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<document id="6497213AAF2735C2A6547C751B578DF8" ID-DOI="10.11646/zootaxa.2391.1.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="5308365" IM.bibliography_approvedBy="diego" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="admin" IM.taxonomicNames_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1629669120741" checkinUser="felipe" docAuthor="Richards, Stephen J., Hoskin, Conrad J., Cunningham, Michael J., Mcdonald, Keith &amp; Donnellan, Stephen C." docDate="2010" docId="03FF878FFFA0200A48B3FB61FB1CB584" docLanguage="en" docName="zt02391p046.pdf" docOrigin="Zootaxa 2391 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2391.1.2" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Nyctimystes loveridgei Neill 1954" docType="treatment" docVersion="1" lastPageNumber="42" masterDocId="FFC6FFF7FFA920034877FFE9FFEFB023" masterDocTitle="Taxonomic re-assessment of the Australian and New Guinean green-eyed treefrogs Litoria eucnemis, L. genimaculata and L. serrata (Anura: Hylidae)" masterLastPageNumber="46" masterPageNumber="33" pageNumber="42" updateTime="1730367965011" updateUser="diego" zenodo-license-document="CLOSED">
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<mods:title id="86802054FCAACCFA499AAFB0F10D1C96">Taxonomic re-assessment of the Australian and New Guinean green-eyed treefrogs Litoria eucnemis, L. genimaculata and L. serrata (Anura: Hylidae)</mods:title>
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<mods:namePart id="BC17C423FB5C01C018237A3B6A6D13B4">Richards, Stephen J.</mods:namePart>
<mods:affiliation id="3689F19D104EF66D2639E90558596EA2">South Australian Museum, North Terrace, Adelaide, 5000, Australia. Email: Steve. Richards @ samuseum. sa. gov. au, and Conservation International, P. O. Box 1024, Atherton, Queensland 4883, Australia.</mods:affiliation>
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<mods:namePart id="B6DDF8F18A3FA4E6E2391575CF891D03">Hoskin, Conrad J.</mods:namePart>
<mods:affiliation id="22AA0DC2E0960C2B04E6FB225A39948C">Research School of Biology, The Australian National University, Canberra, 0200, Australia. Email: conrad. hoskin @ anu. edu. au.</mods:affiliation>
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<mods:namePart id="2B14ED98B1D835FE5868824191B90A15">Cunningham, Michael J.</mods:namePart>
<mods:affiliation id="7884F7DB3EE2BD7F9EF1088B0361D59E">School of Integrative Biology, The University of Queensland, St Lucia, 4072, Qld, Australia. &amp; Department of Zoology, University of the Free State, Private Bag X 13, Phuthaditjhaba, 9866, South Africa. Email: CunninghamMJ @ qwa. ufs. ac. za.</mods:affiliation>
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<mods:namePart id="B21B1E531845543EEF2A12EAAD2ECAC2">Mcdonald, Keith</mods:namePart>
<mods:affiliation id="D816CAE283E1C7DA104C147EC7AB3BDC">Queensland Department of Environment and Heritage, Wet Tropics District Office, P. O. Box 834, Atherton, Australia, 4883. Email: keith. mcdonald @ epa. qld. gov. au.</mods:affiliation>
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<mods:namePart id="CAE7AEF1E2C5DCD0114C56F6AE6EE404">Donnellan, Stephen C.</mods:namePart>
<mods:affiliation id="40AD735A8ADA4485E4334874DC6348F0">South Australian Museum, North Terrace, Adelaide, 5000, Australia. Email: Steve. Richards @ samuseum. sa. gov. au, and Conservation International, P. O. Box 1024, Atherton, Queensland 4883, Australia. &amp; Australian Centre for Evolutionary Biology and Biodiversity, University of Adelaide, Adelaide 5005.</mods:affiliation>
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<mods:date id="FF91411CCD04419147F9D67783E5D87F">2010</mods:date>
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<paragraph id="8BE93699FFA0200A48B3FB61FD82B481" blockId="9.[151,1436,152,2007]" box="[196,621,1160,1186]" pageId="9" pageNumber="42">
<taxonomicName id="4C564D1AFFA0200A48B3FB61FD86B481" ID-CoL="485NK" authority="Neill, 1954" authorityName="Neill" authorityYear="1954" box="[196,617,1160,1186]" class="Amphibia" family="Pelodryadidae" genus="Nyctimystes" kingdom="Animalia" order="Anura" pageId="9" pageNumber="42" phylum="Chordata" rank="species" species="loveridgei">
<emphasis id="B922EA8BFFA0200A48B3FB61FE39B482" box="[196,470,1160,1185]" italics="true" pageId="9" pageNumber="42">Nyctimystes loveridgei</emphasis>
<bibRefCitation id="EFC74B68FFA0200A49A9FB61FD86B481" author="Neill, W. T." box="[478,617,1160,1186]" pageId="9" pageNumber="42" pagination="83 - 85" refId="ref6185" refString="Neill, W. T. (1954) A new species of frog, genus Nyctimystes, from Papua. Copeia, 1954, 83 - 85." type="journal article" year="1954">Neill, 1954</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C34C6512FFA0200A4A01FB61FCA1B4E4" pageId="9" pageNumber="42" type="materials_examined">
<paragraph id="8BE93699FFA0200A4A01FB61FCA1B4E4" blockId="9.[151,1436,152,2007]" pageId="9" pageNumber="42">
<materialsCitation id="3B3E3CC4FFA0200A4A01FB61FCA4B4E4" pageId="9" pageNumber="42">
<typeStatus id="54ED883BFFA0200A4A01FB61FD05B481" box="[630,746,1160,1186]" pageId="9" pageNumber="42" type="holotype">Holotype</typeStatus>
: MCZ 27823. Type locality: near
<location id="8E896042FFA0200A4CF2FB61FB39B481" LSID="urn:lsid:plazi:treatment:03FF878FFFA0200A48B3FB61FB1CB584:8E896042FFA0200A4CF2FB61FB39B481" box="[1157,1238,1160,1186]" country="Papua New Guinea" county="We" municipality="Furthermore" name="Taburi" pageId="9" pageNumber="42">Taburi</location>
, a native village about
<quantity id="4CAE9B7CFFA0200A4897FB44FEDAB4E4" box="[224,309,1197,1223]" metricMagnitude="3" metricUnit="m" metricValue="3.218688" pageId="9" pageNumber="42" unit="mi" value="2.0">2 miles</quantity>
southeast of
<location id="8E896042FFA0200A49A5FB44FDB8B4E4" LSID="urn:lsid:plazi:treatment:03FF878FFFA0200A48B3FB61FB1CB584:8E896042FFA0200A49A5FB44FDB8B4E4" box="[466,599,1197,1223]" country="Papua New Guinea" county="We" municipality="Furthermore" name="Rouna Fall" pageId="9" pageNumber="42">Rouna Fall</location>
,
<collectingCountry id="F3417609FFA0200A4A14FB44FCA4B4E4" box="[611,843,1197,1223]" name="Papua New Guinea" pageId="9" pageNumber="42">Papua New Guinea</collectingCountry>
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.
</paragraph>
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<subSubSection id="C34C6512FFA0200A4B21FB44FB1CB584" pageId="9" pageNumber="42" type="discussion">
<paragraph id="8BE93699FFA0200A4B21FB44FB1CB584" blockId="9.[151,1436,152,2007]" pageId="9" pageNumber="42">
<location id="8E896042FFA0200A4B21FB44FC65B4E4" LSID="urn:lsid:plazi:treatment:03FF878FFFA0200A48B3FB61FB1CB584:8E896042FFA0200A4B21FB44FC65B4E4" box="[854,906,1197,1223]" country="Papua New Guinea" county="We" municipality="Furthermore" name="This" pageId="9" pageNumber="42">This</location>
species is currently incorrectly synonymised with
<taxonomicName id="4C564D1AFFA0200A48A5FB3AFE7FB4CF" authorityName="Horst" authorityYear="1883" box="[210,400,1235,1260]" class="Amphibia" family="Hylidae" genus="Hyla" kingdom="Animalia" order="Anura" pageId="9" pageNumber="42" phylum="Chordata" rank="species" species="genimaculata">
<emphasis id="B922EA8BFFA0200A48A5FB3AFE7FB4CF" box="[210,400,1235,1260]" italics="true" pageId="9" pageNumber="42">L. genimaculata</emphasis>
</taxonomicName>
.
<location id="8E896042FFA0200A49E8FB3BFE0EB4CF" LSID="urn:lsid:plazi:treatment:03FF878FFFA0200A48B3FB61FB1CB584:8E896042FFA0200A49E8FB3BFE0EB4CF" box="[415,481,1234,1260]" country="Papua New Guinea" county="We" municipality="Furthermore" name="Adult" pageId="9" pageNumber="42">Adult</location>
males are much larger than
<taxonomicName id="4C564D1AFFA0200A4B5BFB3AFC05B4CF" authorityName="Horst" authorityYear="1883" box="[812,1002,1235,1260]" class="Amphibia" family="Hylidae" genus="Hyla" kingdom="Animalia" order="Anura" pageId="9" pageNumber="42" phylum="Chordata" rank="species" species="genimaculata">
<emphasis id="B922EA8BFFA0200A4B5BFB3AFC05B4CF" box="[812,1002,1235,1260]" italics="true" pageId="9" pageNumber="42">L. genimaculata</emphasis>
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(SVL 39.547.5;
<typeStatus id="54ED883BFFA0200A4CC8FB3BFACAB4CF" box="[1215,1317,1234,1260]" pageId="9" pageNumber="42" type="holotype">holotype</typeStatus>
41.1 mm, SJR pers. obs.), and the original description mentions a triangular heel lappet that is present only in
<taxonomicName id="4C564D1AFFA0200A4DF3FB11FEEBB515" authorityName="Lonnberg" authorityYear="1900" class="Amphibia" family="Hylidae" genus="Hyla" kingdom="Animalia" order="Anura" pageId="9" pageNumber="42" phylum="Chordata" rank="species" species="eucnemis">
<emphasis id="B922EA8BFFA0200A4DF3FB11FEEBB515" italics="true" pageId="9" pageNumber="42">L. eucnemis</emphasis>
</taxonomicName>
(
<figureCitation id="136D2A1CFFA0200A4964FAF4FE7EB514" box="[275,401,1309,1335]" captionStart="FIGURE 4" captionStartId="8.[151,255,1876,1900]" captionTargetBox="[183,1403,198,1841]" captionTargetId="figure-104@8.[179,1404,194,1851]" captionTargetPageId="8" captionText="FIGURE 4. Photographs in life of A) Litoria eucnemis, Muller Range, PNG, B) L. exophthalmia, Namosado, PNG, C) L. genimaculata, Waigeo Island, Indonesia, D) L. serrata, Wooroonooran National Park, Australia, and of preserved specimens of E) holotype of L. genimaculata, RMNH 4420, F) hand of L. genimaculata holotype showing extent of finger webbing, G) holotype of Hyla rhacophorus, ZMA 5823, and H) holotype of Nyctimystes loveridgei, MCZ 27823 including (below) close-up of large heel flap. All scale bars = 10 mm" figureDoi="http://doi.org/10.5281/zenodo.5308375" httpUri="https://zenodo.org/record/5308375/files/figure.png" pageId="9" pageNumber="42">Figure 4H</figureCitation>
).
<collectingMunicipality id="6B8DACE3FFA0200A49D2FAF4FDD4B514" box="[421,571,1309,1335]" pageId="9" pageNumber="42">Furthermore</collectingMunicipality>
, the original description (
<bibRefCitation id="EFC74B68FFA0200A4B11FAF4FC09B514" author="Neill, W. T." box="[870,998,1309,1335]" pageId="9" pageNumber="42" pagination="83 - 85" refId="ref6185" refString="Neill, W. T. (1954) A new species of frog, genus Nyctimystes, from Papua. Copeia, 1954, 83 - 85." type="journal article" year="1954">Neill 1954</bibRefCitation>
) notes that the call is a series of five or six raucous notes, like laughter which could describe the quite loud series of waah, waah growls produced by
<taxonomicName id="4C564D1AFFA0200A494FFA81FE46B5A2" authorityName="Lonnberg" authorityYear="1900" box="[312,425,1384,1409]" class="Amphibia" family="Hylidae" genus="Hyla" kingdom="Animalia" order="Anura" pageId="9" pageNumber="42" phylum="Chordata" rank="species" species="eucnemis">
<emphasis id="B922EA8BFFA0200A494FFA81FE46B5A2" box="[312,425,1384,1409]" italics="true" pageId="9" pageNumber="42">eucnemis</emphasis>
</taxonomicName>
but not the soft ticking notes produced by
<taxonomicName id="4C564D1AFFA0200A4BA1FA81FB72B5A2" authorityName="Horst" authorityYear="1883" box="[982,1181,1384,1409]" class="Amphibia" family="Hylidae" genus="Hyla" kingdom="Animalia" order="Anura" pageId="9" pageNumber="42" phylum="Chordata" rank="species" species="genimaculata">
<emphasis id="B922EA8BFFA0200A4BA1FA81FB72B5A2" box="[982,1181,1384,1409]" italics="true" pageId="9" pageNumber="42">L. genimaculata</emphasis>
</taxonomicName>
.
<collectingCounty id="62884E15FFA0200A4CD8FA81FB39B5A1" box="[1199,1238,1384,1410]" pageId="9" pageNumber="42">We</collectingCounty>
here remove
<taxonomicName id="4C564D1AFFA0200A4DF7FA81FEE1B585" authorityName="Neill" authorityYear="1954" class="Amphibia" family="Pelodryadidae" genus="Nyctimystes" kingdom="Animalia" order="Anura" pageId="9" pageNumber="42" phylum="Chordata" rank="species" species="loveridgei">
<emphasis id="B922EA8BFFA0200A4DF7FA81FEE1B585" italics="true" pageId="9" pageNumber="42">N. loveridgei</emphasis>
</taxonomicName>
from the synonymy of
<taxonomicName id="4C564D1AFFA0200A4A56FA64FD0FB585" authorityName="Horst" authorityYear="1883" box="[545,736,1421,1446]" class="Amphibia" family="Hylidae" genus="Hyla" kingdom="Animalia" order="Anura" pageId="9" pageNumber="42" phylum="Chordata" rank="species" species="genimaculata">
<emphasis id="B922EA8BFFA0200A4A56FA64FD0FB585" box="[545,736,1421,1446]" italics="true" pageId="9" pageNumber="42">L. genimaculata</emphasis>
</taxonomicName>
and place it in the synonymy of
<taxonomicName id="4C564D1AFFA0200A4C17FA64FB03B585" authorityName="Lonnberg" authorityYear="1900" box="[1120,1260,1421,1446]" class="Amphibia" family="Hylidae" genus="Hyla" kingdom="Animalia" order="Anura" pageId="9" pageNumber="42" phylum="Chordata" rank="species" species="eucnemis">
<emphasis id="B922EA8BFFA0200A4C17FA64FB03B585" box="[1120,1260,1421,1446]" italics="true" pageId="9" pageNumber="42">L. eucnemis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
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