208 lines
18 KiB
XML
208 lines
18 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.878.37403" ID-GBIF-Dataset="83f095d1-8441-49cd-be4b-71f0ddfa311c" ID-PMC="PMC6791902" ID-Pensoft-Pub="1313-2970-878-73" ID-Pensoft-UUID="C93F9CBB14D25FEAB2ACCBC3E6282D28" ID-PubMed="31632177" ID-ZooBank="192214DE1D38467BA577ECD16EC5EAB5" ModsDocID="1313-2970-878-73" checkinTime="1570497196116" checkinUser="pensoft" docAuthor="Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo & Balke, Michael" docDate="2019" docId="757E69DBAB555E01AA5EDD3AA356177C" docLanguage="en" docName="ZooKeys 878: 73-143" docOrigin="ZooKeys 878" docSource="http://dx.doi.org/10.3897/zookeys.878.37403" docTitle="Exocelina jaseminae Balke 1998" docType="treatment" docVersion="6" id="C93F9CBB14D25FEAB2ACCBC3E6282D28" lastPageNumber="73" masterDocId="C93F9CBB14D25FEAB2ACCBC3E6282D28" masterDocTitle="Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae)" masterLastPageNumber="143" masterPageNumber="73" pageNumber="73" updateTime="1668167854570" updateUser="ExternalLinkService">
|
||
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo>
|
||
<mods:title>Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Shaverdo, Helena</mods:namePart>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Surbakti, Suriani</mods:namePart>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Warikar, Evie L.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Sagata, Katayo</mods:namePart>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Balke, Michael</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource>text</mods:typeOfResource>
|
||
<mods:relatedItem type="host">
|
||
<mods:titleInfo>
|
||
<mods:title>ZooKeys</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part>
|
||
<mods:date>2019</mods:date>
|
||
<mods:detail type="volume">
|
||
<mods:number>878</mods:number>
|
||
</mods:detail>
|
||
<mods:extent unit="page">
|
||
<mods:start>73</mods:start>
|
||
<mods:end>143</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location>
|
||
<mods:url>http://dx.doi.org/10.3897/zookeys.878.37403</mods:url>
|
||
</mods:location>
|
||
<mods:classification>journal article</mods:classification>
|
||
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.878.37403</mods:identifier>
|
||
<mods:identifier type="Pensoft-Pub">1313-2970-878-73</mods:identifier>
|
||
<mods:identifier type="ZooBank">192214DE1D38467BA577ECD16EC5EAB5</mods:identifier>
|
||
<mods:identifier type="Pensoft-UUID">C93F9CBB14D25FEAB2ACCBC3E6282D28</mods:identifier>
|
||
</mods:mods>
|
||
<treatment ID-GBIF-Taxon="159654834" LSID="urn:lsid:plazi:treatment:757E69DBAB555E01AA5EDD3AA356177C" httpUri="http://treatment.plazi.org/id/757E69DBAB555E01AA5EDD3AA356177C" lastPageNumber="73" pageId="0" pageNumber="73">
|
||
<subSubSection pageId="0" pageNumber="73" type="nomenclature">
|
||
<paragraph pageId="0" pageNumber="73">
|
||
11.
|
||
<taxonomicName LSID="757E69DB-AB55-5E01-AA5E-DD3AA356177C" authority="Balke, 1998" authorityName="Balke" authorityYear="1998" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina jaseminae" order="Coleoptera" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="jaseminae">Exocelina jaseminae (Balke, 1998)</taxonomicName>
|
||
<figureCitation captionStart="Figures 26–29" captionStartId="F14" captionText="Figures 26 - 29. Habitus and colouration 26 Exocelina aseki sp. nov. 27 E. jaseminae (Balke, 1998) 28 E. kailaki sp. nov. 29 E. pseudojaseminae sp. nov." figureDoi="10.3897/zookeys.878.37403.figures26-29" httpUri="https://binary.pensoft.net/fig/344982" pageId="0" pageNumber="73" tableDoi="10.3897/zookeys.878.37403.figures26-29">Figs 27</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 30, 31" captionStartId="F15" captionText="Figures 30, 31. 30 Exocelina aseki sp. nov. 31 E. jaseminae (Balke, 1998) A median lobe in ventral view B median lobe in lateral view C paramere in external view D male protarsomeres 4 - 5 in ventral view." figureDoi="10.3897/zookeys.878.37403.figures30-31" httpUri="https://binary.pensoft.net/fig/344983" pageId="0" pageNumber="73" tableDoi="10.3897/zookeys.878.37403.figures30-31">31</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="73" type="reference_group">
|
||
<paragraph pageId="0" pageNumber="73">
|
||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus (Papuadytes) jaseminae" order="Coleoptera" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="jaseminae" subGenus="Papuadytes">Copelatus (Papuadytes) jaseminae</taxonomicName>
|
||
Balke, 1998: 331;
|
||
<bibRefCitation pageId="0" pageNumber="73" refId="B9">Nilsson 2001</bibRefCitation>
|
||
: 77 (catalogue).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="73">
|
||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Papuadytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Papuadytes jaseminae" order="Coleoptera" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="jaseminae">Papuadytes jaseminae</taxonomicName>
|
||
(Balke, 1998):
|
||
<bibRefCitation author="Nilsson, AN" journalOrPublisher="Koleopterologische Rundschau" pageId="0" pageNumber="73" pagination="55 - 74" refId="B11" refString="Nilsson, AN, Fery, H, 2006. World Catalogue of Dytiscidae - corrections and additions, 3 (Coleoptera: Dytiscidae). Koleopterologische Rundschau 76: 55 - 74" title="World Catalogue of Dytiscidae - corrections and additions, 3 (Coleoptera: Dytiscidae)." volume="76" year="2006">Nilsson and Fery 2006</bibRefCitation>
|
||
: 56 (comb. nov.).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="73">
|
||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina jaseminae" order="Coleoptera" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="jaseminae">Exocelina jaseminae</taxonomicName>
|
||
(Balke, 1998):
|
||
<bibRefCitation author="Nilsson, AN" journalOrPublisher="Latissimus" pageId="0" pageNumber="73" pagination="33 - 34" refId="B10" refString="Nilsson, AN, 2007. Exocelina Broun, 1886, is the valid name of Papuadytes Balke, 1998. Latissimus 23: 33 - 34" title="Exocelina Broun, 1886, is the valid name of Papuadytes Balke, 1998." volume="23" year="2007">Nilsson 2007</bibRefCitation>
|
||
: 33 (comb. nov.).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="73">
|
||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina jaseminae" order="Coleoptera" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="jaseminae">Exocelina jaseminae</taxonomicName>
|
||
MB1382:
|
||
<bibRefCitation DOI="https://doi.org/10.1038/ncomms5001" author="Toussaint, EFA" journalOrPublisher="Nature Communications" pageId="0" pageNumber="73" pagination="1 - 10" refId="B22" refString="Toussaint, EFA, Hall, R, Monaghan, MT, Sagata, K, Ibalim, S, Shaverdo, HV, Vogler, AP, Pons, J, Balke, M, 2014. The towering orogeny of New Guinea as a trigger for arthropod megadiversity. Nature Communications 1: 1 - 10" title="The towering orogeny of New Guinea as a trigger for arthropod megadiversity." url="https://doi.org/10.1038/ncomms5001" volume="1" year="2014">Toussaint et al. 2014</bibRefCitation>
|
||
: supplementary figs 1-4, tab. 2;
|
||
<bibRefCitation DOI="https://doi.org/10.1038/srep16016" author="Toussaint, EFA" journalOrPublisher="Nature Communications" pageId="0" pageNumber="73" refId="B23" refString="Toussaint, EFA, Henrich, L, Shaverdo, H, Balke, M, 2015. . https://doi.org/10.1038/srep16016" url="https://doi.org/10.1038/srep16016" year="2015">Toussaint et al. 2015</bibRefCitation>
|
||
: supplementary figs S1, S2, tab. S3, and information S5, S6.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="73" type="type locality">
|
||
<paragraph pageId="0" pageNumber="73">Type locality.</paragraph>
|
||
<paragraph pageId="0" pageNumber="73">
|
||
Papua New Guinea: Morobe Province, Herzog Range, Wagau (Vagau), ca.
|
||
<geoCoordinate degrees="06" direction="south" minutes="48" orientation="latitude" precision="925" value="-6.8">06°48'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="146" direction="east" minutes="48" orientation="longitude" precision="925" value="146.8">146°48'E</geoCoordinate>
|
||
, ca. 1300 m a.s.l.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="73" type="type material">
|
||
<paragraph pageId="0" pageNumber="73">Type material studied.</paragraph>
|
||
<paragraph pageId="0" pageNumber="73">
|
||
<emphasis italics="true" pageId="0" pageNumber="73">Holotype</emphasis>
|
||
: male "Stn. No. 150", "NEW GUINEA: Morobe Dist., Herzog Mts., Vagau, C.4,000ft. 4
|
||
<normalizedToken originalValue="–17.i.1965”">-17.i.1965"</normalizedToken>
|
||
, "M. E. Bacchus. B. M. 1965-120",
|
||
<normalizedToken originalValue="“HOLOTYPUS”">"HOLOTYPUS"</normalizedToken>
|
||
[red], "
|
||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus" order="Coleoptera" pageId="0" pageNumber="73" phylum="Arthropoda" rank="genus">Copelatus</taxonomicName>
|
||
<taxonomicName lsidName="jaseminae" pageId="0" pageNumber="73" rank="species" species="jaseminae">jaseminae</taxonomicName>
|
||
sp. nov. Balke des. 1997" [red] (BMNH).
|
||
<emphasis italics="true" pageId="0" pageNumber="73">Paratypes</emphasis>
|
||
: 2 males, 3 females with the same label as the holotype and additionally with a red label "Paratypus
|
||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus" order="Coleoptera" pageId="0" pageNumber="73" phylum="Arthropoda" rank="genus">Copelatus</taxonomicName>
|
||
<taxonomicName lsidName="jaseminae" pageId="0" pageNumber="73" rank="species" species="jaseminae">jaseminae</taxonomicName>
|
||
sp.n. Balke des. 1997", one of the males with an additional label "measured J. Parkin 85" (BMNH, NHMW). 1 male with the same label as the holotype, the red paratype label missing (BMNH).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="73" type="materials_examined">
|
||
<paragraph pageId="0" pageNumber="73">Additional material.</paragraph>
|
||
<paragraph pageId="0" pageNumber="73">
|
||
<emphasis bold="true" pageId="0" pageNumber="73">Morobe</emphasis>
|
||
: 1 male "Stn. No. 149A", "NEW GUINEA: Morobe Dist., Herzog Mts., Vagau, C.4,000ft. 4
|
||
<normalizedToken originalValue="–17.i.1965”">-17.i.1965"</normalizedToken>
|
||
, "M. E. Bacchus. B. M. 1965-120", "Paratypus
|
||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus" order="Coleoptera" pageId="0" pageNumber="73" phylum="Arthropoda" rank="genus">Copelatus</taxonomicName>
|
||
<taxonomicName lsidName="monae" pageId="0" pageNumber="73" rank="species" species="monae">monae</taxonomicName>
|
||
sp.n. Balke des. 1997" [red], "
|
||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina jaseminae" order="Coleoptera" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="jaseminae">Exocelina jaseminae</taxonomicName>
|
||
(Balke) det. H.Shaverdo 2014" (BMNH). 2 males "Papua New Guinea: Morobe, Wagau, Herzog Mts., 1150m, 19.xi.2006, 06.51.067S 146.48.068E, Balke & Kinibel (PNG 102)", one male with an additional green label "DNA M.Balke 1382" (ZSM). 1 male, 2 females "Papua New Guinea: Morobe, Wagau, Herzog Mts., 1150m, 19.xi.2006, 06.51.067S 146.48.068E, Balke & Kinibel (PNG 103)" (ZSM). 2 males "Papua New Guinea: Gulf [sic!], Menyamya, Mt Inji 1700m, 14.xi.2006 nr 07.14.813S 146.01.330E Balke & Kinibel (PNG 96)" (ZSM). 9 males "PAPUA N. G.: Morobe Prov. Aseki, Oiwa, 1600-1700 m, 11.-12.3.1998 leg. A. Riedel" (NHMW). 6 males, 2 females "Papua New Guinea: Gulf [Morobe], Marawaka, Andakombe towards Morobe, 1100m, 12.xi.200, 07.09.766S, 145.46.333E, Balke & Kinibel (PNG 92)" (NHMW, ZSM).
|
||
<emphasis bold="true" pageId="0" pageNumber="73">Eastern Highlands</emphasis>
|
||
: 38 males, 38 females "Papua New Guinea: Eastern Highlands, Marawaka, Ande, 1700m, 8.xi.2005, 07.01.697S 145.49.807E, Balke & Kinibel (PNG 86)", one male with an additional green label "DNA M.Balke 1365" (NHMW, ZSM). 49 males, 40 females "Papua New Guinea: Eastern Highlands, Marawaka, Ande, 1700-1800m, 9.xi.2006, 07.01.697S 145.49.807E, Balke & Kinibel (PNG 87)" (NHMW, ZSM).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="73" type="diagnosis">
|
||
<paragraph pageId="0" pageNumber="73">Diagnosis.</paragraph>
|
||
<paragraph pageId="0" pageNumber="73">
|
||
For complete description, see
|
||
<bibRefCitation pageId="0" pageNumber="73" refId="B2">Balke (1998</bibRefCitation>
|
||
: 331). Beetle medium-sized: TL-H 3.55-4.1 mm; oblong-oval; brown to dark brown, with reddish to reddish brown pronotal sides or pronotum and often head; shiny, with very fine, on elytra often almost invisible punctation and weakly impressed microreticulation; pronotum with lateral bead (
|
||
<figureCitation captionStart="Figures 26–29" captionStartId="F14" captionText="Figures 26 - 29. Habitus and colouration 26 Exocelina aseki sp. nov. 27 E. jaseminae (Balke, 1998) 28 E. kailaki sp. nov. 29 E. pseudojaseminae sp. nov." figureDoi="10.3897/zookeys.878.37403.figures26-29" httpUri="https://binary.pensoft.net/fig/344982" pageId="0" pageNumber="73" tableDoi="10.3897/zookeys.878.37403.figures26-29">Fig. 27</figureCitation>
|
||
); male protarsomere 4 with anterolateral seta thin, evenly curved, smaller than or equal to more laterally situated large seta; male protarsomere 5 ventrally with anterior band of more than 60 and posterior row of 13-15 relatively long setae (
|
||
<figureCitation captionStart="Figures 30, 31" captionStartId="F15" captionText="Figures 30, 31. 30 Exocelina aseki sp. nov. 31 E. jaseminae (Balke, 1998) A median lobe in ventral view B median lobe in lateral view C paramere in external view D male protarsomeres 4 - 5 in ventral view." figureDoi="10.3897/zookeys.878.37403.figures30-31" httpUri="https://binary.pensoft.net/fig/344983" pageId="0" pageNumber="73" tableDoi="10.3897/zookeys.878.37403.figures30-31">Fig. 31D</figureCitation>
|
||
); median lobe in lateral view with apical lobes distinctly developed and rounded,
|
||
<normalizedToken originalValue="“nose”">"nose"</normalizedToken>
|
||
small but distinct; paramere slightly concave on dorsal side and with long, dense, thin setae, situated along dorsal margin distinctly divided to dense and strong subdistal setae and sparser proximal ones, setae in middle short and fine (
|
||
<figureCitation captionStart="Figures 30, 31" captionStartId="F15" captionText="Figures 30, 31. 30 Exocelina aseki sp. nov. 31 E. jaseminae (Balke, 1998) A median lobe in ventral view B median lobe in lateral view C paramere in external view D male protarsomeres 4 - 5 in ventral view." figureDoi="10.3897/zookeys.878.37403.figures30-31" httpUri="https://binary.pensoft.net/fig/344983" pageId="0" pageNumber="73" tableDoi="10.3897/zookeys.878.37403.figures30-31">
|
||
Fig. 31
|
||
<normalizedToken originalValue="A–C">A-C</normalizedToken>
|
||
</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="73" type="affinities">
|
||
<paragraph pageId="0" pageNumber="73">Affinities.</paragraph>
|
||
<paragraph pageId="0" pageNumber="73">
|
||
In the area of its distribution,
|
||
<taxonomicName lsidName="E. jaseminae" pageId="0" pageNumber="73" rank="species" species="jaseminae">
|
||
<emphasis italics="true" pageId="0" pageNumber="73">E. jaseminae</emphasis>
|
||
</taxonomicName>
|
||
co-occurs with numerous species of the
|
||
<taxonomicName lsidName="E. ekari" pageId="0" pageNumber="73" rank="species" species="ekari">
|
||
<emphasis italics="true" pageId="0" pageNumber="73">E. ekari</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName lsidName="E. ullrichi" pageId="0" pageNumber="73" rank="species" species="ullrichi">
|
||
<emphasis italics="true" pageId="0" pageNumber="73">E. ullrichi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName lsidName="E. broschii" pageId="0" pageNumber="73" rank="species" species="broschii">
|
||
<emphasis italics="true" pageId="0" pageNumber="73">E. broschii</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName lsidName="E. danae" pageId="0" pageNumber="73" rank="species" species="danae">
|
||
<emphasis italics="true" pageId="0" pageNumber="73">E. danae</emphasis>
|
||
</taxonomicName>
|
||
groups. From them, this species can be distinguished by its size, dorsal colouration, surface sculpture, simple male antennae, presence of pronotal bead, and mainly by the shape of its median lobe. In its external appearance,
|
||
<taxonomicName lsidName="E. jaseminae" pageId="0" pageNumber="73" rank="species" species="jaseminae">
|
||
<emphasis italics="true" pageId="0" pageNumber="73">E. jaseminae</emphasis>
|
||
</taxonomicName>
|
||
is especially similar to
|
||
<taxonomicName lsidName="E. monae" pageId="0" pageNumber="73" rank="species" species="monae">
|
||
<emphasis italics="true" pageId="0" pageNumber="73">E. monae</emphasis>
|
||
</taxonomicName>
|
||
(Balke, 1998), from which can be distinguished by the shape of its median lobe. For the affinities within the group, see the
|
||
<normalizedToken originalValue="“Key”">"Key"</normalizedToken>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="73" type="distribution">
|
||
<paragraph pageId="0" pageNumber="73">Distribution.</paragraph>
|
||
<paragraph pageId="0" pageNumber="73">
|
||
Papua New Guinea: Eastern Highlands and Morobe Provinces (
|
||
<figureCitation captionStart="Figure 34" captionStartId="F17" captionText="Figure 34. Map of the eastern part of New Guinea showing distribution of the species of the E. jaseminae group." figureDoi="10.3897/zookeys.878.37403.figure34" httpUri="https://binary.pensoft.net/fig/344985" pageId="0" pageNumber="73" tableDoi="10.3897/zookeys.878.37403.figure34">Fig. 34</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |