162 lines
21 KiB
XML
162 lines
21 KiB
XML
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<mods:title id="C481ABD4FBD325891B645E00CE5E912E">Review of the central African leaf chafer genus Entypophana Moser, 1913 (Coleoptera: Scarabaeidae: Melolonthinae: Schizonychini)</mods:title>
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<mods:namePart id="4D0A896CAF06331C69D7A10EE52407AD">Sehnal, Richard</mods:namePart>
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<mods:date id="D2389E34326EDBAC70198B4851D7B0D0">2017</mods:date>
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<mods:number id="91E90EC4EA478229DC0A32F3847FDF57">4258</mods:number>
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<treatment id="03FF87E2E115FFA218F7FD28FB67F972" ID-DOI="http://doi.org/10.5281/zenodo.6034944" ID-GBIF-Taxon="128810295" ID-Zenodo-Dep="6034944" LSID="urn:lsid:plazi:treatment:03FF87E2E115FFA218F7FD28FB67F972" httpUri="http://treatment.plazi.org/id/03FF87E2E115FFA218F7FD28FB67F972" lastPageNumber="158" pageId="1" pageNumber="158">
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<subSubSection id="C34C657FE115FFA218F7FD28FE3DFD78" box="[151,472,727,754]" pageId="1" pageNumber="158" type="nomenclature">
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<paragraph id="8BE936F4E115FFA218F7FD28FE3DFD78" blockId="1.[151,472,727,754]" box="[151,472,727,754]" pageId="1" pageNumber="158">
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<heading id="D0A18198E115FFA218F7FD28FE3DFD78" bold="true" box="[151,472,727,754]" fontSize="11" level="1" pageId="1" pageNumber="158" reason="1">
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<taxonomicName id="4C564D77E115FFA218F7FD28FE3DFD78" authority="Moser, 1913" authorityName="Moser" authorityYear="1913" box="[151,472,727,754]" class="Insecta" family="Melolonthidae" genus="Entypophana" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="158" phylum="Arthropoda" rank="genus">
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<emphasis id="B922EAE6E115FFA218F7FD28FE3DFD78" bold="true" box="[151,472,727,754]" pageId="1" pageNumber="158">
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<emphasis id="B922EAE6E115FFA218F7FD28FEDFFD7B" bold="true" box="[151,314,727,753]" italics="true" pageId="1" pageNumber="158">Entypophana</emphasis>
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<bibRefCitation id="EFC74B05E115FFA21921FD27FE3DFD78" author="Moser" box="[321,472,728,754]" pageId="1" pageNumber="158" refString="Moser, J. (1913) Neue afrikanische Schizonychinen. Annales de la Societe Entomologique de Belgique, 57, 283 - 296." type="journal article" year="1913">Moser, 1913</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34C657FE115FFA218F7FCE0FE7AFCD9" pageId="1" pageNumber="158" type="reference_group">
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<paragraph id="8BE936F4E115FFA218F7FCE0FE7AFCD9" blockId="1.[151,1436,798,851]" pageId="1" pageNumber="158">
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<treatmentCitationGroup id="AB4611DAE115FFA218F7FCE0FE7AFCD9" pageId="1" pageNumber="158">
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<taxonomicName id="4C564D77E115FFA218F7FCE0FE0FFCBE" authority="Moser, 1913: 294" authorityName="Moser" authorityPageNumber="294" authorityYear="1913" box="[151,490,798,820]" class="Insecta" family="Melolonthidae" genus="Entypophana" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="158" phylum="Arthropoda" rank="genus">
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<emphasis id="B922EAE6E115FFA218F7FCE0FEFBFCBE" box="[151,286,799,820]" italics="true" pageId="1" pageNumber="158">Entypophana</emphasis>
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<treatmentCitation id="0AF710E5E115FFA2194BFCE0FE0FFCBE" author="Moser" box="[299,490,798,820]" page="294" pageId="1" pageNumber="158" year="1913">
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<bibRefCitation id="EFC74B05E115FFA2194BFCE0FE56FCBE" author="Moser" box="[299,435,798,820]" pageId="1" pageNumber="158" refString="Moser, J. (1913) Neue afrikanische Schizonychinen. Annales de la Societe Entomologique de Belgique, 57, 283 - 296." type="journal article" year="1913">Moser, 1913</bibRefCitation>
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: 294
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</treatmentCitation>
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</taxonomicName>
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(description);
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<treatmentCitation id="0AF710E5E115FFA21AECFCE0FCB9FCBE" author="Burgeon" box="[652,860,798,820]" page="262" pageId="1" pageNumber="158" year="1946">
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<bibRefCitation id="EFC74B05E115FFA21AECFCE0FCC0FCBE" author="Burgeon" box="[652,805,798,820]" pageId="1" pageNumber="158" refString="Burgeon, L. (1946) Melolonthini et Pachydemini du Congo belge. Revue de Zoologie et de Botanique Africains, 39, 230 - 273." type="journal article" year="1946">Burgeon 1946</bibRefCitation>
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: 262
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||
</treatmentCitation>
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(review);
|
||
<treatmentCitation id="0AF710E5E115FFA21BB2FCE1FB7FFCBE" author="Lacroix" box="[978,1178,798,820]" page="208" pageId="1" pageNumber="158" year="2010">
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<bibRefCitation id="EFC74B05E115FFA21BB2FCE1FB80FCBE" author="Lacroix" box="[978,1125,798,820]" pageId="1" pageNumber="158" refString="Lacroix, M. (2010) Melolonthinae afrotropicaux (Scarabaeoidea, Melolonthidae). Genera et catalogue commente. Editions Marc Lacroix, Paris, 277 pp., 192 pls." type="book" year="2010">Lacroix 2010</bibRefCitation>
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: 208
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||
</treatmentCitation>
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(catalogue);
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<treatmentCitation id="0AF710E5E115FFA21D4DFCE1FE7EFCD9" author="Lacroix" page="723" pageId="1" pageNumber="158" year="2012">
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<bibRefCitation id="EFC74B05E115FFA21D4DFCE1FE8CFCD9" author="Lacroix" pageId="1" pageNumber="158" refString="Lacroix, M. & Montreuil, O. (2012) Revision du genre Entyposis Kolbe, 1894 (Insecta, Coleoptera, Scarabaeoidea, Melolonthidae). Zoosystema, 34, 721 - 735." type="journal article" year="2012">Lacroix & Montreuil 2012</bibRefCitation>
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: 723
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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<subSubSection id="C34C657FE115FFA218F7FC80FC2BFC12" box="[151,974,895,920]" pageId="1" pageNumber="158" type="materials_examined">
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<paragraph id="8BE936F4E115FFA218F7FC80FC2BFC12" blockId="1.[151,1437,895,1785]" box="[151,974,895,920]" pageId="1" pageNumber="158">
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<emphasis id="B922EAE6E115FFA218F7FC80FED7FC12" bold="true" box="[151,306,895,920]" pageId="1" pageNumber="158">Type species.</emphasis>
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<taxonomicName id="4C564D77E115FFA21959FC7FFD5AFC12" authority="Moser, 1913" authorityName="Moser" authorityYear="1913" box="[313,703,896,920]" class="Insecta" family="Melolonthidae" genus="Entypophana" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="158" phylum="Arthropoda" rank="species" species="apicata">
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<emphasis id="B922EAE6E115FFA21959FC7FFDCCFC1D" box="[313,553,896,920]" italics="true" pageId="1" pageNumber="158">Entypophana apicata</emphasis>
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<bibRefCitation id="EFC74B05E115FFA21A50FC7FFD5AFC12" author="Moser" box="[560,703,896,920]" pageId="1" pageNumber="158" refString="Moser, J. (1913) Neue afrikanische Schizonychinen. Annales de la Societe Entomologique de Belgique, 57, 283 - 296." type="journal article" year="1913">Moser, 1913</bibRefCitation>
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</taxonomicName>
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, by present designation.
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</paragraph>
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<subSubSection id="C34C657FE115FFA218A7FC5BFCE0F9CF" pageId="1" pageNumber="158" type="diagnosis">
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<paragraph id="8BE936F4E115FFA218A7FC5BFA78FA52" blockId="1.[151,1437,895,1785]" pageId="1" pageNumber="158">
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<emphasis id="B922EAE6E115FFA218A7FC5BFEA4FC37" bold="true" box="[199,321,932,957]" pageId="1" pageNumber="158">Diagnosis.</emphasis>
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<taxonomicName id="4C564D77E115FFA21928FC5BFE38FC36" box="[328,477,932,956]" class="Insecta" family="Melolonthidae" genus="Entypophana" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="158" phylum="Arthropoda" rank="genus">
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<emphasis id="B922EAE6E115FFA21928FC5BFE38FC36" box="[328,477,932,956]" italics="true" pageId="1" pageNumber="158">Entypophana</emphasis>
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</taxonomicName>
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||
differs from other schizonychine genera by combination of the following characters: Body elongate (length 14.9–20.0 mm), almost parallel-sided, weakly to moderately convex. Labrum transverse, deeply bilobed, lobes rounded. Clypeus transverse, anterior angles rounded. Frontoclypeal carina elevated, medially sinuate forward. Frontoclypeal suture straight or gently undulate. Occipital carina always present, prominent, medially elevated, laterally diminishing. Head including clypeus with species-diagnostic macrosetose punctation. Eyes small (combined width of both eyes is less than maximum width of frons between eyes) to large (combined width of both eyes exceeds maximum width of frons between eyes). Antenna with 10 antennomeres, club with three antennomeres. Terminal maxillary palpomere elongate, club-shaped. Pronotum transverse, widest behind midlength, with anteromedial depression. Front margin straight with angles extended forward. Lateral margins crenulate and macrosetose. Base bordered, medially gently arched toward elytra, with smooth margin paralleled by a row of coarse punctures and hind angles broadly rounded. Crest delimiting anteromedial depression prominent, with top and frontal edges impunctate and remaining parts with species-diagnostic macrosetose punctation. Elytra weakly to moderately convex, parallel-sided, rounded apically, apical angle approximately rectangular. Striae absent. Macropterous. Protibia narrow, tridentate, terminal spine inserted against medial tooth. Claws bifid, with ventrobasal teeth (
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<emphasis id="B922EAE6E115FFA21A53FA62FDBEFA3E" box="[563,603,1437,1460]" italics="true" pageId="1" pageNumber="158">e.g.</emphasis>
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,
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<figureCitation id="136D2A71E115FFA21A0BFA63FD0AFA3E" box="[619,751,1436,1461]" captionStart="FIGURES 9" captionStartId="13.[151,264,925,948]" captionTargetBox="[159,1429,206,892]" captionTargetId="figure@13.[151,1436,195,904]" captionTargetPageId="13" captionText="FIGURES 9 A – F. Inner mesotarsal claw. A, Entypophana apicata Moser, 1913, holotype; B, Entypophana biapicata Moser, 1913, lectotype; C, Entypophana hulstaerti Burgeon, 1946, lectotype; D, Entypophana lujai Moser, 1917, lectotype; E, Entypophana maynei Burgeon, 1946, lectotype; F, Entypophana njiapanda new species, holotype. Scale bar: 1.0 mm." httpUri="https://zenodo.org/record/569451/files/figure.png" pageId="1" pageNumber="158">Figs. 9A–F</figureCitation>
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). Parameres symmetrical, narrow or curved in lateral aspect, blunt or extended apically in dorsal or lateral aspect; apex with tuft of long, yellow macrosetae (
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<figureCitation id="136D2A71E115FFA21CA2FA40FAA7FA52" box="[1218,1346,1471,1496]" captionStart="FIGURES 7" captionStartId="11.[151,264,1701,1724]" captionTargetBox="[185,1411,848,1668]" captionTargetId="figure@11.[177,1414,841,1674]" captionTargetPageId="11" captionText="FIGURES 7 A – F. Aedeagus in dorsal view. A, Entypophana apicata Moser, 1913, holotype; B, Entypophana biapicata Moser, 1913, lectotype; C, Entypophana hulstaerti Burgeon, 1946, lectotype; D, Entypophana lujai Moser, 1917, lectotype; E, Entypophana maynei Burgeon, 1946, lectotype; F, Entypophana njiapanda new species, holotype. Scale bar: 1.0 mm." httpUri="https://zenodo.org/record/569449/files/figure.png" pageId="1" pageNumber="158">Figs. 7A–F</figureCitation>
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,
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<figureCitation id="136D2A71E115FFA21D2CFA3FFA75FA52" box="[1356,1424,1472,1496]" captionStart="FIGURES 8" captionStartId="12.[151,264,1326,1349]" captionTargetBox="[170,1427,419,1302]" captionTargetId="figure@12.[151,1437,411,1305]" captionTargetPageId="12" captionText="FIGURES 8 A – F. Aedeagus in lateral view. A, Entypophana apicata Moser, 1913, holotype; B, Entypophana biapicata Moser, 1913, lectotype; C, Entypophana hulstaerti Burgeon, 1946, lectotype; D, Entypophana lujai Moser, 1917, lectotype; E, Entypophana maynei Burgeon, 1946, lectotype; F, Entypophana njiapanda new species, holotype. Scale bar: 1.0 mm." httpUri="https://zenodo.org/record/569450/files/figure.png" pageId="1" pageNumber="158">8A–F</figureCitation>
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).
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</paragraph>
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<paragraph id="8BE936F4E115FFA218A7FA1BFCE0F9CF" blockId="1.[151,1437,895,1785]" pageId="1" pageNumber="158">
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<emphasis id="B922EAE6E115FFA218A7FA1BFE4AFA77" bold="true" box="[199,431,1508,1533]" pageId="1" pageNumber="158">Sexual dimorphism</emphasis>
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. Female differs from male in the following characters: body slightly broadened posteriad; antennal club straight, considerably shorter; eyes small (width of both eyes combined shorter than maximum width of frons between eyes); anteromedial depression flatter.
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C34C657FE115FFA218A7F9B0FF52F906" pageId="1" pageNumber="158" type="description">
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<paragraph id="8BE936F4E115FFA218A7F9B0FF52F906" blockId="1.[151,1437,895,1785]" pageId="1" pageNumber="158">
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<materialsCitation id="3B3E3CA9E115FFA218A7F9B0FF52F906" ID-GBIF-Occurrence="1499294090" country="Democratic Republic of the Congo, Tanzania" location="Central Africa" pageId="1" pageNumber="158" specimenCount="1">
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||
<emphasis id="B922EAE6E115FFA218A7F9B0FE0AF9E2" bold="true" box="[199,495,1615,1640]" pageId="1" pageNumber="158">Geographic distribution.</emphasis>
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||
<location id="8E89602FE115FFA21995F9B0FD73F9E2" LSID="urn:lsid:plazi:treatment:03FF87E2E115FFA218F7FD28FB67F972:8E89602FE115FFA21995F9B0FD73F9E2" box="[501,662,1615,1640]" country="Democratic Republic of the Congo, Tanzania" name="Central Africa" pageId="1" pageNumber="158">Central Africa</location>
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||
(
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||
<collectingCountry id="F3417664E115FFA21AC5F9B0FB42F9E2" box="[677,1191,1615,1640]" isEnumeration="true" pageId="1" pageNumber="158">Democratic Republic of the Congo, Tanzania</collectingCountry>
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||
,
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||
<collectingCountry id="F3417664E115FFA21CD2F9B0FAEEF9E2" box="[1202,1291,1615,1640]" name="Uganda" pageId="1" pageNumber="158">Uganda</collectingCountry>
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||
) (
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||
<figureCitation id="136D2A71E115FFA21D42F9B0FF4FF906" captionStart="FIGURES 10" captionStartId="13.[151,264,1927,1950]" captionTargetBox="[151,1435,1059,1906]" captionTargetId="figure@13.[151,1435,1059,1906]" captionTargetPageId="13" captionText="FIGURES 10 A – F. Map with currently known geographical distribution. A, Entypophana apicata Moser, 1913; B, Entypophana biapicata Moser, 1913; C, Entypophana hulstaerti Burgeon, 1946; D, Entypophana lujai Moser, 1917; E, Entypophana maynei Burgeon, 1946; F, Entypophana njiapanda new species." httpUri="https://zenodo.org/record/569452/files/figure.png" pageId="1" pageNumber="158">Fig. 10 A– F</figureCitation>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34C657FE115FFA218A7F968FB67F972" pageId="1" pageNumber="158" type="discussion">
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<paragraph id="8BE936F4E115FFA218A7F968FB67F972" blockId="1.[151,1437,895,1785]" pageId="1" pageNumber="158">
|
||
<emphasis id="B922EAE6E115FFA218A7F968FED5F93A" bold="true" box="[199,304,1687,1712]" pageId="1" pageNumber="158">Remark.</emphasis>
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||
<bibRefCitation id="EFC74B05E115FFA21958F966FE37F93A" author="Moser" box="[312,466,1688,1712]" pageId="1" pageNumber="158" refString="Moser, J. (1913) Neue afrikanische Schizonychinen. Annales de la Societe Entomologique de Belgique, 57, 283 - 296." type="journal article" year="1913">Moser (1913)</bibRefCitation>
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||
established his new genus
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<taxonomicName id="4C564D77E115FFA21B68F967FC78F93A" box="[776,925,1688,1712]" class="Insecta" family="Melolonthidae" genus="Entypophana" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="158" phylum="Arthropoda" rank="genus">
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<emphasis id="B922EAE6E115FFA21B68F967FC78F93A" box="[776,925,1688,1712]" italics="true" pageId="1" pageNumber="158">Entypophana</emphasis>
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||
</taxonomicName>
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||
for two species:
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<taxonomicName id="4C564D77E115FFA21C01F966FA8EF93A" authority="Moser, 1913" authorityName="Moser" authorityYear="1913" box="[1121,1387,1688,1712]" class="Insecta" family="Melolonthidae" genus="Entypophana" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="158" phylum="Arthropoda" rank="species" species="apicata">
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<emphasis id="B922EAE6E115FFA21C01F966FB32F925" box="[1121,1239,1688,1712]" italics="true" pageId="1" pageNumber="158">E. apicata</emphasis>
|
||
<bibRefCitation id="EFC74B05E115FFA21CBEF966FA8EF93A" author="Moser" box="[1246,1387,1688,1712]" pageId="1" pageNumber="158" refString="Moser, J. (1913) Neue afrikanische Schizonychinen. Annales de la Societe Entomologique de Belgique, 57, 283 - 296." type="journal article" year="1913">Moser, 1913</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C564D77E115FFA218F7F942FE21F95E" authority="Moser, 1913" authorityName="Moser" authorityYear="1913" box="[151,452,1724,1748]" class="Insecta" family="Melolonthidae" genus="Entypophana" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="158" phylum="Arthropoda" rank="species" species="biapicata">
|
||
<emphasis id="B922EAE6E115FFA218F7F942FEC0F95E" box="[151,293,1724,1748]" italics="true" pageId="1" pageNumber="158">E. biapicata</emphasis>
|
||
<bibRefCitation id="EFC74B05E115FFA21951F942FE21F95E" author="Moser" box="[305,452,1724,1748]" pageId="1" pageNumber="158" refString="Moser, J. (1913) Neue afrikanische Schizonychinen. Annales de la Societe Entomologique de Belgique, 57, 283 - 296." type="journal article" year="1913">Moser, 1913</bibRefCitation>
|
||
</taxonomicName>
|
||
, but neither has been designated as the
|
||
<typeStatus id="54ED8856E115FFA21BC5F942FC30F95F" box="[933,981,1725,1749]" pageId="1" pageNumber="158">type</typeStatus>
|
||
species of the genus. To remedy this,
|
||
<taxonomicName id="4C564D77E115FFA218F7F91FFDFEF972" authority="Moser, 1913" authorityName="Moser" authorityYear="1913" box="[151,539,1760,1784]" class="Insecta" family="Melolonthidae" genus="Entypophana" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="158" phylum="Arthropoda" rank="species" species="apicata">
|
||
<emphasis id="B922EAE6E115FFA218F7F91FFE62F97D" box="[151,391,1760,1784]" italics="true" pageId="1" pageNumber="158">Entypophana apicata</emphasis>
|
||
<bibRefCitation id="EFC74B05E115FFA219EEF91EFDFEF972" author="Moser" box="[398,539,1760,1784]" pageId="1" pageNumber="158" refString="Moser, J. (1913) Neue afrikanische Schizonychinen. Annales de la Societe Entomologique de Belgique, 57, 283 - 296." type="journal article" year="1913">Moser, 1913</bibRefCitation>
|
||
</taxonomicName>
|
||
is here designated as the
|
||
<typeStatus id="54ED8856E115FFA21B59F91EFC8CF973" box="[825,873,1761,1785]" pageId="1" pageNumber="158">type</typeStatus>
|
||
species of
|
||
<taxonomicName id="4C564D77E115FFA21B86F91FFB9EF972" box="[998,1147,1760,1784]" class="Insecta" family="Melolonthidae" genus="Entypophana" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="158" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B922EAE6E115FFA21B86F91FFB9EF972" box="[998,1147,1760,1784]" italics="true" pageId="1" pageNumber="158">Entypophana</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |