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<document id="C06ACDE1E90E1A7877E7872F9618C46C" ID-DOI="10.5281/zenodo.222736" ID-GBIF-Dataset="59197a35-ca2a-4ed2-8a1b-f756ce67e1c5" ID-ISSN="1175-5326" ID-Zenodo-Dep="222736" ID-ZooBank="B04BEFE6-86E0-4F43-A578-DAEFCA7FCDB6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460567221551" checkinUser="plazi" docAuthor="Martins, Itamar A. & Zaher, Hussam" docDate="2013" docId="03EC58232A69AE64C9ACFD36FD0894D5" docLanguage="en" docName="zt03599p188.pdf" docOrigin="Zootaxa 3599 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Holoaden suarezi Martins & Zaher, 2013, sp. nov." docType="treatment" docVersion="8" lastPageNumber="185" masterDocId="FFD5205B2A68AE63C93BFFB9FFCF9324" masterDocTitle="A new species of the highland frog genus Holoaden (Amphibia, Strabomantidae) from cloud forests of southeastern Brazil" masterLastPageNumber="188" masterPageNumber="178" pageNumber="179" updateTime="1698321268696" updateUser="plazi">
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<mods:mods id="8259AA94F5AD92220B028A2FD30850AB" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="E25A7BC5F54516BEDC2CF9751F3955A7">
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<mods:title id="F2168A14ABAF9281E99BA13FD86F41E4">A new species of the highland frog genus Holoaden (Amphibia, Strabomantidae) from cloud forests of southeastern Brazil</mods:title>
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</mods:titleInfo>
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<mods:name id="68C24E90FB60B5C658CE8838136A7D99" type="personal">
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<mods:roleTerm id="2CB260B22693A10042DD34B862791398">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="C17C62B63FF0880F975E1C47E5B345B7">Martins, Itamar A.</mods:namePart>
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<mods:name id="6C3B54AC02D2C7F9076DAB6077CF9FA6" type="personal">
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<mods:roleTerm id="609EE6D84DEE50562F409E31D83489F8">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="36DC8893620569AE6839728C77088358">Zaher, Hussam</mods:namePart>
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<mods:typeOfResource id="B19394DB570193D00631AA8F03292572">text</mods:typeOfResource>
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<mods:title id="2D589C1188685A6A3F73513B3488E670">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="BAA37450CFCD5EEB1BE7912FE85D432F">
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<mods:date id="33828847D0E5DBA589FE55A6D154D50A">2013</mods:date>
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<mods:detail id="D8F27B101D9BB793F6C5D7560F2BC9E4" type="volume">
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<mods:number id="8E961292C15245A53AFD7DDCEDD45E58">3599</mods:number>
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<mods:detail id="05252579E99B65121ED2728488439B15" type="issue">
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<mods:number id="BE5EAD5FCBF7FC4DF570D89BE4954798">2</mods:number>
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<mods:start id="E01C41C830A6B7BB7C6904C3CCA15ED8">178</mods:start>
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<mods:end id="29530B8F567A631622D83466AB6E5465">188</mods:end>
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<mods:classification id="F0DFC6D4580A66A4D797EA9701CF067E">journal article</mods:classification>
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<mods:identifier id="4CFF4E8B72E6E357A763772E70380432" type="DOI">10.5281/zenodo.222736</mods:identifier>
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<mods:identifier id="85A47C09CADAAFA3F6E88883943CE9EB" type="GBIF-Dataset">59197a35-ca2a-4ed2-8a1b-f756ce67e1c5</mods:identifier>
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<mods:identifier id="4D7A4F2A6C503AB54CC69AEE68F6DB6A" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="16502039AA07925DC240BC65D4026C49" type="Zenodo-Dep">222736</mods:identifier>
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<treatment id="03EC58232A69AE64C9ACFD36FD0894D5" ID-DOI="http://doi.org/10.5281/zenodo.5628864" ID-GBIF-Taxon="119557823" ID-Zenodo-Dep="5628864" LSID="urn:lsid:plazi:treatment:03EC58232A69AE64C9ACFD36FD0894D5" httpUri="http://treatment.plazi.org/id/03EC58232A69AE64C9ACFD36FD0894D5" lastPageId="7" lastPageNumber="185" pageId="1" pageNumber="179">
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<subSubSection id="C35FBABE2A69AE62C9ACFD36FE0391EE" pageId="1" pageNumber="179" type="nomenclature">
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<paragraph id="8BFAE9352A69AE62C9ACFD36FE1F918D" blockId="1.[151,464,655,714]" box="[151,464,655,681]" pageId="1" pageNumber="179">
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<heading id="D0B25E592A69AE62C9ACFD36FE1F918D" bold="true" box="[151,464,655,681]" fontSize="11" level="1" pageId="1" pageNumber="179" reason="1">
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<emphasis id="B93135272A69AE62C9ACFD36FE1F918D" bold="true" box="[151,464,655,681]" pageId="1" pageNumber="179">
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<taxonomicName id="4C4592B62A69AE62C9ACFD36FEA3918D" box="[151,364,655,681]" class="Amphibia" family="Strabomantidae" genus="Holoaden" kingdom="Animalia" order="Anura" pageId="1" pageNumber="179" phylum="Chordata" rank="species" species="suarezi" status="sp. nov.">
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||
<emphasis id="B93135272A69AE62C9ACFD36FEA3918D" bold="true" box="[151,364,655,681]" italics="true" pageId="1" pageNumber="179">Holoaden suarezi</emphasis>
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||
</taxonomicName>
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||
<taxonomicNameLabel id="A202885C2A69AE62C848FD36FE1F918D" box="[371,464,655,681]" pageId="1" pageNumber="179" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
|
||
</heading>
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</paragraph>
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<paragraph id="8BFAE9352A69AE62C9ACFD0BFE0391EE" blockId="1.[151,464,655,714]" box="[151,460,690,714]" pageId="1" pageNumber="179">
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<figureCitation id="137EF5B02A69AE62C9ACFD0BFF1091EE" box="[151,223,690,714]" captionStart="FIGURE 1" captionStartId="2.[151,250,1951,1974]" captionTargetBox="[198,1401,151,1915]" captionTargetId="figure@2.[198,1403,149,1930]" captionTargetPageId="2" captionText="FIGURE 1. Dorsal and ventral views of the body: A and B, holotype of Holoaden suarezi sp. nov. (SVL 42.6 mm, MZUSP 131872) C and D, holotype of Holoaden pholeter (SVL 44.6 mm, MNRJ 51475); E and F, lectotype of Holoaden luederwaldti (SVL 40.8 mm, MZUSP 87)." httpUri="https://zenodo.org/record/222738/files/figure.png" pageId="1" pageNumber="179">Figs. 1</figureCitation>
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A, B; 2A, B; 3A; 4A
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||
</paragraph>
|
||
</subSubSection>
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<subSubSection id="C35FBABE2A69AE62C9ACFD4DFC479761" pageId="1" pageNumber="179" type="materials_examined">
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<paragraph id="8BFAE9352A69AE62C9ACFD4DFBF89075" blockId="1.[151,1437,756,2028]" pageId="1" pageNumber="179">
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<typeStatus id="54FE57972A69AE62C9ACFD4DFECB9028" box="[151,260,756,780]" pageId="1" pageNumber="179" type="holotype">
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<emphasis id="B93135272A69AE62C9ACFD4DFECB9028" bold="true" box="[151,260,756,780]" pageId="1" pageNumber="179">Holotype</emphasis>
|
||
</typeStatus>
|
||
.
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||
<collectionCode id="ED5471F02A69AE62C82AFD4DFEA19028" box="[273,366,756,780]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="1" pageNumber="179">MZUSP</collectionCode>
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131872, adult female collected at Estação Ecológica de Bananal (
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<geoCoordinate id="EE718FF22A69AE62CD75FD4DFB279028" box="[1102,1256,756,780]" direction="south" orientation="latitude" pageId="1" pageNumber="179" precision="15" value="-22.795">22° 47’ 42” S</geoCoordinate>
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,
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<geoCoordinate id="EE718FF22A69AE62CDCEFD4DFA579028" box="[1269,1432,756,780]" direction="west" orientation="longitude" pageId="1" pageNumber="179" precision="15" value="-44.359997">44° 21’ 36” W</geoCoordinate>
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,
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<quantity id="4CBD44D02A69AE62C9ACFCAFFF22900A" box="[151,237,790,814]" metricMagnitude="3" metricUnit="m" metricValue="1.22" pageId="1" pageNumber="179" unit="m" value="1220.0">1220 m</quantity>
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above sea level), municipality of Bananal, State of São Paulo,
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<collectingCountry id="F352A9A52A69AE62CA83FCAFFC30900A" box="[952,1023,790,814]" name="Brazil" pageId="1" pageNumber="179">Brazil</collectingCountry>
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||
, on
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<date id="FFFBCFF52A69AE62CD14FCAFFAC9900A" box="[1071,1286,790,814]" pageId="1" pageNumber="179" value="2003-12-18">18 December 2003</date>
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by H. Zaher, A. P. Suarez, E. A. Domenico, T. Camolez, R. Clemente, R Masiero, and P. Nunes.
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||
</paragraph>
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||
<paragraph id="8BFAE9352A69AE62C9FCFCE5FC479761" blockId="1.[151,1437,756,2028]" pageId="1" pageNumber="179">
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||
<typeStatus id="54FE57972A69AE62C9FCFCE5FEF09050" box="[199,319,860,884]" pageId="1" pageNumber="179" type="paratype">
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||
<emphasis id="B93135272A69AE62C9FCFCE5FEF09050" bold="true" box="[199,319,860,884]" pageId="1" pageNumber="179">Paratypes</emphasis>
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||
</typeStatus>
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||
.
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<collectionCode id="ED5471F02A69AE62C875FCE5FE649050" box="[334,427,860,884]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="1" pageNumber="179">MZUSP</collectionCode>
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23794, adult female collected at Estação Biológica de Boracéia (
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<geoCoordinate id="EE718FF22A69AE62CDADFCE5FACA9050" box="[1174,1285,860,884]" direction="south" orientation="latitude" pageId="1" pageNumber="179" precision="925" value="-23.633333">23° 38’ S</geoCoordinate>
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,
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<geoCoordinate id="EE718FF22A69AE62CC28FCE5FA409050" box="[1299,1423,860,884]" direction="west" orientation="longitude" pageId="1" pageNumber="179" precision="925" value="-45.866665">45° 52’ W</geoCoordinate>
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), municipality of Salesópolis, State of São Paulo, on
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<date id="FFFBCFF52A69AE62CBE1FCC7FC4990B2" box="[730,902,894,918]" pageId="1" pageNumber="179" value="1963-03-23">23 March 1963</date>
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||
by A. S. Rand and P. Rand.
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<collectionCode id="ED5471F02A69AE62CDF2FCC7FAE990B2" box="[1225,1318,894,918]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="1" pageNumber="179">MZUSP</collectionCode>
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94415, an adult male collected at Campo de Fruticultura da Bocaina (presently known as Ponte Alta, inside of Parque Nacional da Serra da Bocaina) (
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<geoCoordinate id="EE718FF22A69AE62C8C0FC7DFDA890F8" box="[507,615,964,988]" direction="south" orientation="latitude" pageId="1" pageNumber="179" precision="925" value="-22.7">22° 42’ S</geoCoordinate>
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,
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<geoCoordinate id="EE718FF22A69AE62CB48FC7DFD2490F8" box="[627,747,964,988]" direction="west" orientation="longitude" pageId="1" pageNumber="179" precision="925" value="-44.633335">44° 38’ W</geoCoordinate>
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), Municipality of São José do Barreiro, state of São Paulo, on 0
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<date id="FFFBCFF52A69AE62C99DFC5FFEBF90DA" box="[166,368,998,1022]" pageId="1" pageNumber="179" value="1965-11-03">3 November 1965</date>
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by W. C. A. Bokermann (ex
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||
<collectionCode id="ED5471F02A69AE62CB87FC5FFCDF90DA" box="[700,784,998,1022]" pageId="1" pageNumber="179">WCAB</collectionCode>
|
||
37146);
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||
<collectionCode id="ED5471F02A69AE62CA4CFC5FFC1B90DA" box="[887,980,998,1022]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="1" pageNumber="179">MZUSP</collectionCode>
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138348, adult male collected at Estação Biológica de Boracéia (
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<geoCoordinate id="EE718FF22A69AE62C890FBB0FDD09705" box="[427,543,1033,1057]" direction="south" orientation="latitude" pageId="1" pageNumber="179" precision="925" value="-23.633333">23° 38’ S</geoCoordinate>
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,
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<geoCoordinate id="EE718FF22A69AE62CB0BFBB0FD7F9705" box="[560,688,1033,1057]" direction="west" orientation="longitude" pageId="1" pageNumber="179" precision="925" value="-45.866665">45° 52’ W</geoCoordinate>
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), Municipality of Salesópolis, State of São Paulo,
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<collectingCountry id="F352A9A52A69AE62CC38FBB0FA859705" box="[1283,1354,1033,1057]" name="Brazil" pageId="1" pageNumber="179">Brazil</collectingCountry>
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||
, on 0
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<date id="FFFBCFF52A69AE62CCB5FBB0FE839760" pageId="1" pageNumber="179" value="2003-11-06">6 November 2003</date>
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by M. T. Rodrigues, V. Verdade, and M. C. Lange.
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||
</paragraph>
|
||
</subSubSection>
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||
<subSubSection id="C35FBABE2A69AE62C9FCFBF7FB2E94C8" pageId="1" pageNumber="179" type="diagnosis">
|
||
<paragraph id="8BFAE9352A69AE62C9FCFBF7FDA09612" blockId="1.[151,1437,756,2028]" pageId="1" pageNumber="179">
|
||
<emphasis id="B93135272A69AE62C9FCFBF7FEF59743" bold="true" box="[199,314,1102,1127]" pageId="1" pageNumber="179">Diagnosis</emphasis>
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||
.
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<taxonomicName id="4C4592B62A69AE62C871FBF7FDDD9742" box="[330,530,1102,1126]" class="Amphibia" family="Strabomantidae" genus="Holoaden" kingdom="Animalia" order="Anura" pageId="1" pageNumber="179" phylum="Chordata" rank="species" species="suarezi">
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<emphasis id="B93135272A69AE62C871FBF7FDDD9742" box="[330,530,1102,1126]" italics="true" pageId="1" pageNumber="179">Holoaden suarezi</emphasis>
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</taxonomicName>
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||
can be recognized by the following combination of characters: (1) a moderate body size (female
|
||
<quantity id="4CBD44D02A69AE62C858FBC8FDC997AD" box="[355,518,1137,1161]" metricMagnitude="-2" metricUnit="m" metricValue="4.34" metricValueMax="4.42" metricValueMin="4.26" pageId="1" pageNumber="179" unit="mm" value="43.4" valueMax="44.2" valueMin="42.6">42.6–44.2 mm</quantity>
|
||
SVL; male
|
||
<quantity id="4CBD44D02A69AE62CBB7FBC8FCE197AD" box="[652,814,1137,1161]" metricMagnitude="-2" metricUnit="m" metricValue="3.785" metricValueMax="3.85" metricValueMin="3.72" pageId="1" pageNumber="179" unit="mm" value="37.85" valueMax="38.5" valueMin="37.2">37.2–38.5 mm</quantity>
|
||
SVL) indicating sexual dimorphism in size; (2) dorsum and head textures densely glandular (macroglands), with glands reaching the internasal region; (3) thigh and tibia with glandular texture; (4) forearms long and very slender; (5) legs and toes long and slender; (6) toe tips rounded, fingers not webbed nor fringed; (7) few supernumerary tubercles on the palmar region, (8) outer metatarsal tubercle elliptical, small; (9) texture of outer tarsal and sole of foot smooth; (10) iris black in living specimens; (11) tympanic membrane poorly or not visible.
|
||
</paragraph>
|
||
<paragraph id="8BFAE9352A69AE62C9FCFAF8FB2E94C8" blockId="1.[151,1437,756,2028]" pageId="1" pageNumber="179">
|
||
<emphasis id="B93135272A69AE62C9FCFAF8FD89967E" bold="true" box="[199,582,1345,1370]" pageId="1" pageNumber="179">Comparisons with other species</emphasis>
|
||
.
|
||
<taxonomicName id="4C4592B62A69AE62CB6DFAF8FCD2967D" box="[598,797,1345,1369]" class="Amphibia" family="Strabomantidae" genus="Holoaden" kingdom="Animalia" order="Anura" pageId="1" pageNumber="179" phylum="Chordata" rank="species" species="suarezi">
|
||
<emphasis id="B93135272A69AE62CB6DFAF8FCD2967D" box="[598,797,1345,1369]" italics="true" pageId="1" pageNumber="179">Holoaden suarezi</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C4592B62A69AE62CA8DFAFBFBEC967D" box="[950,1059,1345,1369]" class="Amphibia" family="Strabomantidae" genus="Holoaden" kingdom="Animalia" order="Anura" pageId="1" pageNumber="179" phylum="Chordata" rank="species" species="bradei">
|
||
<emphasis id="B93135272A69AE62CA8DFAFBFBEC967D" box="[950,1059,1345,1369]" italics="true" pageId="1" pageNumber="179">H. bradei</emphasis>
|
||
</taxonomicName>
|
||
by a larger body size (
|
||
<taxonomicName id="4C4592B62A69AE62CC16FAF8FF289658" class="Amphibia" family="Strabomantidae" genus="Holoaden" kingdom="Animalia" order="Anura" pageId="1" pageNumber="179" phylum="Chordata" rank="species" species="suarezi">
|
||
<emphasis id="B93135272A69AE62CC16FAF8FF289658" italics="true" pageId="1" pageNumber="179">Holoaden suarezi</emphasis>
|
||
</taxonomicName>
|
||
= males
|
||
<quantity id="4CBD44D02A69AE62C876FADDFE3F9658" box="[333,496,1380,1404]" metricMagnitude="-2" metricUnit="m" metricValue="3.785" metricValueMax="3.85" metricValueMin="3.72" pageId="1" pageNumber="179" unit="mm" value="37.85" valueMax="38.5" valueMin="37.2">37.2–38.5 mm</quantity>
|
||
SVL; females
|
||
<quantity id="4CBD44D02A69AE62CBA3FADDFCF49658" box="[664,827,1380,1404]" metricMagnitude="-2" metricUnit="m" metricValue="4.34" metricValueMax="4.42" metricValueMin="4.26" pageId="1" pageNumber="179" unit="mm" value="43.4" valueMax="44.2" valueMin="42.6">42.6–44.2 mm</quantity>
|
||
SVL;
|
||
<taxonomicName id="4C4592B62A69AE62CABDFADCFC3E9658" box="[902,1009,1380,1404]" class="Amphibia" family="Strabomantidae" genus="Holoaden" kingdom="Animalia" order="Anura" pageId="1" pageNumber="179" phylum="Chordata" rank="species" species="bradei">
|
||
<emphasis id="B93135272A69AE62CABDFADCFC3E9658" box="[902,1009,1380,1404]" italics="true" pageId="1" pageNumber="179">H. bradei</emphasis>
|
||
</taxonomicName>
|
||
= males
|
||
<quantity id="4CBD44D02A69AE62CD6CFADDFB359658" box="[1111,1274,1380,1404]" metricMagnitude="-2" metricUnit="m" metricValue="1.925" metricValueMax="2.84" metricValueMin="1.01" pageId="1" pageNumber="179" unit="mm" value="19.25" valueMax="28.4" valueMin="10.1">10.1–28.4 mm</quantity>
|
||
SVL; females
|
||
<quantity id="4CBD44D02A69AE62C9ACFA3FFEF296BA" box="[151,317,1414,1438]" metricMagnitude="-2" metricUnit="m" metricValue="3.1599999999999997" metricValueMax="3.6799999999999997" metricValueMin="2.6399999999999997" pageId="1" pageNumber="179" unit="mm" value="31.599999999999998" valueMax="36.8" valueMin="26.4">26.4–36.8 mm</quantity>
|
||
SVL) (
|
||
<tableCitation id="C6C7DC8E2A69AE62C8A3FA3FFE2096BA" box="[408,495,1414,1438]" captionStart="TABLE 1" captionStartId="4.[151,239,151,175]" captionTargetBox="[165,1420,240,912]" captionTargetPageId="4" captionText="TABLE 1. Measurements (in millimeters) of Holoaden suarezi sp. nov., Holoaden bradei, Holoaden luederwaldti, and Holoaden pholeter (Pombal et al., 2008). * Measurements include juveniles that were not sexed." httpUri="http://table.plazi.org/id/DF3AB9BD2A6CAE67C9ACFF2EFB5293EB" pageId="1" pageNumber="179" tableUuid="DF3AB9BD2A6CAE67C9ACFF2EFB5293EB">Table 1</tableCitation>
|
||
), longer limbs (
|
||
<taxonomicName id="4C4592B62A69AE62CB8BFA3FFCB696BA" box="[688,889,1414,1438]" class="Amphibia" family="Strabomantidae" genus="Holoaden" kingdom="Animalia" order="Anura" pageId="1" pageNumber="179" phylum="Chordata" rank="species" species="suarezi">
|
||
<emphasis id="B93135272A69AE62CB8BFA3FFCB696BA" box="[688,889,1414,1438]" italics="true" pageId="1" pageNumber="179">Holoaden suarezi</emphasis>
|
||
</taxonomicName>
|
||
= forearms
|
||
<quantity id="4CBD44D02A69AE62CD37FA3FFB7E96BA" box="[1036,1201,1414,1438]" metricMagnitude="-2" metricUnit="m" metricValue="1.17" metricValueMax="1.22" metricValueMin="1.1199999999999999" pageId="1" pageNumber="179" unit="mm" value="11.7" valueMax="12.2" valueMin="11.2">11.2–12.2 mm</quantity>
|
||
and 13.0–
|
||
<quantity id="4CBD44D02A69AE62CC0BFA3FFA5896BA" box="[1328,1431,1414,1438]" metricMagnitude="-2" metricUnit="m" metricValue="1.31" pageId="1" pageNumber="179" unit="mm" value="13.1">13.1 mm</quantity>
|
||
, thighs 17.8–18.0 mm and
|
||
<quantity id="4CBD44D02A69AE62C8E5FA10FD4396E5" box="[478,652,1449,1473]" metricMagnitude="-2" metricUnit="m" metricValue="1.9799999999999998" metricValueMax="1.9899999999999998" metricValueMin="1.97" pageId="1" pageNumber="179" unit="mm" value="19.799999999999997" valueMax="19.9" valueMin="19.7">19.7–19.9 mm</quantity>
|
||
, tibia
|
||
<quantity id="4CBD44D02A69AE62CBDAFA10FC4296E5" box="[737,909,1449,1473]" metricMagnitude="-2" metricUnit="m" metricValue="1.7299999999999998" metricValueMax="1.77" metricValueMin="1.69" pageId="1" pageNumber="179" unit="mm" value="17.299999999999997" valueMax="17.7" valueMin="16.9">16.9–17.7 mm</quantity>
|
||
and
|
||
<quantity id="4CBD44D02A69AE62CAEFFA10FB4F96E5" box="[980,1152,1449,1473]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" metricValueMax="1.8699999999999999" metricValueMin="1.83" pageId="1" pageNumber="179" unit="mm" value="18.5" valueMax="18.7" valueMin="18.3">18.3–18.7 mm</quantity>
|
||
in males and females, respectively;
|
||
<taxonomicName id="4C4592B62A69AE62C816FA74FE5696C0" box="[301,409,1484,1508]" class="Amphibia" family="Strabomantidae" genus="Holoaden" kingdom="Animalia" order="Anura" pageId="1" pageNumber="179" phylum="Chordata" rank="species" species="bradei">
|
||
<emphasis id="B93135272A69AE62C816FA74FE5696C0" box="[301,409,1484,1508]" italics="true" pageId="1" pageNumber="179">H. bradei</emphasis>
|
||
</taxonomicName>
|
||
= forearms
|
||
<quantity id="4CBD44D02A69AE62CB1FFA75FD6496C0" box="[548,683,1484,1508]" metricMagnitude="-3" metricUnit="m" metricValue="4.45" metricValueMax="6.7" metricValueMin="2.2" pageId="1" pageNumber="179" unit="mm" value="4.45" valueMax="6.7" valueMin="2.2">2.2–6.7 mm</quantity>
|
||
and
|
||
<quantity id="4CBD44D02A69AE62CBD8FA75FCA396C0" box="[739,876,1484,1508]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" metricValueMax="7.5" metricValueMin="5.3" pageId="1" pageNumber="179" unit="mm" value="6.4" valueMax="7.5" valueMin="5.3">5.3–7.5 mm</quantity>
|
||
, thighs
|
||
<quantity id="4CBD44D02A69AE62CAFFFA75FB9696C0" box="[964,1113,1484,1508]" metricMagnitude="-3" metricUnit="m" metricValue="8.1" metricValueMax="11.9" metricValueMin="4.3" pageId="1" pageNumber="179" unit="mm" value="8.1" valueMax="11.9" valueMin="4.3">4.3–11.9 mm</quantity>
|
||
and 5.9–12.1, tibia
|
||
<quantity id="4CBD44D02A69AE62CC01FA75FF0C9522" metricMagnitude="-3" metricUnit="m" metricValue="7.25" metricValueMax="10.9" metricValueMin="3.6" pageId="1" pageNumber="179" unit="mm" value="7.25" valueMax="10.9" valueMin="3.6">3.6–10.9 mm</quantity>
|
||
and 6.0–
|
||
<quantity id="4CBD44D02A69AE62C817FA57FE419522" box="[300,398,1518,1542]" metricMagnitude="-2" metricUnit="m" metricValue="1.21" pageId="1" pageNumber="179" unit="mm" value="12.1">12.1 mm</quantity>
|
||
in males and females, respectively) (
|
||
<tableCitation id="C6C7DC8E2A69AE62CA16FA57FC4E9522" box="[813,897,1518,1542]" captionStart="TABLE 1" captionStartId="4.[151,239,151,175]" captionTargetBox="[165,1420,240,912]" captionTargetPageId="4" captionText="TABLE 1. Measurements (in millimeters) of Holoaden suarezi sp. nov., Holoaden bradei, Holoaden luederwaldti, and Holoaden pholeter (Pombal et al., 2008). * Measurements include juveniles that were not sexed." httpUri="http://table.plazi.org/id/DF3AB9BD2A6CAE67C9ACFF2EFB5293EB" pageId="1" pageNumber="179" tableUuid="DF3AB9BD2A6CAE67C9ACFF2EFB5293EB">Table 1</tableCitation>
|
||
), and a uniformly dark brown coloration (upper surface of dorsum olive brown with irregular and dispersed black marks in
|
||
<taxonomicName id="4C4592B62A69AE62CAC7F9ABFBA6950D" box="[1020,1129,1553,1577]" class="Amphibia" family="Strabomantidae" genus="Holoaden" kingdom="Animalia" order="Anura" pageId="1" pageNumber="179" phylum="Chordata" rank="species" species="bradei">
|
||
<emphasis id="B93135272A69AE62CAC7F9ABFBA6950D" box="[1020,1129,1553,1577]" italics="true" pageId="1" pageNumber="179">H. bradei</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<taxonomicName id="4C4592B62A69AE62CDB8F9A8FA85950D" box="[1155,1354,1553,1577]" class="Amphibia" family="Strabomantidae" genus="Holoaden" kingdom="Animalia" order="Anura" pageId="1" pageNumber="179" phylum="Chordata" rank="species" species="suarezi">
|
||
<emphasis id="B93135272A69AE62CDB8F9A8FA85950D" box="[1155,1354,1553,1577]" italics="true" pageId="1" pageNumber="179">Holoaden suarezi</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C4592B62A69AE62C9EDF98CFE479568" box="[214,392,1588,1612]" class="Amphibia" family="Strabomantidae" genus="Holoaden" kingdom="Animalia" order="Anura" pageId="1" pageNumber="179" phylum="Chordata" rank="species" species="luederwaldti">
|
||
<emphasis id="B93135272A69AE62C9EDF98CFE479568" box="[214,392,1588,1612]" italics="true" pageId="1" pageNumber="179">H. luederwaldti</emphasis>
|
||
</taxonomicName>
|
||
by an uniformly dark brown coloration (dorsal surface brown to dark purple-brown or redbrown and ventral surface cream in
|
||
<taxonomicName id="4C4592B62A69AE62CB73F9E1FD33954A" box="[584,764,1622,1647]" class="Amphibia" family="Strabomantidae" genus="Holoaden" kingdom="Animalia" order="Anura" pageId="1" pageNumber="179" phylum="Chordata" rank="species" species="luederwaldti">
|
||
<emphasis id="B93135272A69AE62CB73F9E1FD33954A" box="[584,764,1622,1647]" italics="true" pageId="1" pageNumber="179">H. luederwaldti</emphasis>
|
||
</taxonomicName>
|
||
), longer forearms, thighs, and tibia (
|
||
<taxonomicName id="4C4592B62A69AE62CD8DF9EFFA4F954A" box="[1206,1408,1622,1646]" class="Amphibia" family="Strabomantidae" genus="Holoaden" kingdom="Animalia" order="Anura" pageId="1" pageNumber="179" phylum="Chordata" rank="species" species="suarezi">
|
||
<emphasis id="B93135272A69AE62CD8DF9EFFA4F954A" box="[1206,1408,1622,1646]" italics="true" pageId="1" pageNumber="179">Holoaden suarezi</emphasis>
|
||
</taxonomicName>
|
||
= forearms
|
||
<quantity id="4CBD44D02A69AE62C83EF9C0FE6595B5" box="[261,426,1657,1681]" metricMagnitude="-2" metricUnit="m" metricValue="1.17" metricValueMax="1.22" metricValueMin="1.1199999999999999" pageId="1" pageNumber="179" unit="mm" value="11.7" valueMax="12.2" valueMin="11.2">11.2–12.2 mm</quantity>
|
||
and 13.0–
|
||
<quantity id="4CBD44D02A69AE62CB13F9C0FD4095B5" box="[552,655,1657,1681]" metricMagnitude="-2" metricUnit="m" metricValue="1.31" pageId="1" pageNumber="179" unit="mm" value="13.1">13.1 mm</quantity>
|
||
, thighs 17.8–18.0 mm and
|
||
<quantity id="4CBD44D02A69AE62CAEBF9C0FBB795B5" box="[976,1144,1657,1681]" metricMagnitude="-2" metricUnit="m" metricValue="1.9799999999999998" metricValueMax="1.9899999999999998" metricValueMin="1.97" pageId="1" pageNumber="179" unit="mm" value="19.799999999999997" valueMax="19.9" valueMin="19.7">19.7–19.9 mm</quantity>
|
||
, tibia
|
||
<quantity id="4CBD44D02A69AE62CDF8F9C0FAA595B5" box="[1219,1386,1657,1681]" metricMagnitude="-2" metricUnit="m" metricValue="1.7299999999999998" metricValueMax="1.77" metricValueMin="1.69" pageId="1" pageNumber="179" unit="mm" value="17.299999999999997" valueMax="17.7" valueMin="16.9">16.9–17.7 mm</quantity>
|
||
and
|
||
<quantity id="4CBD44D02A69AE62C9ACF925FEF29590" box="[151,317,1692,1716]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" metricValueMax="1.8699999999999999" metricValueMin="1.83" pageId="1" pageNumber="179" unit="mm" value="18.5" valueMax="18.7" valueMin="18.3">18.3–18.7 mm</quantity>
|
||
in males and females, respectively;
|
||
<taxonomicName id="4C4592B62A69AE62CBDEF924FC579590" box="[741,920,1692,1716]" class="Amphibia" family="Strabomantidae" genus="Holoaden" kingdom="Animalia" order="Anura" pageId="1" pageNumber="179" phylum="Chordata" rank="species" species="luederwaldti">
|
||
<emphasis id="B93135272A69AE62CBDEF924FC579590" box="[741,920,1692,1716]" italics="true" pageId="1" pageNumber="179">H. luederwaldti</emphasis>
|
||
</taxonomicName>
|
||
= forearms
|
||
<quantity id="4CBD44D02A69AE62CD11F925FB0E9590" box="[1066,1217,1692,1716]" metricMagnitude="-3" metricUnit="m" metricValue="9.75" metricValueMax="11.6" metricValueMin="7.9" pageId="1" pageNumber="179" unit="mm" value="9.75" valueMax="11.6" valueMin="7.9">7.9–11.6 mm</quantity>
|
||
and
|
||
<quantity id="4CBD44D02A69AE62CDC6F925FA589590" box="[1277,1431,1692,1716]" metricMagnitude="-2" metricUnit="m" metricValue="1.06" metricValueMax="1.21" metricValueMin="0.9099999999999999" pageId="1" pageNumber="179" unit="mm" value="10.6" valueMax="12.1" valueMin="9.1">9.1–12.1 mm</quantity>
|
||
, thighs
|
||
<quantity id="4CBD44D02A69AE62C9D9F907FE4C95F2" box="[226,387,1726,1750]" metricMagnitude="-2" metricUnit="m" metricValue="1.46" metricValueMax="1.75" metricValueMin="1.17" pageId="1" pageNumber="179" unit="mm" value="14.6" valueMax="17.5" valueMin="11.7">11.7–17.5 mm</quantity>
|
||
and
|
||
<quantity id="4CBD44D02A69AE62C882F907FD9295F2" box="[441,605,1726,1750]" metricMagnitude="-2" metricUnit="m" metricValue="1.685" metricValueMax="1.8199999999999998" metricValueMin="1.55" pageId="1" pageNumber="179" unit="mm" value="16.85" valueMax="18.2" valueMin="15.5">15.5–18.2 mm</quantity>
|
||
, tibia 8.0–17.0 mm and
|
||
<quantity id="4CBD44D02A69AE62CA50F907FBC295F2" box="[875,1037,1726,1750]" metricMagnitude="-2" metricUnit="m" metricValue="1.3699999999999999" metricValueMax="1.72" metricValueMin="1.02" pageId="1" pageNumber="179" unit="mm" value="13.7" valueMax="17.2" valueMin="10.2">10.2–17.2 mm</quantity>
|
||
in males and females, respectively) (
|
||
<tableCitation id="C6C7DC8E2A69AE62C9A5F958FF3D95DD" box="[158,242,1761,1785]" captionStart="TABLE 1" captionStartId="4.[151,239,151,175]" captionTargetBox="[165,1420,240,912]" captionTargetPageId="4" captionText="TABLE 1. Measurements (in millimeters) of Holoaden suarezi sp. nov., Holoaden bradei, Holoaden luederwaldti, and Holoaden pholeter (Pombal et al., 2008). * Measurements include juveniles that were not sexed." httpUri="http://table.plazi.org/id/DF3AB9BD2A6CAE67C9ACFF2EFB5293EB" pageId="1" pageNumber="179" tableUuid="DF3AB9BD2A6CAE67C9ACFF2EFB5293EB">Table 1</tableCitation>
|
||
), a longer length of the head that corresponds to 79% of its width (65% of head width in
|
||
<taxonomicName id="4C4592B62A69AE62CDE7F95BFA4495DD" box="[1244,1419,1761,1785]" class="Amphibia" family="Strabomantidae" genus="Holoaden" kingdom="Animalia" order="Anura" pageId="1" pageNumber="179" phylum="Chordata" rank="species" species="luederwaldti">
|
||
<emphasis id="B93135272A69AE62CDE7F95BFA4495DD" box="[1244,1419,1761,1785]" italics="true" pageId="1" pageNumber="179">H. luederwaldti</emphasis>
|
||
</taxonomicName>
|
||
), and a black iris coloration in live specimens (brown iris in
|
||
<taxonomicName id="4C4592B62A69AE62CA76F8BCFC309438" box="[845,1023,1796,1820]" class="Amphibia" family="Strabomantidae" genus="Holoaden" kingdom="Animalia" order="Anura" pageId="1" pageNumber="179" phylum="Chordata" rank="species" species="luederwaldti">
|
||
<emphasis id="B93135272A69AE62CA76F8BCFC309438" box="[845,1023,1796,1820]" italics="true" pageId="1" pageNumber="179">H. luederwaldti</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<taxonomicName id="4C4592B62A69AE62CD20F8BDFB2C9438" box="[1051,1251,1796,1820]" class="Amphibia" family="Strabomantidae" genus="Holoaden" kingdom="Animalia" order="Anura" pageId="1" pageNumber="179" phylum="Chordata" rank="species" species="suarezi">
|
||
<emphasis id="B93135272A69AE62CD20F8BDFB2C9438" box="[1051,1251,1796,1820]" italics="true" pageId="1" pageNumber="179">Holoaden suarezi</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C4592B62A69AE62CCBAF8BCFF3A941A" class="Amphibia" family="Strabomantidae" genus="Holoaden" kingdom="Animalia" order="Anura" pageId="1" pageNumber="179" phylum="Chordata" rank="species" species="pholeter">
|
||
<emphasis id="B93135272A69AE62CCBAF8BCFF3A941A" italics="true" pageId="1" pageNumber="179">H. pholeter</emphasis>
|
||
</taxonomicName>
|
||
an uniformly dark grey ventral surface of the body (ventral surface of the body dark brown with a light abdominal blotch of variable size in
|
||
<taxonomicName id="4C4592B62A69AE62CB04F8F3FD0E9445" box="[575,705,1865,1889]" class="Amphibia" family="Strabomantidae" genus="Holoaden" kingdom="Animalia" order="Anura" pageId="1" pageNumber="179" phylum="Chordata" rank="species" species="pholeter">
|
||
<emphasis id="B93135272A69AE62CB04F8F3FD0E9445" box="[575,705,1865,1889]" italics="true" pageId="1" pageNumber="179">H. pholeter</emphasis>
|
||
</taxonomicName>
|
||
) (
|
||
<figureCitation id="137EF5B02A69AE62CBE7F8F0FCEF9445" box="[732,800,1865,1889]" captionStart="FIGURE 1" captionStartId="2.[151,250,1951,1974]" captionTargetBox="[198,1401,151,1915]" captionTargetId="figure@2.[198,1403,149,1930]" captionTargetPageId="2" captionText="FIGURE 1. Dorsal and ventral views of the body: A and B, holotype of Holoaden suarezi sp. nov. (SVL 42.6 mm, MZUSP 131872) C and D, holotype of Holoaden pholeter (SVL 44.6 mm, MNRJ 51475); E and F, lectotype of Holoaden luederwaldti (SVL 40.8 mm, MZUSP 87)." httpUri="https://zenodo.org/record/222738/files/figure.png" pageId="1" pageNumber="179">Fig. 1</figureCitation>
|
||
), dense glandular granules on the dorsal surface of the head and body that extend to the internasal region (glandular granules extend to the inter-orbital region in
|
||
<taxonomicName id="4C4592B62A69AE62CCBAF8D4FF3B9482" class="Amphibia" family="Strabomantidae" genus="Holoaden" kingdom="Animalia" order="Anura" pageId="1" pageNumber="179" phylum="Chordata" rank="species" species="pholeter">
|
||
<emphasis id="B93135272A69AE62CCBAF8D4FF3B9482" italics="true" pageId="1" pageNumber="179">H. pholeter</emphasis>
|
||
</taxonomicName>
|
||
), thigh and tibia also covered by glandular granules (thigh and tibia smooth in
|
||
<taxonomicName id="4C4592B62A69AE62CDA9F829FADB9482" box="[1170,1300,1934,1959]" class="Amphibia" family="Strabomantidae" genus="Holoaden" kingdom="Animalia" order="Anura" pageId="1" pageNumber="179" phylum="Chordata" rank="species" species="pholeter">
|
||
<emphasis id="B93135272A69AE62CDA9F829FADB9482" box="[1170,1300,1934,1959]" italics="true" pageId="1" pageNumber="179">H. pholeter</emphasis>
|
||
</taxonomicName>
|
||
) (
|
||
<figureCitation id="137EF5B02A69AE62CC0BF837FABB9482" box="[1328,1396,1934,1958]" captionStart="FIGURE 1" captionStartId="2.[151,250,1951,1974]" captionTargetBox="[198,1401,151,1915]" captionTargetId="figure@2.[198,1403,149,1930]" captionTargetPageId="2" captionText="FIGURE 1. Dorsal and ventral views of the body: A and B, holotype of Holoaden suarezi sp. nov. (SVL 42.6 mm, MZUSP 131872) C and D, holotype of Holoaden pholeter (SVL 44.6 mm, MNRJ 51475); E and F, lectotype of Holoaden luederwaldti (SVL 40.8 mm, MZUSP 87)." httpUri="https://zenodo.org/record/222738/files/figure.png" pageId="1" pageNumber="179">Fig. 1</figureCitation>
|
||
), a distinct and strait canthus rostralis (canthus rostralis weakly marked and slightly concave in
|
||
<taxonomicName id="4C4592B62A69AE62CD89F80BFAFC94ED" box="[1202,1331,1969,1993]" class="Amphibia" family="Strabomantidae" genus="Holoaden" kingdom="Animalia" order="Anura" pageId="1" pageNumber="179" phylum="Chordata" rank="species" species="pholeter">
|
||
<emphasis id="B93135272A69AE62CD89F80BFAFC94ED" box="[1202,1331,1969,1993]" italics="true" pageId="1" pageNumber="179">H. pholeter</emphasis>
|
||
</taxonomicName>
|
||
) (
|
||
<figureCitation id="137EF5B02A69AE62CC76F808FA5394ED" box="[1357,1436,1969,1993]" captionStart="FIGURE 1" captionStartId="2.[151,250,1951,1974]" captionTargetBox="[198,1401,151,1915]" captionTargetId="figure@2.[198,1403,149,1930]" captionTargetPageId="2" captionText="FIGURE 1. Dorsal and ventral views of the body: A and B, holotype of Holoaden suarezi sp. nov. (SVL 42.6 mm, MZUSP 131872) C and D, holotype of Holoaden pholeter (SVL 44.6 mm, MNRJ 51475); E and F, lectotype of Holoaden luederwaldti (SVL 40.8 mm, MZUSP 87)." httpUri="https://zenodo.org/record/222738/files/figure.png" pageId="1" pageNumber="179">Figs. 1</figureCitation>
|
||
and
|
||
<figureCitation id="137EF5B02A69AE62C9FDF86DFF1994C8" box="[198,214,2004,2028]" captionStart="FIGURE 2" captionStartId="3.[151,250,1976,1999]" captionTargetBox="[172,1414,179,1955]" captionTargetId="figure@3.[172,1416,179,1956]" captionTargetPageId="3" captionText="FIGURE 2. Dorsal and lateral views of the head. A and B, holotype of Holoaden suarezi sp. nov.; C and D, holotype of Holoaden pholeter; E and F, lectotype of Holoaden luederwaldti." httpUri="https://zenodo.org/record/222739/files/figure.png" pageId="1" pageNumber="179">2</figureCitation>
|
||
), and a shorter length of the head that corresponds to 79% of its width (94% in
|
||
<taxonomicName id="4C4592B62A69AE62CD6AF86CFB1F94C8" box="[1105,1232,2004,2028]" class="Amphibia" family="Strabomantidae" genus="Holoaden" kingdom="Animalia" order="Anura" pageId="1" pageNumber="179" phylum="Chordata" rank="species" species="pholeter">
|
||
<emphasis id="B93135272A69AE62CD6AF86CFB1F94C8" box="[1105,1232,2004,2028]" italics="true" pageId="1" pageNumber="179">H. pholeter</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF3AB9BD2A6AAE61C9ACF826FE7094D7" httpUri="https://zenodo.org/record/222738/files/figure.png" pageId="2" pageNumber="180" targetBox="[198,1401,151,1915]" targetPageId="2">
|
||
<paragraph id="8BFAE9352A6AAE61C9ACF826FE7094D7" blockId="2.[151,1436,1951,2035]" pageId="2" pageNumber="180">
|
||
<emphasis id="B93135272A6AAE61C9ACF826FEDB9490" bold="true" box="[151,276,1951,1974]" pageId="2" pageNumber="180">FIGURE 1.</emphasis>
|
||
Dorsal and ventral views of the body: A and B, holotype of
|
||
<taxonomicName id="4C4592B62A6AAE61CABDF826FBF59491" box="[902,1082,1951,1973]" class="Amphibia" family="Strabomantidae" genus="Holoaden" kingdom="Animalia" order="Anura" pageId="2" pageNumber="180" phylum="Chordata" rank="species" species="suarezi" status="sp. nov.">
|
||
<emphasis id="B93135272A6AAE61CABDF826FBF59491" box="[902,1082,1951,1973]" italics="true" pageId="2" pageNumber="180">Holoaden suarezi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A202885C2A6AAE61CD7AF819FB5E9491" box="[1089,1169,1952,1973]" pageId="2" pageNumber="180" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(SVL 42.6 mm, MZUSP 131872) C and D, holotype of
|
||
<taxonomicName id="4C4592B62A6AAE61C8F5F807FD4394F0" box="[462,652,1982,2004]" class="Amphibia" family="Strabomantidae" genus="Holoaden" kingdom="Animalia" order="Anura" pageId="2" pageNumber="180" phylum="Chordata" rank="species" species="pholeter">
|
||
<emphasis id="B93135272A6AAE61C8F5F807FD4394F0" box="[462,652,1982,2004]" italics="true" pageId="2" pageNumber="180">Holoaden pholeter</emphasis>
|
||
</taxonomicName>
|
||
(SVL 44.6 mm, MNRJ 51475); E and F, lectotype of
|
||
<taxonomicName id="4C4592B62A6AAE61CD89F807FA5394F0" box="[1202,1436,1982,2004]" class="Amphibia" family="Strabomantidae" genus="Holoaden" kingdom="Animalia" order="Anura" pageId="2" pageNumber="180" phylum="Chordata" rank="species" species="luederwaldti">
|
||
<emphasis id="B93135272A6AAE61CD89F807FA5394F0" box="[1202,1436,1982,2004]" italics="true" pageId="2" pageNumber="180">Holoaden luederwaldti</emphasis>
|
||
</taxonomicName>
|
||
(SVL 40.8 mm, MZUSP 87).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF3AB9BD2A6BAE60C9ACF801FCE594D5" httpUri="https://zenodo.org/record/222739/files/figure.png" pageId="3" pageNumber="181" targetBox="[172,1414,179,1955]" targetPageId="3">
|
||
<paragraph id="8BFAE9352A6BAE60C9ACF801FCE594D5" blockId="3.[151,1436,1976,2033]" pageId="3" pageNumber="181">
|
||
<emphasis id="B93135272A6BAE60C9ACF801FED994EA" bold="true" box="[151,278,1976,1999]" pageId="3" pageNumber="181">FIGURE 2.</emphasis>
|
||
Dorsal and lateral views of the head. A and B, holotype of
|
||
<taxonomicName id="4C4592B62A6BAE60CAA1F800FB8094EB" box="[922,1103,1977,1999]" class="Amphibia" family="Strabomantidae" genus="Holoaden" kingdom="Animalia" order="Anura" pageId="3" pageNumber="181" phylum="Chordata" rank="species" species="suarezi" status="sp. nov.">
|
||
<emphasis id="B93135272A6BAE60CAA1F800FB8094EB" box="[922,1103,1977,1999]" italics="true" pageId="3" pageNumber="181">Holoaden suarezi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A202885C2A6BAE60CD62F803FB6294EB" box="[1113,1197,1978,1999]" pageId="3" pageNumber="181" rank="species">sp. nov.</taxonomicNameLabel>
|
||
; C and D, holotype of
|
||
<taxonomicName id="4C4592B62A6BAE60C9ACF862FE9A94D5" box="[151,341,2011,2033]" class="Amphibia" family="Strabomantidae" genus="Holoaden" kingdom="Animalia" order="Anura" pageId="3" pageNumber="181" phylum="Chordata" rank="species" species="pholeter">
|
||
<emphasis id="B93135272A6BAE60C9ACF862FE9A94D5" box="[151,341,2011,2033]" italics="true" pageId="3" pageNumber="181">Holoaden pholeter</emphasis>
|
||
</taxonomicName>
|
||
; E and F, lectotype of
|
||
<taxonomicName id="4C4592B62A6BAE60CB00F862FCEA94D5" box="[571,805,2011,2033]" class="Amphibia" family="Strabomantidae" genus="Holoaden" kingdom="Animalia" order="Anura" pageId="3" pageNumber="181" phylum="Chordata" rank="species" species="luederwaldti">
|
||
<emphasis id="B93135272A6BAE60CB00F862FCEA94D5" box="[571,805,2011,2033]" italics="true" pageId="3" pageNumber="181">Holoaden luederwaldti</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF3AB9BD2A6CAE67C9ACFF2EFB5293EB" ID-Table-UUID="DF3AB9BD2A6CAE67C9ACFF2EFB5293EB" httpUri="http://table.plazi.org/id/DF3AB9BD2A6CAE67C9ACFF2EFB5293EB" pageId="4" pageNumber="182" targetBox="[165,1420,240,912]" targetIsTable="true" targetPageId="4">
|
||
<paragraph id="8BFAE9352A6CAE67C9ACFF2EFB5293EB" blockId="4.[151,1436,151,207]" pageId="4" pageNumber="182">
|
||
<emphasis id="B93135272A6CAE67C9ACFF2EFEC4938B" bold="true" box="[151,267,151,175]" pageId="4" pageNumber="182">TABLE 1.</emphasis>
|
||
Measurements (in millimeters) of
|
||
<taxonomicName id="4C4592B62A6CAE67CBBFFF21FC8F938B" box="[644,832,152,175]" class="Amphibia" family="Strabomantidae" genus="Holoaden" kingdom="Animalia" order="Anura" pageId="4" pageNumber="182" phylum="Chordata" rank="species" species="suarezi" status="sp. nov.">
|
||
<emphasis id="B93135272A6CAE67CBBFFF21FC8F938B" box="[644,832,152,175]" italics="true" pageId="4" pageNumber="182">Holoaden suarezi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A202885C2A6CAE67CA72FF21FC50938B" box="[841,927,152,175]" pageId="4" pageNumber="182" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C4592B62A6CAE67CA91FF21FBAF938B" box="[938,1120,152,175]" class="Amphibia" family="Strabomantidae" genus="Holoaden" kingdom="Animalia" order="Anura" pageId="4" pageNumber="182" phylum="Chordata" rank="species" species="bradei">
|
||
<emphasis id="B93135272A6CAE67CA91FF21FBAF938B" box="[938,1120,152,175]" italics="true" pageId="4" pageNumber="182">Holoaden bradei</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4592B62A6CAE67CD54FF21FAA8938B" box="[1135,1383,152,175]" class="Amphibia" family="Strabomantidae" genus="Holoaden" kingdom="Animalia" order="Anura" pageId="4" pageNumber="182" phylum="Chordata" rank="species" species="luederwaldti">
|
||
<emphasis id="B93135272A6CAE67CD54FF21FAA8938B" box="[1135,1383,152,175]" italics="true" pageId="4" pageNumber="182">Holoaden luederwaldti</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C4592B62A6CAE67C9ACFF01FD8493EB" authority="Pombal et al., 2008" authorityName="Pombal et al." authorityYear="2008" box="[151,587,184,207]" class="Amphibia" family="Strabomantidae" genus="Holoaden" kingdom="Animalia" order="Anura" pageId="4" pageNumber="182" phylum="Chordata" rank="species" species="pholeter">
|
||
<emphasis id="B93135272A6CAE67C9ACFF01FEAF93EB" box="[151,352,184,207]" italics="true" pageId="4" pageNumber="182">Holoaden pholeter</emphasis>
|
||
(Pombal
|
||
<emphasis id="B93135272A6CAE67C8FCFF01FE3993EB" box="[455,502,184,207]" italics="true" pageId="4" pageNumber="182">et al</emphasis>
|
||
., 2008)
|
||
</taxonomicName>
|
||
. * Measurements include juveniles that were not sexed.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BFAE9352A6CAE67C885FF48FA4390AB" pageId="4" pageNumber="182">
|
||
<table id="F9451B952A6C519CC99EFF49FA4390B4" box="[165,1420,240,912]" gridcols="5" gridrows="12" pageId="4" pageNumber="182">
|
||
<tr id="3575EB772A6C519CC99EFF49FA439222" box="[165,1420,240,262]" gridrow="0" pageId="4" pageNumber="182" rowspan-0="1">
|
||
<th id="76A4820B2A6C519CC89AFF49FD5F9222" box="[417,656,240,262]" gridcol="1" gridrow="0" pageId="4" pageNumber="182">
|
||
<taxonomicName id="4C4592B62A6CAE67C885FF48FDE99222" box="[446,550,241,262]" class="Amphibia" family="Strabomantidae" genus="Holoaden" kingdom="Animalia" order="Anura" pageId="4" pageNumber="182" phylum="Chordata" rank="species" species="suarezi">
|
||
<emphasis id="B93135272A6CAE67C885FF48FDE99222" box="[446,550,241,262]" italics="true" pageId="4" pageNumber="182">H. suarezi</emphasis>
|
||
</taxonomicName>
|
||
sp nov
|
||
</th>
|
||
<th id="76A4820B2A6C519CCB8BFF49FC6F9222" box="[688,928,240,262]" gridcol="2" gridrow="0" pageId="4" pageNumber="182">
|
||
<taxonomicName id="4C4592B62A6CAE67CBC2FF48FC969222" box="[761,857,240,262]" class="Amphibia" family="Strabomantidae" genus="Holoaden" kingdom="Animalia" order="Anura" pageId="4" pageNumber="182" phylum="Chordata" rank="species" species="bradei">
|
||
<emphasis id="B93135272A6CAE67CBC2FF48FC969222" box="[761,857,240,262]" italics="true" pageId="4" pageNumber="182">H. bradei</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="76A4820B2A6C519CCAFEFF49FB7A9222" box="[965,1205,240,262]" gridcol="3" gridrow="0" pageId="4" pageNumber="182">
|
||
<taxonomicName id="4C4592B62A6CAE67CAD6FF48FB439222" box="[1005,1164,240,262]" class="Amphibia" family="Strabomantidae" genus="Holoaden" kingdom="Animalia" order="Anura" pageId="4" pageNumber="182" phylum="Chordata" rank="species" species="luederwaldti">
|
||
<emphasis id="B93135272A6CAE67CAD6FF48FB439222" box="[1005,1164,240,262]" italics="true" pageId="4" pageNumber="182">H. luederwaldti</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="76A4820B2A6C519CCDECFF49FA439222" box="[1239,1420,240,262]" gridcol="4" gridrow="0" pageId="4" pageNumber="182">
|
||
<taxonomicName id="4C4592B62A6CAE67CDC3FF48FAA49222" box="[1272,1387,240,262]" class="Amphibia" family="Strabomantidae" genus="Holoaden" kingdom="Animalia" order="Anura" pageId="4" pageNumber="182" phylum="Chordata" rank="species" species="pholeter">
|
||
<emphasis id="B93135272A6CAE67CDC3FF48FAA49222" box="[1272,1387,240,262]" italics="true" pageId="4" pageNumber="182">H. pholeter</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
</tr>
|
||
<tr id="3575EB772A6C519CC99EFEA5FA43925B" box="[165,1420,284,383]" gridrow="1" pageId="4" pageNumber="182">
|
||
<th id="76A4820B2A6C519CC99EFEA5FEB6925B" box="[165,377,284,383]" gridcol="0" gridrow="1" pageId="4" pageNumber="182">Snout-vent length</th>
|
||
<td id="76A4820B2A6C519CC89AFEA5FD5F925B" box="[417,656,284,383]" gridcol="1" gridrow="1" pageId="4" pageNumber="182">Male Female (n=2) (n=2) 37.2; 38.5 42.6; 44.2</td>
|
||
<td id="76A4820B2A6C519CCB8BFEA5FC6F925B" box="[688,928,284,383]" gridcol="2" gridrow="1" pageId="4" pageNumber="182">Male Female (n=134)* (n=18) 10.1–28.4 26.4–36.8</td>
|
||
<td id="76A4820B2A6C519CCAFEFEA5FB7A925B" box="[965,1205,284,383]" gridcol="3" gridrow="1" pageId="4" pageNumber="182">Male Female (n=26) (n=8) 27.1–39.9 39.3–42.8</td>
|
||
<td id="76A4820B2A6C519CCDECFEA5FA43925B" box="[1239,1420,284,383]" gridcol="4" gridrow="1" pageId="4" pageNumber="182">Mal Female e (1) (n=2) 41.6 44.6; 47.7</td>
|
||
</tr>
|
||
<tr id="3575EB772A6C519CC99EFE2CFA43928F" box="[165,1420,405,427]" gridrow="2" pageId="4" pageNumber="182">
|
||
<th id="76A4820B2A6C519CC99EFE2CFEB6928F" box="[165,377,405,427]" gridcol="0" gridrow="2" pageId="4" pageNumber="182">Head width</th>
|
||
<td id="76A4820B2A6C519CC89AFE2CFD5F928F" box="[417,656,405,427]" gridcol="1" gridrow="2" pageId="4" pageNumber="182">14.6; 15.9 16.5; 17.1</td>
|
||
<td id="76A4820B2A6C519CCB8BFE2CFC6F928F" box="[688,928,405,427]" gridcol="2" gridrow="2" pageId="4" pageNumber="182">3.7–10.3 8.5–12.2</td>
|
||
<td id="76A4820B2A6C519CCAFEFE2CFB7A928F" box="[965,1205,405,427]" gridcol="3" gridrow="2" pageId="4" pageNumber="182">10.1–16.0 13.7–16.3</td>
|
||
<td id="76A4820B2A6C519CCDECFE2CFA43928F" box="[1239,1420,405,427]" gridcol="4" gridrow="2" pageId="4" pageNumber="182">16.4 16.3; 19.3</td>
|
||
</tr>
|
||
<tr id="3575EB772A6C519CC99EFE78FA4392F3" box="[165,1420,449,471]" gridrow="3" pageId="4" pageNumber="182">
|
||
<th id="76A4820B2A6C519CC99EFE78FEB692F3" box="[165,377,449,471]" gridcol="0" gridrow="3" pageId="4" pageNumber="182">Head length</th>
|
||
<td id="76A4820B2A6C519CC89AFE78FD5F92F3" box="[417,656,449,471]" gridcol="1" gridrow="3" pageId="4" pageNumber="182">13.1; 13.6 13.0; 13.1</td>
|
||
<td id="76A4820B2A6C519CCB8BFE78FC6F92F3" box="[688,928,449,471]" gridcol="2" gridrow="3" pageId="4" pageNumber="182">2.7–6.9 4.6–8.3</td>
|
||
<td id="76A4820B2A6C519CCAFEFE78FB7A92F3" box="[965,1205,449,471]" gridcol="3" gridrow="3" pageId="4" pageNumber="182">6.6–11.0 8.3–11.3</td>
|
||
<td id="76A4820B2A6C519CCDECFE78FA4392F3" box="[1239,1420,449,471]" gridcol="4" gridrow="3" pageId="4" pageNumber="182">14.7 15.2; 18.4</td>
|
||
</tr>
|
||
<tr id="3575EB772A6C519CC99EFE54FA439127" box="[165,1420,493,515]" gridrow="4" pageId="4" pageNumber="182">
|
||
<th id="76A4820B2A6C519CC99EFE54FEB69127" box="[165,377,493,515]" gridcol="0" gridrow="4" pageId="4" pageNumber="182">Eye diameter</th>
|
||
<td id="76A4820B2A6C519CC89AFE54FD5F9127" box="[417,656,493,515]" gridcol="1" gridrow="4" pageId="4" pageNumber="182">4,6; 4.8 5.1; 5.6</td>
|
||
<td id="76A4820B2A6C519CCB8BFE54FC6F9127" box="[688,928,493,515]" gridcol="2" gridrow="4" pageId="4" pageNumber="182">1.2–3.2 2.0–3.5</td>
|
||
<td id="76A4820B2A6C519CCAFEFE54FB7A9127" box="[965,1205,493,515]" gridcol="3" gridrow="4" pageId="4" pageNumber="182">4.2- 6.1 5.1–6.2</td>
|
||
<td id="76A4820B2A6C519CCDECFE54FA439127" box="[1239,1420,493,515]" gridcol="4" gridrow="4" pageId="4" pageNumber="182">4.7 5.4; 6.6</td>
|
||
</tr>
|
||
<tr id="3575EB772A6C519CC99EFDA0FA43910B" box="[165,1420,537,559]" gridrow="5" pageId="4" pageNumber="182">
|
||
<th id="76A4820B2A6C519CC99EFDA0FEB6910B" box="[165,377,537,559]" gridcol="0" gridrow="5" pageId="4" pageNumber="182">Eye-nostril distance</th>
|
||
<td id="76A4820B2A6C519CC89AFDA0FD5F910B" box="[417,656,537,559]" gridcol="1" gridrow="5" pageId="4" pageNumber="182">3.5; 3.5 3.9; 4.2</td>
|
||
<td id="76A4820B2A6C519CCB8BFDA0FC6F910B" box="[688,928,537,559]" gridcol="2" gridrow="5" pageId="4" pageNumber="182">0.7–2.0 1.2–2.6</td>
|
||
<td id="76A4820B2A6C519CCAFEFDA0FB7A910B" box="[965,1205,537,559]" gridcol="3" gridrow="5" pageId="4" pageNumber="182">3.0–4.4 3.1–4.5</td>
|
||
<td id="76A4820B2A6C519CCDECFDA0FA43910B" box="[1239,1420,537,559]" gridcol="4" gridrow="5" pageId="4" pageNumber="182">3.7 4.6; 4.4</td>
|
||
</tr>
|
||
<tr id="3575EB772A6C519CC99EFDFCFA43917F" box="[165,1420,581,603]" gridrow="6" pageId="4" pageNumber="182">
|
||
<th id="76A4820B2A6C519CC99EFDFCFEB6917F" box="[165,377,581,603]" gridcol="0" gridrow="6" pageId="4" pageNumber="182">Inter-orbital distance</th>
|
||
<td id="76A4820B2A6C519CC89AFDFCFD5F917F" box="[417,656,581,603]" gridcol="1" gridrow="6" pageId="4" pageNumber="182">6.0; 6.4 6.7; 6.9;</td>
|
||
<td id="76A4820B2A6C519CCB8BFDFCFC6F917F" box="[688,928,581,603]" gridcol="2" gridrow="6" pageId="4" pageNumber="182">1.3–5.1 4.4–5.5</td>
|
||
<td id="76A4820B2A6C519CCAFEFDFCFB7A917F" box="[965,1205,581,603]" gridcol="3" gridrow="6" pageId="4" pageNumber="182">7.1–11.9 7.2–10.5</td>
|
||
<td id="76A4820B2A6C519CCDECFDFCFA43917F" box="[1239,1420,581,603]" gridcol="4" gridrow="6" pageId="4" pageNumber="182">4.2 5.9; 6.2</td>
|
||
</tr>
|
||
<tr id="3575EB772A6C519CC99EFDC8FA4391A3" box="[165,1420,625,647]" gridrow="7" pageId="4" pageNumber="182">
|
||
<th id="76A4820B2A6C519CC99EFDC8FEB691A3" box="[165,377,625,647]" gridcol="0" gridrow="7" pageId="4" pageNumber="182">Internasal distance</th>
|
||
<td id="76A4820B2A6C519CC89AFDC8FD5F91A3" box="[417,656,625,647]" gridcol="1" gridrow="7" pageId="4" pageNumber="182">2.9; 3.0 3.1; 3.3</td>
|
||
<td id="76A4820B2A6C519CCB8BFDC8FC6F91A3" box="[688,928,625,647]" gridcol="2" gridrow="7" pageId="4" pageNumber="182">1.0–2.4 2.1–2.8</td>
|
||
<td id="76A4820B2A6C519CCAFEFDC8FB7A91A3" box="[965,1205,625,647]" gridcol="3" gridrow="7" pageId="4" pageNumber="182">3.6–5.1 4.4–5.2</td>
|
||
<td id="76A4820B2A6C519CCDECFDC8FA4391A3" box="[1239,1420,625,647]" gridcol="4" gridrow="7" pageId="4" pageNumber="182">3.1 3.6; 3.4</td>
|
||
</tr>
|
||
<tr id="3575EB772A6C519CC99EFD24FA439197" box="[165,1420,669,691]" gridrow="8" pageId="4" pageNumber="182">
|
||
<th id="76A4820B2A6C519CC99EFD24FEB69197" box="[165,377,669,691]" gridcol="0" gridrow="8" pageId="4" pageNumber="182">Arm length</th>
|
||
<td id="76A4820B2A6C519CC89AFD24FD5F9197" box="[417,656,669,691]" gridcol="1" gridrow="8" pageId="4" pageNumber="182">8.1; 9.5 9.0; 9.4</td>
|
||
<td id="76A4820B2A6C519CCB8BFD24FC6F9197" box="[688,928,669,691]" gridcol="2" gridrow="8" pageId="4" pageNumber="182">1.8–5.6 4.1–6.6</td>
|
||
<td id="76A4820B2A6C519CCAFEFD24FB7A9197" box="[965,1205,669,691]" gridcol="3" gridrow="8" pageId="4" pageNumber="182">5.7–8.4 7.1–8.8</td>
|
||
<td id="76A4820B2A6C519CCDECFD24FA439197" box="[1239,1420,669,691]" gridcol="4" gridrow="8" pageId="4" pageNumber="182">– –</td>
|
||
</tr>
|
||
<tr id="3575EB772A6C519CC99EFD73FA43902F" box="[165,1420,714,779]" gridrow="9" pageId="4" pageNumber="182">
|
||
<th id="76A4820B2A6C519CC99EFD73FEB6902F" box="[165,377,714,779]" gridcol="0" gridrow="9" pageId="4" pageNumber="182">Forearm length Femur length</th>
|
||
<td id="76A4820B2A6C519CC89AFD73FD5F902F" box="[417,656,714,779]" gridcol="1" gridrow="9" pageId="4" pageNumber="182">11.2; 12.2 13.0; 13.1 17.8; 18.0 19.7; 19.9</td>
|
||
<td id="76A4820B2A6C519CCB8BFD73FC6F902F" box="[688,928,714,779]" gridcol="2" gridrow="9" pageId="4" pageNumber="182">2.2–6.7 5.3–7.5 4.3–11.9 5.9–12.1</td>
|
||
<td id="76A4820B2A6C519CCAFEFD73FB7A902F" box="[965,1205,714,779]" gridcol="3" gridrow="9" pageId="4" pageNumber="182">7.9–11.6 9.0–12.1 11.7–17.5 15.5–18.2</td>
|
||
<td id="76A4820B2A6C519CCDECFD73FA43902F" box="[1239,1420,714,779]" gridcol="4" gridrow="9" pageId="4" pageNumber="182">11.9 12.9; 12.8 17.7 19.9; 20.6</td>
|
||
</tr>
|
||
<tr id="3575EB772A6C519CC99EFC9BFA439047" box="[165,1420,802,867]" gridrow="10" pageId="4" pageNumber="182">
|
||
<th id="76A4820B2A6C519CC99EFC9BFEB69047" box="[165,377,802,867]" gridcol="0" gridrow="10" pageId="4" pageNumber="182">Tibia length Foot length</th>
|
||
<td id="76A4820B2A6C519CC89AFC9BFD5F9047" box="[417,656,802,867]" gridcol="1" gridrow="10" pageId="4" pageNumber="182">16.9; 17.7 18.3; 18.7 16.6; 17.5 17.9; 18.2</td>
|
||
<td id="76A4820B2A6C519CCB8BFC9BFC6F9047" box="[688,928,802,867]" gridcol="2" gridrow="10" pageId="4" pageNumber="182">3.6–10.9 6.0–12.1 3.7–12.7 10.9–14.5</td>
|
||
<td id="76A4820B2A6C519CCAFEFC9BFB7A9047" box="[965,1205,802,867]" gridcol="3" gridrow="10" pageId="4" pageNumber="182">8.0–17.0 10.2–17.2 10.6–16.9 14.9–17.5</td>
|
||
<td id="76A4820B2A6C519CCDECFC9BFA439047" box="[1239,1420,802,867]" gridcol="4" gridrow="10" pageId="4" pageNumber="182">18.0 19.0; 19.3 17.7 17.7; 19.5</td>
|
||
</tr>
|
||
<tr id="3575EB772A6C519CC99EFCC3FA4390B4" box="[165,1420,890,912]" gridrow="11" pageId="4" pageNumber="182">
|
||
<th id="76A4820B2A6C519CC99EFCC3FEB690B4" box="[165,377,890,912]" gridcol="0" gridrow="11" pageId="4" pageNumber="182">Hand length</th>
|
||
<td id="76A4820B2A6C519CC89AFCC3FD5F90B4" box="[417,656,890,912]" gridcol="1" gridrow="11" pageId="4" pageNumber="182">9.2; 10.4 10.7; 10.9</td>
|
||
<td id="76A4820B2A6C519CCB8BFCC3FC6F90B4" box="[688,928,890,912]" gridcol="2" gridrow="11" pageId="4" pageNumber="182">2.1–7.2 4.7–7.8</td>
|
||
<td id="76A4820B2A6C519CCAFEFCC3FB7A90B4" box="[965,1205,890,912]" gridcol="3" gridrow="11" pageId="4" pageNumber="182">6.6–11.0 9.1–11.0</td>
|
||
<td id="76A4820B2A6C519CCDECFCC3FA4390B4" box="[1239,1420,890,912]" gridcol="4" gridrow="11" pageId="4" pageNumber="182">11.0 11.2; 12.0</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<subSubSection id="C35FBABE2A6CAE67C9FCFC68FD1C94FD" pageId="4" pageNumber="182" type="description">
|
||
<paragraph id="8BFAE9352A6CAE67C9FCFC68FE489569" blockId="4.[151,1436,977,2009]" pageId="4" pageNumber="182">
|
||
<emphasis id="B93135272A6CAE67C9FCFC68FDD490CE" bold="true" box="[199,539,977,1002]" pageId="4" pageNumber="182">
|
||
Description of the
|
||
<typeStatus id="54FE57972A6CAE67C88EFC6BFDD490CE" box="[437,539,978,1002]" pageId="4" pageNumber="182" type="holotype">holotype</typeStatus>
|
||
</emphasis>
|
||
. Female with oocytes in the initial stage of development; The oocytes are unpigmented; Body robust; head large, wider than long (
|
||
<figureCitation id="137EF5B02A6CAE67CA27FC4CFCA59729" box="[796,874,1013,1037]" captionStart="FIGURE 1" captionStartId="2.[151,250,1951,1974]" captionTargetBox="[198,1401,151,1915]" captionTargetId="figure@2.[198,1403,149,1930]" captionTargetPageId="2" captionText="FIGURE 1. Dorsal and ventral views of the body: A and B, holotype of Holoaden suarezi sp. nov. (SVL 42.6 mm, MZUSP 131872) C and D, holotype of Holoaden pholeter (SVL 44.6 mm, MNRJ 51475); E and F, lectotype of Holoaden luederwaldti (SVL 40.8 mm, MZUSP 87)." httpUri="https://zenodo.org/record/222738/files/figure.png" pageId="4" pageNumber="182">Figs. 1</figureCitation>
|
||
and
|
||
<figureCitation id="137EF5B02A6CAE67CA98FC4CFC7C9729" box="[931,947,1013,1037]" captionStart="FIGURE 2" captionStartId="3.[151,250,1976,1999]" captionTargetBox="[172,1414,179,1955]" captionTargetId="figure@3.[172,1416,179,1956]" captionTargetPageId="3" captionText="FIGURE 2. Dorsal and lateral views of the head. A and B, holotype of Holoaden suarezi sp. nov.; C and D, holotype of Holoaden pholeter; E and F, lectotype of Holoaden luederwaldti." httpUri="https://zenodo.org/record/222739/files/figure.png" pageId="4" pageNumber="182">2</figureCitation>
|
||
), head length 30% of SVL and head width 38% of SVL; head length corresponds to 79% of head width; snout nearly rounded in dorsal view and almost rounded in lateral view (
|
||
<figureCitation id="137EF5B02A6CAE67C890FB84FE219771" box="[427,494,1085,1109]" captionStart="FIGURE 2" captionStartId="3.[151,250,1976,1999]" captionTargetBox="[172,1414,179,1955]" captionTargetId="figure@3.[172,1416,179,1956]" captionTargetPageId="3" captionText="FIGURE 2. Dorsal and lateral views of the head. A and B, holotype of Holoaden suarezi sp. nov.; C and D, holotype of Holoaden pholeter; E and F, lectotype of Holoaden luederwaldti." httpUri="https://zenodo.org/record/222739/files/figure.png" pageId="4" pageNumber="182">Fig. 2</figureCitation>
|
||
A, B); nostril elliptical and slightly protuberant, directed laterally; canthus rostralis distinct, straight; loreal region concave; eye large, protruding, lateral and slightly forward, eye diameter 30% of head length; pupil round; tympanic not visible externally or poorly visible; a glandular supratympanic fold extending from the corner of the eye to the insertion of the forearm (
|
||
<figureCitation id="137EF5B02A6CAE67CA9FFB10FC2897E5" box="[932,999,1193,1217]" captionStart="FIGURE 2" captionStartId="3.[151,250,1976,1999]" captionTargetBox="[172,1414,179,1955]" captionTargetId="figure@3.[172,1416,179,1956]" captionTargetPageId="3" captionText="FIGURE 2. Dorsal and lateral views of the head. A and B, holotype of Holoaden suarezi sp. nov.; C and D, holotype of Holoaden pholeter; E and F, lectotype of Holoaden luederwaldti." httpUri="https://zenodo.org/record/222739/files/figure.png" pageId="4" pageNumber="182">Fig. 2</figureCitation>
|
||
A); tongue large; two short transverse series of vomerine teeth behind and between choanae; choanae elliptical; numerous teeth on maxilla and premaxillae; arms very slender; forearms slender; fingers long and slender, forearm length 30% of SVL; finger lengths II≤IV<I<III; hand length 24% of SVL; finger tips rounded; fingers not webbed nor fringed; subarticular tubercles single, rounded; inner metacarpal tubercle large and nearly elliptical; outer metacarpal tubercle ovoid; few supernumerary tubercles on the palmar region (
|
||
<figureCitation id="137EF5B02A6CAE67CBE3FAE4FCD59651" box="[728,794,1373,1397]" captionStart="FIGURE 3" captionStartId="5.[151,250,1249,1272]" captionTargetBox="[329,1257,151,1228]" captionTargetId="figure@5.[329,1257,137,1228]" captionTargetPageId="5" captionText="FIGURE 3. Ventral view of hands and fingers. (A) holotype of Holoaden suarezi sp. nov.; (B) holotype of Holoaden pholeter; (C) lectotype and (D) topotype of Holoaden luederwaldti." httpUri="https://zenodo.org/record/222740/files/figure.png" pageId="4" pageNumber="182">Fig. 3</figureCitation>
|
||
A); legs long and slender, thigh and tibia length 46% and 44% of SVL, respectively; toes very long and slender, toe lengths I<II<V<III<IV; toe tips rounded, with poorly expanded disc; toes not webbed nor fringed; subarticular tubercles single, elliptical or round; inner metatarsal tubercle elliptical; outer metatarsal tubercle elliptical, small; outer tarsal and sole of foot texture smooth (
|
||
<figureCitation id="137EF5B02A6CAE67CC15FA70FABF96C5" box="[1326,1392,1481,1505]" captionStart="FIGURE 4" captionStartId="6.[151,250,1964,1987]" captionTargetBox="[332,1252,193,1942]" captionTargetId="figure@6.[332,1256,193,1944]" captionTargetPageId="6" captionText="FIGURE 4. Ventral view of feet and toes. (A) holotype of Holoaden suarezi sp. nov.; (B) lectotype and (D) topotype of Holoaden luederwaldti; (C) holotype of Holoaden pholeter." httpUri="https://zenodo.org/record/222741/files/figure.png" pageId="4" pageNumber="182">Fig. 4</figureCitation>
|
||
A); foot length 41.6% of SVL; thigh and tibia with glandular texture; dorsal surface of head and body with glandular texture; the glandular granulation extends to the internasal region (
|
||
<figureCitation id="137EF5B02A6CAE67CA9FF9A8FC25950D" box="[932,1002,1553,1577]" captionStart="FIGURE 2" captionStartId="3.[151,250,1976,1999]" captionTargetBox="[172,1414,179,1955]" captionTargetId="figure@3.[172,1416,179,1956]" captionTargetPageId="3" captionText="FIGURE 2. Dorsal and lateral views of the head. A and B, holotype of Holoaden suarezi sp. nov.; C and D, holotype of Holoaden pholeter; E and F, lectotype of Holoaden luederwaldti." httpUri="https://zenodo.org/record/222739/files/figure.png" pageId="4" pageNumber="182">Fig. 2</figureCitation>
|
||
B); arm and forearm smooth; venter smooth (
|
||
<figureCitation id="137EF5B02A6CAE67C9C2F98CFEF49569" box="[249,315,1589,1613]" captionStart="FIGURE 1" captionStartId="2.[151,250,1951,1974]" captionTargetBox="[198,1401,151,1915]" captionTargetId="figure@2.[198,1403,149,1930]" captionTargetPageId="2" captionText="FIGURE 1. Dorsal and ventral views of the body: A and B, holotype of Holoaden suarezi sp. nov. (SVL 42.6 mm, MZUSP 131872) C and D, holotype of Holoaden pholeter (SVL 44.6 mm, MNRJ 51475); E and F, lectotype of Holoaden luederwaldti (SVL 40.8 mm, MZUSP 87)." httpUri="https://zenodo.org/record/222738/files/figure.png" pageId="4" pageNumber="182">Fig. 1</figureCitation>
|
||
A, B).
|
||
</paragraph>
|
||
<paragraph id="8BFAE9352A6CAE67C9FCF9E0FE29959D" blockId="4.[151,1436,977,2009]" pageId="4" pageNumber="182">
|
||
<emphasis id="B93135272A6CAE67C9FCF9E0FD499556" bold="true" box="[199,646,1625,1650]" pageId="4" pageNumber="182">
|
||
Color of the
|
||
<typeStatus id="54FE57972A6CAE67C858F9E3FE059556" box="[355,458,1626,1650]" pageId="4" pageNumber="182" type="holotype">holotype</typeStatus>
|
||
in preservative
|
||
</emphasis>
|
||
. Dorsal surfaces of the head, dorsum, limbs, and flanks dark brown; lateral and ventral surfaces clear brown; palmar and plantar tubercles and tips of digits light brown, similar to ventral coloration (
|
||
<figureCitation id="137EF5B02A6CAE67C857F918FE16959D" box="[364,473,1697,1721]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="2.[151,250,1951,1974]" captionStartId-1="3.[151,250,1976,1999]" captionStartId-2="5.[151,250,1249,1272]" captionStartId-3="6.[151,250,1964,1987]" captionTargetBox-0="[198,1401,151,1915]" captionTargetBox-1="[172,1414,179,1955]" captionTargetBox-2="[329,1257,151,1228]" captionTargetBox-3="[332,1252,193,1942]" captionTargetId-0="figure@2.[198,1403,149,1930]" captionTargetId-1="figure@3.[172,1416,179,1956]" captionTargetId-2="figure@5.[329,1257,137,1228]" captionTargetId-3="figure@6.[332,1256,193,1944]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="5" captionTargetPageId-3="6" captionText-0="FIGURE 1. Dorsal and ventral views of the body: A and B, holotype of Holoaden suarezi sp. nov. (SVL 42.6 mm, MZUSP 131872) C and D, holotype of Holoaden pholeter (SVL 44.6 mm, MNRJ 51475); E and F, lectotype of Holoaden luederwaldti (SVL 40.8 mm, MZUSP 87)." captionText-1="FIGURE 2. Dorsal and lateral views of the head. A and B, holotype of Holoaden suarezi sp. nov.; C and D, holotype of Holoaden pholeter; E and F, lectotype of Holoaden luederwaldti." captionText-2="FIGURE 3. Ventral view of hands and fingers. (A) holotype of Holoaden suarezi sp. nov.; (B) holotype of Holoaden pholeter; (C) lectotype and (D) topotype of Holoaden luederwaldti." captionText-3="FIGURE 4. Ventral view of feet and toes. (A) holotype of Holoaden suarezi sp. nov.; (B) lectotype and (D) topotype of Holoaden luederwaldti; (C) holotype of Holoaden pholeter." httpUri-0="https://zenodo.org/record/222738/files/figure.png" httpUri-1="https://zenodo.org/record/222739/files/figure.png" httpUri-2="https://zenodo.org/record/222740/files/figure.png" httpUri-3="https://zenodo.org/record/222741/files/figure.png" pageId="4" pageNumber="182">Figs. 1–4</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BFAE9352A6CAE67C9FCF97FFD009425" blockId="4.[151,1436,977,2009]" pageId="4" pageNumber="182">
|
||
<emphasis id="B93135272A6CAE67C9FCF97FFDE095FA" bold="true" box="[199,559,1733,1758]" pageId="4" pageNumber="182">
|
||
Measurements of the
|
||
<typeStatus id="54FE57972A6CAE67C8F2F97FFDE095FA" box="[457,559,1734,1758]" pageId="4" pageNumber="182" type="holotype">holotype</typeStatus>
|
||
</emphasis>
|
||
. SVL 42.6; HL 13.1; HW 16.5; ED 5.1; IOD 6.7;
|
||
<collectingCountry id="F352A9A52A6CAE67CD5DF97FFB6F95F9" box="[1126,1184,1734,1757]" name="India" pageId="4" pageNumber="182">END</collectingCountry>
|
||
3.9;
|
||
<collectingCountry id="F352A9A52A6CAE67CDE2F97FFAC495F9" box="[1241,1291,1734,1757]" name="India" pageId="4" pageNumber="182">IND</collectingCountry>
|
||
3.1; AL 9.4; FAL 13.0; HAL 10.7; FL 19.7; TL18.7; FOL 18.2.
|
||
</paragraph>
|
||
<paragraph id="8BFAE9352A6CAE67C9FCF8B4FC549449" blockId="4.[151,1436,977,2009]" pageId="4" pageNumber="182">
|
||
<emphasis id="B93135272A6CAE67C9FCF8B4FE959402" bold="true" box="[199,346,1805,1830]" pageId="4" pageNumber="182">Color in life</emphasis>
|
||
. Dorsal surfaces of the head, body, and limbs deeply dark brown. The glandular portion of the lateral surface dark brown; flanks and ventral surfaces of the body and limbs slightly lighter than dorsum; palmar and plantar tubercles and tips of digits light gray; eyes black (
|
||
<figureCitation id="137EF5B02A6CAE67CA72F8ECFC429449" box="[841,909,1877,1901]" captionStart="FIGURE 5" captionStartId="7.[151,250,1291,1314]" captionTargetBox="[151,1435,193,1269]" captionTargetId="figure@7.[151,1436,193,1270]" captionTargetPageId="7" captionText="FIGURE 5. Living specimen of Holoaden suarezi sp. nov. (MZUSP 138348) (A) lateral, (B) ventral, (C) frontal and (D) dorsal views. Estação Ecológica de Boracéia, State of São Paulo, Brazil. Photo: Martha C. Lange." httpUri="https://zenodo.org/record/222742/files/figure.png" pageId="4" pageNumber="182">Fig. 5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BFAE9352A6CAE67C9FCF8C0FD1C94FD" blockId="4.[151,1436,977,2009]" pageId="4" pageNumber="182">
|
||
<emphasis id="B93135272A6CAE67C9FCF8C0FEFA94B6" bold="true" box="[199,309,1913,1938]" pageId="4" pageNumber="182">Variation</emphasis>
|
||
. Males were smaller and with more robust forearms than females (
|
||
<tableCitation id="C6C7DC8E2A6CAE67CD02F8C0FB5494B5" box="[1081,1179,1913,1937]" captionStart="TABLE 1" captionStartId="4.[151,239,151,175]" captionTargetBox="[165,1420,240,912]" captionTargetPageId="4" captionText="TABLE 1. Measurements (in millimeters) of Holoaden suarezi sp. nov., Holoaden bradei, Holoaden luederwaldti, and Holoaden pholeter (Pombal et al., 2008). * Measurements include juveniles that were not sexed." httpUri="http://table.plazi.org/id/DF3AB9BD2A6CAE67C9ACFF2EFB5293EB" pageId="4" pageNumber="182" tableUuid="DF3AB9BD2A6CAE67C9ACFF2EFB5293EB">Tables 1</tableCitation>
|
||
and
|
||
<tableCitation id="C6C7DC8E2A6CAE67CDEEF8C0FB2994B5" box="[1237,1254,1913,1937]" captionStart="TABLE 2" captionStartId="5.[151,239,1340,1364]" captionTargetBox="[165,1370,1395,2015]" captionTargetPageId="5" captionText="TABLE 2. Measurements (in millimeters) of paratypes of Holoaden suarezi sp. nov." httpUri="http://table.plazi.org/id/DF3AB9BD2A6DAE66C9ACFA85FBD39677" pageId="4" pageNumber="182" tableUuid="DF3AB9BD2A6DAE66C9ACFA85FBD39677">2</tableCitation>
|
||
). Otherwise, no conspicuous qualitative differences were found between the available specimens, which were very similar to the
|
||
<typeStatus id="54FE57972A6CAE67C9ACF878FF3694FD" box="[151,249,1985,2009]" pageId="4" pageNumber="182" type="holotype">holotype</typeStatus>
|
||
in external morphology and color pattern.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF3AB9BD2A6DAE66C9ACFB58FD2E9632" httpUri="https://zenodo.org/record/222740/files/figure.png" pageId="5" pageNumber="183" targetBox="[329,1257,151,1228]" targetPageId="5">
|
||
<paragraph id="8BFAE9352A6DAE66C9ACFB58FD2E9632" blockId="5.[151,1436,1249,1303]" pageId="5" pageNumber="183">
|
||
<emphasis id="B93135272A6DAE66C9ACFB58FEDD97D3" bold="true" box="[151,274,1249,1272]" pageId="5" pageNumber="183">FIGURE 3.</emphasis>
|
||
Ventral view of hands and fingers. (A) holotype of
|
||
<taxonomicName id="4C4592B62A6DAE66CA24FB5BFC1D97DC" box="[799,978,1250,1272]" class="Amphibia" family="Strabomantidae" genus="Holoaden" kingdom="Animalia" order="Anura" pageId="5" pageNumber="183" phylum="Chordata" rank="species" species="suarezi" status="sp. nov.">
|
||
<emphasis id="B93135272A6DAE66CA24FB5BFC1D97DC" box="[799,978,1250,1272]" italics="true" pageId="5" pageNumber="183">Holoaden suarezi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A202885C2A6DAE66CAE3FB5AFBE597DC" box="[984,1066,1251,1272]" pageId="5" pageNumber="183" rank="species">sp. nov.</taxonomicNameLabel>
|
||
; (B) holotype of
|
||
<taxonomicName id="4C4592B62A6DAE66CDEDFB5BFA5A97DC" box="[1238,1429,1250,1272]" class="Amphibia" family="Strabomantidae" genus="Holoaden" kingdom="Animalia" order="Anura" pageId="5" pageNumber="183" phylum="Chordata" rank="species" species="pholeter">
|
||
<emphasis id="B93135272A6DAE66CDEDFB5BFA5A97DC" box="[1238,1429,1250,1272]" italics="true" pageId="5" pageNumber="183">Holoaden pholeter</emphasis>
|
||
</taxonomicName>
|
||
; (C) lectotype and (D) topotype of
|
||
<taxonomicName id="4C4592B62A6DAE66C8CAFAB9FD149632" box="[497,731,1280,1302]" class="Amphibia" family="Strabomantidae" genus="Holoaden" kingdom="Animalia" order="Anura" pageId="5" pageNumber="183" phylum="Chordata" rank="species" species="luederwaldti">
|
||
<emphasis id="B93135272A6DAE66C8CAFAB9FD149632" box="[497,731,1280,1302]" italics="true" pageId="5" pageNumber="183">Holoaden luederwaldti</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF3AB9BD2A6DAE66C9ACFA85FBD39677" ID-Table-UUID="DF3AB9BD2A6DAE66C9ACFA85FBD39677" box="[151,1052,1340,1364]" httpUri="http://table.plazi.org/id/DF3AB9BD2A6DAE66C9ACFA85FBD39677" pageId="5" pageNumber="183" targetBox="[165,1370,1395,2015]" targetIsTable="true" targetPageId="5">
|
||
<paragraph id="8BFAE9352A6DAE66C9ACFA85FBD39677" blockId="5.[151,1052,1340,1364]" box="[151,1052,1340,1364]" pageId="5" pageNumber="183">
|
||
<emphasis id="B93135272A6DAE66C9ACFA85FEC59670" bold="true" box="[151,266,1340,1364]" pageId="5" pageNumber="183">TABLE 2.</emphasis>
|
||
Measurements (in millimeters) of paratypes of
|
||
<taxonomicName id="4C4592B62A6DAE66CA3DFA85FC0D9677" box="[774,962,1340,1363]" class="Amphibia" family="Strabomantidae" genus="Holoaden" kingdom="Animalia" order="Anura" pageId="5" pageNumber="183" phylum="Chordata" rank="species" species="suarezi" status="sp. nov.">
|
||
<emphasis id="B93135272A6DAE66CA3DFA85FC0D9677" box="[774,962,1340,1363]" italics="true" pageId="5" pageNumber="183">Holoaden suarezi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A202885C2A6DAE66CAF2FA85FBD39677" box="[969,1052,1340,1363]" pageId="5" pageNumber="183" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BFAE9352A6DAE66CA03FACAFAEE94FB" pageId="5" pageNumber="183">
|
||
<table id="F9451B952A6D519CC99EFACAFA9594FB" box="[165,1370,1395,2015]" gridcols="3" gridrows="15" pageId="5" pageNumber="183">
|
||
<tr id="3575EB772A6D519CC99EFACAFA9596AD" box="[165,1370,1395,1417]" gridrow="0" pageId="5" pageNumber="183" rowspan-0="1">
|
||
<th id="76A4820B2A6D519CCB51FACAFBF996AD" box="[618,1078,1395,1417]" gridcol="1" gridrow="0" pageId="5" pageNumber="183">Male</th>
|
||
<th id="76A4820B2A6D519CCD85FACAFA9596AD" box="[1214,1370,1395,1417]" gridcol="2" gridrow="0" pageId="5" pageNumber="183">Female</th>
|
||
</tr>
|
||
<tr id="3575EB772A6D519CC99EFA27FA959690" box="[165,1370,1438,1460]" gridrow="1" pageId="5" pageNumber="183" rowspan-0="1">
|
||
<td id="76A4820B2A6D519CCB51FA27FBF99690" box="[618,1078,1438,1460]" gridcol="1" gridrow="1" pageId="5" pageNumber="183">MZUSP-94415 MZUSP-138348</td>
|
||
<td id="76A4820B2A6D519CCD85FA27FA959690" box="[1214,1370,1438,1460]" gridcol="2" gridrow="1" pageId="5" pageNumber="183">MZUSP-23794</td>
|
||
</tr>
|
||
<tr id="3575EB772A6D519CC99EFA70FA9596FB" box="[165,1370,1481,1503]" gridrow="2" pageId="5" pageNumber="183">
|
||
<th id="76A4820B2A6D519CC99EFA70FEB696FB" box="[165,377,1481,1503]" gridcol="0" gridrow="2" pageId="5" pageNumber="183">Snout-vent length</th>
|
||
<td id="76A4820B2A6D519CCB51FA70FBF996FB" box="[618,1078,1481,1503]" gridcol="1" gridrow="2" pageId="5" pageNumber="183">38.5 37.2</td>
|
||
<td id="76A4820B2A6D519CCD85FA70FA9596FB" box="[1214,1370,1481,1503]" gridcol="2" gridrow="2" pageId="5" pageNumber="183">44.2</td>
|
||
</tr>
|
||
<tr id="3575EB772A6D519CC99EFA4AFA95952D" box="[165,1370,1523,1545]" gridrow="3" pageId="5" pageNumber="183">
|
||
<th id="76A4820B2A6D519CC99EFA4AFEB6952D" box="[165,377,1523,1545]" gridcol="0" gridrow="3" pageId="5" pageNumber="183">Head width</th>
|
||
<td id="76A4820B2A6D519CCB51FA4AFBF9952D" box="[618,1078,1523,1545]" gridcol="1" gridrow="3" pageId="5" pageNumber="183">15.9 14.6</td>
|
||
<td id="76A4820B2A6D519CCD85FA4AFA95952D" box="[1214,1370,1523,1545]" gridcol="2" gridrow="3" pageId="5" pageNumber="183">17.1</td>
|
||
</tr>
|
||
<tr id="3575EB772A6D519CC99EF9A7FA959510" box="[165,1370,1566,1588]" gridrow="4" pageId="5" pageNumber="183">
|
||
<th id="76A4820B2A6D519CC99EF9A7FEB69510" box="[165,377,1566,1588]" gridcol="0" gridrow="4" pageId="5" pageNumber="183">Head length</th>
|
||
<td id="76A4820B2A6D519CCB51F9A7FBF99510" box="[618,1078,1566,1588]" gridcol="1" gridrow="4" pageId="5" pageNumber="183">13.6 13.3</td>
|
||
<td id="76A4820B2A6D519CCD85F9A7FA959510" box="[1214,1370,1566,1588]" gridcol="2" gridrow="4" pageId="5" pageNumber="183">13.0</td>
|
||
</tr>
|
||
<tr id="3575EB772A6D519CC99EF9F0FA95957B" box="[165,1370,1609,1631]" gridrow="5" pageId="5" pageNumber="183">
|
||
<th id="76A4820B2A6D519CC99EF9F0FEB6957B" box="[165,377,1609,1631]" gridcol="0" gridrow="5" pageId="5" pageNumber="183">Eye diameter</th>
|
||
<td id="76A4820B2A6D519CCB51F9F0FBF9957B" box="[618,1078,1609,1631]" gridcol="1" gridrow="5" pageId="5" pageNumber="183">4.8 4.6</td>
|
||
<td id="76A4820B2A6D519CCD85F9F0FA95957B" box="[1214,1370,1609,1631]" gridcol="2" gridrow="5" pageId="5" pageNumber="183">5.6</td>
|
||
</tr>
|
||
<tr id="3575EB772A6D519CC99EF9CAFA9595AD" box="[165,1370,1651,1673]" gridrow="6" pageId="5" pageNumber="183">
|
||
<th id="76A4820B2A6D519CC99EF9CAFEB695AD" box="[165,377,1651,1673]" gridcol="0" gridrow="6" pageId="5" pageNumber="183">Eye-nostril distance</th>
|
||
<td id="76A4820B2A6D519CCB51F9CAFBF995AD" box="[618,1078,1651,1673]" gridcol="1" gridrow="6" pageId="5" pageNumber="183">3.5 3.5</td>
|
||
<td id="76A4820B2A6D519CCD85F9CAFA9595AD" box="[1214,1370,1651,1673]" gridcol="2" gridrow="6" pageId="5" pageNumber="183">4.2</td>
|
||
</tr>
|
||
<tr id="3575EB772A6D519CC99EF927FA959590" box="[165,1370,1694,1716]" gridrow="7" pageId="5" pageNumber="183">
|
||
<th id="76A4820B2A6D519CC99EF927FEB69590" box="[165,377,1694,1716]" gridcol="0" gridrow="7" pageId="5" pageNumber="183">Inter-orbital distance</th>
|
||
<td id="76A4820B2A6D519CCB51F927FBF99590" box="[618,1078,1694,1716]" gridcol="1" gridrow="7" pageId="5" pageNumber="183">6.4 6.0</td>
|
||
<td id="76A4820B2A6D519CCD85F927FA959590" box="[1214,1370,1694,1716]" gridcol="2" gridrow="7" pageId="5" pageNumber="183">6.9</td>
|
||
</tr>
|
||
<tr id="3575EB772A6D519CC99EF970FA9595FB" box="[165,1370,1737,1759]" gridrow="8" pageId="5" pageNumber="183">
|
||
<th id="76A4820B2A6D519CC99EF970FEB695FB" box="[165,377,1737,1759]" gridcol="0" gridrow="8" pageId="5" pageNumber="183">Internasal distance</th>
|
||
<td id="76A4820B2A6D519CCB51F970FBF995FB" box="[618,1078,1737,1759]" gridcol="1" gridrow="8" pageId="5" pageNumber="183">3.0 2.9</td>
|
||
<td id="76A4820B2A6D519CCD85F970FA9595FB" box="[1214,1370,1737,1759]" gridcol="2" gridrow="8" pageId="5" pageNumber="183">3.3</td>
|
||
</tr>
|
||
<tr id="3575EB772A6D519CC99EF94AFA95942D" box="[165,1370,1779,1801]" gridrow="9" pageId="5" pageNumber="183">
|
||
<th id="76A4820B2A6D519CC99EF94AFEB6942D" box="[165,377,1779,1801]" gridcol="0" gridrow="9" pageId="5" pageNumber="183">Arm length</th>
|
||
<td id="76A4820B2A6D519CCB51F94AFBF9942D" box="[618,1078,1779,1801]" gridcol="1" gridrow="9" pageId="5" pageNumber="183">9.5 8.1</td>
|
||
<td id="76A4820B2A6D519CCD85F94AFA95942D" box="[1214,1370,1779,1801]" gridcol="2" gridrow="9" pageId="5" pageNumber="183">9.0</td>
|
||
</tr>
|
||
<tr id="3575EB772A6D519CC99EF8A7FA959410" box="[165,1370,1822,1844]" gridrow="10" pageId="5" pageNumber="183">
|
||
<th id="76A4820B2A6D519CC99EF8A7FEB69410" box="[165,377,1822,1844]" gridcol="0" gridrow="10" pageId="5" pageNumber="183">Forearm length</th>
|
||
<td id="76A4820B2A6D519CCB51F8A7FBF99410" box="[618,1078,1822,1844]" gridcol="1" gridrow="10" pageId="5" pageNumber="183">11.2 12.2</td>
|
||
<td id="76A4820B2A6D519CCD85F8A7FA959410" box="[1214,1370,1822,1844]" gridcol="2" gridrow="10" pageId="5" pageNumber="183">13.1</td>
|
||
</tr>
|
||
<tr id="3575EB772A6D519CC99EF8F0FA95947B" box="[165,1370,1865,1887]" gridrow="11" pageId="5" pageNumber="183">
|
||
<th id="76A4820B2A6D519CC99EF8F0FEB6947B" box="[165,377,1865,1887]" gridcol="0" gridrow="11" pageId="5" pageNumber="183">Femur length</th>
|
||
<td id="76A4820B2A6D519CCB51F8F0FBF9947B" box="[618,1078,1865,1887]" gridcol="1" gridrow="11" pageId="5" pageNumber="183">17.8 18.0</td>
|
||
<td id="76A4820B2A6D519CCD85F8F0FA95947B" box="[1214,1370,1865,1887]" gridcol="2" gridrow="11" pageId="5" pageNumber="183">19.9</td>
|
||
</tr>
|
||
<tr id="3575EB772A6D519CC99EF8CAFA9594AD" box="[165,1370,1907,1929]" gridrow="12" pageId="5" pageNumber="183">
|
||
<th id="76A4820B2A6D519CC99EF8CAFEB694AD" box="[165,377,1907,1929]" gridcol="0" gridrow="12" pageId="5" pageNumber="183">Tibia length</th>
|
||
<td id="76A4820B2A6D519CCB51F8CAFBF994AD" box="[618,1078,1907,1929]" gridcol="1" gridrow="12" pageId="5" pageNumber="183">16.9 17.7</td>
|
||
<td id="76A4820B2A6D519CCD85F8CAFA9594AD" box="[1214,1370,1907,1929]" gridcol="2" gridrow="12" pageId="5" pageNumber="183">18.3</td>
|
||
</tr>
|
||
<tr id="3575EB772A6D519CC99EF827FA959490" box="[165,1370,1950,1972]" gridrow="13" pageId="5" pageNumber="183">
|
||
<th id="76A4820B2A6D519CC99EF827FEB69490" box="[165,377,1950,1972]" gridcol="0" gridrow="13" pageId="5" pageNumber="183">Foot length</th>
|
||
<td id="76A4820B2A6D519CCB51F827FBF99490" box="[618,1078,1950,1972]" gridcol="1" gridrow="13" pageId="5" pageNumber="183">16.6 17.5</td>
|
||
<td id="76A4820B2A6D519CCD85F827FA959490" box="[1214,1370,1950,1972]" gridcol="2" gridrow="13" pageId="5" pageNumber="183">17.9</td>
|
||
</tr>
|
||
<tr id="3575EB772A6D519CC99EF870FA9594FB" box="[165,1370,1993,2015]" gridrow="14" pageId="5" pageNumber="183">
|
||
<th id="76A4820B2A6D519CC99EF870FEB694FB" box="[165,377,1993,2015]" gridcol="0" gridrow="14" pageId="5" pageNumber="183">Hand length</th>
|
||
<td id="76A4820B2A6D519CCB51F870FBF994FB" box="[618,1078,1993,2015]" gridcol="1" gridrow="14" pageId="5" pageNumber="183">10.4 9.2</td>
|
||
<td id="76A4820B2A6D519CCD85F870FA9594FB" box="[1214,1370,1993,2015]" gridcol="2" gridrow="14" pageId="5" pageNumber="183">10.9</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<caption id="DF3AB9BD2A6EAE65C9ACF815FD3B94C6" httpUri="https://zenodo.org/record/222741/files/figure.png" pageId="6" pageNumber="184" targetBox="[332,1252,193,1942]" targetPageId="6">
|
||
<paragraph id="8BFAE9352A6EAE65C9ACF815FD3B94C6" blockId="6.[151,1436,1964,2018]" pageId="6" pageNumber="184">
|
||
<emphasis id="B93135272A6EAE65C9ACF815FED994E6" bold="true" box="[151,278,1964,1987]" pageId="6" pageNumber="184">FIGURE 4.</emphasis>
|
||
Ventral view of feet and toes. (A) holotype of
|
||
<taxonomicName id="4C4592B62A6EAE65CA2FF814FC0594E7" box="[788,970,1965,1987]" class="Amphibia" family="Strabomantidae" genus="Holoaden" kingdom="Animalia" order="Anura" pageId="6" pageNumber="184" phylum="Chordata" rank="species" species="suarezi" status="sp. nov.">
|
||
<emphasis id="B93135272A6EAE65CA2FF814FC0594E7" box="[788,970,1965,1987]" italics="true" pageId="6" pageNumber="184">Holoaden suarezi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A202885C2A6EAE65CAEFF817FBE694E7" box="[980,1065,1966,1987]" pageId="6" pageNumber="184" rank="species">sp. nov.</taxonomicNameLabel>
|
||
; (B) lectotype and (D) topotype of
|
||
<taxonomicName id="4C4592B62A6EAE65C9ACF875FE4E94C6" box="[151,385,1996,2018]" class="Amphibia" family="Strabomantidae" genus="Holoaden" kingdom="Animalia" order="Anura" pageId="6" pageNumber="184" phylum="Chordata" rank="species" species="luederwaldti">
|
||
<emphasis id="B93135272A6EAE65C9ACF875FE4E94C6" box="[151,385,1996,2018]" italics="true" pageId="6" pageNumber="184">Holoaden luederwaldti</emphasis>
|
||
</taxonomicName>
|
||
; (C) holotype of
|
||
<taxonomicName id="4C4592B62A6EAE65CB0AF875FD2194C6" box="[561,750,1996,2018]" class="Amphibia" family="Strabomantidae" genus="Holoaden" kingdom="Animalia" order="Anura" pageId="6" pageNumber="184" phylum="Chordata" rank="species" species="pholeter">
|
||
<emphasis id="B93135272A6EAE65CB0AF875FD2194C6" box="[561,750,1996,2018]" italics="true" pageId="6" pageNumber="184">Holoaden pholeter</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF3AB9BD2A6FAE64C9ACFAB2FBFE9660" httpUri="https://zenodo.org/record/222742/files/figure.png" pageId="7" pageNumber="185" targetBox="[151,1435,193,1269]" targetPageId="7">
|
||
<paragraph id="8BFAE9352A6FAE64C9ACFAB2FBFE9660" blockId="7.[151,1436,1291,1348]" pageId="7" pageNumber="185">
|
||
<emphasis id="B93135272A6FAE64C9ACFAB2FEDD9605" bold="true" box="[151,274,1291,1314]" pageId="7" pageNumber="185">FIGURE 5.</emphasis>
|
||
Living specimen of
|
||
<taxonomicName id="4C4592B62A6FAE64C8D9FAB5FD5B9606" box="[482,660,1292,1314]" class="Amphibia" family="Strabomantidae" genus="Holoaden" kingdom="Animalia" order="Anura" pageId="7" pageNumber="185" phylum="Chordata" rank="species" species="suarezi" status="sp. nov.">
|
||
<emphasis id="B93135272A6FAE64C8D9FAB5FD5B9606" box="[482,660,1292,1314]" italics="true" pageId="7" pageNumber="185">Holoaden suarezi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A202885C2A6FAE64CBA1FAB4FD279606" box="[666,744,1293,1314]" pageId="7" pageNumber="185" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(MZUSP 138348) (A) lateral, (B) ventral, (C) frontal and (D) dorsal views. Estação Ecológica de Boracéia, State of São Paulo, Brazil. Photo: Martha C. Lange.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C35FBABE2A6FAE64C9FCFACDFC0195F5" pageId="7" pageNumber="185" type="distribution">
|
||
<paragraph id="8BFAE9352A6FAE64C9FCFACDFF7A9539" blockId="7.[151,1436,1396,2033]" pageId="7" pageNumber="185">
|
||
<emphasis id="B93135272A6FAE64C9FCFACDFE9696A9" bold="true" box="[199,345,1396,1421]" pageId="7" pageNumber="185">Distribution</emphasis>
|
||
. So far,
|
||
<taxonomicName id="4C4592B62A6FAE64C88CFACCFDB396A9" box="[439,636,1397,1421]" class="Amphibia" family="Strabomantidae" genus="Holoaden" kingdom="Animalia" order="Anura" pageId="7" pageNumber="185" phylum="Chordata" rank="species" species="suarezi">
|
||
<emphasis id="B93135272A6FAE64C88CFACCFDB396A9" box="[439,636,1397,1421]" italics="true" pageId="7" pageNumber="185">Holoaden suarezi</emphasis>
|
||
</taxonomicName>
|
||
was recorded in three localities of high altitude (between 900 and
|
||
<quantity id="4CBD44D02A6FAE64CC5FFACCFF619695" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="7" pageNumber="185" unit="m" value="1300.0">1300 m</quantity>
|
||
) in the State of São Paulo, situated in the Atlantic Forest Domain (
|
||
<emphasis id="B93135272A6FAE64CA8FFA23FC3D9695" box="[948,1010,1434,1457]" italics="true" pageId="7" pageNumber="185">sensu</emphasis>
|
||
Ab’Saber 1977) of the Serra do Mar mountain range. The three localities are situated in the municipalities of Bananal (Estação Ecológica de Bananal), São José do Barreiro (Parque Nacional da Serra da Bocaina), and Salesópolis (Estação Biológica de Boracéia) (
|
||
<figureCitation id="137EF5B02A6FAE64CC55FA58FF689539" captionStart="FIGURE 6" captionStartId="8.[151,250,1141,1164]" captionTargetBox="[160,1431,203,1110]" captionTargetId="figure@8.[151,1436,193,1120]" captionTargetPageId="8" captionText="FIGURE 6. Geographic distribution of the genus Holoaden along the Atlantic forest of southeastern Brazil. Asterisk = type locality of H. suarezi sp. nov.; triangle = other localities of H. suarezi in the Serra do Mar; circle = Holoaden luederwaldti (Martins 2010); star = Holoaden bradei (Pombal et al. 2008); square = Holoaden pholeter (Pombal et al. 2008)." httpUri="https://zenodo.org/record/222743/files/figure.png" pageId="7" pageNumber="185">Fig. 6</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BFAE9352A6FAE64C9FCF990FC0195F5" blockId="7.[151,1436,1396,2033]" pageId="7" pageNumber="185">
|
||
Currently,
|
||
<taxonomicName id="4C4592B62A6FAE64C87BF990FDC89565" box="[320,519,1577,1601]" class="Amphibia" family="Strabomantidae" genus="Holoaden" kingdom="Animalia" order="Anura" pageId="7" pageNumber="185" phylum="Chordata" rank="species" species="suarezi">
|
||
<emphasis id="B93135272A6FAE64C87BF990FDC89565" box="[320,519,1577,1601]" italics="true" pageId="7" pageNumber="185">Holoaden suarezi</emphasis>
|
||
</taxonomicName>
|
||
is endemic of the Serra do Mar in the State of São Paulo, and is geographically isolated from
|
||
<taxonomicName id="4C4592B62A6FAE64C802F9F7FE239541" box="[313,492,1613,1637]" class="Amphibia" family="Strabomantidae" genus="Holoaden" kingdom="Animalia" order="Anura" pageId="7" pageNumber="185" phylum="Chordata" rank="species" species="luederwaldti">
|
||
<emphasis id="B93135272A6FAE64C802F9F7FE239541" box="[313,492,1613,1637]" italics="true" pageId="7" pageNumber="185">H. luederwaldti</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4592B62A6FAE64CB13F9F7FD589541" box="[552,663,1613,1637]" class="Amphibia" family="Strabomantidae" genus="Holoaden" kingdom="Animalia" order="Anura" pageId="7" pageNumber="185" phylum="Chordata" rank="species" species="bradei">
|
||
<emphasis id="B93135272A6FAE64CB13F9F7FD589541" box="[552,663,1613,1637]" italics="true" pageId="7" pageNumber="185">H. bradei</emphasis>
|
||
</taxonomicName>
|
||
, which are endemic species of the Serra da Mantiqueira mountain range. Although
|
||
<taxonomicName id="4C4592B62A6FAE64C86FF9CBFE1C95AD" box="[340,467,1649,1673]" class="Amphibia" family="Strabomantidae" genus="Holoaden" kingdom="Animalia" order="Anura" pageId="7" pageNumber="185" phylum="Chordata" rank="species" species="pholeter">
|
||
<emphasis id="B93135272A6FAE64C86FF9CBFE1C95AD" box="[340,467,1649,1673]" italics="true" pageId="7" pageNumber="185">H. pholeter</emphasis>
|
||
</taxonomicName>
|
||
also occurs in the Atlantic Forest Domain of the Serra do Mar mountain range, it is an endemic species of the Serra dos Órgãos that corresponds to a geographically isolated eastern portion of the Serra do Mar in the State of Rio de Janeiro (Pombal
|
||
<emphasis id="B93135272A6FAE64CB9AF903FD1E95F5" box="[673,721,1721,1745]" italics="true" pageId="7" pageNumber="185">et al</emphasis>
|
||
. 2008;
|
||
<bibRefCitation id="EFD494C42A6FAE64CA1CF900FC0D95F5" author="Martins" box="[807,962,1721,1745]" pageId="7" pageNumber="185" refString="Martins, I. A. (2010) Natural history of Holoaden luederwaldti (Amphibia: Strabomantidae: Holoadeninae) in southeastern of Brazil. Zoologia, 27, 40 - 46." type="journal article" year="2010">Martins 2010</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35FBABE2A6FAE64C9FCF964FD0894D5" pageId="7" pageNumber="185" type="etymology">
|
||
<paragraph id="8BFAE9352A6FAE64C9FCF964FE349419" blockId="7.[151,1436,1396,2033]" pageId="7" pageNumber="185">
|
||
<emphasis id="B93135272A6FAE64C9FCF964FE8995D1" bold="true" box="[199,326,1757,1781]" pageId="7" pageNumber="185">Etymology</emphasis>
|
||
. A patronym honoring José Roberto Alves Suarez, Forest Ranger from the Instituto Florestal of São Paulo who dedicated most of his long-time career to the preservation of São Paulo’s high altitude Atlantic forest remnants and their fauna.
|
||
</paragraph>
|
||
<paragraph id="8BFAE9352A6FAE64C9FCF8F0FD0894D5" blockId="7.[151,1436,1396,2033]" pageId="7" pageNumber="185">
|
||
<emphasis id="B93135272A6FAE64C9FCF8F0FEB19445" bold="true" box="[199,382,1864,1889]" pageId="7" pageNumber="185">Natural history</emphasis>
|
||
. The
|
||
<typeStatus id="54FE57972A6FAE64C8FBF8F0FDED9445" box="[448,546,1865,1889]" pageId="7" pageNumber="185" type="holotype">holotype</typeStatus>
|
||
and
|
||
<typeStatus id="54FE57972A6FAE64CB62F8F3FD769446" box="[601,697,1866,1890]" pageId="7" pageNumber="185" type="paratype">paratype</typeStatus>
|
||
MZUSP 138348 were captured in pitfall traps during the months of highest precipitations (
|
||
<date id="FFFBCFF52A6FAE64C88EF8D4FCD294A1" box="[437,797,1901,1925]" pageId="7" pageNumber="185" value="2003-11" valueMax="2003-12-31" valueMin="2003-11-00">November and December 2003</date>
|
||
).
|
||
<bibRefCitation id="EFD494C42A6FAE64CA0BF8D7FBC894A1" box="[816,1031,1901,1926]" pageId="7" pageNumber="185" refString="Heyer, W. R., Rand, A. S., Cruz, C. A. G., Peixoto, O. L. & Nelson, C. E. (1990) Frogs of Boraceia. Arquivos de Zoologia, 31, 231 - 410." type="journal article">
|
||
Heyer
|
||
<emphasis id="B93135272A6FAE64CA47F8D7FC6194A1" box="[892,942,1901,1925]" italics="true" pageId="7" pageNumber="185">et al</emphasis>
|
||
. (1990)
|
||
</bibRefCitation>
|
||
mentioned that MZUSP 23794 was taken from the ground on a forest trail at night. MZUSP 23794 contained a small number of large and unpigmented eggs. Similar eggs were reported in
|
||
<taxonomicName id="4C4592B62A6FAE64CB73F80FFC9E94E9" authority="Lutz 1958" authorityName="Lutz" authorityYear="1958" box="[584,849,1973,1997]" class="Amphibia" family="Strabomantidae" genus="Holoaden" kingdom="Animalia" order="Anura" pageId="7" pageNumber="185" phylum="Chordata" rank="species" species="bradei">
|
||
<emphasis id="B93135272A6FAE64CB73F80FFD7794E9" box="[584,696,1973,1997]" italics="true" pageId="7" pageNumber="185">H. bradei</emphasis>
|
||
(
|
||
<bibRefCitation id="EFD494C42A6FAE64CBF7F80CFC8794E9" author="Lutz" box="[716,840,1973,1997]" pageId="7" pageNumber="185" refString="Lutz, B. (1958) Anfibios novos e raros das serras costeiras do Brasil. Memorias do Instituto Oswaldo Cruz, Rio de Janeiro, 56, 382 - 389." type="journal article" year="1958">Lutz 1958</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4592B62A6FAE64CA59F80FFB1C94E9" authority="Martins 2010" authorityName="Martins" authorityYear="2010" box="[866,1235,1973,1997]" class="Amphibia" family="Strabomantidae" genus="Holoaden" kingdom="Animalia" order="Anura" pageId="7" pageNumber="185" phylum="Chordata" rank="species" species="luederwaldti">
|
||
<emphasis id="B93135272A6FAE64CA59F80FFBD994E9" box="[866,1046,1973,1997]" italics="true" pageId="7" pageNumber="185">H. luederwaldti</emphasis>
|
||
(
|
||
<bibRefCitation id="EFD494C42A6FAE64CD12F80CFB0594E9" author="Martins" box="[1065,1226,1973,1997]" pageId="7" pageNumber="185" refString="Martins, I. A. (2010) Natural history of Holoaden luederwaldti (Amphibia: Strabomantidae: Holoadeninae) in southeastern of Brazil. Zoologia, 27, 40 - 46." type="journal article" year="2010">Martins 2010</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C4592B62A6FAE64CC2CF80FFE4F94D5" authority="Pombal et al. 2008" authorityName="Pombal et al." authorityYear="2008" class="Amphibia" family="Strabomantidae" genus="Holoaden" kingdom="Animalia" order="Anura" pageId="7" pageNumber="185" phylum="Chordata" rank="species" species="pholeter">
|
||
<emphasis id="B93135272A6FAE64CC2CF80FFA5394E9" box="[1303,1436,1973,1997]" italics="true" pageId="7" pageNumber="185">H. pholeter</emphasis>
|
||
(Pombal
|
||
<emphasis id="B93135272A6FAE64C9C7F863FEFA94D5" box="[252,309,2009,2033]" italics="true" pageId="7" pageNumber="185">et al.</emphasis>
|
||
2008)
|
||
</taxonomicName>
|
||
. Its vocalization is unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |