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<document ID-DOI="http://doi.org/10.5281/zenodo.3790295" ID-GBIF-Dataset="ca35c50f-b0c5-4032-b06d-f71b00710c8d" ID-GBIF-Taxon="163751260" ID-ISSN="13132970" ID-Zenodo-Dep="3790295" ID-ZooBank="C26E1A82-0DD4-48EF-865C-9D8AA788B739" approvalRequired="188" approvalRequired_for_taxonomicNames="59" approvalRequired_for_textStreams="129" checkinTime="1587990637238" checkinUser="plazi" docAuthor="Crabo, Lars &amp; Lafontaine, Donald" docDate="2009" docId="75513F417B7FFFD9FF02EE2A92A9FD9F" docLanguage="en" docName="ZK_article_2110.pdf" docOrigin="ZooKeys 30 (30)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@648ded35" docStyleName="zookeys.2008.journal_article" docTitle="Lasionycta phaea" docType="treatment" docVersion="5" lastPageId="15" lastPageNumber="16" masterDocId="896847397B71FFD6FF8EE829912DFFF8" masterDocTitle="A Revision of Lasionycta Aurivillius (Lepidoptera: Noctuidae) for North America and notes on Eurasian species, with descriptions of 17 new species, 6 new subspecies, a new genus, and two new species of Tricholita Grote" masterLastPageNumber="156" masterPageNumber="1" pageId="14" pageNumber="15" updateTime="1643569142744" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
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<mods:title>A Revision of Lasionycta Aurivillius (Lepidoptera: Noctuidae) for North America and notes on Eurasian species, with descriptions of 17 new species, 6 new subspecies, a new genus, and two new species of Tricholita Grote</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Crabo, Lars</mods:namePart>
<mods:affiliation>Washington State University, Bellingham, United States of America</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lafontaine, Donald</mods:namePart>
<mods:affiliation>Agriculture and Agri-Food Canada, Ottawa, Canada</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2009</mods:date>
<mods:detail type="pubDate">
<mods:number>2009-12-18</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>30</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>30</mods:number>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.3897/zookeys.30.308</mods:identifier>
<mods:identifier type="GBIF-Dataset">ca35c50f-b0c5-4032-b06d-f71b00710c8d</mods:identifier>
<mods:identifier type="ISSN">13132970</mods:identifier>
<mods:identifier type="Zenodo-Dep">576576</mods:identifier>
<mods:identifier type="ZooBank">C26E1A82-0DD4-48EF-865C-9D8AA788B739</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3790295" ID-GBIF-Taxon="163751260" ID-Zenodo-Dep="3790295" LSID="urn:lsid:plazi:treatment:75513F417B7FFFD9FF02EE2A92A9FD9F" httpUri="http://treatment.plazi.org/id/75513F417B7FFFD9FF02EE2A92A9FD9F" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
<subSubSection lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15" type="nomenclature">
<paragraph blockId="14.[140,517,1539,1566]" box="[140,517,1539,1566]" pageId="14" pageNumber="15">
<heading box="[140,517,1539,1566]" fontSize="11" level="2" pageId="14" pageNumber="15" reason="3">
<emphasis bold="true" box="[140,517,1539,1566]" pageId="14" pageNumber="15">
<taxonomicName baseAuthorityName="Hampson" baseAuthorityYear="1905" box="[140,346,1539,1566]" class="Insecta" family="Noctuidae" genus="Lasionycta" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="phaea">
<emphasis bold="true" box="[140,346,1539,1566]" italics="true" pageId="14" pageNumber="15">Lasionycta phaea</emphasis>
</taxonomicName>
species-group
</emphasis>
</heading>
</paragraph>
<paragraph blockId="14.[140,1108,1610,1672]" lastBlockId="15.[140,1108,167,616]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
The male genitalia of the
<taxonomicName baseAuthorityName="Hampson" baseAuthorityYear="1905" box="[407,495,1610,1636]" class="Insecta" family="Noctuidae" genus="Lasionycta" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="phaea">
<emphasis box="[407,423,1611,1636]" italics="true" pageId="14" pageNumber="15">L</emphasis>
.
<emphasis box="[434,495,1610,1636]" italics="true" pageId="14" pageNumber="15">phaea</emphasis>
</taxonomicName>
species-group resemble those of the
<taxonomicName baseAuthorityName="Herrich-Schaffer" baseAuthorityYear="1852" box="[881,1020,1610,1636]" class="Insecta" family="Noctuidae" genus="Lasionycta" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="skraelingia">
<emphasis box="[881,897,1611,1636]" italics="true" pageId="14" pageNumber="15">L</emphasis>
.
<emphasis box="[909,1020,1610,1636]" italics="true" pageId="14" pageNumber="15">skraelingia</emphasis>
</taxonomicName>
speciesgroup, but the uncus is broadly flattened and spatulate with a squared-off apex and the digitus is longer, extending below the valve. Females have a distinctly different bursa than those of the
<taxonomicName baseAuthorityName="Herrich-Schaffer" baseAuthorityYear="1852" box="[279,422,202,228]" class="Insecta" family="Noctuidae" genus="Lasionycta" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="skraelingia">
<emphasis box="[279,295,203,228]" italics="true" pageId="15" pageNumber="16">L</emphasis>
.
<emphasis box="[311,422,202,228]" italics="true" pageId="15" pageNumber="16">skraelingia</emphasis>
</taxonomicName>
species-group. Th e corpus bursae is ovoid, not rounded, with less sclerotized signa. Th e appendix bursae is attached to the dorsal corpus bursae rather than to the left side and extends dorsally and to the left. Its surface is grooved producing a weak spiral of approximately 1.5 turns. Th e male antenna is beadlike (
<taxonomicName baseAuthorityName="Hampson" baseAuthorityYear="1905" box="[909,1001,307,334]" class="Insecta" family="Noctuidae" genus="Lasionycta" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="phaea">
<emphasis box="[909,925,308,333]" italics="true" pageId="15" pageNumber="16">L</emphasis>
.
<emphasis box="[940,1001,307,333]" italics="true" pageId="15" pageNumber="16">phaea</emphasis>
</taxonomicName>
) to biserrate (1.5× as wide as shaft in two Asian species (
<bibRefCitation author="Lafontaine JD &amp; Kononenko VS" box="[657,1026,343,369]" pageId="15" pageNumber="16" pagination="903 - 916" refId="ref65105" refString="Lafontaine JD, Kononenko VS (1988) A revision of the Lasionycta skraelingia Herrich-Schaffer) species complex (Lepidoptera: Noctuidae). Th e Canadian Entomologist 120: 903 - 916." type="journal article" year="1988">Lafontaine and Kononenko 1988</bibRefCitation>
)).
</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="16" type="description">
<paragraph blockId="15.[140,1108,167,616]" pageId="15" pageNumber="16">
<taxonomicName baseAuthorityName="Hampson" baseAuthorityYear="1905" box="[185,360,378,404]" class="Insecta" family="Noctuidae" genus="Lasionycta" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="phaea">
<emphasis box="[185,360,378,404]" italics="true" pageId="15" pageNumber="16">Lasionycta phaea</emphasis>
</taxonomicName>
is the only North American member of the species-group. Three additional species,
<taxonomicName authority="Lafontaine &amp; Kononenko" authorityName="Lafontaine &amp; Kononenko (Lafontaine and Kononenko" authorityYear="1988" box="[294,694,413,440]" class="Insecta" family="Noctuidae" genus="Lasionycta" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="alpicola">
<emphasis box="[294,310,414,439]" italics="true" pageId="15" pageNumber="16">L</emphasis>
.
<emphasis box="[323,403,413,439]" italics="true" pageId="15" pageNumber="16">alpicola</emphasis>
Lafontaine &amp; Kononenko
</taxonomicName>
,
<taxonomicName authority="Kononenko" authorityName="Kononenko" box="[704,967,413,440]" class="Insecta" family="Noctuidae" genus="Lasionycta" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="buraetica">
<emphasis box="[704,719,414,439]" italics="true" pageId="15" pageNumber="16">L</emphasis>
.
<emphasis box="[733,828,413,439]" italics="true" pageId="15" pageNumber="16">buraetica</emphasis>
Kononenko
</taxonomicName>
, and
<taxonomicName authority="Kononenko" authorityName="Kononenko" class="Insecta" family="Noctuidae" genus="Lasionycta" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="corax">
<emphasis box="[1024,1040,414,439]" italics="true" pageId="15" pageNumber="16">L</emphasis>
.
<emphasis box="[1053,1107,414,439]" italics="true" pageId="15" pageNumber="16">corax</emphasis>
Kononenko
</taxonomicName>
, are found in central and western Asia (
<bibRefCitation author="Lafontaine JD &amp; Kononenko VS" box="[698,1067,448,474]" pageId="15" pageNumber="16" pagination="903 - 916" refId="ref65105" refString="Lafontaine JD, Kononenko VS (1988) A revision of the Lasionycta skraelingia Herrich-Schaffer) species complex (Lepidoptera: Noctuidae). Th e Canadian Entomologist 120: 903 - 916." type="journal article" year="1988">Lafontaine and Kononenko 1988</bibRefCitation>
).
</paragraph>
<paragraph blockId="15.[140,1108,167,616]" pageId="15" pageNumber="16">
The species in this species-group were included in the
<taxonomicName baseAuthorityName="Herrich-Schaffer" baseAuthorityYear="1852" box="[800,947,483,510]" class="Insecta" family="Noctuidae" genus="Lasionycta" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="skraelingia">
<emphasis box="[800,816,484,509]" italics="true" pageId="15" pageNumber="16">L</emphasis>
.
<emphasis box="[831,947,483,509]" italics="true" pageId="15" pageNumber="16">skraelingia</emphasis>
</taxonomicName>
species-group by
<bibRefCitation author="Lafontaine JD &amp; Kononenko VS" box="[178,588,519,545]" pageId="15" pageNumber="16" pagination="903 - 916" refId="ref65105" refString="Lafontaine JD, Kononenko VS (1988) A revision of the Lasionycta skraelingia Herrich-Schaffer) species complex (Lepidoptera: Noctuidae). Th e Canadian Entomologist 120: 903 - 916." type="journal article" year="1988">Lafontaine and Kononenko (1988)</bibRefCitation>
. In addition to the structural differences between the species-groups,
<taxonomicName baseAuthorityName="Hampson" baseAuthorityYear="1905" box="[436,531,554,580]" class="Insecta" family="Noctuidae" genus="Lasionycta" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="phaea">
<emphasis box="[436,452,555,580]" italics="true" pageId="15" pageNumber="16">L</emphasis>
.
<emphasis box="[468,531,554,580]" italics="true" pageId="15" pageNumber="16">phaea</emphasis>
</taxonomicName>
fails to group with
<taxonomicName baseAuthorityName="Herrich-Schaffer" baseAuthorityYear="1852" box="[757,905,554,580]" class="Insecta" family="Noctuidae" genus="Lasionycta" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="skraelingia">
<emphasis box="[757,773,555,580]" italics="true" pageId="15" pageNumber="16">L</emphasis>
.
<emphasis box="[789,905,554,580]" italics="true" pageId="15" pageNumber="16">skraelingia</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Lafontaine" authorityYear="1988" box="[963,1069,554,580]" class="Insecta" family="Noctuidae" genus="Lasionycta" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="taigata">
<emphasis box="[963,979,555,580]" italics="true" pageId="15" pageNumber="16">L</emphasis>
.
<emphasis box="[994,1069,555,580]" italics="true" pageId="15" pageNumber="16">taigata</emphasis>
</taxonomicName>
on CO1 distance analysis, instead clustering near
<taxonomicName box="[665,780,589,616]" class="Insecta" family="Noctuidae" genus="Lasionycta" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="secedens">
<emphasis box="[665,681,590,615]" italics="true" pageId="15" pageNumber="16">L</emphasis>
.
<emphasis box="[696,780,589,615]" italics="true" pageId="15" pageNumber="16">secedens</emphasis>
</taxonomicName>
(
<figureCitation box="[795,888,589,615]" captionStart="Figure 247" captionStartId="141.[140,212,1494,1516]" captionTargetBox="[192,1083,225,1446]" captionTargetId="figure@141.[159,1089,165,1475]" captionTargetPageId="141" captionText="Figure 247. Neighbor-joining CO1 tree of North American Lasionycta, Psammopolia, Tricholita ferrisi and Eurasian Lasionhada proxima and Eriopygodes imbecilla. Incomplete haplotypes with 600657 base pairs are denoted with (*) and those with 550599 base pairs with (**). The letter and number code after each species is a unique haplotype identifier followed by the number of samples in parentheses. No data is available for Psammopolia insolens, P. sala, or P. ochracea. The illustrated species show a member of each species-group and are scaled to the same size. The Lasionycta leucocycla species-group is shown in Fig. 248." figureDoi="http://doi.org/10.5281/zenodo.3769245" httpUri="https://zenodo.org/record/3769245/files/figure.png" pageId="15" pageNumber="16">Fig 247</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>