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<mods:title id="8BE0CE820149F224332EF4875771B488">Revision of the Siobla annulicornis, acutiscutella and sheni groups (Hymenoptera: Tenthredinidae)</mods:title>
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<mods:namePart id="6E87242655E0707E9AFF489603BFE082">Niu, Gengyun</mods:namePart>
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<subSubSection id="C34C6535FF893145FF409062920AFD76" box="[151,352,629,655]" pageId="1" pageNumber="46" type="nomenclature">
<paragraph id="8BE936BEFF893145FF409062920AFD76" blockId="1.[151,352,629,655]" box="[151,352,629,655]" pageId="1" pageNumber="46">
<heading id="D0A181D2FF893145FF409062920AFD76" bold="true" box="[151,352,629,655]" fontSize="11" level="1" pageId="1" pageNumber="46" reason="1">
<taxonomicName id="4C564D3DFF893145FF409062920AFD76" authority="Cameron" authorityName="Cameron" box="[151,352,629,655]" class="Insecta" family="Tenthredinidae" genus="Siobla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="genus">
<emphasis id="B922EAACFF893145FF409062920AFD76" bold="true" box="[151,352,629,655]" pageId="1" pageNumber="46">
<emphasis id="B922EAACFF893145FF409062938EFD76" bold="true" box="[151,228,629,655]" italics="true" pageId="1" pageNumber="46">Siobla</emphasis>
<collectingCountry id="F341762EFF893145FF3C9062920AFD76" box="[235,352,629,655]" name="Cameroon" pageId="1" pageNumber="46">Cameron</collectingCountry>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34C6535FF893145FF4090AA9223FCAD" pageId="1" pageNumber="46" type="reference_group">
<paragraph id="8BE936BEFF893145FF4090AA926EFD0D" blockId="1.[151,1436,701,852]" pageId="1" pageNumber="46">
<treatmentCitationGroup id="AB461190FF893145FF4090AA926EFD0D" pageId="1" pageNumber="46">
<taxonomicName id="4C564D3DFF893145FF4090AA92D8FD2D" authority="Cameron, 1877: 88" authorityName="Cameron" authorityPageNumber="88" authorityYear="1877" box="[151,434,701,724]" class="Insecta" family="Tenthredinidae" genus="Siobla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="genus">
<emphasis id="B922EAACFF893145FF4090AA93B1FD2D" box="[151,219,701,724]" italics="true" pageId="1" pageNumber="46">Siobla</emphasis>
<treatmentCitation id="0AF710AFFF893145FF3590AA92D8FD2D" author="Cameron" box="[226,434,701,724]" page="88" pageId="1" pageNumber="46" year="1877">
<bibRefCitation id="EFC74B4FFF893145FF3590AA92E1FD2D" author="Cameron" box="[226,395,701,724]" pageId="1" pageNumber="46" refString="Cameron, P. (1877) Descriptions of new genera and species of East Indian Tenthredinidae. (The) Transactions of the Entomological Society of London, 2, 88 - 90." type="journal article" year="1877">
<collectingCountry id="F341762EFF893145FF3590AA922DFD2D" box="[226,327,701,724]" name="Cameroon" pageId="1" pageNumber="46">Cameron</collectingCountry>
, 1877
</bibRefCitation>
: 88
</treatmentCitation>
</taxonomicName>
89.
<typeStatus id="54ED881CFF893145FE3090AA9176FD2D" box="[487,540,701,724]" pageId="1" pageNumber="46">Type</typeStatus>
species:
<taxonomicName id="4C564D3DFF893145FDA890AA908FFD2D" authority="Cameron, 1877" authorityName="Cameron" authorityYear="1877" box="[639,997,701,724]" class="Insecta" family="Tenthredinidae" genus="Siobla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="mooreana">
<emphasis id="B922EAACFF893145FDA890AA905FFD2D" box="[639,821,701,724]" italics="true" pageId="1" pageNumber="46">Siobla mooreana</emphasis>
<bibRefCitation id="EFC74B4FFF893145FCEA90AA908FFD2D" author="Cameron" box="[829,997,701,724]" pageId="1" pageNumber="46" refString="Cameron, P. (1877) Descriptions of new genera and species of East Indian Tenthredinidae. (The) Transactions of the Entomological Society of London, 2, 88 - 90." type="journal article" year="1877">
<collectingCountry id="F341762EFF893145FCEA90AA90C8FD2D" box="[829,930,701,724]" name="Cameroon" pageId="1" pageNumber="46">Cameron</collectingCountry>
, 1877
</bibRefCitation>
</taxonomicName>
, by subsequent designation of
<bibRefCitation id="EFC74B4FFF893145FAE690AA926AFD0D" author="Ashmead" pageId="1" pageNumber="46" refString="Ashmead, W. H. (1898) Classification of the horntails and sawflies, or the sub-order Phytophaga (Paper No. 7. - Conclusion). The Canadian Entomologist, 30, 309 - 310." type="journal article" year="1898">Ashmead, 1898</bibRefCitation>
.
