treatments-xml/data/75/51/3F/75513F417B5BFFFAFF02ED5E92EDFAFC.xml

<document ID-DOI="http://doi.org/10.5281/zenodo.3790238" ID-GBIF-Dataset="ca35c50f-b0c5-4032-b06d-f71b00710c8d" ID-GBIF-Taxon="163751281" ID-ISSN="1313–2970" ID-Zenodo-Dep="3790238" ID-ZooBank="C26E1A82-0DD4-48EF-865C-9D8AA788B739" LSID="urn:lsid:zoobank.org:act:74F2198E-C747-46B9-9F9B-8D8783E51CF5" approvalRequired="188" approvalRequired_for_taxonomicNames="59" approvalRequired_for_textStreams="129" checkinTime="1587990637238" checkinUser="plazi" docAuthor="Crabo, Lars &amp; Lafontaine, Donald" docDate="2009" docId="75513F417B5BFFFAFF02ED5E92EDFAFC" docLanguage="en" docName="ZK_article_2110.pdf" docOrigin="ZooKeys 30 (30)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@648ded35" docStyleName="zookeys.2008.journal_article" docTitle="Lasionycta frigida Crabo &amp; Lafontaine 2009, sp. n." docType="treatment" docUuid="74F2198E-C747-46B9-9F9B-8D8783E51CF5" docUuidSource="ZooBank" docVersion="6" lastPageId="44" lastPageNumber="45" masterDocId="896847397B71FFD6FF8EE829912DFFF8" masterDocTitle="A Revision of Lasionycta Aurivillius (Lepidoptera: Noctuidae) for North America and notes on Eurasian species, with descriptions of 17 new species, 6 new subspecies, a new genus, and two new species of Tricholita Grote" masterLastPageNumber="156" masterPageNumber="1" pageId="42" pageNumber="43" updateTime="1643569142744" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A Revision of Lasionycta Aurivillius (Lepidoptera: Noctuidae) for North America and notes on Eurasian species, with descriptions of 17 new species, 6 new subspecies, a new genus, and two new species of Tricholita Grote</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Crabo, Lars</mods:namePart>
<mods:affiliation>Washington State University, Bellingham, United States of America</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lafontaine, Donald</mods:namePart>
<mods:affiliation>Agriculture and Agri-Food Canada, Ottawa, Canada</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2009</mods:date>
<mods:detail type="pubDate">
<mods:number>2009-12-18</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>30</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>30</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>156</mods:end>
</mods:extent>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.3897/zookeys.30.308</mods:identifier>
<mods:identifier type="GBIF-Dataset">ca35c50f-b0c5-4032-b06d-f71b00710c8d</mods:identifier>
<mods:identifier type="ISSN">1313–2970</mods:identifier>
<mods:identifier type="Zenodo-Dep">576576</mods:identifier>
<mods:identifier type="ZooBank">C26E1A82-0DD4-48EF-865C-9D8AA788B739</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3790238" ID-GBIF-Taxon="163751281" ID-Zenodo-Dep="3790238" LSID="urn:lsid:zoobank.org:act:74F2198E-C747-46B9-9F9B-8D8783E51CF5" httpUri="http://treatment.plazi.org/id/75513F417B5BFFFAFF02ED5E92EDFAFC" lastPageId="44" lastPageNumber="45" pageId="42" pageNumber="43">
<subSubSection pageId="42" pageNumber="43" type="nomenclature">
<paragraph blockId="42.[140,962,1398,1496]" box="[140,686,1398,1425]" pageId="42" pageNumber="43">
<heading box="[140,686,1398,1425]" fontSize="11" level="2" pageId="42" pageNumber="43" reason="3">
<emphasis bold="true" box="[140,686,1398,1425]" pageId="42" pageNumber="43">
<taxonomicName authority="Crabo &amp; Lafontaine" authorityName="Crabo &amp; Lafontaine" authorityYear="2009" box="[140,608,1398,1425]" class="Insecta" family="Noctuidae" genus="Lasionycta" kingdom="Animalia" order="Lepidoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="frigida" status="sp. n.">
<emphasis bold="true" box="[140,356,1398,1425]" italics="true" pageId="42" pageNumber="43">
<emphasis box="[140,268,1399,1425]" inLineHeading="true" pageId="42" pageNumber="43" reason="1">Lasionycta</emphasis>
frigida
</emphasis>
Crabo &amp; Lafontaine
</taxonomicName>
, 
<taxonomicNameLabel box="[621,686,1399,1425]" pageId="42" pageNumber="43" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<uri box="[140,962,1434,1461]" pageId="42" pageNumber="43">
urn:lsid:zoobank.org:act: 
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</uri>
</paragraph>
</subSubSection>
<subSubSection box="[140,429,1469,1496]" pageId="42" pageNumber="43" type="description">
<paragraph blockId="42.[140,962,1398,1496]" box="[140,429,1469,1496]" pageId="42" pageNumber="43">Figs 34, 154, 210. Map 7</paragraph>
</subSubSection>
<subSubSection lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43" type="materials_examined">
<paragraph blockId="42.[140,1108,1539,1673]" pageId="42" pageNumber="43">
<emphasis bold="true" box="[140,315,1539,1565]" pageId="42" pageNumber="43">
<emphasis box="[140,198,1539,1565]" inLineHeading="true" pageId="42" pageNumber="43" reason="1">Type</emphasis>
material.
</emphasis>
<materialsCitation ID-GBIF-Occurrence="2608941606" collectingDate="2001-07-14" collectionCode="CNC" collectorName="G. G. Anweiler &amp; Genitalia &amp; Databased &amp; Noctuoidea" county="Canada" elevation="1640" location="Barcodes of Life Project" municipality="Prospect Creek" pageId="42" pageNumber="43" specimenCount="1" stateProvince="Alberta" typeStatus="holotype">
<emphasis bold="true" box="[325,438,1539,1565]" pageId="42" pageNumber="43">
<typeStatus box="[325,438,1539,1565]" pageId="42" pageNumber="43">Holotype</typeStatus>
</emphasis>
♁. 
<collectingCounty box="[486,575,1539,1565]" pageId="42" pageNumber="43">Canada</collectingCounty>
, 
<collectingRegion box="[590,676,1539,1565]" country="Canada" name="Alberta" pageId="42" pageNumber="43">Alberta</collectingRegion>
, mouth of 
<collectingMunicipality box="[812,989,1539,1566]" pageId="42" pageNumber="43">Prospect Creek</collectingMunicipality>
, 
<quantity box="[1004,1099,1539,1566]" metricMagnitude="3" metricUnit="m" metricValue="1.64" pageId="42" pageNumber="43" unit="m" value="1640.0">
<elevation box="[1004,1099,1539,1566]" metricMagnitude="3" metricUnit="m" metricValue="1.64" pageId="42" pageNumber="43" unit="m" value="1640.0">1640 m</elevation>
</quantity>
, 
<collectingMethod box="[140,240,1575,1601]" pageId="42" pageNumber="43">UV trap</collectingMethod>
, 
<date box="[254,409,1575,1601]" pageId="42" pageNumber="43" value="2001-07-14">
<collectingDate box="[254,409,1575,1601]" pageId="42" pageNumber="43" value="2001-07-14">14 July 2001</collectingDate>
</date>
, subalpine spruce-pine-willow forest, 
<collectorName box="[847,1028,1575,1601]" pageId="42" pageNumber="43">G. G. Anweiler</collectorName>
, 
<collectorName pageId="42" pageNumber="43">Genitalia</collectorName>
slide 95848, 
<collectorName box="[346,464,1610,1636]" pageId="42" pageNumber="43">Databased</collectorName>
for 
<collectionCode box="[514,578,1610,1636]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="42" pageNumber="43">CNC</collectionCode>
<collectorName box="[587,719,1610,1636]" pageId="42" pageNumber="43">Noctuoidea</collectorName>
# 6627, 
<location LSID="urn:lsid:plazi:treatment:75513F417B5BFFFAFF02ED5E92EDFAFC:F827D88C7B5BFFFCFCB5EE639562F99C" box="[827,1103,1610,1636]" county="Canada" municipality="Prospect Creek" name="Barcodes of Life Project" pageId="42" pageNumber="43" stateProvince="Alberta">Barcodes of Life Project</location>
, 
<location LSID="urn:lsid:plazi:treatment:75513F417B5BFFFAFF02ED5E92EDFAFC:F827D88C7B5BFFFCFF02EE449051F97F" box="[140,380,1645,1671]" county="Canada" municipality="Prospect Creek" name="University of Guelph" pageId="42" pageNumber="43" stateProvince="Alberta">University of Guelph</location>
, DNA # 
<collectorName box="[484,616,1645,1671]" pageId="42" pageNumber="43">Noctuoidea</collectorName>
6627. 
<collectionCode box="[695,762,1645,1671]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="42" pageNumber="43">CNC</collectionCode>
</materialsCitation>
. 
<emphasis bold="true" box="[773,888,1645,1671]" pageId="42" pageNumber="43">
<typeStatus box="[773,888,1645,1671]" pageId="42" pageNumber="43">Paratypes</typeStatus>
</emphasis>
4 ♁, 
<specimenCount box="[949,985,1645,1672]" pageId="42" pageNumber="43" type="female">3 ♀</specimenCount>
. 
<emphasis bold="true" box="[1009,1100,1645,1671]" pageId="42" pageNumber="43">
<collectingCountry box="[1009,1100,1645,1671]" name="Canada" pageId="42" pageNumber="43">Canada</collectingCountry>
</emphasis>
.
</paragraph>
<paragraph blockId="43.[140,1108,166,1672]" pageId="43" pageNumber="44">
<materialsCitation ID-GBIF-Occurrence="2608941514" box="[140,986,166,194]" collectingDate="1939-07-12" collectorName="K. Bowman &amp; Slide No." location="Nordegg" pageId="43" pageNumber="44" specimenCount="1" specimenCount-female="1" stateProvince="Alberta">
<emphasis bold="true" box="[140,228,166,192]" inLineHeading="true" pageId="43" pageNumber="44" reason="1">
<collectingRegion box="[140,228,166,192]" country="Canada" name="Alberta" pageId="43" pageNumber="44">Alberta</collectingRegion>
</emphasis>
. 
<location LSID="urn:lsid:plazi:treatment:75513F417B5BFFFAFF02ED5E92EDFAFC:F827D88C7B5AFFFDFF7BE88E9076FF39" box="[245,347,167,193]" name="Nordegg" pageId="43" pageNumber="44" stateProvince="Alberta">Nordegg</location>
, 
<date box="[360,513,167,193]" pageId="43" pageNumber="44" value="1939-07-12">
<collectingDate box="[360,513,167,193]" pageId="43" pageNumber="44" value="1939-07-12">12 July 1939</collectingDate>
</date>
, 
<collectorName box="[527,668,167,193]" pageId="43" pageNumber="44">K. Bowman</collectorName>
, 
<collectorName box="[682,790,167,193]" pageId="43" pageNumber="44">Slide No.</collectorName>
8335 (1 ♁, 
<specimenCount box="[936,975,167,193]" pageId="43" pageNumber="44" type="female">1 ♀</specimenCount>
)
</materialsCitation>
; 
<materialsCitation ID-GBIF-Occurrence="2608941613" collectingDate="1999-07-17" collectionCode="CNC" collectorName="Rocky &amp; Chris Schmidt &amp; Genitalia &amp; Databased &amp; Noctuoidea" county="Prospect Creek" elevation="2400" location="University of Guelph" municipality="Barcode of Life Project" pageId="43" pageNumber="44" specimenCount="1" stateProvince="Alberta">
Prospect Mtn above 
<collectingCounty box="[273,447,202,228]" pageId="43" pageNumber="44">Prospect Creek</collectingCounty>
, 
<quantity box="[460,554,202,228]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="43" pageNumber="44" unit="m" value="2400.0">
<elevation box="[460,554,202,228]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="43" pageNumber="44" unit="m" value="2400.0">2400 m</elevation>
</quantity>
, 
<date box="[567,719,202,228]" pageId="43" pageNumber="44" value="1999-07-17">
<collectingDate box="[567,719,202,228]" pageId="43" pageNumber="44" value="1999-07-17">17 July 1999</collectingDate>
</date>
, 
<collectorName box="[732,804,202,228]" pageId="43" pageNumber="44">Rocky</collectorName>
vegetated slopes nectaring 
<collectedFrom box="[140,239,237,264]" pageId="43" pageNumber="44">
on 
<taxonomicName box="[178,239,237,263]" class="Magnoliopsida" family="Polemoniaceae" genus="Phlox" kingdom="Plantae" order="Ericales" pageId="43" pageNumber="44" phylum="Tracheophyta" rank="genus">
<emphasis box="[178,239,237,263]" italics="true" pageId="43" pageNumber="44">Phlox</emphasis>
</taxonomicName>
</collectedFrom>
sp. 
<collectorName box="[289,459,237,263]" pageId="43" pageNumber="44">Chris Schmidt</collectorName>
, 
<collectorName box="[472,576,237,263]" pageId="43" pageNumber="44">Genitalia</collectorName>
slide 95847, 
<collectorName box="[734,852,237,263]" pageId="43" pageNumber="44">Databased</collectorName>
for 
<collectionCode box="[902,966,237,263]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="43" pageNumber="44">CNC</collectionCode>
<collectorName box="[975,1107,237,263]" pageId="43" pageNumber="44">Noctuoidea</collectorName>
# 6608/ 
<collectingMunicipality box="[240,507,272,298]" pageId="43" pageNumber="44">Barcode of Life Project</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:75513F417B5BFFFAFF02ED5E92EDFAFC:F827D88C7B5AFFFDFD86E93993D0FED2" box="[520,765,272,298]" county="Prospect Creek" municipality="Barcode of Life Project" name="University of Guelph" pageId="43" pageNumber="44" stateProvince="Alberta">University of Guelph</location>
, DNA # 
<collectorName box="[875,1007,272,298]" pageId="43" pageNumber="44">Noctuoidea</collectorName>
6608 (1 ♁)
</materialsCitation>
; 
<materialsCitation ID-GBIF-Occurrence="2608941550" collectingDate="1999-07-17" collectorName="B. C. Schmidt" elevation="2300" location="Prospect Mtn" pageId="43" pageNumber="44" specimenCount="1" stateProvince="Alberta">
<location LSID="urn:lsid:plazi:treatment:75513F417B5BFFFAFF02ED5E92EDFAFC:F827D88C7B5AFFFDFF37E91D907BFEB6" box="[185,342,308,334]" name="Prospect Mtn" pageId="43" pageNumber="44" stateProvince="Alberta">Prospect Mtn</location>
