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<mods:title id="8D870648E633D33CA998C0FE63EF5CC3">A new classification of Callianassidae and related families (Crustacea: Decapoda: Axiidea) derived from a molecular phylogeny with morphological support</mods:title>
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<paragraph id="8B9D0B4DF2560835FC98B5B5A9C5FE62" blockId="8.[809,1258,435,457]" box="[809,1258,435,457]" pageId="8" pageNumber="81">
<heading id="D0D5BC21F2560835FC98B5B5A9C5FE62" bold="true" box="[809,1258,435,457]" fontSize="9" level="2" pageId="8" pageNumber="81" reason="2">
<taxonomicName id="4C2270CEF2560835FC98B5B5A9C5FE62" ID-CoL="BMJWX" authority="Manning and Felder, 1991" authorityName="Manning and Felder" authorityYear="1991" box="[809,1258,435,457]" class="Malacostraca" family="Anacalliacidae" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="81" phylum="Arthropoda" rank="family">
<emphasis id="B956D75FF2560835FC98B5B5A9C5FE62" bold="true" box="[809,1258,435,457]" pageId="8" pageNumber="81">
Anacalliacidae
<bibRefCitation id="EFB376BCF2560835FC7EB5B5A9C5FE62" author="Manning, R. B. &amp; Felder, D. L." box="[975,1258,435,457]" pageId="8" pageNumber="81" pagination="764 - 792" refId="ref52690" refString="Manning, R. B., and Felder, D. L. 1991. Revision of the American Callianassidae (Crustacea: Decapoda: Thalassinidea). Proceedings of the Biological Society of Washington 104: 764 - 792. http: // biodiversitylibrary. org / page / 34809466" type="journal article" year="1991">Manning and Felder, 1991</bibRefCitation>
</emphasis>
</taxonomicName>
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<paragraph id="8B9D0B4DF2560835FC98B5E6AE50FE5C" blockId="8.[809,895,480,502]" box="[809,895,480,502]" pageId="8" pageNumber="81">
<figureCitation id="131917C8F2560835FC98B5E6AE50FE5C" box="[809,895,480,502]" captionStart="Figure 5" captionStartId="9.[147,207,1010,1029]" captionTargetBox="[147,1439,231,986]" captionTargetPageId="9" captionText="Figure 5. Diagnostic characters of Anacalliacidae, Anacalliax: a, dorsal anterior carapace, eyestalks, antennules, antennae; b, lateral carapace; c, maxilliped 3; d, male pleopod 1; e, female pleopod 2; f, telson, uropod." figureDoi="http://doi.org/10.5281/zenodo.12214189" httpUri="https://zenodo.org/record/12214189/files/figure.png" pageId="8" pageNumber="81">Figure 5</figureCitation>
</paragraph>
<paragraph id="8B9D0B4DF2560835FCFEB60DA9A3FDFE" blockId="8.[847,1414,523,596]" pageId="8" pageNumber="81">
Anacalliinae
<bibRefCitation id="EFB376BCF2560835FC7BB60DA9C8FDB4" author="Manning, R. B. &amp; Felder, D. L." box="[970,1255,523,543]" pageId="8" pageNumber="81" pagination="764 - 792" refId="ref52690" refString="Manning, R. B., and Felder, D. L. 1991. Revision of the American Callianassidae (Crustacea: Decapoda: Thalassinidea). Proceedings of the Biological Society of Washington 104: 764 - 792. http: // biodiversitylibrary. org / page / 34809466" type="journal article" year="1991">Manning and Felder, 1991: 786</bibRefCitation>
.
<taxonomicName id="4C2270CEF2560835FCFEB620AEF7FD93" box="[847,984,550,569]" pageId="8" pageNumber="81" rank="subFamily" subFamily="Anacalliacinae">Anacalliacinae</taxonomicName>
.—
<bibRefCitation id="EFB376BCF2560835FC43B620A9BBFD93" author="Sakai, K." box="[1010,1172,550,569]" pageId="8" pageNumber="81" pagination="1 - 152" refId="ref55068" refString="Sakai, K. 1999 a. Synopsis of the family Callianassidae, with keys to subfamilies, genera and species, and the description of new taxa (Crustacea: Decapoda: Thalassinidea). Zoologische Verhandelingen 326: 1 - 152." type="journal article" year="1999">Sakai, 1999a: 126</bibRefCitation>
.—
<bibRefCitation id="EFB376BCF2560835FB1FB620A8ADFD93" author="Sakai, K." box="[1198,1410,550,569]" pageId="8" pageNumber="81" pagination="1 - 285" refId="ref55408" refString="Sakai, K. 2005 b. Callianassoidea of the world (Decapoda: Thalassinidea). Crustaceana Monographs 4: 1 - 285." type="journal article" year="2005">Sakai, 2005b: 208210</bibRefCitation>
.
<taxonomicName id="4C2270CEF2560835FCFEB647AEF7FDFE" authorityName="Manning and Felder" authorityYear="1991" box="[847,984,577,596]" class="Malacostraca" family="Anacalliacidae" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="81" phylum="Arthropoda" rank="family">Anacalliacidae</taxonomicName>
.—
<bibRefCitation id="EFB376BCF2560835FC43B647A9A7FDFE" author="Sakai, K." box="[1010,1160,577,596]" pageId="8" pageNumber="81" pagination="1 - 616" refId="ref55522" refString="Sakai, K. 2011. Axioidea of the world and a reconsideration of the Callianassoidea (Decapoda, Thalassinidea, Callianassida). Crustaceana Monographs 13: 1 - 616." type="journal article" year="2011">Sakai, 2011: 341</bibRefCitation>
.
</paragraph>
<paragraph id="8B9D0B4DF2560835FC98B66AA82CFD2A" blockId="8.[809,1283,619,641]" box="[809,1283,619,641]" pageId="8" pageNumber="81">
<emphasis id="B956D75FF2560835FC98B66AAEB5FD2B" box="[809,922,620,641]" italics="true" pageId="8" pageNumber="81">
<typeStatus id="5499B5EFF2560835FC98B66AAE77FD2B" box="[809,856,620,641]" pageId="8" pageNumber="81">Type</typeStatus>
genus
</emphasis>
.
<taxonomicName id="4C2270CEF2560835FC17B66DA9D1FD2A" authority="de Saint Laurent, 1973" authorityName="de Saint Laurent" authorityYear="1973" box="[934,1278,619,641]" class="Malacostraca" family="Anacalliacidae" genus="Anacalliax" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="81" phylum="Arthropoda" rank="genus">
<emphasis id="B956D75FF2560835FC17B66DA93EFD2A" box="[934,1041,619,640]" italics="true" pageId="8" pageNumber="81">Anacalliax</emphasis>
de
<bibRefCitation id="EFB376BCF2560835FB87B66DA9D1FD2A" author="Saint Laurent, M. de" box="[1078,1278,619,641]" pageId="8" pageNumber="81" pagination="513 - 516" refId="ref54586" refString="Saint Laurent, M. de 1973. Sur la systematique et la phylogenie des Thalassinidea: definition des familles des Callianassidae et des Upogebiidae et diagnose de cinq genres nouveaux. Comptes Rendus Hebdomadaires de Seances de l'Academie des Sciences, Paris (Ser. D) 277: 513 - 516." type="journal article" year="1973">Saint Laurent, 1973</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="8B9D0B4DF2560835FC98B69EA94CFA48" blockId="8.[808,1442,663,1506]" pageId="8" pageNumber="81">
<emphasis id="B956D75FF2560835FC98B69EAEA0FD07" box="[809,911,664,685]" italics="true" pageId="8" pageNumber="81">Diagnosis</emphasis>
.
<emphasis id="B956D75FF2560835FC14B69EAEA8FD61" bold="true" italics="true" pageId="8" pageNumber="81">Rostrum flat, short, triangular, shorter than eyestalks</emphasis>
; median carina on rostrum only; gastric carinae absent; cervical groove well defined; suture between ocular lobe and end of linea thalassinica horizontal in lateral view; anterior branchiostegal margin sinusoidal or semicircular; anterior branchiostegal lobe simple, scarcely calcified, merging smoothly with anterodorsal branchiostegal angle and anterolateral margin of carapace; posterior margin of carapace without lateral lobes, pleomere 1 without anterolateral lobes, weakly chitinised. Eyestalks flattened, contiguous, with subdistal dorsal cornea. Antennal scaphocerite elongate. Maxilla scaphognathite without long seta on posterior lobe extending into branchial chamber. Maxilliped 1 epipod with acute anterior lobe lying alongside exopod. Maxilliped 3 propodus longer than wide, not prominently lobed on lower margin; dactylus slender, digitiform, with setae irregularly spaced along all margins. Cheliped merus lower margin spinose; major cheliped palm oval in cross-section, barely crested above or below. Pereopod 3 propodus broad, with proximal lobe on lower margin, without distal spiniform setae on lateral face (often with 1 distal spiniform seta on lower margin). Pereopod 5 minutely chelate or subchelate. Female pleopod 2 rami narrower and with more reduced setation than pleopods 35; endopod 25 times as long as wide. Pleopods 35 with oblique peduncles, endopods oval, exopods attached laterally, not proximally lobed, shorter than and barely overlapping endopods; appendices internae reduced and almost embedded in mesial margin of endopod.
<emphasis id="B956D75FF2560835FB56B1A9A973FA48" bold="true" italics="true" pageId="8" pageNumber="81">Uropodal exopod with margin divided by notch</emphasis>
.
</paragraph>
<paragraph id="8B9D0B4DF2560835FC98B1FFA809F9E0" blockId="8.[808,1440,1529,1903]" pageId="8" pageNumber="81">
<emphasis id="B956D75FF2560835FC98B1FFAEADF9A4" box="[809,898,1529,1550]" italics="true" pageId="8" pageNumber="81">Remarks</emphasis>
. The only genus,
<taxonomicName id="4C2270CEF2560835FB8DB1FFA8B2F9A4" authority="de Saint Laurent, 1973" authorityName="de Saint Laurent" authorityYear="1973" box="[1084,1437,1529,1551]" class="Malacostraca" family="Anacalliacidae" genus="Anacalliax" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="81" phylum="Arthropoda" rank="genus">
<emphasis id="B956D75FF2560835FB8DB1FFA983F9A4" box="[1084,1196,1529,1550]" italics="true" pageId="8" pageNumber="81">Anacalliax</emphasis>
de
<bibRefCitation id="EFB376BCF2560835FB60B1FFA8B2F9A4" author="Saint Laurent, M. de" box="[1233,1437,1529,1551]" pageId="8" pageNumber="81" pagination="513 - 516" refId="ref54586" refString="Saint Laurent, M. de 1973. Sur la systematique et la phylogenie des Thalassinidea: definition des familles des Callianassidae et des Upogebiidae et diagnose de cinq genres nouveaux. Comptes Rendus Hebdomadaires de Seances de l'Academie des Sciences, Paris (Ser. D) 277: 513 - 516." type="journal article" year="1973">Saint Laurent, 1973</bibRefCitation>
</taxonomicName>
, is recognised by the unique combination of a short flat rostrum and the uropodal exopod having a marginal notch.
</paragraph>
<paragraph id="8B9D0B4DF2560834FCFEB257AC8FFB56" blockId="8.[808,1440,1529,1903]" lastBlockId="9.[146,780,1129,1681]" lastPageId="9" lastPageNumber="82" pageId="8" pageNumber="81">
The subfamily
<taxonomicName id="4C2270CEF2560835FC43B257A9A4F9CD" box="[1010,1163,1617,1639]" pageId="8" pageNumber="81" rank="subFamily" subFamily="Anacalliacinae">Anacalliacinae</taxonomicName>
(misspelled Anacalliinae) was erected by
<bibRefCitation id="EFB376BCF2560835FC62B268A9DCF92E" author="Manning, R. B. &amp; Felder, D. L." box="[979,1267,1646,1668]" pageId="8" pageNumber="81" pagination="764 - 792" refId="ref52690" refString="Manning, R. B., and Felder, D. L. 1991. Revision of the American Callianassidae (Crustacea: Decapoda: Thalassinidea). Proceedings of the Biological Society of Washington 104: 764 - 792. http: // biodiversitylibrary. org / page / 34809466" type="journal article" year="1991">Manning and Felder (1991)</bibRefCitation>
as one of three subfamilies of
<taxonomicName id="4C2270CEF2560835FC73B28AA979F908" authorityName="Manning and Felder" authorityYear="1991" box="[962,1110,1676,1698]" class="Malacostraca" family="Ctenochelidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="81" phylum="Arthropoda" rank="family">Ctenochelidae</taxonomicName>
, the others being Ctenochelinae and Callianopsinae.
