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<document id="8F0B1EDBCEAFF63883B37121792A3F5B" ID-CLB-Dataset="28149" ID-DOI="10.11646/zootaxa.4563.2.1" ID-GBIF-Dataset="e6037c5d-a96f-41e5-8e70-71f2659659e2" ID-ISSN="1175-5326" ID-Zenodo-Dep="2601072" ID-ZooBank="CF79B01B-8B5F-4B3A-B642-2CADE4B339AF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553171433896" checkinUser="plazi" docAuthor="Wesener, Thomas" docDate="2019" docId="03EB5506E32F8C12A2EDFD40FD81FBC9" docLanguage="en" docName="zootaxa.4563.2.1.pdf" docOrigin="Zootaxa 4563 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Sphaerobelum schwendingeri Wesener 2019, new species" docType="treatment" docVersion="9" lastPageNumber="239" masterDocId="FFD22D7EE30A8C34A27AFFCFFFE9FFFC" masterDocTitle="First records of giant pill-millipedes from Laos (Diplopoda, Sphaerotheriida, Zephroniidae)" masterLastPageNumber="248" masterPageNumber="201" pageNumber="238" updateTime="1698703176884" updateUser="ExternalLinkService">
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<mods:title id="D11A75C5704DE5B57D44C5D46333B9B9">First records of giant pill-millipedes from Laos (Diplopoda, Sphaerotheriida, Zephroniidae)</mods:title>
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<mods:namePart id="F234FD15EFC164470A6C3A02A9B97E57">Wesener, Thomas</mods:namePart>
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<mods:title id="2006067FFD33745618B298ABC9CFF4A9">Zootaxa</mods:title>
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<mods:part id="1FA8D0E900F102C4F7715C12B0CDC16D">
<mods:date id="56DD10E8228F9932EACA5AA868C0E880">2019</mods:date>
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<mods:number id="9E0129226F644DBBA96B7630C0DB3DB7">2019-03-01</mods:number>
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<mods:identifier id="97EC9970E3F7911B746728954F1936CA" type="DOI">10.11646/zootaxa.4563.2.1</mods:identifier>
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<treatment id="03EB5506E32F8C12A2EDFD40FD81FBC9" ID-DOI="http://doi.org/10.5281/zenodo.5934540" ID-GBIF-Taxon="154474545" ID-Zenodo-Dep="5934540" LSID="urn:lsid:plazi:treatment:03EB5506E32F8C12A2EDFD40FD81FBC9" httpUri="http://treatment.plazi.org/id/03EB5506E32F8C12A2EDFD40FD81FBC9" lastPageId="38" lastPageNumber="239" pageId="37" pageNumber="238">
<subSubSection id="C358B79BE32F8C11A2EDFD40FD7CFD56" box="[151,661,655,682]" pageId="37" pageNumber="238" type="nomenclature">
<paragraph id="8BFDE410E32F8C11A2EDFD40FD7CFD56" blockId="37.[151,661,655,715]" box="[151,661,655,682]" pageId="37" pageNumber="238">
<heading id="D0B5537CE32F8C11A2EDFD40FD7CFD56" bold="true" box="[151,661,655,682]" fontSize="11" level="1" pageId="37" pageNumber="238" reason="1">
<emphasis id="B9363802E32F8C11A2EDFD40FD7CFD56" bold="true" box="[151,661,655,682]" pageId="37" pageNumber="238">
<taxonomicName id="4C429F93E32F8C11A2EDFD40FE17FD55" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[151,510,655,681]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="37" pageNumber="238" phylum="Arthropoda" rank="species" species="schwendingeri" status="sp. nov.">
<emphasis id="B9363802E32F8C11A2EDFD40FE17FD55" bold="true" box="[151,510,655,681]" italics="true" pageId="37" pageNumber="238">Sphaerobelum schwendingeri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2058579E32F8C11A07FFD5FFD7CFD56" box="[517,661,656,682]" pageId="37" pageNumber="238" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C358B79BE32F8C11A2EDFD7DFEEAFD37" box="[151,259,690,715]" pageId="37" pageNumber="238" type="description">
<paragraph id="8BFDE410E32F8C11A2EDFD7DFEEAFD37" blockId="37.[151,661,655,715]" box="[151,259,690,715]" pageId="37" pageNumber="238">
<figureCitation id="1379F895E32F8C11A2EDFD7DFEEAFD37" box="[151,259,690,715]" captionStart="FIGURE 25" captionStartId="41.[151,250,1885,1907]" captionTargetBox="[207,1378,195,1868]" captionTargetId="figure@41.[207,1378,195,1868]" captionTargetPageId="41" captionText="FIGURE 25. Sphaerobelum schwendingeri n. sp. AC female paratype (ZMUC), DG male holotype (MHNG). A: coxa of first right leg with first stigmatic plate; B: 9th left leg; C: coxa and prefemur of second leg with vulva; D: left anterior telopod, anterior view; E: left anterior telopod, posterior view; F: right posterior telopod, anterior view; G: right posterior telopod, posterior view. Abbreviations: Cx = coxa; O = operculum of vulva; Pre = prefemur; St-Pl = stigmatic plate; Syn-Cx = syncoxite. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601125/files/figure.png" pageId="37" pageNumber="238">Figure 25</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C358B79BE32F8C11A2EDFD35FC27FBF3" pageId="37" pageNumber="238" type="materials_examined">
<paragraph id="8BFDE410E32F8C11A2EDFD35FC35FC5F" blockId="37.[151,1437,762,2011]" pageId="37" pageNumber="238">
<emphasis id="B9363802E32F8C11A2EDFD35FE6FFCEF" bold="true" box="[151,390,762,787]" pageId="37" pageNumber="238">Material examined:</emphasis>
Type material. 1 M
<typeStatus id="54F95AB2E32F8C11A007FD35FD36FCEF" box="[637,735,762,787]" pageId="37" pageNumber="238" type="holotype">holotype</typeStatus>
(MHNG LT 10/03) from
<collectingCountry id="F355A480E32F8C11A669FD34FBA6FCEE" box="[1043,1103,763,786]" name="Laos" pageId="37" pageNumber="238">Laos</collectingCountry>
;,
<materialsCitation id="3B2AEE4DE32F8C11A619FD35FF39FC82" ID-GBIF-Occurrence="2013952576" collectingDate="2010-09-07" collectionCode="MHNG, SMF" collectorName="Tham Pou Kham, W. &amp; P. Schwendinger." country="Laos" elevation="240" latitude="18.925554" location="Province" longLatPrecision="21" longitude="102.399445" municipality="Vang Vieng" pageId="37" pageNumber="238" specimenCode="LT 10" specimenCount="2" stateProvince="Viangchan" typeStatus="paratype">
