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<document id="AE32C659908FEDDCE799FC3C3B2CCE6B" ID-DOI="10.5281/zenodo.276621" ID-GBIF-Dataset="a6eb9c5a-9b0b-47df-a207-6c5ee54acebd" ID-ISSN="1175-5326" ID-Zenodo-Dep="276621" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460225034079" checkinUser="plazi" docAuthor="Latha, Mathews, Cuccodoro, Giulio &amp; Sabu, Thomas K." docDate="2011" docId="03FF87EFFFDCB858FF7235E6E16EF83E" docLanguage="en" docName="zt02745p029.pdf" docOrigin="Zootaxa 2745" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Ochicanthon murthyi Vinod &amp; Sabu, sp. nov." docType="treatment" docVersion="8" lastPageNumber="14" masterDocId="FFC6FF97FFD1B855FFE53643E06BFFEF" masterDocTitle="Taxonomy of the dung beetle genus Ochicanthon Vaz-de-Mello (Coleoptera: Scarabaeidae: Scarabaeinae) of the Indian subcontinent, with notes on distribution patterns and flightlessness" masterLastPageNumber="29" masterPageNumber="1" pageNumber="14" updateTime="1698238191473" updateUser="plazi">
<mods:mods id="BCFE7C0D0ECEEE51D108C15BDC9B9F21" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="8ADE44EA03AF3B83DD622E2B87AD3429">
<mods:title id="2EDFD3C36E361641FBB175B84F7A1531">Taxonomy of the dung beetle genus Ochicanthon Vaz-de-Mello (Coleoptera: Scarabaeidae: Scarabaeinae) of the Indian subcontinent, with notes on distribution patterns and flightlessness</mods:title>
</mods:titleInfo>
<mods:name id="E49EB510B39AD26D8AC0CA3A4C931762" type="personal">
<mods:role id="4003DDBB76F41081D9A87155397AD404">
<mods:roleTerm id="1B70C3C28034BD5052D3124BC26C6CD3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FBBBA1F40E3E36882879629778FA04DA">Latha, Mathews</mods:namePart>
</mods:name>
<mods:name id="F696A074C35664FD7EC9B19E2341AACB" type="personal">
<mods:role id="61397BB7564617BE9D37786B8DCF500D">
<mods:roleTerm id="AE65EC2755F26D6CA5B0FEA21E76CE20">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EC2780AE9831A365ADAA5A2E259FE9F0">Cuccodoro, Giulio</mods:namePart>
</mods:name>
<mods:name id="42E2D6C2602834B2DA4B8B4152E5BB42" type="personal">
<mods:role id="B08E1AC1E387CCD54E1B3AAAC0FADC59">
<mods:roleTerm id="FC3883B0994F34D44C09ACB8C42F4C72">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="871B0DB3A8190B2BF0ACCF05061EFDC6">Sabu, Thomas K.</mods:namePart>
</mods:name>
<mods:typeOfResource id="32010EC2DCF3C984480AE3F3803D9E1E">text</mods:typeOfResource>
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<mods:titleInfo id="B665CA5039C51AC3C7374F67E9F76A62">
<mods:title id="4833EA9E1A167B7EE93901BB7BCE7980">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="82068FA2DF812D5DF6738F765E2A84AD">
<mods:date id="B20268B0C7DDFEEDED15312D822C4395">2011</mods:date>
<mods:detail id="7DCA3A9F90C85A5865DAC65127AEC684" type="volume">
<mods:number id="4748756DCC88C935689670B913765A13">2745</mods:number>
</mods:detail>
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<mods:start id="38F9BBF8197A63D7BEC41C7169B231B3">1</mods:start>
<mods:end id="0DFFDB34ADB2F1398ACD724E5743B227">29</mods:end>
</mods:extent>
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<mods:classification id="B50B321ABF34913F85CC08E82D124D4A">journal article</mods:classification>
<mods:identifier id="E306007859085950611D2937C70BC482" type="DOI">10.5281/zenodo.276621</mods:identifier>
<mods:identifier id="30F84C3BEC67682FBD6BB246D29D5070" type="GBIF-Dataset">a6eb9c5a-9b0b-47df-a207-6c5ee54acebd</mods:identifier>
<mods:identifier id="1DAFA0F3DEAE2733F9292AE3823B5118" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="ACA165260F735282AEE6975E50A147C6" type="Zenodo-Dep">276621</mods:identifier>
</mods:mods>
<treatment id="03FF87EFFFDCB858FF7235E6E16EF83E" ID-DOI="http://doi.org/10.5281/zenodo.6182054" ID-GBIF-Taxon="119386100" ID-Zenodo-Dep="6182054" LSID="urn:lsid:plazi:treatment:03FF87EFFFDCB858FF7235E6E16EF83E" httpUri="http://treatment.plazi.org/id/03FF87EFFFDCB858FF7235E6E16EF83E" lastPageNumber="14" pageId="13" pageNumber="14">
<subSubSection id="C34C6572FFDCB858FF7235E6E14CFC0E" pageId="13" pageNumber="14" type="nomenclature">
<paragraph id="8BE936F9FFDCB858FF7235E6E2C9FC52" blockId="13.[151,674,933,993]" box="[151,674,933,958]" pageId="13" pageNumber="14">
<heading id="D0A18195FFDCB858FF7235E6E2C9FC52" bold="true" box="[151,674,933,958]" fontSize="10" level="2" pageId="13" pageNumber="14" reason="2">
<emphasis id="B922EAEBFFDCB858FF7235E6E2C9FC52" bold="true" box="[151,674,933,958]" pageId="13" pageNumber="14">
<taxonomicName id="4C564D7AFFDCB858FF7235E6E22AFC51" ID-CoL="48CVW" authority="Vinod &amp; Sabu" authorityName="Vinod &amp; Sabu" box="[151,577,933,958]" class="Insecta" family="Scarabaeidae" genus="Ochicanthon" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="murthyi" status="sp. nov.">
<emphasis id="B922EAEBFFDCB858FF7235E6E1FBFC51" bold="true" box="[151,400,933,958]" italics="true" pageId="13" pageNumber="14">Ochicanthon murthyi</emphasis>
Vinod &amp; Sabu
</taxonomicName>
<taxonomicNameLabel id="A2115790FFDCB858FDAD35E6E2C9FC52" box="[584,674,933,957]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE936F9FFDCB858FF72358AE14CFC0E" blockId="13.[151,674,933,993]" box="[151,295,969,993]" pageId="13" pageNumber="14">
(
<figureCitation id="136D2A7CFFDCB858FF7A358AE174FC0E" box="[159,287,969,993]" captionStart="FIGURES 36 – 40" captionStartId="16.[151,264,1979,2002]" captionTargetBox="[190,1409,203,1946]" captionTargetId="figure@16.[190,1411,193,1972]" captionTargetPageId="16" captionText="FIGURES 36 – 40. Ochicanthon murthyi (holotype). Dorsal habitus (36); base of pronotum and elytra (37); mesosternum and metasternum (38); aedeagus, ventral view (39); aedeagus, lateral left view (40). Scale bars = 1 mm." httpUri="https://zenodo.org/record/276631/files/figure.png" pageId="13" pageNumber="14">Figs 36–40</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34C6572FFDCB858FF723252E53DFA12" pageId="13" pageNumber="14" type="description">
<paragraph id="8BE936F9FFDCB858FF723252E34DFA5A" blockId="13.[151,1437,1041,2001]" pageId="13" pageNumber="14">
<emphasis id="B922EAEBFFDCB858FF723252E142FBC5" bold="true" box="[151,297,1041,1066]" pageId="13" pageNumber="14">Description.</emphasis>
Body (
<figureCitation id="136D2A7CFFDCB858FE9F3252E1A0FBC6" box="[378,459,1041,1065]" captionStart="FIGURES 36 – 40" captionStartId="16.[151,264,1979,2002]" captionTargetBox="[190,1409,203,1946]" captionTargetId="figure@16.[190,1411,193,1972]" captionTargetPageId="16" captionText="FIGURES 36 – 40. Ochicanthon murthyi (holotype). Dorsal habitus (36); base of pronotum and elytra (37); mesosternum and metasternum (38); aedeagus, ventral view (39); aedeagus, lateral left view (40). Scale bars = 1 mm." httpUri="https://zenodo.org/record/276631/files/figure.png" pageId="13" pageNumber="14">Fig. 36</figureCitation>
) predominantly black with a faint orange-yellow humeral spot on each elytron. Punctation on dorsal surface of head annular with intervals between punctures not exceeding their diameter. Pronotum in lateral view weakly convex, outline at base almost in line with that of elytral suture; laterobasal paramarginal ridge well marked, extended on slightly more than one third of pronotal length. Pronotal punctation (
<figureCitation id="136D2A7CFFDCB858FB2D323EE570FB7A" box="[1224,1307,1149,1173]" captionStart="FIGURES 36 – 40" captionStartId="16.[151,264,1979,2002]" captionTargetBox="[190,1409,203,1946]" captionTargetId="figure@16.[190,1411,193,1972]" captionTargetPageId="16" captionText="FIGURES 36 – 40. Ochicanthon murthyi (holotype). Dorsal habitus (36); base of pronotum and elytra (37); mesosternum and metasternum (38); aedeagus, ventral view (39); aedeagus, lateral left view (40). Scale bars = 1 mm." httpUri="https://zenodo.org/record/276631/files/figure.png" pageId="13" pageNumber="14">Fig. 37</figureCitation>
) consisting of annular setiferous punctures separated by interval of less than their diameter, becoming somewhat sparser anteromedially; setae almost indistinct, straight and recumbent. Elytra in lateral view moderately convex; suture not raised; striae shallow, wide, consisting of chains of fairly confluent discoidal depressions (
<figureCitation id="136D2A7CFFDCB858FB6732AAE4BEFAEE" box="[1154,1237,1257,1281]" captionStart="FIGURES 36 – 40" captionStartId="16.[151,264,1979,2002]" captionTargetBox="[190,1409,203,1946]" captionTargetId="figure@16.[190,1411,193,1972]" captionTargetPageId="16" captionText="FIGURES 36 – 40. Ochicanthon murthyi (holotype). Dorsal habitus (36); base of pronotum and elytra (37); mesosternum and metasternum (38); aedeagus, ventral view (39); aedeagus, lateral left view (40). Scale bars = 1 mm." httpUri="https://zenodo.org/record/276631/files/figure.png" pageId="13" pageNumber="14">Fig. 37</figureCitation>
); interstriae fairly flat, uneven, and irregularly punctate; background microsculpture smooth; setae fine, straight and recumbent; lateral ridge posteriorly extended to level of apex of third stria. Mesosternum without punctures; meso-metasternal line arcuate; metasternal shield with close annular punctures separated by intervals fairly equal to their diameter (
<figureCitation id="136D2A7CFFDCB858FF7A333AE098FA7E" box="[159,243,1401,1425]" captionStart="FIGURES 36 – 40" captionStartId="16.[151,264,1979,2002]" captionTargetBox="[190,1409,203,1946]" captionTargetId="figure@16.[190,1411,193,1972]" captionTargetPageId="16" captionText="FIGURES 36 – 40. Ochicanthon murthyi (holotype). Dorsal habitus (36); base of pronotum and elytra (37); mesosternum and metasternum (38); aedeagus, ventral view (39); aedeagus, lateral left view (40). Scale bars = 1 mm." httpUri="https://zenodo.org/record/276631/files/figure.png" pageId="13" pageNumber="14">Fig. 38</figureCitation>
). Pygidium densely covered with well-marked annular setiferous punctures becoming finer ventrally. Protrochanter with outer margin angulate
<collectingCountry id="F3417669FFDCB858FDA133DEE217FA5A" box="[580,636,1437,1461]" name="India" pageId="13" pageNumber="14">Hind</collectingCountry>
wings present.
</paragraph>
<paragraph id="8BE936F9FFDCB858FF203382E451FA36" blockId="13.[151,1437,1041,2001]" box="[197,1082,1473,1497]" pageId="13" pageNumber="14">
Male lacking obvious secondary sexual characters. Aedeagus as in
<figureCitation id="136D2A7CFFDCB858FC533382E45DFA36" box="[950,1078,1473,1497]" captionStart="FIGURES 36 – 40" captionStartId="16.[151,264,1979,2002]" captionTargetBox="[190,1409,203,1946]" captionTargetId="figure@16.[190,1411,193,1972]" captionTargetPageId="16" captionText="FIGURES 36 – 40. Ochicanthon murthyi (holotype). Dorsal habitus (36); base of pronotum and elytra (37); mesosternum and metasternum (38); aedeagus, ventral view (39); aedeagus, lateral left view (40). Scale bars = 1 mm." httpUri="https://zenodo.org/record/276631/files/figure.png" pageId="13" pageNumber="14">Figs 39–40</figureCitation>
.
</paragraph>
<paragraph id="8BE936F9FFDCB858FF2033A6E53DFA12" blockId="13.[151,1437,1041,2001]" box="[197,1366,1509,1533]" pageId="13" pageNumber="14">
<emphasis id="B922EAEBFFDCB858FF2033A6E11FFA12" bold="true" box="[197,372,1509,1533]" pageId="13" pageNumber="14">Measurements</emphasis>
(mm; n = 12): TL = 4.5–5.2; BW = 3.0–3.4; PL = 1.2–1.7; PW = 2.8–3.0; EL = 2.6–2.8.
</paragraph>
</subSubSection>
<subSubSection id="C34C6572FFDCB858FF203049E276F95E" pageId="13" pageNumber="14" type="materials_examined">
<paragraph id="8BE936F9FFDCB858FF203049E31DF9AA" blockId="13.[151,1437,1041,2001]" pageId="13" pageNumber="14">
<emphasis id="B922EAEBFFDCB858FF203049E11AF9CD" bold="true" box="[197,369,1545,1570]" pageId="13" pageNumber="14">
<typeStatus id="54ED885BFFDCB858FF203049E095F9CD" box="[197,254,1546,1570]" pageId="13" pageNumber="14">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54ED885BFFDCB858FE9E304AE188F9CE" box="[379,483,1545,1569]" pageId="13" pageNumber="14" type="holotype">Holotype</typeStatus>
(male, in
<collectionCode id="ED47AE3CFFDCB858FDB2304AE2F8F9CE" box="[599,659,1545,1569]" httpUri="http://grbio.org/cool/nwky-2b63" name="National Pusa Collection" pageId="13" pageNumber="14">NPC</collectionCode>
): “
<collectingCountry id="F3417669FFDCB858FD54304AE284F9CE" box="[689,751,1545,1569]" name="India" pageId="13" pageNumber="14">India</collectingCountry>
, Kerala, Wayanad District, Thirunelly,
<quantity id="4CAE9B1CFFDCB858FB4E304AE49EF9CE" box="[1195,1269,1545,1569]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="13" pageNumber="14" unit="m" value="900.0">900 m</quantity>
, moist deciduous forest, dung baited pit fall trap,
<date id="FFE81039FFDCB858FDC2306EE2C2F9AA" box="[551,681,1581,1605]" pageId="13" pageNumber="14" value="2005-09-23">23.IX.2005</date>
, leg. Vinod, K.V.”
</paragraph>
<paragraph id="8BE936F9FFDCB858FF203011E276F95E" blockId="13.[151,1437,1041,2001]" pageId="13" pageNumber="14">
<typeStatus id="54ED885BFFDCB858FF203011E159F985" box="[197,306,1618,1642]" pageId="13" pageNumber="14" type="paratype">Paratypes</typeStatus>
(11): Same data as
<typeStatus id="54ED885BFFDCB858FDE83012E219F986" box="[525,626,1617,1641]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
,
<specimenCount id="9D50FD70FFDCB858FD993012E2B9F986" box="[636,722,1617,1641]" pageId="13" pageNumber="14" type="male">2 males</specimenCount>
and
<specimenCount id="9D50FD70FFDCB858FCED3012E31FF986" box="[776,884,1617,1641]" pageId="13" pageNumber="14" type="female">6 females</specimenCount>
in
<collectionCode id="ED47AE3CFFDCB858FC723012E3A6F986" box="[919,973,1617,1641]" httpUri="http://grbio.org/cool/nwky-2b63" name="National Pusa Collection" pageId="13" pageNumber="14">NPC</collectionCode>
and
<collectionCode id="ED47AE3CFFDCB858FBE63012E458F986" box="[1027,1075,1617,1641]" pageId="13" pageNumber="14">SJC</collectionCode>
; “
<collectingCountry id="F3417669FFDCB858FBAC3012E4ECF986" box="[1097,1159,1617,1641]" name="India" pageId="13" pageNumber="14">India</collectingCountry>
, Kerala,
<collectingCountry id="F3417669FFDCB858FB0F3012E50EF986" box="[1258,1381,1617,1641]" name="India" pageId="13" pageNumber="14">Kozhikode</collectingCountry>
District, Devagiri,
<quantity id="4CAE9B1CFFDCB858FEAD3036E1EEF962" box="[328,389,1653,1677]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="13" pageNumber="14" unit="m" value="50.0">50 m</quantity>
, shrub jungle, dung baited pit fall trap,
<date id="FFE81039FFDCB858FCB33036E389F962" box="[854,994,1653,1677]" pageId="13" pageNumber="14" value="2008-10-14">14. X. 2008</date>
, leg. Jizla, V.P.”
<specimenCount id="9D50FD70FFDCB858FB4F3036E572F962" box="[1194,1305,1653,1677]" pageId="13" pageNumber="14" type="female">2 females</specimenCount>
in ZSI-Ca; “
<collectingCountry id="F3417669FFDCB858FF4430DAE0B4F95E" box="[161,223,1689,1713]" name="India" pageId="13" pageNumber="14">India</collectingCountry>
, Kerala”
<specimenCount id="9D50FD70FFDCB858FEAC30DAE1F4F95E" box="[329,415,1689,1713]" pageId="13" pageNumber="14" type="male">2 males</specimenCount>
in
<collectionCode id="ED47AE3CFFDCB858FE2630DAE272F95E" box="[451,537,1689,1713]" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="13" pageNumber="14">MHNG</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="C34C6572FFDCB858FF2030FEE17DF8F2" pageId="13" pageNumber="14" type="distribution">
<paragraph id="8BE936F9FFDCB858FF2030FEE17DF8F2" blockId="13.[151,1437,1041,2001]" pageId="13" pageNumber="14">
<emphasis id="B922EAEBFFDCB858FF2030FEE22DF939" bold="true" box="[197,582,1725,1750]" pageId="13" pageNumber="14">Distribution and natural history</emphasis>
. South
<collectingCountry id="F3417669FFDCB858FD7B30FEE2B3F93A" box="[670,728,1725,1749]" name="India" pageId="13" pageNumber="14">India</collectingCountry>
(upper montane cloud forests at Thirunelly in the South Western Ghats montane rain forest ecoregion; Calicut-Devagiri college campus in Malabar Coast moist deciduous forest ecoregion).
</paragraph>
</subSubSection>
<subSubSection id="C34C6572FFDCB858FF203169E52AF8AE" box="[197,1345,1833,1858]" pageId="13" pageNumber="14" type="etymology">
<paragraph id="8BE936F9FFDCB858FF203169E52AF8AE" blockId="13.[151,1437,1041,2001]" box="[197,1345,1833,1858]" pageId="13" pageNumber="14">
<emphasis id="B922EAEBFFDCB858FF203169E12FF8AD" bold="true" box="[197,324,1834,1858]" pageId="13" pageNumber="14">Etymology</emphasis>
. This species is named in honor of late Prof. V.A. Murthy, Loyola College, Chennai,
<collectingCountry id="F3417669FFDCB858FAE5316AE556F8AE" box="[1280,1341,1833,1857]" name="India" pageId="13" pageNumber="14">India</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C34C6572FFDCB858FF20310EE16EF83E" pageId="13" pageNumber="14" type="diagnosis">
<paragraph id="8BE936F9FFDCB858FF20310EE16EF83E" blockId="13.[151,1437,1041,2001]" pageId="13" pageNumber="14">
<emphasis id="B922EAEBFFDCB858FF20310EE154F889" bold="true" box="[197,319,1869,1894]" pageId="13" pageNumber="14">Diagnosis.</emphasis>
<taxonomicName id="4C564D7AFFDCB858FEA3310EE25FF88A" box="[326,564,1869,1893]" class="Insecta" family="Scarabaeidae" genus="Ochicanthon" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="murthyi">
<emphasis id="B922EAEBFFDCB858FEA3310EE25FF88A" box="[326,564,1869,1893]" italics="true" pageId="13" pageNumber="14">Ochicanthon murthyi</emphasis>
</taxonomicName>
shares unicolored elytra bearing striae consisting of chains of fairly confluent discoidal depressions only with
<taxonomicName id="4C564D7AFFDCB858FDE13131E2FBF866" box="[516,656,1905,1929]" class="Insecta" family="Scarabaeidae" genus="Ochicanthon" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="tristoides">
<emphasis id="B922EAEBFFDCB858FDE13131E2FBF866" box="[516,656,1905,1929]" italics="true" pageId="13" pageNumber="14">O. tristoides</emphasis>
</taxonomicName>
, from northeast
<collectingCountry id="F3417669FFDCB858FCAB3132E3E0F866" box="[846,907,1905,1929]" name="India" pageId="13" pageNumber="14">India</collectingCountry>
. This species is easily distinguishable from
<taxonomicName id="4C564D7AFFDCB858FA643131E094F842" class="Insecta" family="Scarabaeidae" genus="Ochicanthon" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="tristoides">
<emphasis id="B922EAEBFFDCB858FA643131E094F842" italics="true" pageId="13" pageNumber="14">O. tristoides</emphasis>
</taxonomicName>
by the presence of angulate outer margin of the protrochanter and the variation in the morphology of aedeagus.
</paragraph>
</subSubSection>
</treatment>
</document>
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