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<document id="684FC585347ADFDD744F90B63B0BE5F1" ID-CLB-Dataset="46210" ID-DOI="10.5281/zenodo.278895" ID-GBIF-Dataset="3498da36-33bd-4135-a475-9c978c7f7abc" ID-ISSN="1175-5326" ID-Zenodo-Dep="278895" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460397671527" checkinUser="plazi" docAuthor="Borrero, Francisco J. &amp; Breure, Abraham S. H." docDate="2011" docId="5C37A735FFD4BA4EFF28B4BAF597D34C" docLanguage="en" docName="zt03054p059.pdf" docOrigin="Zootaxa 3054" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Plekocheilus (Eurytus) labiosus Borrero &amp; Breure, 2011, spec. nov." docType="treatment" docVersion="9" lastPageNumber="48" masterDocId="A00EDF4DFFFABA61FFBFB77CF716D113" masterDocTitle="The Amphibulimidae (Mollusca: Gastropoda: Orthalicoidea) from Colombia and adjacent areas" masterLastPageNumber="59" masterPageNumber="1" pageNumber="47" updateTime="1698298882325" updateUser="ExternalLinkService">
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<mods:title id="42B7456AA80ADFCC0CCBF8C18A5665C2">The Amphibulimidae (Mollusca: Gastropoda: Orthalicoidea) from Colombia and adjacent areas</mods:title>
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<mods:namePart id="56B8F24F734929C7D79769F506936821">Borrero, Francisco J.</mods:namePart>
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<mods:namePart id="8E6C8CE4B308DD92C8E2FDF10964FA1B">Breure, Abraham S. H.</mods:namePart>
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<mods:date id="F5B06894F10B7FEA50448E0469C17527">2011</mods:date>
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<treatment id="5C37A735FFD4BA4EFF28B4BAF597D34C" ID-DOI="http://doi.org/10.5281/zenodo.6195132" ID-GBIF-Taxon="119418608" ID-Zenodo-Dep="6195132" LSID="urn:lsid:plazi:treatment:5C37A735FFD4BA4EFF28B4BAF597D34C" httpUri="http://treatment.plazi.org/id/5C37A735FFD4BA4EFF28B4BAF597D34C" lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47">
<subSubSection id="9C8445A8FFD4BA4FFF28B4BAF670D512" pageId="46" pageNumber="47" type="nomenclature">
<paragraph id="D4211623FFD4BA4FFF28B4BAF580D2F3" blockId="46.[151,662,966,1025]" box="[151,662,966,992]" pageId="46" pageNumber="47">
<heading id="8F69A14FFFD4BA4FFF28B4BAF580D2F3" bold="true" box="[151,662,966,992]" fontSize="11" level="1" pageId="46" pageNumber="47" reason="1">
<emphasis id="E6EACA31FFD4BA4FFF28B4BAF580D2F3" bold="true" box="[151,662,966,992]" pageId="46" pageNumber="47">
<taxonomicName id="139E6DA0FFD4BA4FFF28B4BAF501D2F3" box="[151,535,966,992]" class="Gastropoda" family="Orthalicidae" genus="Plekocheilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="labiosus" status="sp. nov." subGenus="Eurytus">
<emphasis id="E6EACA31FFD4BA4FFF28B4BAF624D2F3" bold="true" box="[151,306,966,992]" italics="true" pageId="46" pageNumber="47">Plekocheilus</emphasis>
(
<emphasis id="E6EACA31FFD4BA4FFEFCB4BAF6B5D2F3" bold="true" box="[323,419,966,992]" italics="true" pageId="46" pageNumber="47">Eurytus</emphasis>
)
<emphasis id="E6EACA31FFD4BA4FFE0BB4BAF501D2F3" bold="true" box="[436,535,966,992]" italics="true" pageId="46" pageNumber="47">labiosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FDD9774AFFD4BA4FFDA1B4BAF580D2F3" box="[542,662,966,992]" pageId="46" pageNumber="47" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="D4211623FFD4BA4FFF28B495F670D512" blockId="46.[151,662,966,1025]" box="[151,358,1001,1025]" pageId="46" pageNumber="47">(Figs 12A, 18HI)</paragraph>
</subSubSection>
<subSubSection id="9C8445A8FFD4BA4FFF28B34DF4ECD57E" pageId="46" pageNumber="47" type="diagnosis">
<paragraph id="D4211623FFD4BA4FFF28B34DF4ECD57E" blockId="46.[151,1437,1073,2033]" pageId="46" pageNumber="47">
<emphasis id="E6EACA31FFD4BA4FFF28B34DF607D559" bold="true" box="[151,273,1073,1098]" pageId="46" pageNumber="47">Diagnosis.</emphasis>
A medium-sized
<taxonomicName id="139E6DA0FFD4BA4FFE59B34DF5FDD55A" authority="(Eurytus)" baseAuthorityName="Eurytus" box="[486,747,1073,1097]" class="Gastropoda" family="Orthalicidae" genus="Plekocheilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="46" pageNumber="47" phylum="Mollusca" rank="genus">
<emphasis id="E6EACA31FFD4BA4FFE59B34DF563D55A" box="[486,629,1073,1097]" italics="true" pageId="46" pageNumber="47">Plekocheilus</emphasis>
(
<emphasis id="E6EACA31FFD4BA4FFD34B34EF5F4D55A" box="[651,738,1074,1097]" italics="true" pageId="46" pageNumber="47">Eurytus</emphasis>
)
</taxonomicName>
characterized by a very expanded and reflexed peristome, sculpture of fine longitudinal growth striae, and well defined columellar fold.