̴ 
<locationDeviation box="[364,616,307,334]" location="Cadomin" pageId="43" pageNumber="44">
<quantity box="[364,442,307,334]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="43" pageNumber="44" unit="km" value="10.0">10 km</quantity>
SW Cadomin
</locationDeviation>
, 45–24W5, 
<quantity box="[763,858,308,334]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="43" pageNumber="44" unit="m" value="2300.0">
<elevation box="[763,858,308,334]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="43" pageNumber="44" unit="m" value="2300.0">2300 m</elevation>
</quantity>
, 
<date box="[872,1026,307,334]" pageId="43" pageNumber="44" value="1999-07-17">
<collectingDate box="[872,1026,307,334]" pageId="43" pageNumber="44" value="1999-07-17">17 July 1999</collectingDate>
</date>
, 
<collectorName pageId="43" pageNumber="44">B. C. Schmidt</collectorName>
(2 ♁)
</materialsCitation>
; 
<materialsCitation ID-GBIF-Occurrence="2608941570" collectingDate="2000-07-21" collectorName="Schmidt &amp; Anweiler" location="Prospect Ck" municipality="Cadomin" pageId="43" pageNumber="44" specimenCount="1" specimenCount-female="1" stateProvince="Alberta">
near 
<collectingMunicipality box="[369,480,343,369]" pageId="43" pageNumber="44">Cadomin</collectingMunicipality>
, mouth of 
<location LSID="urn:lsid:plazi:treatment:75513F417B5BFFFAFF02ED5E92EDFAFC:F827D88C7B5AFFFDFDD7E97E93CEFE89" box="[601,739,343,369]" municipality="Cadomin" name="Prospect Ck" pageId="43" pageNumber="44" stateProvince="Alberta">Prospect Ck</location>
̴5500’, 
<date box="[838,986,343,369]" pageId="43" pageNumber="44" value="2000-07-21">
<collectingDate box="[838,986,343,369]" pageId="43" pageNumber="44" value="2000-07-21">21 July 2000</collectingDate>
</date>
, UV light, 
<collectorName box="[140,236,378,404]" pageId="43" pageNumber="44">Schmidt</collectorName>
and 
<collectorName box="[295,396,378,404]" pageId="43" pageNumber="44">Anweiler</collectorName>
(
<specimenCount box="[415,451,378,404]" pageId="43" pageNumber="44" type="female">1 ♀</specimenCount>
)
</materialsCitation>
; 
<materialsCitation ID-GBIF-Occurrence="2608941528" collectingDate="1999-07-18" collectionCode="CNC, LGC" collectorName="B. C. Schmidt" location="Cardinal Divide" municipality="Mt Tripoli" pageId="43" pageNumber="44" specimenCount="1" specimenCount-female="1" stateProvince="Alberta">
<collectingMunicipality box="[476,599,378,404]" pageId="43" pageNumber="44">Mt Tripoli</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:75513F417B5BFFFAFF02ED5E92EDFAFC:F827D88C7B5AFFFDFDEAE9539232FE6C" box="[612,799,378,404]" municipality="Mt Tripoli" name="Cardinal Divide" pageId="43" pageNumber="44" stateProvince="Alberta">Cardinal Divide</location>
, 45–23W5, 
<date box="[944,1095,378,404]" pageId="43" pageNumber="44" value="1999-07-18">
<collectingDate box="[944,1095,378,404]" pageId="43" pageNumber="44" value="1999-07-18">18 July 1999</collectingDate>
</date>
, 
<collectorName box="[140,303,413,440]" pageId="43" pageNumber="44">B. C. Schmidt</collectorName>
(
<specimenCount box="[322,357,413,440]" pageId="43" pageNumber="44" type="female">1 ♀</specimenCount>
). 
<collectionCode box="[381,447,413,439]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="43" pageNumber="44">CNC</collectionCode>
, 
<collectionCode box="[459,521,413,439]" pageId="43" pageNumber="44">LGC</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="43" pageNumber="44" type="etymology">
<paragraph blockId="43.[140,1108,166,1672]" pageId="43" pageNumber="44">
<emphasis bold="true" box="[185,320,448,474]" pageId="43" pageNumber="44">
<emphasis box="[185,315,448,474]" inLineHeading="true" pageId="43" pageNumber="44" reason="1">Etymology</emphasis>
.
</emphasis>
The name is derived from the Latin 
<emphasis box="[734,815,448,474]" italics="true" pageId="43" pageNumber="44">frigidus</emphasis>
meaning cold. It refers to the cold habitats where this species occurs.
</paragraph>
</subSubSection>
<subSubSection pageId="43" pageNumber="44" type="diagnosis">
<paragraph blockId="43.[140,1108,166,1672]" pageId="43" pageNumber="44">
<emphasis bold="true" box="[185,312,518,544]" pageId="43" pageNumber="44">
<emphasis box="[185,307,518,544]" inLineHeading="true" pageId="43" pageNumber="44" reason="1">Diagnosis</emphasis>
.
</emphasis>
<taxonomicName authorityName="Crabo &amp; Lafontaine" authorityYear="2009" box="[322,517,519,545]" class="Insecta" family="Noctuidae" genus="Lasionycta" kingdom="Animalia" order="Lepidoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="frigida">
<emphasis box="[322,517,519,545]" italics="true" pageId="43" pageNumber="44">Lasionycta frigida</emphasis>
</taxonomicName>
is similar to 
<taxonomicName authorityName="Crabo &amp; Lafontaine" authorityYear="2009" baseAuthorityName="Barnes &amp; Benjamin" baseAuthorityYear="1923" box="[674,756,519,545]" class="Insecta" family="Noctuidae" genus="Lasionycta" kingdom="Animalia" order="Lepidoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="poca">
<emphasis box="[674,690,519,544]" italics="true" pageId="43" pageNumber="44">L</emphasis>
. 
<emphasis box="[708,756,519,544]" italics="true" pageId="43" pageNumber="44">poca</emphasis>
</taxonomicName>
and occurs with it in western 
<collectingRegion box="[140,227,554,580]" country="Canada" name="Alberta" pageId="43" pageNumber="44">Alberta</collectingRegion>
. Th ey are structurally indistinguishable, including male antenna and eyes, but can be distinguished by color, hindwing markings, and CO1 barcode. 
<taxonomicName authorityName="Crabo &amp; Lafontaine" authorityYear="2009" class="Insecta" family="Noctuidae" genus="Lasionycta" kingdom="Animalia" order="Lepidoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="frigida">
<emphasis italics="true" pageId="43" pageNumber="44">Lasionycta frigida</emphasis>
</taxonomicName>
has a gray brown forewing with indistinct markings. Th e hindwing is brownish off-white and the postmedial line is paler than the marginal band. 
<taxonomicName authorityName="Crabo &amp; Lafontaine" authorityYear="2009" baseAuthorityName="Barnes &amp; Benjamin" baseAuthorityYear="1923" box="[894,1063,660,685]" class="Insecta" family="Noctuidae" genus="Lasionycta" kingdom="Animalia" order="Lepidoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="poca">
<emphasis box="[894,1063,660,685]" italics="true" pageId="43" pageNumber="44">Lasionycta poca</emphasis>
</taxonomicName>
has a dark-gray forewing with crisper markings and a purer white hindwing with the dark postmedial line and marginal band similar in color. The CO1 sequence of 
<taxonomicName authorityName="Crabo &amp; Lafontaine" authorityYear="2009" box="[1003,1107,730,756]" class="Insecta" family="Noctuidae" genus="Lasionycta" kingdom="Animalia" order="Lepidoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="frigida">
<emphasis box="[1003,1019,731,756]" italics="true" pageId="43" pageNumber="44">L</emphasis>
. 
<emphasis box="[1035,1107,730,756]" italics="true" pageId="43" pageNumber="44">frigida</emphasis>
</taxonomicName>
differs from that of 
<taxonomicName authorityName="Crabo &amp; Lafontaine" authorityYear="2009" baseAuthorityName="Barnes &amp; Benjamin" baseAuthorityYear="1923" box="[361,440,766,792]" class="Insecta" family="Noctuidae" genus="Lasionycta" kingdom="Animalia" order="Lepidoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="poca">
<emphasis box="[361,377,766,791]" italics="true" pageId="43" pageNumber="44">L</emphasis>
. 
<emphasis box="[392,440,766,791]" italics="true" pageId="43" pageNumber="44">poca</emphasis>
</taxonomicName>
by over 2 % and is more similar to those of 
<taxonomicName box="[940,1074,765,792]" class="Insecta" family="Noctuidae" genus="Lasionycta" kingdom="Animalia" order="Lepidoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="leucocycla">
<emphasis box="[940,956,766,791]" italics="true" pageId="43" pageNumber="44">L</emphasis>
. 
<emphasis box="[970,1074,765,791]" italics="true" pageId="43" pageNumber="44">leucocycla</emphasis>
</taxonomicName>
. It is the only species in the sub-group with a combination of a wide male antenna and a 
<taxonomicName box="[140,274,835,862]" class="Insecta" family="Noctuidae" genus="Lasionycta" kingdom="Animalia" order="Lepidoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="leucocycla">
<emphasis box="[140,156,836,861]" italics="true" pageId="43" pageNumber="44">L</emphasis>
. 
<emphasis box="[170,274,835,861]" italics="true" pageId="43" pageNumber="44">leucocycla</emphasis>
</taxonomicName>
-like CO1 haplotype. The other wide-antenna species resembling 
<taxonomicName class="Insecta" family="Noctuidae" genus="Lasionycta" kingdom="Animalia" order="Lepidoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="leucocycla">
<emphasis box="[1014,1030,836,861]" italics="true" pageId="43" pageNumber="44">L</emphasis>
. 
<emphasis italics="true" pageId="43" pageNumber="44">leucocycla</emphasis>
</taxonomicName>
have CO1 DNA more similar to 
<taxonomicName box="[573,693,871,897]" class="Insecta" family="Noctuidae" genus="Scotogramma" kingdom="Animalia" order="Lepidoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="perplexa">
<emphasis box="[573,589,871,896]" italics="true" pageId="43" pageNumber="44">L</emphasis>
. 
<emphasis box="[604,693,871,897]" italics="true" pageId="43" pageNumber="44">perplexa</emphasis>
</taxonomicName>
than 
<taxonomicName box="[759,894,871,897]" class="Insecta" family="Noctuidae" genus="Lasionycta" kingdom="Animalia" order="Lepidoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="leucocycla">
<emphasis box="[759,775,871,896]" italics="true" pageId="43" pageNumber="44">L</emphasis>
. 
<emphasis box="[790,894,871,897]" italics="true" pageId="43" pageNumber="44">leucocycla</emphasis>
</taxonomicName>
(
<figureCitation box="[910,1009,871,897]" captionStart="Figure 248" captionStartId="142.[140,212,1558,1580]" captionTargetBox="[192,1058,178,1498]" captionTargetId="figure@142.[177,1071,165,1539]" captionTargetPageId="142" captionText="Figure 248. Neighbor-joining CO1 tree of the North American Lasionycta leucocycla species-group. Symbols and codes are the same as in Fig. 247. No data is available for Lasionycta carolynae, L. illima, L. leucocycla hampa, L. macleani, L. mono, or L. uniformis handfieldi. The illustrated specimens demonstrate the variation in appearance of the species-group and are scaled to the same size." figureDoi="http://doi.org/10.5281/zenodo.3769247" httpUri="https://zenodo.org/record/3769247/files/figure.png" pageId="43" pageNumber="44">Fig. 248</figureCitation>
).
</paragraph>
<paragraph blockId="43.[140,1108,166,1672]" pageId="43" pageNumber="44">
<taxonomicName authorityName="Crabo &amp; Lafontaine" authorityYear="2009" box="[185,372,906,932]" class="Insecta" family="Noctuidae" genus="Lasionycta" kingdom="Animalia" order="Lepidoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="illima">
<emphasis box="[185,372,906,932]" italics="true" pageId="43" pageNumber="44">Lasionycta illima</emphasis>
</taxonomicName>
, another similar species with similar male antenna found to the north of 
<taxonomicName authorityName="Crabo &amp; Lafontaine" authorityYear="2009" box="[249,356,941,968]" class="Insecta" family="Noctuidae" genus="Lasionycta" kingdom="Animalia" order="Lepidoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="frigida">
<emphasis box="[249,265,942,967]" italics="true" pageId="43" pageNumber="44">L</emphasis>
. 
<emphasis box="[284,356,941,967]" italics="true" pageId="43" pageNumber="44">frigida</emphasis>
</taxonomicName>
, has smaller eyes and a yellower hindwing. 
<taxonomicName authorityName="Crabo &amp; Lafontaine" authorityYear="2009" box="[882,1079,941,967]" class="Insecta" family="Noctuidae" genus="Lasionycta" kingdom="Animalia" order="Lepidoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="frigida">
<emphasis box="[882,1079,941,967]" italics="true" pageId="43" pageNumber="44">Lasionycta frigida</emphasis>
</taxonomicName>
is nearly indistinguishable in superficial appearance to northern 
<collectingRegion box="[857,930,976,1002]" country="Canada" name="Yukon" pageId="43" pageNumber="44">Yukon</collectingRegion>
populations of 
<emphasis box="[140,156,1012,1037]" italics="true" pageId="43" pageNumber="44">L</emphasis>
. 
<taxonomicName box="[172,299,1011,1038]" class="Insecta" family="Noctuidae" genus="Lasionycta" kingdom="Animalia" order="Lepidoptera" pageId="43" pageNumber="143" phylum="Arthropoda" rank="species" species="leucocycla">
<emphasis box="[172,179,1011,1037]" italics="true" pageId="43" pageNumber="44">l</emphasis>
. 
<emphasis box="[195,299,1011,1037]" italics="true" pageId="43" pageNumber="44">leucocycla</emphasis>
</taxonomicName>
(compare Figs 34 and 22) but differs in antennal structure. 
<taxonomicName authorityName="Crabo &amp; Lafontaine" authorityYear="2009" class="Insecta" family="Noctuidae" genus="Lasionycta" kingdom="Animalia" order="Lepidoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="frigida">
<emphasis italics="true" pageId="43" pageNumber="44">Lasionycta frigida</emphasis>
</taxonomicName>
is easily distinguished from the 
<collectingRegion box="[572,655,1047,1073]" country="Canada" name="Alberta" pageId="43" pageNumber="44">Alberta</collectingRegion>
subspecies 
<emphasis box="[784,800,1047,1072]" italics="true" pageId="43" pageNumber="44">L</emphasis>
. 
<emphasis box="[814,821,1047,1073]" italics="true" pageId="43" pageNumber="44">l</emphasis>
. 
<emphasis box="[835,946,1047,1073]" italics="true" pageId="43" pageNumber="44">albertensis</emphasis>
that has a yellow hindwing.
</paragraph>
</subSubSection>
<subSubSection lastPageId="44" lastPageNumber="45" pageId="43" pageNumber="44" type="description">
<paragraph blockId="43.[140,1108,166,1672]" lastBlockId="44.[140,1108,167,1284]" lastPageId="44" lastPageNumber="45" pageId="43" pageNumber="44">
<emphasis bold="true" box="[185,426,1117,1143]" pageId="43" pageNumber="44">
<emphasis box="[185,329,1117,1143]" inLineHeading="true" pageId="43" pageNumber="44" reason="1">Description</emphasis>
. Head –
</emphasis>
Antenna of male weakly bipectinate and fasciculate; individual segments 2.4× shaft width. Antenna of female filiform and ciliate. Dorsal segments dark gray proximally, white distally. Scape white. Eye normal size. Palpus covered with brownish off-white and few dark-gray scales. Frons covered by pale brown-gray hair-like scales. Top of head with equal mixture of dark-gray and white hair-like scales. 
<emphasis bold="true" pageId="43" pageNumber="44">Thorax –</emphasis>
Vestiture a mixture of hair-like white and gray scales and bifurcate white-tipped gray scales, appearing light gray ventrally and hoary gray dorsally with slightly darker gray lines on prothoracic collar and patagium. Legs with equal mixture of gray and luteous off-white scales. Tarsal segments gray with scattered off-white scales, ringed distally with off-white. 
<emphasis bold="true" box="[258,358,1433,1460]" pageId="43" pageNumber="44">Wings –</emphasis>
Forewing length: male 
<quantity box="[631,716,1434,1460]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="43" pageNumber="44" unit="mm" value="12.0">12 mm</quantity>
(expanse 
<quantity box="[830,919,1434,1460]" metricMagnitude="-2" metricUnit="m" metricValue="2.6" pageId="43" pageNumber="44" unit="mm" value="26.0">26 mm</quantity>
); female 
<quantity box="[1023,1107,1434,1460]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="43" pageNumber="44" unit="mm" value="13.0">13 mm</quantity>
(expanse 
<quantity box="[242,327,1469,1496]" metricMagnitude="-2" metricUnit="m" metricValue="2.7" pageId="43" pageNumber="44" unit="mm" value="27.0">27 mm</quantity>
). Ground color of forewing a mixture of light- to dark-gray, white, and luteous scales, appearing slightly uneven dark gray with faint brown tint. Basal, antemedial, and postmedial lines double, dark gray with pale-gray filling. Basal and antemedial lines undulating. Medial shade dark gray, faint, evident near costa and in cell. Postmedial line moderately scalloped, with darker proximal and faint distal components, mildly excurved near cell, otherwise oblique across wing. Subterminal line pale, sinuous, preceded by prominent dark-gray chevrons between veins forming a nearly continuous dark line across the wing. Spots dark gray. Orbicular spot round, filled with light-gray scales peripherally and a dark-gray ocellus centrally. Reniform spot weakly hourglass shaped, filled with light gray peripherally and dark-gray scales centrally. Claviform spot faint, extending half distance between the antemedial and postmedial lines. Terminal area and fringe slightly darker than remainder of wing, fringe weakly checkered. Ventral forewing pale, slightly luteous gray with darker gray shade along costa and in cell. Markings all similar, dark gray. Discal spot moderately large. Postmedial line scalloped, complete across wing. Marginal band moderately sharply demarcated. Dorsal hindwing slightly brownish off-white with scattered dark-gray scales and dark-gray markings. Discal spot ovoid, prominent. Postmedial line undulating, weakly scalloped, more faint than discal spot and marginal band. Marginal band relatively wide, sharply demarcated. Hindwing fringe light gray proximally, brownish off-white distally. Ventral hindwing slightly brownish off-white with scattered dark-gray scales and darker gray markings than on ventral forewing. Discal spot large, ovoid. Postmedial line solid near anterior margin and fading near anal angle, fainter than ventral forewing postmedial line and other hindwing markings. Marginal band darker, narrower, and more sharply defined than ventral forewing marginal band. 
<emphasis bold="true" box="[562,700,765,791]" pageId="44" pageNumber="45">Abdomen –</emphasis>
Covered with pale-luteous off-white and scattered gray scales, appearing light gray. 
<emphasis bold="true" box="[683,880,800,827]" pageId="44" pageNumber="45">
Male genitalia 
<emphasis bold="true" box="[865,880,801,827]" italics="true" pageId="44" pageNumber="45">–</emphasis>
</emphasis>
(Fig. 154). Genital capsule and aedeagus as in the 
<taxonomicName box="[494,629,835,862]" class="Insecta" family="Noctuidae" genus="Lasionycta" kingdom="Animalia" order="Lepidoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="leucocycla">
<emphasis box="[494,510,836,861]" italics="true" pageId="44" pageNumber="45">L</emphasis>
. 
<emphasis box="[526,629,835,861]" italics="true" pageId="44" pageNumber="45">leucocycla</emphasis>
</taxonomicName>
species-group and 
<taxonomicName box="[849,984,835,862]" class="Insecta" family="Noctuidae" genus="Lasionycta" kingdom="Animalia" order="Lepidoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="leucocycla">
<emphasis box="[849,865,836,861]" italics="true" pageId="44" pageNumber="45">L</emphasis>
. 
<emphasis box="[881,984,835,861]" italics="true" pageId="44" pageNumber="45">leucocycla</emphasis>
</taxonomicName>
sub-group descriptions. Valve approximately 5.5× as long as wide. Cucullus typical, corona single except double at apex. Vesica with a single basal cornutus (N = 1). 
<emphasis bold="true" box="[892,1108,905,932]" pageId="44" pageNumber="45">
Female genitalia 
<emphasis bold="true" box="[1093,1108,906,932]" italics="true" pageId="44" pageNumber="45">–</emphasis>
</emphasis>
(Fig. 210). Ovipositor lobe, segment VIII, and bursa copulatrix as in the 
<taxonomicName box="[972,1107,941,968]" class="Insecta" family="Noctuidae" genus="Lasionycta" kingdom="Animalia" order="Lepidoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="leucocycla">
<emphasis box="[972,988,942,967]" italics="true" pageId="44" pageNumber="45">L</emphasis>
. 
<emphasis box="[1004,1107,941,967]" italics="true" pageId="44" pageNumber="45">leucocycla</emphasis>
</taxonomicName>
species-group description. Corpus bursae approximately 1.4× as long as ductus bursae and 0.6× as wide as long. Proximal appendix bursae relatively bulbous.
</paragraph>
</subSubSection>
<subSubSection pageId="44" pageNumber="45" type="distribution">
<paragraph blockId="44.[140,1108,167,1284]" pageId="44" pageNumber="45">
<emphasis bold="true" box="[185,493,1046,1073]" pageId="44" pageNumber="45">
<emphasis box="[185,336,1046,1072]" inLineHeading="true" pageId="44" pageNumber="45" reason="1">Distribution</emphasis>
and biology.
</emphasis>
<taxonomicName authorityName="Crabo &amp; Lafontaine" authorityYear="2009" box="[501,694,1047,1073]" class="Insecta" family="Noctuidae" genus="Lasionycta" kingdom="Animalia" order="Lepidoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="frigida">
<emphasis box="[501,694,1047,1073]" italics="true" pageId="44" pageNumber="45">Lasionycta frigida</emphasis>
</taxonomicName>
has a restricted range in the 
<collectingRegion box="[1025,1108,1047,1073]" country="Canada" name="Alberta" pageId="44" pageNumber="45">Alberta</collectingRegion>
Rocky Mountains. Th e habitat is mixed forest in cold microclimates (C. Schmidt pers. comm.). It is nocturnal and comes to light, although one specimen was collected during the day. All specimens are from mid-July. Th is species is very rare in collections.
</paragraph>
<paragraph blockId="44.[140,1108,167,1284]" pageId="44" pageNumber="45">
The 
<collectingRegion box="[234,317,1187,1213]" country="Canada" name="Alberta" pageId="44" pageNumber="45">Alberta</collectingRegion>
front ranges where 
<taxonomicName authorityName="Crabo &amp; Lafontaine" authorityYear="2009" box="[538,640,1187,1214]" class="Insecta" family="Noctuidae" genus="Lasionycta" kingdom="Animalia" order="Lepidoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="frigida">
<emphasis box="[538,554,1188,1213]" italics="true" pageId="44" pageNumber="45">L</emphasis>
. 
<emphasis box="[568,640,1187,1213]" italics="true" pageId="44" pageNumber="45">frigida</emphasis>
</taxonomicName>
occurs harbors the southernmost populations of several arctic and subarctic 
<taxonomicName box="[548,685,1223,1249]" class="Insecta" kingdom="Animalia" order="Lepidoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="order">Lepidoptera</taxonomicName>
suggesting that 
<taxonomicName authorityName="Crabo &amp; Lafontaine" authorityYear="2009" box="[872,976,1223,1249]" class="Insecta" family="Noctuidae" genus="Lasionycta" kingdom="Animalia" order="Lepidoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="frigida">
<emphasis box="[872,888,1223,1248]" italics="true" pageId="44" pageNumber="45">L</emphasis>
. 
<emphasis box="[904,976,1223,1249]" italics="true" pageId="44" pageNumber="45">frigida</emphasis>
</taxonomicName>
might be a subarctic species. It should be sought farther north in 
<collectingRegion box="[749,822,1258,1284]" country="Canada" name="Yukon" pageId="44" pageNumber="45">Yukon</collectingRegion>
and 
<collectingRegion box="[879,956,1258,1284]" country="United States of America" name="Alaska" pageId="44" pageNumber="45">Alaska</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>