<collectingRegion id="49862AF2E32F8C11A619FD35FB27FCEF" box="[1123,1230,762,787]" country="Laos" name="Viangchan" pageId="37" pageNumber="238">Vientiane</collectingRegion>
<location id="8E9DB2CBE32F8C11A6A3FD35FAA9FCEF" LSID="urn:lsid:plazi:treatment:03EB5506E32F8C12A2EDFD40FD81FBC9:8E9DB2CBE32F8C11A6A3FD35FAA9FCEF" box="[1241,1344,762,787]" country="Laos" latitude="18.925554" longLatPrecision="21" longitude="102.399445" municipality="Vang Vieng" name="Province" pageId="37" pageNumber="238" stateProvince="Viangchan">Province</location>
, trail to
<collectorName id="26B781C6E32F8C11A2EDFCD1FE63FCCA" box="[151,394,798,823]" pageId="37" pageNumber="238">Tham Pou Kham, W.</collectorName>
of
<collectingMunicipality id="6B997E6AE32F8C11A3C9FCD0FDD1FCCB" box="[435,568,798,823]" pageId="37" pageNumber="238">Vang Vieng</collectingMunicipality>
(
<geoCoordinate id="EE7682D7E32F8C11A030FCD1FD38FCCB" box="[586,721,798,823]" direction="north" orientation="latitude" pageId="37" pageNumber="238" precision="15" value="18.925554">N18°55'32&quot;</geoCoordinate>
,
<geoCoordinate id="EE7682D7E32F8C11A0A4FCD1FC99FCCB" box="[734,880,798,823]" direction="east" orientation="longitude" pageId="37" pageNumber="238" precision="15" value="102.399445">102°23'58&quot;E</geoCoordinate>
),
<quantity id="4CBA49F5E32F8C11A1FCFCD0FC38FCCA" box="[902,977,799,823]" metricMagnitude="2" metricUnit="m" metricValue="2.4" pageId="37" pageNumber="238" unit="m" value="240.0">
<elevation id="006F0323E32F8C11A1FCFCD0FC38FCCA" box="[902,977,799,823]" metricMagnitude="2" metricUnit="m" metricValue="2.4" pageId="37" pageNumber="238" unit="m" value="240.0">240 m</elevation>
</quantity>
, secondary forest on limestone hill, leg.
<date id="FFFCC2D0E32F8C11A2EDFC8CFEE2FCA7" box="[151,267,835,859]" pageId="37" pageNumber="238" value="2010-09-07">
<collectingDate id="EFB83B38E32F8C11A2EDFC8CFEE2FCA7" box="[151,267,835,859]" pageId="37" pageNumber="238" value="2010-09-07">7.IX.2010</collectingDate>
</date>
,
<collectorName id="26B781C6E32F8C11A362FC8CFE33FCA7" box="[280,474,834,859]" pageId="37" pageNumber="238">P. Schwendinger.</collectorName>
1 M
<typeStatus id="54F95AB2E32F8C11A060FC8CFD93FCA7" box="[538,634,835,859]" pageId="37" pageNumber="238" type="paratype">paratype</typeStatus>
(
<collectionCode id="ED537CD5E32F8C11A0F0FC8CFD08FCA7" box="[650,737,835,859]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="37" pageNumber="238" type="Museum">MHNG</collectionCode>
<specimenCode id="DBE44C6BE32F8C11A090FC8CFCC7FCA7" box="[746,814,835,859]" collectionCode="MHNG" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="37" pageNumber="238" type="Museum">LT 10</specimenCode>
/03), same data as previous. 1 F
<typeStatus id="54F95AB2E32F8C11A6D8FC8CFAEAFCA7" box="[1186,1283,835,859]" pageId="37" pageNumber="238" type="paratype">paratype</typeStatus>
(
<collectionCode id="ED537CD5E32F8C11A76EFC8CFAB8FCA7" box="[1300,1361,835,859]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="37" pageNumber="238">SMF</collectionCode>
), from
<collectingCountry id="F355A480E32F8C11A2EDFCA8FF39FC82" box="[151,208,871,894]" name="Laos" pageId="37" pageNumber="238">Laos</collectingCountry>
</materialsCitation>
,
<materialsCitation id="3B2AEE4DE32F8C11A2A6FCA9FC30FC5F" ID-GBIF-Occurrence="2013952571" collectingDate="2007-03-13" collectorName="P. Jager &amp; F. Steinmetz" country="Laos" county="Tham Pou Kham" elevation="260" latitude="18.925817" location="Province" longLatPrecision="1" longitude="102.39557" municipality="Vang Vieng" pageId="37" pageNumber="238" specimenCount="1" stateProvince="Viangchan" typeStatus="paratype">
<collectingRegion id="49862AF2E32F8C11A2A6FCA9FEAEFC83" box="[220,327,870,895]" country="Laos" name="Viangchan" pageId="37" pageNumber="238">Vientiane</collectingRegion>
<location id="8E9DB2CBE32F8C11A335FCA9FE5FFC83" LSID="urn:lsid:plazi:treatment:03EB5506E32F8C12A2EDFD40FD81FBC9:8E9DB2CBE32F8C11A335FCA9FE5FFC83" box="[335,438,870,895]" country="Laos" county="Tham Pou Kham" latitude="18.925817" longLatPrecision="1" longitude="102.39557" municipality="Vang Vieng" name="Province" pageId="37" pageNumber="238" stateProvince="Viangchan">Province</location>
,
<collectingMunicipality id="6B997E6AE32F8C11A3BBFCA8FDA1FC83" box="[449,584,870,895]" pageId="37" pageNumber="238">Vang Vieng</collectingMunicipality>
,
<collectingCounty id="629C9C9CE32F8C11A02EFCA9FCFEFC83" box="[596,791,870,895]" pageId="37" pageNumber="238">Tham Pou Kham</collectingCounty>
,
<quantity id="4CBA49F5E32F8C11A159FCA9FC82FC82" box="[803,875,870,895]" metricMagnitude="2" metricUnit="m" metricValue="2.6" pageId="37" pageNumber="238" unit="m" value="260.0">
<elevation id="006F0323E32F8C11A159FCA9FC82FC82" box="[803,875,870,895]" metricMagnitude="2" metricUnit="m" metricValue="2.6" pageId="37" pageNumber="238" unit="m" value="260.0">260 m</elevation>
</quantity>
(
<geoCoordinate id="EE7682D7E32F8C11A100FCA9FBE4FC83" box="[890,1037,870,895]" direction="north" orientation="latitude" pageId="37" pageNumber="238" precision="1" value="18.925817">N18°55.549'</geoCoordinate>
,
<geoCoordinate id="EE7682D7E32F8C11A662FCA8FB5DFC83" box="[1048,1204,871,895]" direction="east" orientation="longitude" pageId="37" pageNumber="238" precision="1" value="102.39557">102°23.734'E</geoCoordinate>
), outside cave, close to cave stream, at rocks, leg. by hand,
<date id="FFFCC2D0E32F8C11A038FC44FD2DFC5F" box="[578,708,907,931]" pageId="37" pageNumber="238" value="2007-03-13">
<collectingDate id="EFB83B38E32F8C11A038FC44FD2DFC5F" box="[578,708,907,931]" pageId="37" pageNumber="238" value="2007-03-13">13.III.2007</collectingDate>
</date>
,
<collectorName id="26B781C6E32F8C11A0AAFC44FCCCFC5F" box="[720,805,907,931]" pageId="37" pageNumber="238">P. Jäger</collectorName>
&amp;
<collectorName id="26B781C6E32F8C11A133FC44FC30FC5F" box="[841,985,906,931]" pageId="37" pageNumber="238">F. Steinmetz</collectorName>
</materialsCitation>
.
</paragraph>
<paragraph id="8BFDE410E32F8C11A2BDFC61FC27FBF3" blockId="37.[151,1437,762,2011]" pageId="37" pageNumber="238">
<emphasis id="B9363802E32F8C11A2BDFC61FDEDFC3B" bold="true" box="[199,516,942,967]" pageId="37" pageNumber="238">Other material examined:</emphasis>
1 M, 1 imm. F (SMF) from
<collectingCountry id="F355A480E32F8C11A121FC60FC7DFC3A" box="[859,916,943,966]" name="Laos" pageId="37" pageNumber="238">Laos</collectingCountry>
,
<collectingRegion id="49862AF2E32F8C11A1D8FC61FBE4FC3B" box="[930,1037,942,967]" country="Laos" name="Viangchan" pageId="37" pageNumber="238">Vientiane</collectingRegion>
Province, Van Vieng, across Nam Song, Tham Chang, in or in front of cave (
<geoCoordinate id="EE7682D7E32F8C11A0DBFC1DFCDDFC17" box="[673,820,978,1003]" direction="north" orientation="latitude" pageId="37" pageNumber="238" precision="1" value="18.909166">N18°54.550'</geoCoordinate>
,
<geoCoordinate id="EE7682D7E32F8C11A139FC1DFC36FC17" box="[835,991,978,1003]" direction="east" orientation="longitude" pageId="37" pageNumber="238" precision="1" value="102.442116">102°26.527'E</geoCoordinate>
),
<quantity id="4CBA49F5E32F8C11A18DFC1CFBAFFC16" box="[1015,1094,979,1003]" metricMagnitude="2" metricUnit="m" metricValue="2.7" pageId="37" pageNumber="238" unit="m" value="270.0">270 m</quantity>
, leg. by hand, P. Jäger &amp; S. Packheiser,
<date id="FFFCC2D0E32F8C11A367FC39FE4DFBF3" box="[285,420,1014,1039]" pageId="37" pageNumber="238" value="2018-07-22">22.Vii.2018</date>
. 1 M (ZFMK MYR8103), same data as previous.
</paragraph>
</subSubSection>
<subSubSection id="C358B79BE32F8C11A2BDFBD5FCBEFAAF" pageId="37" pageNumber="238" type="diagnosis">
<paragraph id="8BFDE410E32F8C11A2BDFBD5FCBEFAAF" blockId="37.[151,1437,762,2011]" pageId="37" pageNumber="238">
<emphasis id="B9363802E32F8C11A2BDFBD5FED3FBCF" bold="true" box="[199,314,1050,1075]" pageId="37" pageNumber="238">Diagnosis</emphasis>
:
<taxonomicName id="4C429F93E32F8C11A333FBD4FDE0FBCF" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[329,521,1051,1075]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="37" pageNumber="238" phylum="Arthropoda" rank="species" species="schwendingeri" status="sp. nov.">
<emphasis id="B9363802E32F8C11A333FBD4FDE0FBCF" box="[329,521,1051,1075]" italics="true" pageId="37" pageNumber="238">S. schwendingeri</emphasis>
</taxonomicName>
<emphasis id="B9363802E32F8C11A06BFBD5FDB8FBCF" bold="true" box="[529,593,1050,1075]" pageId="37" pageNumber="238">
<taxonomicNameLabel id="A2058579E32F8C11A06BFBD5FDB8FBCF" box="[529,593,1050,1075]" pageId="37" pageNumber="238" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
belongs to a group of
<taxonomicName id="4C429F93E32F8C11A12BFBD4FC1BFBCF" authorityName="Verhoeff" authorityYear="1924" box="[849,1010,1051,1075]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="37" pageNumber="238" phylum="Arthropoda" rank="genus">
<emphasis id="B9363802E32F8C11A12BFBD4FC1BFBCF" box="[849,1010,1051,1075]" italics="true" pageId="37" pageNumber="238">Sphaerobelum</emphasis>
</taxonomicName>
species in which the mesal margin of the femur is unmodified (
<figureCitation id="1379F895E32F8C11A3CEFBF1FDF2FBAB" box="[436,539,1086,1111]" captionStart="FIGURE 25" captionStartId="41.[151,250,1885,1907]" captionTargetBox="[207,1378,195,1868]" captionTargetId="figure@41.[207,1378,195,1868]" captionTargetPageId="41" captionText="FIGURE 25. Sphaerobelum schwendingeri n. sp. AC female paratype (ZMUC), DG male holotype (MHNG). A: coxa of first right leg with first stigmatic plate; B: 9th left leg; C: coxa and prefemur of second leg with vulva; D: left anterior telopod, anterior view; E: left anterior telopod, posterior view; F: right posterior telopod, anterior view; G: right posterior telopod, posterior view. Abbreviations: Cx = coxa; O = operculum of vulva; Pre = prefemur; St-Pl = stigmatic plate; Syn-Cx = syncoxite. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601125/files/figure.png" pageId="37" pageNumber="238">Fig. 25B</figureCitation>
).
<taxonomicName id="4C429F93E32F8C11A054FBF0FD07FBAA" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[558,750,1086,1110]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="37" pageNumber="238" phylum="Arthropoda" rank="species" species="schwendingeri" status="sp. nov.">
<emphasis id="B9363802E32F8C11A054FBF0FD07FBAA" box="[558,750,1086,1110]" italics="true" pageId="37" pageNumber="238">S. schwendingeri</emphasis>
</taxonomicName>
<emphasis id="B9363802E32F8C11A08FFBF1FCDAFBAB" bold="true" box="[757,819,1086,1111]" pageId="37" pageNumber="238">
<taxonomicNameLabel id="A2058579E32F8C11A08FFBF1FCDAFBAB" box="[757,819,1086,1111]" pageId="37" pageNumber="238" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
shares only with
<taxonomicName id="4C429F93E32F8C11A183FBF0FB90FBAA" authorityName="Verhoeff" authorityYear="1924" box="[1017,1145,1086,1110]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="37" pageNumber="238" phylum="Arthropoda" rank="species" species="hirsutum">
<emphasis id="B9363802E32F8C11A183FBF0FB90FBAA" box="[1017,1145,1086,1110]" italics="true" pageId="37" pageNumber="238">S. hirsutum</emphasis>
</taxonomicName>
,
<taxonomicName id="4C429F93E32F8C11A6FDFBF0FAC8FBAA" authorityName="Verhoeff" authorityYear="1924" box="[1159,1313,1086,1110]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="37" pageNumber="238" phylum="Arthropoda" rank="species" species="clavigerum">
<emphasis id="B9363802E32F8C11A6FDFBF0FAC8FBAA" box="[1159,1313,1086,1110]" italics="true" pageId="37" pageNumber="238">S. clavigerum</emphasis>
</taxonomicName>
,
<taxonomicName id="4C429F93E32F8C11A755FBF0FA75FBAA" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[1327,1436,1087,1110]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="37" pageNumber="238" phylum="Arthropoda" rank="species" species="nigrum" status="sp. nov.">
<emphasis id="B9363802E32F8C11A755FBF0FA75FBAA" box="[1327,1436,1087,1110]" italics="true" pageId="37" pageNumber="238">S. nigrum</emphasis>
</taxonomicName>
<emphasis id="B9363802E32F8C11A2EDFBADFF31FB87" bold="true" box="[151,216,1122,1147]" pageId="37" pageNumber="238">
<taxonomicNameLabel id="A2058579E32F8C11A2EDFBADFF31FB87" box="[151,216,1122,1147]" pageId="37" pageNumber="238" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4C429F93E32F8C11A292FBACFE6EFB87" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[232,391,1123,1147]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="37" pageNumber="238" phylum="Arthropoda" rank="species" species="splendidum" status="sp. nov.">
<emphasis id="B9363802E32F8C11A292FBACFE6EFB87" box="[232,391,1123,1147]" italics="true" pageId="37" pageNumber="238">S. splendidum</emphasis>
</taxonomicName>
<emphasis id="B9363802E32F8C11A3EBFBADFE3BFB87" bold="true" box="[401,466,1122,1147]" pageId="37" pageNumber="238">
<taxonomicNameLabel id="A2058579E32F8C11A3EBFBADFE3BFB87" box="[401,466,1122,1147]" pageId="37" pageNumber="238" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C429F93E32F8C11A076FBACFD64FB87" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[524,653,1123,1147]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="37" pageNumber="238" phylum="Arthropoda" rank="species" species="laoticum" status="sp. nov.">
<emphasis id="B9363802E32F8C11A076FBACFD64FB87" box="[524,653,1123,1147]" italics="true" pageId="37" pageNumber="238">S. laoticum</emphasis>
</taxonomicName>
<emphasis id="B9363802E32F8C11A0ECFBADFD3EFB87" bold="true" box="[662,727,1122,1147]" pageId="37" pageNumber="238">
<taxonomicNameLabel id="A2058579E32F8C11A0ECFBADFD3EFB87" box="[662,727,1122,1147]" pageId="37" pageNumber="238" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
the presence of a &quot;hook&quot; at telopoditomere 4 of the posterior telopods(
<figureCitation id="1379F895E32F8C11A285FB49FE6CFB63" box="[255,389,1158,1183]" captionStart="FIGURE 25" captionStartId="41.[151,250,1885,1907]" captionTargetBox="[207,1378,195,1868]" captionTargetId="figure@41.[207,1378,195,1868]" captionTargetPageId="41" captionText="FIGURE 25. Sphaerobelum schwendingeri n. sp. AC female paratype (ZMUC), DG male holotype (MHNG). A: coxa of first right leg with first stigmatic plate; B: 9th left leg; C: coxa and prefemur of second leg with vulva; D: left anterior telopod, anterior view; E: left anterior telopod, posterior view; F: right posterior telopod, anterior view; G: right posterior telopod, posterior view. Abbreviations: Cx = coxa; O = operculum of vulva; Pre = prefemur; St-Pl = stigmatic plate; Syn-Cx = syncoxite. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601125/files/figure.png" pageId="37" pageNumber="238">Figs 25F, G</figureCitation>
).
<taxonomicName id="4C429F93E32F8C11A3E2FB48FDBEFB62" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[408,599,1158,1182]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="37" pageNumber="238" phylum="Arthropoda" rank="species" species="schwendingeri" status="sp. nov.">
<emphasis id="B9363802E32F8C11A3E2FB48FDBEFB62" box="[408,599,1158,1182]" italics="true" pageId="37" pageNumber="238">S. schwendingeri</emphasis>
</taxonomicName>
<emphasis id="B9363802E32F8C11A024FB49FD75FB63" bold="true" box="[606,668,1158,1183]" pageId="37" pageNumber="238">
<taxonomicNameLabel id="A2058579E32F8C11A024FB49FD75FB63" box="[606,668,1158,1183]" pageId="37" pageNumber="238" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="4C429F93E32F8C11A14AFB48FC59FB62" authorityName="Verhoeff" authorityYear="1924" box="[816,944,1158,1182]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="37" pageNumber="238" phylum="Arthropoda" rank="species" species="hirsutum">
<emphasis id="B9363802E32F8C11A14AFB48FC59FB62" box="[816,944,1158,1182]" italics="true" pageId="37" pageNumber="238">S. hirsutum</emphasis>
</taxonomicName>
in the presence of glabrous tergites, from
<taxonomicName id="4C429F93E32F8C11A7FDFB48FEFCFB3F" authorityName="Verhoeff" authorityYear="1924" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="37" pageNumber="238" phylum="Arthropoda" rank="species" species="clavigerum">
<emphasis id="B9363802E32F8C11A7FDFB48FEFCFB3F" italics="true" pageId="37" pageNumber="238">S. clavigerum</emphasis>
</taxonomicName>
in the presence of a short locking carina on the anal shield. It differs from
<taxonomicName id="4C429F93E32F8C11A6E8FB64FAEBFB3E" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[1170,1282,1195,1218]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="37" pageNumber="238" phylum="Arthropoda" rank="species" species="nigrum" status="sp. nov.">
<emphasis id="B9363802E32F8C11A6E8FB64FAEBFB3E" box="[1170,1282,1195,1218]" italics="true" pageId="37" pageNumber="238">S. nigrum</emphasis>
</taxonomicName>
<emphasis id="B9363802E32F8C11A776FB65FAA6FB3F" bold="true" box="[1292,1359,1194,1219]" pageId="37" pageNumber="238">
<taxonomicNameLabel id="A2058579E32F8C11A776FB65FAA6FB3F" box="[1292,1359,1194,1219]" pageId="37" pageNumber="238" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
in the presence of a cylindrical antennomere 6, the presence of 810 ventral spines on the tarsi (
<figureCitation id="1379F895E32F8C11A6B9FB01FAC4FB1B" box="[1219,1325,1230,1255]" captionStart="FIGURE 25" captionStartId="41.[151,250,1885,1907]" captionTargetBox="[207,1378,195,1868]" captionTargetId="figure@41.[207,1378,195,1868]" captionTargetPageId="41" captionText="FIGURE 25. Sphaerobelum schwendingeri n. sp. AC female paratype (ZMUC), DG male holotype (MHNG). A: coxa of first right leg with first stigmatic plate; B: 9th left leg; C: coxa and prefemur of second leg with vulva; D: left anterior telopod, anterior view; E: left anterior telopod, posterior view; F: right posterior telopod, anterior view; G: right posterior telopod, posterior view. Abbreviations: Cx = coxa; O = operculum of vulva; Pre = prefemur; St-Pl = stigmatic plate; Syn-Cx = syncoxite. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601125/files/figure.png" pageId="37" pageNumber="238">Fig. 25B</figureCitation>
), and the presence of a strongly curved telopoditomere 4 of the posterior telopod, all characters only shared with
<taxonomicName id="4C429F93E32F8C11A7FDFB3CFEF1FAD2" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="37" pageNumber="238" phylum="Arthropoda" rank="species" species="splendidum" status="sp. nov.">
<emphasis id="B9363802E32F8C11A7FDFB3CFEF1FAD2" italics="true" pageId="37" pageNumber="238">S. splendidum</emphasis>
</taxonomicName>
<emphasis id="B9363802E32F8C11A35AFAD9FEB6FAD3" bold="true" box="[288,351,1302,1327]" pageId="37" pageNumber="238">
<taxonomicNameLabel id="A2058579E32F8C11A35AFAD9FEB6FAD3" box="[288,351,1302,1327]" pageId="37" pageNumber="238" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C429F93E32F8C11A3EDFAD8FDFFFAD2" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[407,534,1302,1326]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="37" pageNumber="238" phylum="Arthropoda" rank="species" species="laoticum" status="sp. nov.">
<emphasis id="B9363802E32F8C11A3EDFAD8FDFFFAD2" box="[407,534,1302,1326]" italics="true" pageId="37" pageNumber="238">S. laoticum</emphasis>
</taxonomicName>
<emphasis id="B9363802E32F8C11A067FAD9FDB4FAD3" bold="true" box="[541,605,1302,1327]" pageId="37" pageNumber="238">
<taxonomicNameLabel id="A2058579E32F8C11A067FAD9FDB4FAD3" box="[541,605,1302,1327]" pageId="37" pageNumber="238" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
For a differentiation of
<taxonomicName id="4C429F93E32F8C11A114FAD8FBC7FAD2" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[878,1070,1302,1326]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="37" pageNumber="238" phylum="Arthropoda" rank="species" species="schwendingeri" status="sp. nov.">
<emphasis id="B9363802E32F8C11A114FAD8FBC7FAD2" box="[878,1070,1302,1326]" italics="true" pageId="37" pageNumber="238">S. schwendingeri</emphasis>
</taxonomicName>
<emphasis id="B9363802E32F8C11A64CFAD9FB9CFAD3" bold="true" box="[1078,1141,1302,1327]" pageId="37" pageNumber="238">
<taxonomicNameLabel id="A2058579E32F8C11A64CFAD9FB9CFAD3" box="[1078,1141,1302,1327]" pageId="37" pageNumber="238" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
with
<taxonomicName id="4C429F93E32F8C11A6CDFAD8FABCFAD2" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[1207,1365,1302,1326]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="37" pageNumber="238" phylum="Arthropoda" rank="species" species="splendidum" status="sp. nov.">
<emphasis id="B9363802E32F8C11A6CDFAD8FABCFAD2" box="[1207,1365,1302,1326]" italics="true" pageId="37" pageNumber="238">S. splendidum</emphasis>
</taxonomicName>
<emphasis id="B9363802E32F8C11A726FAD9FA75FAD3" bold="true" box="[1372,1436,1302,1327]" pageId="37" pageNumber="238">
<taxonomicNameLabel id="A2058579E32F8C11A726FAD9FA75FAD3" box="[1372,1436,1302,1327]" pageId="37" pageNumber="238" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
see above. For a differentiation to
<taxonomicName id="4C429F93E32F8C11A06DFAF4FD7CFAAF" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[535,661,1339,1363]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="37" pageNumber="238" phylum="Arthropoda" rank="species" species="laoticum" status="sp. nov.">
<emphasis id="B9363802E32F8C11A06DFAF4FD7CFAAF" box="[535,661,1339,1363]" italics="true" pageId="37" pageNumber="238">S. laoticum</emphasis>
</taxonomicName>
<emphasis id="B9363802E32F8C11A0E6FAF5FD32FAAF" bold="true" box="[668,731,1338,1363]" pageId="37" pageNumber="238">
<taxonomicNameLabel id="A2058579E32F8C11A0E6FAF5FD32FAAF" box="[668,731,1338,1363]" pageId="37" pageNumber="238" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
see above.
</paragraph>
</subSubSection>
<subSubSection id="C358B79BE32F8C12A2BDFA91FBC8FC11" lastPageId="38" lastPageNumber="239" pageId="37" pageNumber="238" type="description">
<paragraph id="8BFDE410E32F8C11A2BDFA91FC44FA8A" blockId="37.[151,1437,762,2011]" box="[199,941,1374,1399]" pageId="37" pageNumber="238">
<emphasis id="B9363802E32F8C11A2BDFA91FEB8FA8B" bold="true" box="[199,337,1374,1399]" pageId="37" pageNumber="238">Description</emphasis>
: based on male
<typeStatus id="54F95AB2E32F8C11A07DFA91FD80FA8B" box="[519,617,1374,1399]" pageId="37" pageNumber="238" type="holotype">holotype</typeStatus>
and
<typeStatus id="54F95AB2E32F8C11A0DAFA90FCE2FA8B" box="[672,779,1375,1399]" pageId="37" pageNumber="238" type="paratype">paratypes</typeStatus>
of both sexes.
</paragraph>
<paragraph id="8BFDE410E32F8C11A2BDFA4DFB51F9FB" blockId="37.[151,1437,762,2011]" pageId="37" pageNumber="238">
<emphasis id="B9363802E32F8C11A2BDFA4DFE9FFA67" bold="true" box="[199,374,1410,1435]" pageId="37" pageNumber="238">Measurements</emphasis>
:
<typeStatus id="54F95AB2E32F8C11A3FEFA4DFE0FFA67" box="[388,486,1410,1435]" pageId="37" pageNumber="238" type="holotype">holotype</typeStatus>
male: body length
<quantity id="4CBA49F5E32F8C11A0C7FA4CFCCDFA66" box="[701,804,1411,1435]" metricMagnitude="-2" metricUnit="m" metricValue="2.73" pageId="37" pageNumber="238" unit="mm" value="27.3">27.3 mm</quantity>
. Width, of thoracic shield =
<quantity id="4CBA49F5E32F8C11A625FA4CFB2AFA66" box="[1119,1219,1411,1435]" metricMagnitude="-2" metricUnit="m" metricValue="1.33" pageId="37" pageNumber="238" unit="mm" value="13.3">13.3 mm</quantity>
(= widest). Height, of thoracic shield =
<quantity id="4CBA49F5E32F8C11A305FA68FE3EFA42" box="[383,471,1447,1471]" metricMagnitude="-3" metricUnit="m" metricValue="7.4" pageId="37" pageNumber="238" unit="mm" value="7.4">7.4 mm</quantity>
high (= highest). Female: Body length: broken and rolled up, ca.
<quantity id="4CBA49F5E32F8C11A6B5FA68FACAFA42" box="[1231,1315,1447,1471]" metricMagnitude="-2" metricUnit="m" metricValue="3.7" pageId="37" pageNumber="238" unit="mm" value="37.0">37 mm</quantity>
. Width, of thoracic shield =
<quantity id="4CBA49F5E32F8C11A32DFA04FE4CFA1E" box="[343,421,1483,1507]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="37" pageNumber="238" unit="mm" value="18.0">18 mm</quantity>
(unclear of broadest, broken).
<emphasis id="B9363802E32F8C11A086FA05FC94FA1F" bold="true" box="[764,893,1482,1507]" pageId="37" pageNumber="238">Coloration</emphasis>
: in preserved specimens tergites black, posterior third olive. Head and collum black, anterior half of head green. Antennae and legs orange-red.
</paragraph>
<paragraph id="8BFDE410E32F8C11A2BDF9DDFA74F9B3" blockId="37.[151,1437,762,2011]" pageId="37" pageNumber="238">
<emphasis id="B9363802E32F8C11A2BDF9DDFEEEF9D7" bold="true" box="[199,263,1554,1579]" pageId="37" pageNumber="238">Head</emphasis>
: Eyes with ca. 70 ocelli. Last antennomere thickened. Apical disc with 73/71 apical cones (male), or 36/ 37 (female). All antennomeres covered by long setae. Organ of Tömösváry located inside antennal groove.
</paragraph>
<paragraph id="8BFDE410E32F8C11A2EDF995FEECF96B" blockId="37.[151,1437,762,2011]" pageId="37" pageNumber="238">
<emphasis id="B9363802E32F8C11A2EDF995FE8FF98F" bold="true" box="[151,358,1626,1651]" pageId="37" pageNumber="238">Gnathochilarium</emphasis>
: structure typical of the order. Palpi sensory cones located in several clusters.
<emphasis id="B9363802E32F8C11A690F995FA81F98F" bold="true" box="[1258,1384,1626,1651]" pageId="37" pageNumber="238">Mandibles</emphasis>
: not dissected.
</paragraph>
<paragraph id="8BFDE410E32F8C12A2BDF96DFB6CFEBC" blockId="37.[151,1437,762,2011]" lastBlockId="38.[151,1437,151,1077]" lastPageId="38" lastPageNumber="239" pageId="37" pageNumber="238">
<emphasis id="B9363802E32F8C11A2BDF96DFE6AF947" bold="true" box="[199,387,1698,1723]" pageId="37" pageNumber="238">Stigmatic plates</emphasis>
: first stigmatic plate widely rounded, apex well-rounded, slightly curved towards coxa 1 (
<figureCitation id="1379F895E32F8C11A715F96DFF25F923" captionStart="FIGURE 25" captionStartId="41.[151,250,1885,1907]" captionTargetBox="[207,1378,195,1868]" captionTargetId="figure@41.[207,1378,195,1868]" captionTargetPageId="41" captionText="FIGURE 25. Sphaerobelum schwendingeri n. sp. AC female paratype (ZMUC), DG male holotype (MHNG). A: coxa of first right leg with first stigmatic plate; B: 9th left leg; C: coxa and prefemur of second leg with vulva; D: left anterior telopod, anterior view; E: left anterior telopod, posterior view; F: right posterior telopod, anterior view; G: right posterior telopod, posterior view. Abbreviations: Cx = coxa; O = operculum of vulva; Pre = prefemur; St-Pl = stigmatic plate; Syn-Cx = syncoxite. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601125/files/figure.png" pageId="37" pageNumber="238">Fig. 25A</figureCitation>
).
<emphasis id="B9363802E32F8C11A29AF909FE60F923" bold="true" box="[224,393,1734,1759]" pageId="37" pageNumber="238">Laterotergites</emphasis>
: laterotergite 1 and 2 slightly elongated into a short, broad process.
<emphasis id="B9363802E32F8C11A6E2F909FB19F923" bold="true" box="[1176,1264,1734,1759]" pageId="37" pageNumber="238">Collum</emphasis>
: with glabrous surface, margins with few isolated setae.
<emphasis id="B9363802E32F8C11A0F8F925FCD6F8FF" bold="true" box="[642,831,1770,1795]" pageId="37" pageNumber="238">Thoracic shield</emphasis>
: surface glabrous, but covered with small pits like tergites, setae only in grooves. Shallow grooves beset with numerous long setae, slope towards groove without keels.
<emphasis id="B9363802E32F8C11A299F8FDFEABF8B7" bold="true" box="[227,322,1842,1867]" pageId="37" pageNumber="238">Tergites</emphasis>
: glabrous, shiny. Anterior third with a few setae only visible when the specimen is rolled up, producing a golden sheen. Tips of paratergites of midbody tergites projecting posteriorly.
<emphasis id="B9363802E32F8C11A6C2F899FAA7F893" bold="true" box="[1208,1358,1878,1903]" pageId="37" pageNumber="238">Endotergum</emphasis>
: inner section without any setae or spines. Middle area with a single row of sparse elliptical, cuticular impressions, distance between impressions larger than their diameter. Apically 3-4 dense rows of short marginal bristles, the longest just protruding up to half of length to tergal margin. Bristles not smooth, but with numerous small spicules.
<emphasis id="B9363802E32C8C12A2EDFF58FEC9FF4C" bold="true" box="[151,288,151,176]" pageId="38" pageNumber="239">Anal shield</emphasis>
: sexually dimorphic, in female slightly flattened, in male weakly bell-shaped. Glabrous. Underside without locking carinae.
<emphasis id="B9363802E32C8C12A3CBFF73FE00FF29" bold="true" box="[433,489,188,213]" pageId="38" pageNumber="239">Legs</emphasis>
: leg-pair 1 with 2 ventral spines, leg-pair 2 with 6, leg-pair 3 with 6. First two legpairs without an apical spine. Leg-pairs 421 with 68 ventral spines and one dorso-apical spine. In leg 9 femur 1.7 times, tarsus 3.8 times longer than wide (
<figureCitation id="1379F895E32C8C12A004FECBFD00FEE1" box="[638,745,260,285]" captionStart="FIGURE 25" captionStartId="41.[151,250,1885,1907]" captionTargetBox="[207,1378,195,1868]" captionTargetId="figure@41.[207,1378,195,1868]" captionTargetPageId="41" captionText="FIGURE 25. Sphaerobelum schwendingeri n. sp. AC female paratype (ZMUC), DG male holotype (MHNG). A: coxa of first right leg with first stigmatic plate; B: 9th left leg; C: coxa and prefemur of second leg with vulva; D: left anterior telopod, anterior view; E: left anterior telopod, posterior view; F: right posterior telopod, anterior view; G: right posterior telopod, posterior view. Abbreviations: Cx = coxa; O = operculum of vulva; Pre = prefemur; St-Pl = stigmatic plate; Syn-Cx = syncoxite. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601125/files/figure.png" pageId="38" pageNumber="239">Fig. 25B</figureCitation>
). All podomeres densely setose. Coxa with a well rounded process. Coxa process absent at first leg (
<figureCitation id="1379F895E32C8C12A01FFEE8FD22FEBC" box="[613,715,295,320]" captionStart="FIGURE 25" captionStartId="41.[151,250,1885,1907]" captionTargetBox="[207,1378,195,1868]" captionTargetId="figure@41.[207,1378,195,1868]" captionTargetPageId="41" captionText="FIGURE 25. Sphaerobelum schwendingeri n. sp. AC female paratype (ZMUC), DG male holotype (MHNG). A: coxa of first right leg with first stigmatic plate; B: 9th left leg; C: coxa and prefemur of second leg with vulva; D: left anterior telopod, anterior view; E: left anterior telopod, posterior view; F: right posterior telopod, anterior view; G: right posterior telopod, posterior view. Abbreviations: Cx = coxa; O = operculum of vulva; Pre = prefemur; St-Pl = stigmatic plate; Syn-Cx = syncoxite. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601125/files/figure.png" pageId="38" pageNumber="239">Fig. 25A</figureCitation>
) and sharp-edged at second (
<figureCitation id="1379F895E32C8C12A668FEE8FB91FEBC" box="[1042,1144,295,320]" captionStart="FIGURE 25" captionStartId="41.[151,250,1885,1907]" captionTargetBox="[207,1378,195,1868]" captionTargetId="figure@41.[207,1378,195,1868]" captionTargetPageId="41" captionText="FIGURE 25. Sphaerobelum schwendingeri n. sp. AC female paratype (ZMUC), DG male holotype (MHNG). A: coxa of first right leg with first stigmatic plate; B: 9th left leg; C: coxa and prefemur of second leg with vulva; D: left anterior telopod, anterior view; E: left anterior telopod, posterior view; F: right posterior telopod, anterior view; G: right posterior telopod, posterior view. Abbreviations: Cx = coxa; O = operculum of vulva; Pre = prefemur; St-Pl = stigmatic plate; Syn-Cx = syncoxite. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601125/files/figure.png" pageId="38" pageNumber="239">Fig. 25C</figureCitation>
).
</paragraph>
<paragraph id="8BFDE410E32C8C12A2BDFE83FB64FE51" blockId="38.[151,1437,151,1077]" pageId="38" pageNumber="239">
<emphasis id="B9363802E32C8C12A2BDFE83FE13FE99" bold="true" box="[199,506,332,357]" pageId="38" pageNumber="239">Female sexual characters</emphasis>
: Vulva large, covering mesal 2/3 of coxa, extending mesally to anterior third of prefemur (
<figureCitation id="1379F895E32C8C12A36AFEA0FE92FE74" box="[272,379,367,392]" captionStart="FIGURE 25" captionStartId="41.[151,250,1885,1907]" captionTargetBox="[207,1378,195,1868]" captionTargetId="figure@41.[207,1378,195,1868]" captionTargetPageId="41" captionText="FIGURE 25. Sphaerobelum schwendingeri n. sp. AC female paratype (ZMUC), DG male holotype (MHNG). A: coxa of first right leg with first stigmatic plate; B: 9th left leg; C: coxa and prefemur of second leg with vulva; D: left anterior telopod, anterior view; E: left anterior telopod, posterior view; F: right posterior telopod, anterior view; G: right posterior telopod, posterior view. Abbreviations: Cx = coxa; O = operculum of vulva; Pre = prefemur; St-Pl = stigmatic plate; Syn-Cx = syncoxite. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601125/files/figure.png" pageId="38" pageNumber="239">Fig. 25C</figureCitation>
). Operculum rounded, central part recessed, mesal margin broadly projecting into a sharpedged triangle slightly as higher than remaining operculum.
<emphasis id="B9363802E32C8C12A141FE5BFC36FE51" bold="true" box="[827,991,404,429]" pageId="38" pageNumber="239">Subanal plate</emphasis>
: not examined.
</paragraph>
<paragraph id="8BFDE410E32C8C12A2BDFE78FBC8FC11" blockId="38.[151,1437,151,1077]" pageId="38" pageNumber="239">
<emphasis id="B9363802E32C8C12A2BDFE78FE0BFE2C" bold="true" box="[199,482,439,464]" pageId="38" pageNumber="239">Male sexual characters</emphasis>
: gonopore large, covered by small, inconspicuous plate.
<emphasis id="B9363802E32C8C12A6FAFE78FAB1FE2C" bold="true" box="[1152,1368,439,464]" pageId="38" pageNumber="239">Anterior telopods</emphasis>
(
<figureCitation id="1379F895E32C8C12A710FE78FF11FE08" captionStart="FIGURE 25" captionStartId="41.[151,250,1885,1907]" captionTargetBox="[207,1378,195,1868]" captionTargetId="figure@41.[207,1378,195,1868]" captionTargetPageId="41" captionText="FIGURE 25. Sphaerobelum schwendingeri n. sp. AC female paratype (ZMUC), DG male holotype (MHNG). A: coxa of first right leg with first stigmatic plate; B: 9th left leg; C: coxa and prefemur of second leg with vulva; D: left anterior telopod, anterior view; E: left anterior telopod, posterior view; F: right posterior telopod, anterior view; G: right posterior telopod, posterior view. Abbreviations: Cx = coxa; O = operculum of vulva; Pre = prefemur; St-Pl = stigmatic plate; Syn-Cx = syncoxite. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601125/files/figure.png" pageId="38" pageNumber="239">Figs 25D, E</figureCitation>
): Consisting of three telopoditomere above syncoxite. Last telopoditomeres completely fused. Telopoditomere 1 rectangular, slightly longer than wide. Telopoditomere 2 large, as long as telopoditomere 3. Process of telopoditomere 2 located posteriorly, apex visible in anterior view. Process of telopoditomere 2 wide, projecting to length of telopoditomere 3, strongly curved, apically with a well-rounded tip. Telopoditomere 3 massive, cylindrical, straight, apically slightly tapering. Posterior side with three large spines. In anterior view all telopoditomeres covered by long setae, in posterior view telopoditomeres 2 and 3 only with marginal setae.
<emphasis id="B9363802E32C8C12A2EDFD7BFE9CFD31" bold="true" box="[151,373,692,717]" pageId="38" pageNumber="239">Posterior telopods</emphasis>
(
<figureCitation id="1379F895E32C8C12A3F3FD7BFDF2FD30" box="[393,539,692,717]" captionStart="FIGURE 25" captionStartId="41.[151,250,1885,1907]" captionTargetBox="[207,1378,195,1868]" captionTargetId="figure@41.[207,1378,195,1868]" captionTargetPageId="41" captionText="FIGURE 25. Sphaerobelum schwendingeri n. sp. AC female paratype (ZMUC), DG male holotype (MHNG). A: coxa of first right leg with first stigmatic plate; B: 9th left leg; C: coxa and prefemur of second leg with vulva; D: left anterior telopod, anterior view; E: left anterior telopod, posterior view; F: right posterior telopod, anterior view; G: right posterior telopod, posterior view. Abbreviations: Cx = coxa; O = operculum of vulva; Pre = prefemur; St-Pl = stigmatic plate; Syn-Cx = syncoxite. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601125/files/figure.png" pageId="38" pageNumber="239">Figs 25F, G</figureCitation>
): telopoditomere 1 rectangular, slightly longer than wide. Immovable finger (process of telopoditomere 2) shorter than movable finger, consisting of telopoditomeres 3 and 4. Immovable finger with a spine in anterior view, a characteristic, distally swollen apex, slightly swollen. Apically broader than base, at tip in anterior view with a few large, crenulated teeth. Telopoditomere 3 elongated, at midpoint with a triangular extension carrying a spine towards immovable finger. Telopoditomere 4 tapering, anterior half strongly curved 90° towards immovable finger, which it overlaps. At inner margin with two larger spines. Entire telopoditomere 1 at both sides covered with long setae. Telopoditomere 2 anterior side completely covered by setae except for tip of immovable finger, posterior side glabrous except for lateral margin. Telopoditomere 3 only basally in anterior view with setae, remaining part, as well as telopoditomere 4 glabrous.
</paragraph>
</subSubSection>
<subSubSection id="C358B79BE32C8C12A2BDFC38FD81FBC9" pageId="38" pageNumber="239" type="etymology">
<paragraph id="8BFDE410E32C8C12A2BDFC38FD81FBC9" blockId="38.[151,1437,151,1077]" pageId="38" pageNumber="239">
<emphasis id="B9363802E32C8C12A2BDFC38FE48FBEC" bold="true" box="[199,417,1015,1040]" pageId="38" pageNumber="239">Derivatio nominis</emphasis>
: schwendingeri, noun in apposition, after the Arachnologist Peter Schwendinger, great collector of arthropods in Southeast Asia.
</paragraph>
</subSubSection>
</treatment>
</document>