</paragraph>
</subSubSection>
<subSubSection id="9C8445A8FFD4BA4FFF7AB305F559D7E6" pageId="46" pageNumber="47" type="description">
<paragraph id="D4211623FFD4BA4FFF7AB305F648D7BE" blockId="46.[151,1437,1073,2033]" pageId="46" pageNumber="47">
<emphasis id="E6EACA31FFD4BA4FFF7AB305F640D581" bold="true" box="[197,342,1145,1170]" pageId="46" pageNumber="47">Description.</emphasis>
Shell
<quantity id="1366BBC6FFD4BA4FFE21B305F515D582" box="[414,515,1145,1169]" metricMagnitude="-2" metricUnit="m" metricValue="4.02" pageId="46" pageNumber="47" unit="mm" value="40.2">40.2 mm</quantity>
, about 1.63 times longer than wide, imperforate, ovate, sides strongly convex, relatively thick. Periostracum very thin, honey in color. Shell color pattern consisting of irregularly arranged light brown blotches, but these do not coalesce into a pattern. Surface (hardly) shining, sculptured with more or less regularly spaced, fine axial growth striae (37 to a centimeter in the ventral portion of the last whorl). Each stria consists of series of short sections or elongated granules, regularly interrupted by lower and shorter longitudinal sections, giving the impression that the striae are “beaded,” quite noticeable under magnification. The lower sections are also arranged in a uniform manner, so a spiral pattern of grooves is formed, only evident under magnification. This pattern is present in all sub-apical whorls, but is stronger in the body whorl. Protoconch partly eroded, but with what may be a granular sculpture, thus not inconsistent with that typical of
<taxonomicName id="139E6DA0FFD4BA4FFBFFB2E5F7CDD4C6" authority="Breure 1979" authorityName="Breure" authorityYear="1979" class="Gastropoda" family="Orthalicidae" genus="Plekocheilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="46" pageNumber="47" phylum="Mollusca" rank="subGenus" subGenus="Eurytus">
<emphasis id="E6EACA31FFD4BA4FFBFFB2E5F3D9D4A2" box="[1088,1231,1433,1457]" italics="true" pageId="46" pageNumber="47">Plekocheilus</emphasis>
(
<emphasis id="E6EACA31FFD4BA4FFB60B2E6F220D4A2" box="[1247,1334,1434,1457]" italics="true" pageId="46" pageNumber="47">Eurytus</emphasis>
) (
<bibRefCitation id="B00F6BD2FFD4BA4FFAF1B2E5F7C5D4C6" author="Breure" pageId="46" pageNumber="47" refString="Breure, A. S. H. (1979) Systematics, phylogeny and zoogeography of Bulimulinae (Mollusca). Zoologische Verhandelingen, 168, 1 - 215." type="journal article" year="1979">Breure 1979</bibRefCitation>
)
</taxonomicName>
. Whorls 5.5, convex, more so the last whorl; suture simple, impressed, hardly descending, going up again before the attachment of the outer lip to the last whorl. Aperture sub-ovate, about 1.6 times as long as wide and about 0.69 times the total shell height; color within light brown, with a few irregular spots of the same color as those in the external coloration. Columellar margin straight initially, then bending to an angle of about 140° between the upper and lower portions; a well defined columellar fold is present midway. Peristome very expanded and reflexed, but not thickened; the widest portion approximately 1/6 of the aperture width. Parietal wall with a thin, white callus.