<bibRefCitation id="EFB376BCF2560835FBBDB2AFA815F914" author="Manning, R. B. &amp; Felder, D. L." box="[1036,1338,1705,1727]" pageId="8" pageNumber="81" pagination="764 - 792" refId="ref52690" refString="Manning, R. B., and Felder, D. L. 1991. Revision of the American Callianassidae (Crustacea: Decapoda: Thalassinidea). Proceedings of the Biological Society of Washington 104: 764 - 792. http: // biodiversitylibrary. org / page / 34809466" type="journal article" year="1991">Manning and Felder (1991)</bibRefCitation>
included
<taxonomicName id="4C2270CEF2560835FC98B2C0AE8DF971" authorityName="de Saint Laurent" authorityYear="1979" box="[809,930,1734,1755]" class="Malacostraca" family="Paracalliacidae" genus="Paracalliax" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="81" phylum="Arthropoda" rank="genus">
<emphasis id="B956D75FF2560835FC98B2C0AE8DF971" box="[809,930,1734,1755]" italics="true" pageId="8" pageNumber="81">Paracalliax</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2270CEF2560835FC0BB2C1A90EF976" authorityName="de Saint Laurent" authorityYear="1973" box="[954,1057,1735,1756]" class="Malacostraca" family="Callianassidae" genus="Gourretia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="81" phylum="Arthropoda" rank="genus">
<emphasis id="B956D75FF2560835FC0BB2C1A90EF976" box="[954,1057,1735,1756]" italics="true" pageId="8" pageNumber="81">Gourretia</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2270CEF2560835FBD9B2C1A9F5F976" authorityName="Manning and Felder" authorityYear="1991" box="[1128,1242,1735,1756]" class="Malacostraca" family="Callianassidae" genus="Dawsonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="81" phylum="Arthropoda" rank="genus">
<emphasis id="B956D75FF2560835FBD9B2C1A9F5F976" box="[1128,1242,1735,1756]" italics="true" pageId="8" pageNumber="81">Dawsonius</emphasis>
</taxonomicName>
in the nominate subfamily and only their
<typeStatus id="5499B5EFF2560835FB88B2E2A949F953" box="[1081,1126,1764,1785]" pageId="8" pageNumber="81">type</typeStatus>
genus in the other two. This arrangement reflects in a single family, with all four basally derived families recognised in Robles et al.s (in press) analysis as a paraphyletic grouping.
<bibRefCitation id="EFB376BCF2560835FBF2B33AA9FDF8F8" author="Sakai, K." box="[1091,1234,1852,1874]" pageId="8" pageNumber="81" pagination="1 - 285" refId="ref55408" refString="Sakai, K. 2005 b. Callianassoidea of the world (Decapoda: Thalassinidea). Crustaceana Monographs 4: 1 - 285." type="journal article" year="2005">Sakai (2005b)</bibRefCitation>
treated the taxon as a subfamily of
<taxonomicName id="4C2270CEF2560835FC71B35FA97AF8C5" authorityName="Dana" authorityYear="1852" box="[960,1109,1881,1903]" class="Malacostraca" family="Callianassidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="81" phylum="Arthropoda" rank="family">Callianassidae</taxonomicName>
while retaining
<taxonomicName id="4C2270CEF2560835FB4AB35FA851F8C5" authorityName="Sakai" authorityYear="1999" box="[1275,1406,1881,1903]" class="Malacostraca" family="Gourretiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="81" phylum="Arthropoda" rank="family">Gourretiidae</taxonomicName>
for other subfamilies.
<bibRefCitation id="EFB376BCF2570834FEE6B06FACF6FBD4" author="Sakai, K." box="[343,473,1129,1151]" pageId="9" pageNumber="82" pagination="1 - 616" refId="ref55522" refString="Sakai, K. 2011. Axioidea of the world and a reconsideration of the Callianassoidea (Decapoda, Thalassinidea, Callianassida). Crustaceana Monographs 13: 1 - 616." type="journal article" year="2011">Sakai (2011)</bibRefCitation>
elevated the taxon to family level. The family comprises a single genus. Two genera erected by
<bibRefCitation id="EFB376BCF2570834FF03B0A1AC1EFB16" author="Sakai, K." box="[178,305,1191,1213]" pageId="9" pageNumber="82" pagination="1 - 616" refId="ref55522" refString="Sakai, K. 2011. Axioidea of the world and a reconsideration of the Callianassoidea (Decapoda, Thalassinidea, Callianassida). Crustaceana Monographs 13: 1 - 616." type="journal article" year="2011">Sakai (2011)</bibRefCitation>
are herein synonymised,
<taxonomicName id="4C2270CEF2570834FD8CB0A1AFF8FB16" authorityName="Sakai" authorityYear="2011" box="[573,727,1191,1212]" class="Malacostraca" family="Anacalliacidae" genus="Anacalliaopsis" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="82" phylum="Arthropoda" rank="genus">
<emphasis id="B956D75FF2570834FD8CB0A1AFF8FB16" box="[573,727,1191,1212]" italics="true" pageId="9" pageNumber="82">Anacalliaopsis</emphasis>
</taxonomicName>
with
<taxonomicName id="4C2270CEF2570834FF25B0C0AC2BFB71" authorityName="de Saint Laurent" authorityYear="1973" box="[148,260,1222,1243]" class="Malacostraca" family="Anacalliacidae" genus="Anacalliax" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="82" phylum="Arthropoda" rank="genus">
<emphasis id="B956D75FF2570834FF25B0C0AC2BFB71" box="[148,260,1222,1243]" italics="true" pageId="9" pageNumber="82">Anacalliax</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C2270CEF2570834FEC4B0C0ACDDFB71" authorityName="Sakai" authorityYear="2011" box="[373,498,1222,1243]" class="Malacostraca" family="Callichiridae" genus="Capecalliax" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="82" phylum="Arthropoda" rank="genus">
<emphasis id="B956D75FF2570834FEC4B0C0ACDDFB71" box="[373,498,1222,1243]" italics="true" pageId="9" pageNumber="82">Capecalliax</emphasis>
</taxonomicName>
with
<taxonomicName id="4C2270CEF2570834FDD2B0C0AE25FB71" authorityName="Sakai" authorityYear="2011" box="[611,778,1222,1243]" class="Malacostraca" family="Callianassidae" genus="Balsscallichirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="82" phylum="Arthropoda" rank="genus">
<emphasis id="B956D75FF2570834FDD2B0C0AE25FB71" box="[611,778,1222,1243]" italics="true" pageId="9" pageNumber="82">Balsscallichirus</emphasis>
</taxonomicName>
(
<taxonomicName id="4C2270CEF2570834FF2BB0E0AC0CFB56" authorityName="Manning and Felder" authorityYear="1991" box="[154,291,1254,1276]" class="Malacostraca" family="Callichiridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="82" phylum="Arthropoda" rank="family">Callichiridae</taxonomicName>
; see below).
</paragraph>
<caption id="DF5D5BC5F2570834FF22B7F4AE01FBB5" ID-DOI="http://doi.org/10.5281/zenodo.12214189" ID-Zenodo-Dep="12214189" httpUri="https://zenodo.org/record/12214189/files/figure.png" pageId="9" pageNumber="82" startId="9.[147,207,1010,1029]" targetBox="[147,1439,231,986]" targetPageId="9" targetType="figure">
<paragraph id="8B9D0B4DF2570834FF22B7F4AE01FBB5" blockId="9.[147,1448,1010,1056]" pageId="9" pageNumber="82">
Figure 5. Diagnostic characters of
<taxonomicName id="4C2270CEF2570834FE61B7F4AF74FBAF" authorityName="Manning and Felder" authorityYear="1991" box="[464,603,1010,1029]" class="Malacostraca" family="Anacalliacidae" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="82" phylum="Arthropoda" rank="family">Anacalliacidae</taxonomicName>
,
<taxonomicName id="4C2270CEF2570834FDD5B7F4AFE7FBAF" authorityName="de Saint Laurent" authorityYear="1973" box="[612,712,1010,1029]" class="Malacostraca" family="Anacalliacidae" genus="Anacalliax" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="82" phylum="Arthropoda" rank="genus">
<emphasis id="B956D75FF2570834FDD5B7F4AFE7FBAF" box="[612,712,1010,1029]" italics="true" pageId="9" pageNumber="82">Anacalliax</emphasis>
</taxonomicName>
: a, dorsal anterior carapace, eyestalks, antennules, antennae; b, lateral carapace; c, maxilliped 3; d, male pleopod 1; e, female pleopod 2; f, telson, uropod.
</paragraph>
</caption>
<paragraph id="8B9D0B4DF2570834FF08B103AF0CF999" blockId="9.[146,780,1129,1681]" pageId="9" pageNumber="82">
The similarity of the “dorsal oval” to that of
<taxonomicName id="4C2270CEF2570834FD21B103AE23FAB0" authorityName="Leach" authorityYear="1814" box="[656,780,1285,1306]" class="Malacostraca" family="Callianassidae" genus="Callianassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="82" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B956D75FF2570834FD21B103AE23FAB0" box="[656,780,1285,1306]" italics="true" pageId="9" pageNumber="82">Callianassa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2656A24F2570834FF22B122AD9CFA93" box="[147,179,1316,1337]" pageId="9" pageNumber="82" sensu="stricto">s.s.</taxonomicNameLabel>
was highlighted in
<bibRefCitation id="EFB376BCF2570834FECFB122AF23FA93" author="Sakai, K." box="[382,524,1316,1338]" pageId="9" pageNumber="82" pagination="1 - 616" refId="ref55522" refString="Sakai, K. 2011. Axioidea of the world and a reconsideration of the Callianassoidea (Decapoda, Thalassinidea, Callianassida). Crustaceana Monographs 13: 1 - 616." type="journal article" year="2011">Sakais (2011)</bibRefCitation>
discussion of this group, but this character grades to such an extent throughout all families that we were unable to use it at any level. The broad (almost axiid-like) article 2 of the male pleopod 1, presence of an appendix masculina on the male pleopod 2, absence of any tooth on the merus of the cheliped, absence of a dorsal plate on the uropodal exopod and absence of a distal lobe on the epipod of maxilliped 1 suggest a basal position for this family, as realised by
<bibRefCitation id="EFB376BCF2570834FEBBB218AF32F999" author="Manning, R. B. &amp; Felder, D. L." box="[266,541,1566,1588]" pageId="9" pageNumber="82" pagination="764 - 792" refId="ref52690" refString="Manning, R. B., and Felder, D. L. 1991. Revision of the American Callianassidae (Crustacea: Decapoda: Thalassinidea). Proceedings of the Biological Society of Washington 104: 764 - 792. http: // biodiversitylibrary. org / page / 34809466" type="journal article" year="1991">Manning and Felder (1991)</bibRefCitation>
.
</paragraph>
<paragraph id="8B9D0B4DF2570834FF08B23BAC76F93B" blockId="9.[146,780,1129,1681]" pageId="9" pageNumber="82">
The family is known only from the
<typeStatus id="5499B5EFF2570834FD91B23BAF62F9F8" box="[544,589,1597,1618]" pageId="9" pageNumber="82">type</typeStatus>
material of its two species. No specimen was available for inclusion in the molecular analysis.
</paragraph>
</subSubSection>
</treatment>
</document>