is shared with
<taxonomicName box="[913,1019,1301,1325]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="65" pageNumber="465" phylum="Arthropoda" rank="species" species="mihun" status="sp. n.">
<emphasis box="[913,1019,1301,1325]" italics="true" pageId="65" pageNumber="465">J. mihun</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1037,1105,1301,1325]" pageId="65" pageNumber="465" rank="species">sp. n.</taxonomicNameLabel>
described below. This argument is used here to maintain
<taxonomicName box="[722,817,1335,1359]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="65" pageNumber="465" phylum="Arthropoda" rank="genus">
<emphasis box="[722,817,1335,1359]" italics="true" pageId="65" pageNumber="465">Jacaena</emphasis>
</taxonomicName>
as a valid genus.
</paragraph>
<paragraph blockId="65.[46,1104,1393,1453]" pageId="65" pageNumber="465">
<emphasis box="[47,263,1396,1420]" italics="true" pageId="65" pageNumber="465">Described species</emphasis>
.—
<taxonomicName authority="Thorell, 1897" authorityName="Thorell" authorityYear="1897" box="[309,700,1396,1420]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="65" pageNumber="465" phylum="Arthropoda" rank="species" species="distincta">
<emphasis box="[309,519,1396,1420]" italics="true" pageId="65" pageNumber="465">Jacaena distincta</emphasis>
Thorell, 1897
</taxonomicName>
, ♀, Myanmar (Burma);
<emphasis box="[1000,1019,1396,1420]" italics="true" pageId="65" pageNumber="465">J.</emphasis>
<taxonomicName box="[1033,1103,1396,1420]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="65" pageNumber="465" phylum="Arthropoda" rank="species" species="mihun" status="sp. n.">mihun</taxonomicName>
<taxonomicNameLabel box="[46,110,1429,1453]" pageId="65" pageNumber="465" rank="species">
sp.
<emphasis box="[88,105,1429,1453]" italics="true" pageId="65" pageNumber="465">n</emphasis>
.
</taxonomicNameLabel>
, ♂♀, C and N Thailand, this volume.
</paragraph>
</subSubSection>
<subSubSection pageId="65" pageNumber="465" type="diagnosis">
<paragraph blockId="65.[46,1107,1486,1651]" pageId="65" pageNumber="465">
<emphasis box="[46,164,1489,1513]" italics="true" pageId="65" pageNumber="465">Diagnosis</emphasis>
.—
<taxonomicName box="[206,302,1489,1513]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="65" pageNumber="465" phylum="Arthropoda" rank="genus">
<emphasis box="[206,302,1489,1513]" italics="true" pageId="65" pageNumber="465">Jacaena</emphasis>
</taxonomicName>
is similar in most respects to
<emphasis box="[661,774,1489,1513]" italics="true" pageId="65" pageNumber="465">
<taxonomicName box="[661,770,1489,1513]" class="Arachnida" family="Liocranidae" genus="Sesieutes" kingdom="Animalia" order="Araneae" pageId="65" pageNumber="465" phylum="Arthropoda" rank="genus">Sesieutes</taxonomicName>
.
</emphasis>
It is distinct from
<taxonomicName box="[999,1104,1489,1513]" class="Arachnida" family="Liocranidae" genus="Sesieutes" kingdom="Animalia" order="Araneae" pageId="65" pageNumber="465" phylum="Arthropoda" rank="genus">
<emphasis box="[999,1104,1489,1513]" italics="true" pageId="65" pageNumber="465">Sesieutes</emphasis>
</taxonomicName>
in the construction of the vulva: this consists of a pair of seemingly non-differentiated, meandering and coiling tubes without distinct spermathecae. In the type species the male is unknown; in
<taxonomicName box="[231,327,1592,1616]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="65" pageNumber="465" phylum="Arthropoda" rank="genus">
<emphasis box="[231,327,1592,1616]" italics="true" pageId="65" pageNumber="465">Jacaena</emphasis>
</taxonomicName>
<taxonomicName box="[337,407,1592,1616]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="65" pageNumber="465" phylum="Arthropoda" rank="species" species="mihun">mihun</taxonomicName>
males, described below, the embolus is transformed into a short broadened rigid appendage. The clypeal lobe is reduced.
</paragraph>
<paragraph blockId="65.[47,965,1687,1714]" box="[48,965,1690,1714]" pageId="65" pageNumber="465">
<emphasis box="[48,91,1690,1714]" italics="true" pageId="65" pageNumber="465">Old</emphasis>
material
<emphasis box="[207,291,1690,1714]" italics="true" pageId="65" pageNumber="465">studied</emphasis>
.—
<emphasis box="[333,545,1690,1714]" italics="true" pageId="65" pageNumber="465">
<taxonomicName box="[333,541,1690,1714]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="65" pageNumber="465" phylum="Arthropoda" rank="species" species="distincta">Jacaena distincta</taxonomicName>
,
</emphasis>
1 ♀, Burma, Tenasserim,
<collectionCode box="[857,961,1690,1714]" collectionName="MCSNG" pageId="65" pageNumber="465">MCSNG</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection box="[46,484,1752,1776]" pageId="65" pageNumber="465" type="biology_ecology">
<paragraph blockId="65.[46,484,1749,1776]" box="[46,484,1752,1776]" pageId="65" pageNumber="465">
<heading box="[46,484,1752,1776]" fontSize="12" level="5" pageId="65" pageNumber="465" reason="1">
<emphasis box="[46,270,1752,1776]" italics="true" pageId="65" pageNumber="465">Data on Behaviour</emphasis>
.— Ground-living.
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[47,782,1814,1838]" pageId="65" pageNumber="465" type="distribution">
<paragraph blockId="65.[47,782,1812,1838]" box="[47,782,1814,1838]" pageId="65" pageNumber="465">
<emphasis box="[47,189,1814,1838]" italics="true" pageId="65" pageNumber="465">Distribution</emphasis>
.— Southeastern Myanmar (Burma) and Thailand.
</paragraph>
</subSubSection>
</treatment>
</document>