</paragraph>
<paragraph id="D4211623FFD4BA4FFF7AB1C5F559D7E6" blockId="46.[151,1437,1073,2033]" pageId="46" pageNumber="47">
Dimensions in mm:
<typeStatus id="0B25A881FFD4BA4FFE13B1C5F518D7C2" box="[428,526,1721,1745]" pageId="46" pageNumber="47" type="holotype">holotype</typeStatus>
H 40.2, D 24.7, HA 27.7, WA 17.2, LW 34.5, 5.5 whorls.
<typeStatus id="0B25A881FFD4BA4FFB14B1C6F21BD7C1" box="[1195,1293,1722,1746]" pageId="46" pageNumber="47" type="paratype">Paratype</typeStatus>
H 74, D 43, HA 59.2, WA 33, LW 64.7, 5.5 whorls.
</paragraph>
</subSubSection>
<subSubSection id="9C8445A8FFD4BA4FFF7AB07EF623D696" pageId="46" pageNumber="47" type="materials_examined">
<paragraph id="D4211623FFD4BA4FFF7AB07EF614D62E" blockId="46.[151,1437,1073,2033]" pageId="46" pageNumber="47">
<emphasis id="E6EACA31FFD4BA4FFF7AB07EF677D609" bold="true" box="[197,353,1793,1818]" pageId="46" pageNumber="47">
<typeStatus id="0B25A881FFD4BA4FFF7AB07EF7E8D609" box="[197,254,1794,1818]" pageId="46" pageNumber="47">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="AC8956B3FFD4BA4FFED5B07DF6CDD60A" box="[362,475,1793,1817]" name="Colombia" pageId="46" pageNumber="47">Colombia</collectingCountry>
, Cundinamarca, Parque Nacional Chingaza, Embalse de Chuza [
<geoCoordinate id="B1AA70E4FFD4BA4FFB12B07DF206D60A" box="[1197,1296,1793,1817]" direction="north" orientation="latitude" pageId="46" pageNumber="47" precision="925" value="4.616667">04°37N</geoCoordinate>
<geoCoordinate id="B1AA70E4FFD4BA4FFAA8B07DF287D60A" box="[1303,1425,1793,1817]" direction="west" orientation="longitude" pageId="46" pageNumber="47" precision="925" value="-73.71667">073°43W</geoCoordinate>
], ~
<quantity id="1366BBC6FFD4BA4FFF1BB059F7E8D62E" box="[164,254,1829,1853]" metricMagnitude="3" metricUnit="m" metricValue="3.1" pageId="46" pageNumber="47" unit="m" value="3100.0">3100 m</quantity>
.
</paragraph>
<paragraph id="D4211623FFD4BA4FFF7AB036F623D696" blockId="46.[151,1437,1073,2033]" pageId="46" pageNumber="47">
<emphasis id="E6EACA31FFD4BA4FFF7AB036F661D671" bold="true" box="[197,375,1865,1890]" pageId="46" pageNumber="47">
<typeStatus id="0B25A881FFD4BA4FFF7AB036F616D671" box="[197,256,1866,1890]" pageId="46" pageNumber="47">Type</typeStatus>
material.
</emphasis>
<typeStatus id="0B25A881FFD4BA4FFE3DB035F6FCD672" box="[386,490,1865,1889]" pageId="46" pageNumber="47" type="holotype">Holotype</typeStatus>
<collectionCode id="B28F8EE6FFD4BA4FFE4AB036F539D672" box="[501,559,1866,1889]" pageId="46" pageNumber="47">UVZ</collectionCode>
97034, C. Alvarez leg.,
<date id="A02030E3FFD4BA4FFCE8B035F4BFD672" box="[855,937,1865,1889]" pageId="46" pageNumber="47" value="1997-02">ii.1997</date>
.
<typeStatus id="0B25A881FFD4BA4FFC07B036F309D671" box="[952,1055,1866,1890]" pageId="46" pageNumber="47" type="paratype">Paratype</typeStatus>
:
<collectionCode id="B28F8EE6FFD4BA4FFB92B036F392D672" box="[1069,1156,1866,1889]" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="46" pageNumber="47">UMMZ</collectionCode>
13843, “
<collectingCountry id="AC8956B3FFD4BA4FFB4DB035F27ED672" box="[1266,1384,1865,1889]" name="Colombia" pageId="46" pageNumber="47">Colombia</collectingCountry>
,” ex Stearns Colln.
</paragraph>
</subSubSection>
<subSubSection id="9C8445A8FFD4BA4EFF7AB0EEF367D304" lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47" type="discussion">
<paragraph id="D4211623FFD4BA4FFF7AB0EEF33ED6E2" blockId="46.[151,1437,1073,2033]" pageId="46" pageNumber="47">
<emphasis id="E6EACA31FFD4BA4FFF7AB0EEF62FD6B9" bold="true" box="[197,313,1938,1962]" pageId="46" pageNumber="47">Remarks.</emphasis>
This new taxon resembles
<taxonomicName id="139E6DA0FFD4BA4FFDD8B0EEF432D6BA" box="[615,804,1937,1961]" class="Gastropoda" family="Orthalicidae" genus="Plekocheilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="pulicarius" subGenus="Eurytus">
<emphasis id="E6EACA31FFD4BA4FFDD8B0EEF563D6BA" box="[615,629,1938,1961]" italics="true" pageId="46" pageNumber="47">P</emphasis>
. (
<emphasis id="E6EACA31FFD4BA4FFD33B0EEF58BD6BA" box="[652,669,1938,1961]" italics="true" pageId="46" pageNumber="47">E</emphasis>
.)
<emphasis id="E6EACA31FFD4BA4FFD0CB0EDF432D6BA" box="[691,804,1937,1961]" italics="true" pageId="46" pageNumber="47">pulicarius</emphasis>
</taxonomicName>
in general shape, size and sculpture.
<taxonomicName id="139E6DA0FFD4BA4FFB7DB0EEF258D6BA" box="[1218,1358,1937,1961]" class="Gastropoda" family="Orthalicidae" genus="Plekocheilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="pulicarius">
<emphasis id="E6EACA31FFD4BA4FFB7DB0EEF258D6BA" box="[1218,1358,1937,1961]" italics="true" pageId="46" pageNumber="47">P. pulicarius</emphasis>
</taxonomicName>
exhibits a wide variation in coloration, the lighter forms being similar to
<taxonomicName id="139E6DA0FFD4BA4FFC2CB0CAF31AD6DE" box="[915,1036,1973,1997]" class="Gastropoda" family="Orthalicidae" genus="Plekocheilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="labiosus">
<emphasis id="E6EACA31FFD4BA4FFC2CB0CAF31AD6DE" box="[915,1036,1973,1997]" italics="true" pageId="46" pageNumber="47">P. labiosus</emphasis>
</taxonomicName>
. It is easily distinguishable from
<taxonomicName id="139E6DA0FFD4BA4FFA38B0CAF61ED6E2" class="Gastropoda" family="Orthalicidae" genus="Plekocheilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="pulicarius">
<emphasis id="E6EACA31FFD4BA4FFA38B0CAF61ED6E2" italics="true" pageId="46" pageNumber="47">P. pulicarius</emphasis>
</taxonomicName>
in that
<taxonomicName id="139E6DA0FFD4BA4FFEE2B0A6F6C1D6E2" box="[349,471,2009,2033]" class="Gastropoda" family="Orthalicidae" genus="Plekocheilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="labiosus">
<emphasis id="E6EACA31FFD4BA4FFEE2B0A6F6C1D6E2" box="[349,471,2009,2033]" italics="true" pageId="46" pageNumber="47">P. labiosus</emphasis>
</taxonomicName>
has a considerably wider, more expanded peristome.
</paragraph>
<paragraph id="D4211623FFD5BA4EFF7AB7EBF425D0E7" blockId="47.[151,1437,151,607]" pageId="47" pageNumber="48">
The
<typeStatus id="0B25A881FFD5BA4EFF47B7EBF64CD1BC" box="[248,346,151,175]" pageId="47" pageNumber="48" type="holotype">holotype</typeStatus>
is a partially worn shell, bleached mostly dorsally and laterally, lacking part of the periostracum. The
<typeStatus id="0B25A881FFD5BA4EFF75B7C1F63CD1C6" box="[202,298,189,213]" pageId="47" pageNumber="48" type="paratype">paratype</typeStatus>
is in better condition, with the periostracum nearly complete, but lacks precise locality information. The
<typeStatus id="0B25A881FFD5BA4EFF74B79CF63AD1EB" box="[203,300,224,248]" pageId="47" pageNumber="48" type="paratype">paratype</typeStatus>
is substantially larger than the
<typeStatus id="0B25A881FFD5BA4EFD2AB7A3F5ECD1E4" box="[661,762,223,247]" pageId="47" pageNumber="48" type="holotype">holotype</typeStatus>
, but the shape of the shells and ratios among the measurements presented above are remarkably similar. Describing this species on the basis of only two shells and a single locality is justified on the basis of the unique features exhibited by this new taxon. No other species of
<taxonomicName id="139E6DA0FFD5BA4EFAB2B654F28AD053" box="[1293,1436,296,320]" class="Gastropoda" family="Orthalicidae" genus="Plekocheilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="47" pageNumber="48" phylum="Mollusca" rank="genus">
<emphasis id="E6EACA31FFD5BA4EFAB2B654F28AD053" box="[1293,1436,296,320]" italics="true" pageId="47" pageNumber="48">Plekocheilus</emphasis>
</taxonomicName>
from northwestern South
<collectingCountry id="AC8956B3FFD5BA4EFE04B630F50AD077" box="[443,540,332,356]" name="United States of America" pageId="47" pageNumber="48">America</collectingCountry>
has a similar dramatically expanded and reflexed peristome, making it unique and novel. The
<typeStatus id="0B25A881FFD5BA4EFEF8B60CF661D09B" box="[327,375,368,392]" pageId="47" pageNumber="48">type</typeStatus>
locality is not exactly precise, as the specimen was collected dead, presumably on the side of Embalse Chuza, and the elevation given is approximate.
<taxonomicName id="139E6DA0FFD5BA4EFCA9B6E8F378D0BF" box="[790,1134,404,428]" class="Gastropoda" family="Orthalicidae" genus="Plekocheilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="47" pageNumber="48" phylum="Mollusca" rank="species" species="succineoides" subGenus="Aeropictus">
<emphasis id="E6EACA31FFD5BA4EFCA9B6E8F4B3D0BF" box="[790,933,404,428]" italics="true" pageId="47" pageNumber="48">Plekocheilus</emphasis>
(
<emphasis id="E6EACA31FFD5BA4EFC09B6E8F4D8D0BF" box="[950,974,404,428]" italics="true" pageId="47" pageNumber="48">A.</emphasis>
)
<emphasis id="E6EACA31FFD5BA4EFC60B6E8F378D0BF" box="[991,1134,404,428]" italics="true" pageId="47" pageNumber="48">succineoides</emphasis>
</taxonomicName>
co-occurs with
<taxonomicName id="139E6DA0FFD5BA4EFA9CB6E9F28AD0BF" box="[1315,1436,404,428]" class="Gastropoda" family="Orthalicidae" genus="Plekocheilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="47" pageNumber="48" phylum="Mollusca" rank="species" species="labiosus">
<emphasis id="E6EACA31FFD5BA4EFA9CB6E9F28AD0BF" box="[1315,1436,404,428]" italics="true" pageId="47" pageNumber="48">P. labiosus</emphasis>
</taxonomicName>
at this general locality; this species has a wide distribution mainly in the Altiplano Cundi-Boyacense; the two species are easily separated by shape, sculpture and coloration.
</paragraph>
<paragraph id="D4211623FFD5BA4EFF7AB57CF367D304" blockId="47.[151,1437,151,607]" box="[197,1137,511,536]" pageId="47" pageNumber="48">
<emphasis id="E6EACA31FFD5BA4EFF7AB57CF63FD30B" bold="true" box="[197,297,512,536]" pageId="47" pageNumber="48">Ecology.</emphasis>
The
<typeStatus id="0B25A881FFD5BA4EFEDDB683F6D2D304" box="[354,452,511,535]" pageId="47" pageNumber="48" type="holotype">holotype</typeStatus>
was found “between stones, with grasses and isolated trees.”
</paragraph>
</subSubSection>
<subSubSection id="9C8445A8FFD5BA4EFF7AB558F597D34C" pageId="47" pageNumber="48" type="etymology">
<paragraph id="D4211623FFD5BA4EFF7AB558F597D34C" blockId="47.[151,1437,151,607]" pageId="47" pageNumber="48">
<emphasis id="E6EACA31FFD5BA4EFF7AB558F65DD32F" bold="true" box="[197,331,548,572]" pageId="47" pageNumber="48">Etymology.</emphasis>
<taxonomicName id="139E6DA0FFD5BA4EFEECB558F51ED32F" authority="Latin" authorityName="Latin" box="[339,520,548,572]" class="Gastropoda" family="Orthalicidae" genus="Plekocheilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="47" pageNumber="48" phylum="Mollusca" rank="species" species="labiosus">
<emphasis id="E6EACA31FFD5BA4EFEECB558F6A7D32F" box="[339,433,548,572]" italics="true" pageId="47" pageNumber="48">labiosus</emphasis>
(Latin)
</taxonomicName>
, having large lips. The name refers to the remarkably expanded peristome of this species. The specific epithet is an adjective.
</paragraph>
</subSubSection>
</treatment>
</document>