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</paragraph>
<paragraph id="8BE936BEFF893145FF4090EA9223FCAD" blockId="1.[151,1436,701,852]" pageId="1" pageNumber="46">
<emphasis id="B922EAACFF893145FF4090EA925CFCED" box="[151,310,765,788]" italics="true" pageId="1" pageNumber="46">Encarsioneura</emphasis>
<bibRefCitation id="EFC74B4FFF893145FEEB90EA92A5FCED" author="Konow" box="[316,463,765,788]" pageId="1" pageNumber="46" refString="Konow, F. W. (1890) Tenthredinidae Europae. (In German and Latin). Deutsche Entomologische Zeitschrift, 1890 (2), 225 - 240." type="journal article" year="1890">Konow, 1890</bibRefCitation>
: 240.
<typeStatus id="54ED881CFF893145FDD890EA912EFCED" box="[527,580,765,788]" pageId="1" pageNumber="46">Type</typeStatus>
species:
<taxonomicName id="4C564D3DFF893145FD7290EA9089FCED" authority="Klug, 1817" authorityName="Klug" authorityYear="1817" box="[677,995,765,788]" class="Insecta" family="Psychidae" genus="Tenthredo" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="sturmii">
<emphasis id="B922EAACFF893145FD7290EA9008FCED" box="[677,866,765,788]" italics="true" pageId="1" pageNumber="46">Tenthredo sturmii</emphasis>
<bibRefCitation id="EFC74B4FFF893145FCBF90EA9089FCED" author="Klug" box="[872,995,765,788]" pageId="1" pageNumber="46" refString="Klug, F. (1817) Die Blattwespen nach ihren Gattungen und Arten zusammengestellt. (In German). Der Gesellschaft Naturforschender Freunde zu Berlin Magazin fur die neuesten Entdeckungen in der gesamten Naturkunde. 8 [1814] (2), 116." type="journal article" year="1817">Klug, 1817</bibRefCitation>
</taxonomicName>
, by monotypy.
<emphasis id="B922EAACFF893145FF40910A927FFCCD" box="[151,277,797,820]" italics="true" pageId="1" pageNumber="46">Megasiobla</emphasis>
Dovnar-Zapolskij, 1930: 86.
<typeStatus id="54ED881CFF893145FDA7910A91CFFCCD" box="[624,677,797,820]" pageId="1" pageNumber="46">Type</typeStatus>
species:
<emphasis id="B922EAACFF893145FCC1910A909FFCCD" box="[790,1013,797,820]" italics="true" pageId="1" pageNumber="46">
Megasiobla
<taxonomicName id="4C564D3DFF893145FC75910A909FFCCD" authority="Dovnar-Zapolskij, 1930" authorityName="Dovnar-Zapolskij" authorityYear="1930" box="[930,1013,797,820]" class="Insecta" family="Tenthredinidae" genus="Siobla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="zenaida">zenaida</taxonomicName>
</emphasis>
Dovnar-Zapolskij, 1930, by original designation.
</paragraph>
</subSubSection>
<subSubSection id="C34C6535FF893145FF40919590AEFBCB" pageId="1" pageNumber="46" type="diagnosis">
<paragraph id="8BE936BEFF893145FF40919590AEFBCB" blockId="1.[151,1437,898,2007]" pageId="1" pageNumber="46">
<emphasis id="B922EAACFF893145FF409195924CFC65" bold="true" box="[151,294,898,924]" pageId="1" pageNumber="46">Diagnosis:</emphasis>
Head and thorax densely punctured; anterior margin of clypeus subtruncate; mandibles asymmetrical, left mandible bidentate, right mandible simple, without inner tooth or basal lobe; metabasitarsomere compressed laterally; pedicel of antenna longer than wide; inner tooth of claw much longer than outer tooth; anal cell of fore wing with an oblique crossvein slightly beyond middle; lancet slender, distinctly sclerotized, usually without distinct annular suture.
</paragraph>
</subSubSection>
<subSubSection id="C34C6535FF893146FF12962A9260FAF8" lastPageId="2" lastPageNumber="47" pageId="1" pageNumber="46" type="description">
<paragraph id="8BE936BEFF893145FF12962A900FF82E" blockId="1.[151,1437,898,2007]" pageId="1" pageNumber="46">
<emphasis id="B922EAACFF893145FF12962A920EFBAE" bold="true" box="[197,356,1085,1111]" pageId="1" pageNumber="46">Description:</emphasis>
Body robust, head and thorax usually densely punctured; clypeus small, anterior margin subtruncate, sometimes feebly and roundly protruding, or with a small incision; labrum round at apex; mandibles short and stout, asymmetrical, left one with a distinct inner tooth at about middle, right one simple, without inner tooth or basal lobe; malar space about 0.51.5x diameter of ocellus in female, or narrower in male; eyes large or medium-sized, inner margins straight, strongly converging downwards; supra-antennal tubercles usually not elevated; middle fovea large, lateral fovea distinct, open anteriorly; frons weakly defined; ocellar triangle small; head behind eyes in dorsal view not elongated, lateral sides converging posteriorly; occipital carina sharp and complete; antenna about as long as or longer than abdomen but clearly shorter than thorax and abdomen together, pedicel longer than wide, third antennomere much longer than fourth antennomere; anterior collar of pronotum narrow with broadest part about two times diameter of ocellus; ventral margins of propleura narrowly and roundly meeting; anterior margin of mesoscutellum truncate, furrow before mesoscutellum very deep, transverse; metanotum with a deep and broad basin before metascutellum, distance between cenchri about two times width of a cenchrus; metapostnotum strongly constricted, linear on meson; epicnemium absent, mesepisternum without ventral thorn but with a sharp anterior marginal carina, mesepimeron broad, with a transverse ridge at middle, metepimeron narrow with a small dorsal lobe, posterior corner not extending posteriorly; first abdominal tergite with distinct middle suture and a small blotch; inner tibial spur of fore leg bifurcate; hind coxa normal, not strongly elongated; apex of hind femur extends beyond apex of abdomen; hind tibia longer than hind femur and second trochanter together, inner tibial spur about as long as or shorter than half length of metabasitarsomere; metabasitarsomere strongly compressed laterally, distinctly shorter than following four tarsomeres together; claw without basal lobe, inner tooth much longer than outer tooth; fore wing with cu-a meeting cell 1M at about basal 1/7, apex of vein R distinctly turned down, vein R+M much shorter than cu-a, first abscissa of vein M parallel with 1mcu, anal cell with an oblique cross vein slightly beyond middle; hind wing with cell Rs and M closed, anal cell shortly petiolate, marginal vein in male absent; lancet slender, distinctly sclerotized, usually without distinct annular sutures; penis valve simple, very narrow or elliptical.
</paragraph>
<paragraph id="8BE936BEFF8A3146FF12928F92B3FEBE" blockId="2.[151,1437,152,514]" pageId="2" pageNumber="47">
Whilst some species are apparently narrowly oligophagous (e.g.
<taxonomicName id="4C564D3DFF8A3146FC1F928F9750FF48" box="[968,1082,152,177]" class="Insecta" family="Psychidae" genus="Tenthredo" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="sturmii">
<emphasis id="B922EAACFF8A3146FC1F928F9750FF48" box="[968,1082,152,177]" italics="true" pageId="2" pageNumber="47">S. sturmii</emphasis>
</taxonomicName>
on
<taxonomicName id="4C564D3DFF8A3146FBBF928F97B7FF48" box="[1128,1245,152,177]" class="Magnoliopsida" family="Balsaminaceae" genus="Impatiens" kingdom="Plantae" order="Ericales" pageId="2" pageNumber="47" phylum="Tracheophyta" rank="genus">
<emphasis id="B922EAACFF8A3146FBBF928F97B7FF48" box="[1128,1245,152,177]" italics="true" pageId="2" pageNumber="47">Impatiens</emphasis>
</taxonomicName>
species), others may be polyphagous, as seems to be indicated by records of
<taxonomicName id="4C564D3DFF8A3146FCB192AA9092FF2F" box="[870,1016,189,214]" class="Insecta" family="Tenthredinidae" genus="Siobla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="ruficornis">
<emphasis id="B922EAACFF8A3146FCB192AA9092FF2F" box="[870,1016,189,214]" italics="true" pageId="2" pageNumber="47">S. ruficornis</emphasis>
</taxonomicName>
on plants of various families in the foodplant list by
<bibRefCitation id="EFC74B4FFF8A3146FEB692F5912BFF05" author="Verzhutskii" box="[353,577,226,252]" pageId="2" pageNumber="47" refString="Verzhutskii, B. N. (1981) Rastitel'nojadnye nasekomye v jekosistemah Vostochnoj Sibiri (pilil'shhiki i rogohvosty). [Herbivorous insects in ecosystems of East Siberia (sawflies and wood-wasps)]. (In Russian), Nauka, Novosibirsk, pp. 107 - 109." type="book chapter" year="1981">Verzhutskii (1981)</bibRefCitation>
.
<bibRefCitation id="EFC74B4FFF8A3146FD9992F59182FF05" author="Liston" box="[590,744,226,252]" pageId="2" pageNumber="47" refString="Liston, A. D. (1995) Compendium of European Sawflies. List of species, modern nomenclature, distribution, foodplants, identification literature. Chalastos Forestry, Gottfrieding, pp. 94 - 95." type="book chapter" year="1995">Liston (1995</bibRefCitation>
,
<bibRefCitation id="EFC74B4FFF8A3146FD2292F5905EFF05" author="Liston" box="[757,820,226,252]" pageId="2" pageNumber="47" refString="Liston, A. D. (1997) Hostplant list for European and North African Megalodontoidea and Tenthredinoidea (Hym.). Sawfly News, 1 (3), 38 - 41." type="journal article" year="1997">1997</bibRefCitation>
),
<bibRefCitation id="EFC74B4FFF8A3146FC9D92F5909AFF05" author="Taeger" box="[842,1008,226,252]" pageId="2" pageNumber="47" refString="Taeger, A., Altenhofer, E., Blank, S. M., Jansen, E., Kraus, M., Pschorn-Walcher, H. &amp; Ritzau, C. (1998) Kommentare zur Biologie, Verbreitung und Gefahrdung der Pflanzenwespen Deutschlands (Hymenoptera, Symphyta). (In German). In: Taeger, A. &amp; Blank, S. M. (Eds.), Pflanzenwespen Deutschlands (Hymenoptera, Symphya). Kommentierte Bestandsaufnahme. Goecke &amp; Evers, Keltern, 118." type="book chapter" year="1998">Taeger (1998)</bibRefCitation>
<bibRefCitation id="EFC74B4FFF8A3146FC2F92F597C1FF05" author="Lacourt" box="[1016,1195,226,252]" pageId="2" pageNumber="47" refString="Lacourt, J. (1999) Repertoire des Tenthredinidae ouest-palearctiques (Hymenoptera, Symphyta). Memoires de la SEF, 3, 248." type="journal article" year="1999">Lacourt (1999)</bibRefCitation>
and
<bibRefCitation id="EFC74B4FFF8A3146FB3292F596F6FF05" author="Schmitz" box="[1253,1436,226,252]" pageId="2" pageNumber="47" refString="Schmitz, G. (2007) Neue Nachweise von monophagen Herbivoren am Neophyten Impatiens glandulifera: Siobla sturmi (Klug, 1817) (Hymenoptera: Tenthredinidae) und Xanthoroe biriviata (Borkhausen, 1794) (Lepidoptera: Geometridae). (In German, abstract in German and English). Entomologische Zeitschrift, 117 (2), 60 - 62." type="journal article" year="2007">Schmitz (2007)</bibRefCitation>
mentioned hostplants of
<taxonomicName id="4C564D3DFF8A3146FE17931F9167FED8" box="[448,525,264,289]" class="Insecta" family="Tenthredinidae" genus="Siobla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="genus">
<emphasis id="B922EAACFF8A3146FE17931F9167FED8" box="[448,525,264,289]" italics="true" pageId="2" pageNumber="47">Siobla</emphasis>
</taxonomicName>
species.
<bibRefCitation id="EFC74B4FFF8A3146FDAB931F9039FEDB" author="Nakamura" box="[636,851,264,290]" pageId="2" pageNumber="47" refString="Nakamura, S. (1962) New host plants of Siobla ferox Smith (Tenthredinidae). (In Japanese). Kontyu, 30 (2), 86." type="journal article" year="1962">Nakamura (1962)</bibRefCitation>
, Takeuchi (1949) and
<bibRefCitation id="EFC74B4FFF8A3146FBB6931F9641FEDB" box="[1121,1323,264,290]" pageId="2" pageNumber="47" refString="Lee, J. W., Ryu, S. M., Quan, Y. T. &amp; Jung, J. C. (2000) Economic Insects of Korea 2. Hymenoptera (Symphyta: Tenthredinidae). (In Korean, abstract in Korean and English). National Institute of Agricultural Science and Technology, Suwon, pp. 117 - 121, 189 - 221." type="journal article">
Lee
<emphasis id="B922EAACFF8A3146FB41931F97A1FED8" box="[1174,1227,264,289]" italics="true" pageId="2" pageNumber="47">et al</emphasis>
. (2000)
</bibRefCitation>
recorded some hostplants of
<taxonomicName id="4C564D3DFF8A3146FEAD933A92B8FEBF" box="[378,466,301,326]" class="Insecta" family="Tenthredinidae" genus="Siobla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="ferox">
<emphasis id="B922EAACFF8A3146FEAD933A92B8FEBF" box="[378,466,301,326]" italics="true" pageId="2" pageNumber="47">S. ferox</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BE936BEFF8A3146FF1293459168FDFB" blockId="2.[151,1437,152,514]" pageId="2" pageNumber="47">
The limited existing data suggest that most
<taxonomicName id="4C564D3DFF8A3146FD1B9345907DFE92" box="[716,791,338,363]" class="Insecta" family="Tenthredinidae" genus="Siobla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="genus">
<emphasis id="B922EAACFF8A3146FD1B9345907DFE92" box="[716,791,338,363]" italics="true" pageId="2" pageNumber="47">Siobla</emphasis>
</taxonomicName>
are univoltine. Overwintering is in a leathery cocoon: emergence may sometimes take place after two winters (
<bibRefCitation id="EFC74B4FFF8A3146FC83936F976BFE6B" author="Malaise" box="[852,1025,376,402]" pageId="2" pageNumber="47" refString="Malaise, R. (1931) Blattwespen aus Wladiwostock und anderen Teilen Ostasiens. (In German). Entomologisk Tidskrift, 52, 97 - 159." type="journal article" year="1931">Malaise 1931</bibRefCitation>
). Larvae of
<taxonomicName id="4C564D3DFF8A3146FB40936F978CFE68" box="[1175,1254,376,401]" class="Insecta" family="Tenthredinidae" genus="Siobla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="genus">
<emphasis id="B922EAACFF8A3146FB40936F978CFE68" box="[1175,1254,376,401]" italics="true" pageId="2" pageNumber="47">Siobla</emphasis>
</taxonomicName>
are unusual in appearance, because they possess large, thorn-shaped, fleshy processes (described by some authors as “spines”) on the dorsum of thorax and abdomen (
<bibRefCitation id="EFC74B4FFF8A3146FD3A93D59096FE25" author="Lorenz" box="[749,1020,450,476]" pageId="2" pageNumber="47" refString="Lorenz, H. &amp; Kraus, M. (1957) Die Larvalsystematik der Blattwespen (Tenthredinoidea und Megalodontoidea). (In German). Abhandlungen zur Larvalsystematik der Insekten, 1, 52 - 53." type="journal article" year="1957">Lorenz &amp; Kraus 1957</bibRefCitation>
;
<bibRefCitation id="EFC74B4FFF8A3146FBDE93D597F1FE25" author="Ahola" box="[1033,1179,450,476]" pageId="2" pageNumber="47" refString="Ahola, M. (2001) Redescription of the larva of Siobla ruficornis (Gimmerthal, 1834) (Hymenoptera, Tenthredinidae) from East Finland. Entomologica Fennica, 12, 28 - 32." type="journal article" year="2001">Ahola 2001</bibRefCitation>
). After the last larval moult, these processes are lost.
</paragraph>
<paragraph id="8BE936BEFF8A3146FF4096D29260FAF8" blockId="2.[151,1436,1221,1281]" pageId="2" pageNumber="47">
<emphasis id="B922EAACFF8A3146FF4096D29262FB24" bold="true" box="[151,264,1221,1245]" pageId="2" pageNumber="47">PLATE 1.</emphasis>
<taxonomicName id="4C564D3DFF8A3146FED896D2917AFB25" authority="Wei, 2005" authorityName="Wei" authorityYear="2005" box="[271,528,1221,1244]" class="Insecta" family="Tenthredinidae" genus="Siobla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="zuoae">
<emphasis id="B922EAACFF8A3146FED896D292F3FB25" box="[271,409,1221,1244]" italics="true" pageId="2" pageNumber="47">Siobla zuoae</emphasis>
Wei, 2005
</taxonomicName>
. (a). Veins of fore wing; (b). Veins of hind wing; (c). Cells of fore wing; (d). Cells of hind wing.
</paragraph>
</subSubSection>
</treatment>
</document>