507 lines
62 KiB
XML
507 lines
62 KiB
XML
<document id="3AEE9E4F9043B7CF4B305D8BF24E80B8" ID-CLB-Dataset="28149" ID-DOI="10.11646/zootaxa.4563.2.1" ID-GBIF-Dataset="e6037c5d-a96f-41e5-8e70-71f2659659e2" ID-ISSN="1175-5326" ID-Zenodo-Dep="2601072" ID-ZooBank="CF79B01B-8B5F-4B3A-B642-2CADE4B339AF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553171433896" checkinUser="plazi" docAuthor="Wesener, Thomas" docDate="2019" docId="03EB5506E32A8C11A2EDF8E6FF1BFDC1" docLanguage="en" docName="zootaxa.4563.2.1.pdf" docOrigin="Zootaxa 4563 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Sphaerobelum laoticum Wesener 2019, new species" docType="treatment" docVersion="9" lastPageNumber="238" masterDocId="FFD22D7EE30A8C34A27AFFCFFFE9FFFC" masterDocTitle="First records of giant pill-millipedes from Laos (Diplopoda, Sphaerotheriida, Zephroniidae)" masterLastPageNumber="248" masterPageNumber="201" pageNumber="233" updateTime="1698703176884" updateUser="ExternalLinkService">
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<mods:mods id="35406DBD45BDBAA904F4170ED92A0A51" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="08F6D18AE912759E082F6B48FBBD7A5A">
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<mods:title id="A9A423CB94DB1806A487982C336429AD">First records of giant pill-millipedes from Laos (Diplopoda, Sphaerotheriida, Zephroniidae)</mods:title>
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<mods:name id="A9C03CC9954560ECE5C29624DEE683F5" type="personal">
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<mods:namePart id="6B0216FF4B6275DDC3CB9814696152A9">Wesener, Thomas</mods:namePart>
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<mods:typeOfResource id="AC1296C76931C1EF7FC477FA62B06991">text</mods:typeOfResource>
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<mods:title id="366AC16552A846643E38C7B847CDCA83">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="CD05C026D676F3C00481E7774FB8661E">
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<mods:date id="9ECC19C7729827AA749755D27CF88E60">2019</mods:date>
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<mods:number id="52DFCDF2EB71AA8350EF584D24A6595E">2019-03-01</mods:number>
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<mods:number id="A142692E5A8196C4F8D8F4DBD7568267">4563</mods:number>
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<mods:number id="2A5879F3C64AA0ACA92C2A91F351BEA1">2</mods:number>
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<mods:identifier id="E9E638CFE515587706F123BEB25416A3" type="DOI">10.11646/zootaxa.4563.2.1</mods:identifier>
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<treatment id="03EB5506E32A8C11A2EDF8E6FF1BFDC1" ID-DOI="http://doi.org/10.5281/zenodo.5934538" ID-GBIF-Taxon="154474546" ID-Zenodo-Dep="5934538" LSID="urn:lsid:plazi:treatment:03EB5506E32A8C11A2EDF8E6FF1BFDC1" httpUri="http://treatment.plazi.org/id/03EB5506E32A8C11A2EDF8E6FF1BFDC1" lastPageId="37" lastPageNumber="238" pageId="32" pageNumber="233">
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<subSubSection id="C358B79BE32A8C14A2EDF8E6FDA6F8BF" box="[151,591,1833,1859]" pageId="32" pageNumber="233" type="nomenclature">
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<paragraph id="8BFDE410E32A8C14A2EDF8E6FDA6F8BF" blockId="32.[151,591,1833,1893]" box="[151,591,1833,1859]" pageId="32" pageNumber="233">
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<heading id="D0B5537CE32A8C14A2EDF8E6FDA6F8BF" bold="true" box="[151,591,1833,1859]" fontSize="11" level="1" pageId="32" pageNumber="233" reason="1">
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<emphasis id="B9363802E32A8C14A2EDF8E6FDA6F8BF" bold="true" box="[151,591,1833,1859]" pageId="32" pageNumber="233">
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||
<taxonomicName id="4C429F93E32A8C14A2EDF8E6FE51F8BF" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[151,440,1833,1859]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="32" pageNumber="233" phylum="Arthropoda" rank="species" species="laoticum" status="sp. nov.">
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<emphasis id="B9363802E32A8C14A2EDF8E6FE51F8BF" bold="true" box="[151,440,1833,1859]" italics="true" pageId="32" pageNumber="233">Sphaerobelum laoticum</emphasis>
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||
</taxonomicName>
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<taxonomicNameLabel id="A2058579E32A8C14A3C5F8E6FDA6F8BF" box="[447,591,1833,1859]" pageId="32" pageNumber="233" rank="species">new species</taxonomicNameLabel>
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||
</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C358B79BE32A8C14A2EDF883FE8DF898" box="[151,356,1868,1893]" pageId="32" pageNumber="233" type="description">
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<paragraph id="8BFDE410E32A8C14A2EDF883FE8DF898" blockId="32.[151,591,1833,1893]" box="[151,356,1868,1893]" pageId="32" pageNumber="233">
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<figureCitation id="1379F895E32A8C14A2EDF883FEE2F899" box="[151,267,1868,1893]" captionStart="FIGURE 6" captionStartId="11.[151,250,1916,1938]" captionTargetBox="[150,1423,987,1897]" captionTargetId="figure@11.[150,1436,983,1903]" captionTargetPageId="11" captionText="FIGURE 6. Habitus multi-layer stacked photographs of Sphaerobelum species. A: Sphaerobelum bolavensis n. sp.; B: Sphaerobelum phouloei n. sp.; C: Sphaerobelum denticulatum n. sp.; D: Sphaerobelum spinatum n. sp.; E: Sphaerobelum lachneeis n. sp.; F: Sphaerobelum peterjaegeri n. sp.; G: Sphaerobelum nigrum n. sp.; H: Sphaerobelum splendidum n. sp.; I: Sphaerobelum laoticum n. sp. Not to scale." httpUri="https://zenodo.org/record/2601084/files/figure.png" pageId="32" pageNumber="233">
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Figures
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<date id="FFFCC2D0E32A8C14A288F883FEE2F899" box="[242,267,1868,1893]" pageId="32" pageNumber="233">6I</date>
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</figureCitation>
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,
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<figureCitation id="1379F895E32A8C14A36DF883FEDCF898" box="[279,309,1868,1892]" captionStart="FIGURE 23" captionStartId="34.[151,250,1968,1990]" captionTargetBox="[210,1381,193,1950]" captionTargetId="figure@34.[210,1381,193,1950]" captionTargetPageId="34" captionText="FIGURE 23. Sphaerobelum laoticum n. sp., holotype (SMF); A: coxa of first left leg with first stigmatic plate; B: 9th left leg. Abbreviations: Cx = coxa; St-Pl = stigmatic plate. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601121/files/figure.png" pageId="32" pageNumber="233">23</figureCitation>
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,
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<figureCitation id="1379F895E32A8C14A33BF883FE89F898" box="[321,352,1868,1892]" captionStart="FIGURE 24" captionStartId="35.[151,250,1945,1967]" captionTargetBox="[161,1430,193,1928]" captionTargetId="figure@35.[161,1430,193,1928]" captionTargetPageId="35" captionText="FIGURE 24. Sphaerobelum laoticum n. sp., holotype (SMF); A: coxa of second left leg with gonopore; B: left anterior telopod, anterior view; C: left anterior telopod, posterior view; D: left anterior telopod, lateral view; E: left posterior telopod, anterior view; F: left posterior telopod, posterior view. Abbreviations: Cx = coxa; Syn-Cx = syncoxite. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601123/files/figure.png" pageId="32" pageNumber="233">24</figureCitation>
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.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C358B79BE32A8C10A2EDF85BFB6CFEE7" lastPageId="36" lastPageNumber="237" pageId="32" pageNumber="233" type="materials_examined">
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||
<paragraph id="8BFDE410E32A8C14A2EDF85BFED7F809" blockId="32.[151,1436,1940,2037]" pageId="32" pageNumber="233">
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||
<emphasis id="B9363802E32A8C14A2EDF85BFE6AF851" bold="true" box="[151,387,1940,1965]" pageId="32" pageNumber="233">Material examined:</emphasis>
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Type material. 1 M
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<typeStatus id="54F95AB2E32A8C14A010F85BFD25F851" box="[618,716,1940,1965]" pageId="32" pageNumber="233" type="holotype">holotype</typeStatus>
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(SMF) from
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<collectingCountry id="F355A480E32A8C14A11EF85AFC74F850" box="[868,925,1941,1964]" name="Laos" pageId="32" pageNumber="233">Laos</collectingCountry>
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,
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<collectingRegion id="49862AF2E32A8C14A1D3F85BFBFDF851" box="[937,1044,1940,1965]" country="Laos" name="Viangchan" pageId="32" pageNumber="233">Vientiane</collectingRegion>
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Province, Vang Vieng, W. of Nam Song (
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<geoCoordinate id="EE7682D7E32A8C14A29AF877FE63F82D" box="[224,394,1976,2001]" direction="north" orientation="latitude" pageId="32" pageNumber="233" precision="1" value="18.929682">N18°55'46.86"</geoCoordinate>
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,
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<geoCoordinate id="EE7682D7E32A8C14A3EDF877FDA5F82D" box="[407,588,1976,2001]" direction="east" orientation="longitude" pageId="32" pageNumber="233" precision="1" value="102.34912">102°20'56.82"E</geoCoordinate>
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),
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<quantity id="4CBA49F5E32A8C14A01AF877FD43F82C" box="[608,682,1976,2000]" metricMagnitude="2" metricUnit="m" metricValue="3.2399999999999998" pageId="32" pageNumber="233" unit="m" value="324.0">324 m</quantity>
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, Tham Nam Or Khem, in cave, leg. by hand,
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<date id="FFFCC2D0E32A8C14A6D5F877FAD3F82C" box="[1199,1338,1976,2000]" pageId="32" pageNumber="233" value="2014-07-28">28.VII.2014</date>
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, P. Jäger & S. Münnich.
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</paragraph>
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<caption id="DF3DB498E32B8C15A2EDF84DFBD1F808" httpUri="https://zenodo.org/record/2601119/files/figure.png" pageId="33" pageNumber="234" startId="33.[151,250,1922,1944]" targetBox="[168,1421,193,1901]" targetPageId="33">
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<paragraph id="8BFDE410E32B8C15A2EDF84DFBD1F808" blockId="33.[151,1437,1922,2036]" pageId="33" pageNumber="234">
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<emphasis id="B9363802E32B8C15A2EDF84DFEF3F864" bold="true" box="[151,282,1922,1944]" pageId="33" pageNumber="234">FIGURE 22</emphasis>
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.
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<taxonomicName id="4C429F93E32B8C15A352F84DFDDCF864" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[296,565,1922,1944]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="33" pageNumber="234" phylum="Arthropoda" rank="species" species="splendidum" status="sp. nov.">
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||
<emphasis id="B9363802E32B8C15A352F84DFDDCF864" box="[296,565,1922,1944]" italics="true" pageId="33" pageNumber="234">Sphaerobelum splendidum</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2058579E32B8C15A046F84DFD9EF864" box="[572,631,1922,1944]" pageId="33" pageNumber="234" rank="species">
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<emphasis id="B9363802E32B8C15A046F84DFD9EF864" bold="true" box="[572,631,1922,1944]" pageId="33" pageNumber="234">n. sp.</emphasis>
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</taxonomicNameLabel>
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, holotype (SMF);
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<emphasis id="B9363802E32B8C15A14CF84DFCA1F864" bold="true" box="[822,840,1922,1944]" pageId="33" pageNumber="234">A</emphasis>
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: coxa of first left leg with first stigmatic plate;
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<emphasis id="B9363802E32B8C15A749F84DFAADF864" bold="true" box="[1331,1348,1922,1944]" pageId="33" pageNumber="234">B</emphasis>
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: 9th left leg;
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<emphasis id="B9363802E32B8C15A2B9F86EFF3CF84B" bold="true" box="[195,213,1953,1975]" pageId="33" pageNumber="234">C</emphasis>
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: coxa of second left leg with gonopore;
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<emphasis id="B9363802E32B8C15A016F86EFD97F84B" bold="true" box="[620,638,1953,1975]" pageId="33" pageNumber="234">D</emphasis>
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: left anterior telopod, anterior view;
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<emphasis id="B9363802E32B8C15A189F86EFBE5F84B" bold="true" box="[1011,1036,1953,1975]" pageId="33" pageNumber="234">E:</emphasis>
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left anterior telopod, posterior view;
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<emphasis id="B9363802E32B8C15A7FFF86EFA7DF84B" bold="true" box="[1413,1428,1953,1975]" pageId="33" pageNumber="234">F</emphasis>
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: left anterior telopod, lateral view;
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<emphasis id="B9363802E32B8C15A075F870FDCAF829" bold="true" box="[527,547,1983,2005]" pageId="33" pageNumber="234">G</emphasis>
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: right posterior telopod, anterior view;
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<emphasis id="B9363802E32B8C15A1ADF870FC02F829" bold="true" box="[983,1003,1983,2005]" pageId="33" pageNumber="234">H</emphasis>
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: right posterior telopod, posterior view. Abbreviations:
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<emphasis id="B9363802E32B8C15A34EF811FEBAF808" bold="true" box="[308,339,2014,2036]" pageId="33" pageNumber="234">Cx</emphasis>
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= coxa;
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<emphasis id="B9363802E32B8C15A3D1F811FE37F808" bold="true" box="[427,478,2014,2036]" pageId="33" pageNumber="234">St-Pl</emphasis>
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= stigmatic plate;
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<emphasis id="B9363802E32B8C15A0E6F811FD05F808" bold="true" box="[668,748,2014,2036]" pageId="33" pageNumber="234">Syn-Cx</emphasis>
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= syncoxite. Scale bars = 1 mm.
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</paragraph>
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</caption>
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<caption id="DF3DB498E3288C16A2EDF87FFCB6F818" httpUri="https://zenodo.org/record/2601121/files/figure.png" pageId="34" pageNumber="235" startId="34.[151,250,1968,1990]" targetBox="[210,1381,193,1950]" targetPageId="34">
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<paragraph id="8BFDE410E3288C16A2EDF87FFCB6F818" blockId="34.[151,1437,1968,2020]" pageId="34" pageNumber="235">
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<emphasis id="B9363802E3288C16A2EDF87FFEF0F83A" bold="true" box="[151,281,1968,1990]" pageId="34" pageNumber="235">FIGURE 23</emphasis>
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.
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<taxonomicName id="4C429F93E3288C16A35CF87FFDFEF83A" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[294,535,1968,1990]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="34" pageNumber="235" phylum="Arthropoda" rank="species" species="laoticum" status="sp. nov.">
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<emphasis id="B9363802E3288C16A35CF87FFDFEF83A" box="[294,535,1968,1990]" italics="true" pageId="34" pageNumber="235">Sphaerobelum laoticum</emphasis>
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</taxonomicName>
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<emphasis id="B9363802E3288C16A067F87FFDBEF83A" bold="true" box="[541,599,1968,1990]" pageId="34" pageNumber="235">
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<taxonomicNameLabel id="A2058579E3288C16A067F87FFDBEF83A" box="[541,599,1968,1990]" pageId="34" pageNumber="235" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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, holotype (SMF);
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<emphasis id="B9363802E3288C16A16EF87FFCCFF83A" bold="true" box="[788,806,1968,1990]" pageId="34" pageNumber="235">A</emphasis>
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: coxa of first left leg with first stigmatic plate;
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<emphasis id="B9363802E3288C16A773F87FFAF3F83A" bold="true" box="[1289,1306,1968,1990]" pageId="34" pageNumber="235">B</emphasis>
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: 9th left leg. Abbreviations:
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<emphasis id="B9363802E3288C16A34EF801FEBAF818" bold="true" box="[308,339,1998,2020]" pageId="34" pageNumber="235">Cx</emphasis>
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= coxa;
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<emphasis id="B9363802E3288C16A3D1F801FE37F818" bold="true" box="[427,478,1998,2020]" pageId="34" pageNumber="235">St-Pl</emphasis>
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= stigmatic plate. Scale bars = 1 mm.
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</paragraph>
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</caption>
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<caption id="DF3DB498E3298C17A2EDF856FA9CF80F" httpUri="https://zenodo.org/record/2601123/files/figure.png" pageId="35" pageNumber="236" startId="35.[151,250,1945,1967]" targetBox="[161,1430,193,1928]" targetPageId="35">
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||
<paragraph id="8BFDE410E3298C17A2EDF856FA9CF80F" blockId="35.[151,1436,1945,2035]" pageId="35" pageNumber="236">
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||
<emphasis id="B9363802E3298C17A2EDF856FEF5F853" bold="true" box="[151,284,1945,1967]" pageId="35" pageNumber="236">FIGURE 24</emphasis>
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.
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<taxonomicName id="4C429F93E3298C17A354F856FDCBF853" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[302,546,1945,1967]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="35" pageNumber="236" phylum="Arthropoda" rank="species" species="laoticum" status="sp. nov.">
|
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<emphasis id="B9363802E3298C17A354F856FDCBF853" box="[302,546,1945,1967]" italics="true" pageId="35" pageNumber="236">Sphaerobelum laoticum</emphasis>
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||
</taxonomicName>
|
||
<emphasis id="B9363802E3298C17A056F856FD83F853" bold="true" box="[556,618,1945,1967]" pageId="35" pageNumber="236">
|
||
<taxonomicNameLabel id="A2058579E3298C17A056F856FD83F853" box="[556,618,1945,1967]" pageId="35" pageNumber="236" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
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, holotype (SMF);
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<emphasis id="B9363802E3298C17A148F856FCADF853" bold="true" box="[818,836,1945,1967]" pageId="35" pageNumber="236">A</emphasis>
|
||
: coxa of second left leg with gonopore;
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||
<emphasis id="B9363802E3298C17A685F856FAF9F853" bold="true" box="[1279,1296,1945,1967]" pageId="35" pageNumber="236">B</emphasis>
|
||
: left anterior telopod, anterior view;
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||
<emphasis id="B9363802E3298C17A3FCF875FE48F82C" bold="true" box="[390,417,1978,2000]" pageId="35" pageNumber="236">C:</emphasis>
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left anterior telopod, posterior view;
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<emphasis id="B9363802E3298C17A15BF875FCDDF82C" bold="true" box="[801,820,1978,2000]" pageId="35" pageNumber="236">D</emphasis>
|
||
: left anterior telopod, lateral view;
|
||
<emphasis id="B9363802E3298C17A6DAF875FB58F82C" bold="true" box="[1184,1201,1978,2000]" pageId="35" pageNumber="236">E</emphasis>
|
||
: left posterior telopod, anterior view;
|
||
<emphasis id="B9363802E3298C17A350F812FED0F80F" bold="true" box="[298,313,2013,2035]" pageId="35" pageNumber="236">F</emphasis>
|
||
: left posterior telopod, posterior view. Abbreviations:
|
||
<emphasis id="B9363802E3298C17A11BF812FC69F80F" bold="true" box="[865,896,2013,2035]" pageId="35" pageNumber="236">Cx</emphasis>
|
||
= coxa;
|
||
<emphasis id="B9363802E3298C17A1A2F812FBC1F80F" bold="true" box="[984,1064,2013,2035]" pageId="35" pageNumber="236">Syn-Cx</emphasis>
|
||
= syncoxite. Scale bars = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BFDE410E32E8C10A2BDFF58FB6CFEE7" blockId="36.[151,1437,151,2021]" pageId="36" pageNumber="237">
|
||
<emphasis id="B9363802E32E8C10A2BDFF58FDE0FF4C" bold="true" box="[199,521,151,176]" pageId="36" pageNumber="237">Other material examined:</emphasis>
|
||
1 imm. M (SMF) from
|
||
<collectingCountry id="F355A480E32E8C10A14DFF57FC99FF53" box="[823,880,152,175]" name="Laos" pageId="36" pageNumber="237">Laos</collectingCountry>
|
||
, Vientinae Province, Van Vieng, Ban Sok Say (
|
||
<geoCoordinate id="EE7682D7E32E8C10A2E5FF73FED5FF28" box="[159,316,188,212]" direction="north" orientation="latitude" pageId="36" pageNumber="237" precision="1" value="19.022305">N19°1'20.30"</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE7682D7E32E8C10A33DFF74FE15FF28" box="[327,508,187,212]" direction="east" orientation="longitude" pageId="36" pageNumber="237" precision="1" value="102.448364">102°26'54.11"E</geoCoordinate>
|
||
),
|
||
<quantity id="4CBA49F5E32E8C10A075FF74FDB1FF2F" box="[527,600,187,212]" metricMagnitude="2" metricUnit="m" metricValue="2.95" pageId="36" pageNumber="237" unit="m" value="295.0">295 m</quantity>
|
||
, in vegetation, leg. by hand, P. Jäger & S. Packheiser,
|
||
<date id="FFFCC2D0E32E8C10A6CCFF73FAABFF28" box="[1206,1346,188,212]" pageId="36" pageNumber="237" value="2018-07-23">23.VII.2018</date>
|
||
.
|
||
<materialsCitation id="3B2AEE4DE32E8C10A737FF73FB6BFEE7" ID-GBIF-Occurrence="2013952570" collectingDate="2018-07-23" collectionCode="SMF" collectorName="Van Vieng & P. Jager & S. Packheiser" country="Laos" county="Tham Loup" elevation="256" latitude="19.039167" location="Vientinae Province" longLatPrecision="1" longitude="102.42371" municipality="Tham Hoi" pageId="36" pageNumber="237" specimenCount="1">
|
||
1 imm. F (
|
||
<collectionCode id="ED537CD5E32E8C10A2C0FF10FF1CFF0B" box="[186,245,223,247]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="36" pageNumber="237">SMF</collectionCode>
|
||
) from
|
||
<collectingCountry id="F355A480E32E8C10A336FF2FFE6CFF0B" box="[332,389,224,247]" name="Laos" pageId="36" pageNumber="237">Laos</collectingCountry>
|
||
,
|
||
<location id="8E9DB2CBE32E8C10A3EFFF11FD9AFF0B" LSID="urn:lsid:plazi:treatment:03EB5506E32A8C11A2EDF8E6FF1BFDC1:8E9DB2CBE32E8C10A3EFFF11FD9AFF0B" box="[405,627,222,247]" country="Laos" county="Tham Loup" latitude="19.039167" longLatPrecision="1" longitude="102.42371" municipality="Tham Hoi" name="Vientinae Province" pageId="36" pageNumber="237">Vientinae Province</location>
|
||
,
|
||
<location id="8E9DB2CBE32E8C10A0F9FF2FFCE8FF0B" LSID="urn:lsid:plazi:treatment:03EB5506E32A8C11A2EDF8E6FF1BFDC1:8E9DB2CBE32E8C10A0F9FF2FFCE8FF0B" box="[643,769,222,247]" country="Laos" county="Tham Loup" latitude="19.039167" longLatPrecision="1" longitude="102.42371" municipality="Tham Hoi" name="Van Vieng" pageId="36" pageNumber="237">
|
||
<collectorName id="26B781C6E32E8C10A0F9FF2FFCE8FF0B" box="[643,769,222,247]" pageId="36" pageNumber="237">Van Vieng</collectorName>
|
||
</location>
|
||
, N of
|
||
<location id="8E9DB2CBE32E8C10A12FFF2FFC0AFF0B" LSID="urn:lsid:plazi:treatment:03EB5506E32A8C11A2EDF8E6FF1BFDC1:8E9DB2CBE32E8C10A12FFF2FFC0AFF0B" box="[853,995,222,247]" country="Laos" county="Tham Loup" latitude="19.039167" longLatPrecision="1" longitude="102.42371" municipality="Tham Hoi" name="Ban Phoxay" pageId="36" pageNumber="237">Ban Phoxay</location>
|
||
,
|
||
<collectingMunicipality id="6B997E6AE32E8C10A189FF11FB83FF0B" box="[1011,1130,222,247]" pageId="36" pageNumber="237">Tham Hoi</collectingMunicipality>
|
||
and
|
||
<collectingCounty id="629C9C9CE32E8C10A6D1FF11FADEFF0B" box="[1195,1335,222,247]" pageId="36" pageNumber="237">Tham Loup</collectingCounty>
|
||
, in cave (
|
||
<geoCoordinate id="EE7682D7E32E8C10A2E5FECDFED8FEE7" box="[159,305,258,283]" direction="north" orientation="latitude" pageId="36" pageNumber="237" precision="1" value="19.039167">N19°02.350'</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE7682D7E32E8C10A346FECDFE3EFEE7" box="[316,471,258,283]" direction="east" orientation="longitude" pageId="36" pageNumber="237" precision="1" value="102.42371">102°25.423'E</geoCoordinate>
|
||
),
|
||
<quantity id="4CBA49F5E32E8C10A391FECDFDDDFEE6" box="[491,564,258,283]" metricMagnitude="2" metricUnit="m" metricValue="2.56" pageId="36" pageNumber="237" unit="m" value="256.0">
|
||
<elevation id="006F0323E32E8C10A391FECDFDDDFEE6" box="[491,564,258,283]" metricMagnitude="2" metricUnit="m" metricValue="2.56" pageId="36" pageNumber="237" unit="m" value="256.0">256 m</elevation>
|
||
</quantity>
|
||
, leg. by hand,
|
||
<collectorName id="26B781C6E32E8C10A0A2FECCFCC4FEE7" box="[728,813,259,283]" pageId="36" pageNumber="237">P. Jäger</collectorName>
|
||
&
|
||
<collectorName id="26B781C6E32E8C10A12BFECCFC03FEE7" box="[849,1002,258,283]" pageId="36" pageNumber="237">S. Packheiser</collectorName>
|
||
,
|
||
<date id="FFFCC2D0E32E8C10A18FFECCFB6BFEE7" box="[1013,1154,259,283]" pageId="36" pageNumber="237" value="2018-07-23">
|
||
<collectingDate id="EFB83B38E32E8C10A18FFECCFB6BFEE7" box="[1013,1154,259,283]" pageId="36" pageNumber="237" value="2018-07-23">23.VII.2018</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C358B79BE32E8C10A2BDFEEAFCCFFD14" pageId="36" pageNumber="237" type="diagnosis">
|
||
<paragraph id="8BFDE410E32E8C10A2BDFEEAFCCFFD14" blockId="36.[151,1437,151,2021]" pageId="36" pageNumber="237">
|
||
<emphasis id="B9363802E32E8C10A2BDFEEAFED3FEC2" bold="true" box="[199,314,293,318]" pageId="36" pageNumber="237">Diagnosis</emphasis>
|
||
:
|
||
<taxonomicName id="4C429F93E32E8C10A331FEE9FE22FEC2" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[331,459,294,318]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="36" pageNumber="237" phylum="Arthropoda" rank="species" species="laoticum" status="sp. nov.">
|
||
<emphasis id="B9363802E32E8C10A331FEE9FE22FEC2" box="[331,459,294,318]" italics="true" pageId="36" pageNumber="237">S. laoticum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9363802E32E8C10A3AEFEEAFDFCFEC2" bold="true" box="[468,533,293,318]" pageId="36" pageNumber="237">
|
||
<taxonomicNameLabel id="A2058579E32E8C10A3AEFEEAFDFCFEC2" box="[468,533,293,318]" pageId="36" pageNumber="237" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
belongs to a group of
|
||
<taxonomicName id="4C429F93E32E8C10A166FEE9FC54FEC2" authorityName="Verhoeff" authorityYear="1924" box="[796,957,294,318]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="36" pageNumber="237" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9363802E32E8C10A166FEE9FC54FEC2" box="[796,957,294,318]" italics="true" pageId="36" pageNumber="237">Sphaerobelum</emphasis>
|
||
</taxonomicName>
|
||
species in which the mesal margin of the femur is unmodified (
|
||
<figureCitation id="1379F895E32E8C10A3F1FE86FE1BFE9D" box="[395,498,329,354]" captionStart="FIGURE 23" captionStartId="34.[151,250,1968,1990]" captionTargetBox="[210,1381,193,1950]" captionTargetId="figure@34.[210,1381,193,1950]" captionTargetPageId="34" captionText="FIGURE 23. Sphaerobelum laoticum n. sp., holotype (SMF); A: coxa of first left leg with first stigmatic plate; B: 9th left leg. Abbreviations: Cx = coxa; St-Pl = stigmatic plate. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601121/files/figure.png" pageId="36" pageNumber="237">Fig. 23B</figureCitation>
|
||
).
|
||
<taxonomicName id="4C429F93E32E8C10A07FFE86FD6DFE9D" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[517,644,329,353]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="36" pageNumber="237" phylum="Arthropoda" rank="species" species="laoticum" status="sp. nov.">
|
||
<emphasis id="B9363802E32E8C10A07FFE86FD6DFE9D" box="[517,644,329,353]" italics="true" pageId="36" pageNumber="237">S. laoticum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9363802E32E8C10A0F1FE86FD23FE9E" bold="true" box="[651,714,329,354]" pageId="36" pageNumber="237">
|
||
<taxonomicNameLabel id="A2058579E32E8C10A0F1FE86FD23FE9E" box="[651,714,329,354]" pageId="36" pageNumber="237" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
shares only with
|
||
<taxonomicName id="4C429F93E32E8C10A1EAFE86FBF9FE9D" authorityName="Verhoeff" authorityYear="1924" box="[912,1040,329,353]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="36" pageNumber="237" phylum="Arthropoda" rank="species" species="hirsutum">
|
||
<emphasis id="B9363802E32E8C10A1EAFE86FBF9FE9D" box="[912,1040,329,353]" italics="true" pageId="36" pageNumber="237">S. hirsutum</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C429F93E32E8C10A667FE86FB51FE9D" authorityName="Verhoeff" authorityYear="1924" box="[1053,1208,329,353]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="36" pageNumber="237" phylum="Arthropoda" rank="species" species="clavigerum">
|
||
<emphasis id="B9363802E32E8C10A667FE86FB51FE9D" box="[1053,1208,329,353]" italics="true" pageId="36" pageNumber="237">S. clavigerum</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C429F93E32E8C10A6BFFE86FADBFE9D" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[1221,1330,329,353]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="36" pageNumber="237" phylum="Arthropoda" rank="species" species="nigrum" status="sp. nov.">
|
||
<emphasis id="B9363802E32E8C10A6BFFE86FADBFE9D" box="[1221,1330,329,353]" italics="true" pageId="36" pageNumber="237">S. nigrum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2058579E32E8C10A740FE86FA91FE9E" box="[1338,1400,329,354]" pageId="36" pageNumber="237" rank="species">
|
||
<emphasis id="B9363802E32E8C10A740FE86FA91FE9E" bold="true" box="[1338,1400,329,354]" pageId="36" pageNumber="237">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C429F93E32E8C10A7FDFE86FEF1FE79" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="36" pageNumber="237" phylum="Arthropoda" rank="species" species="splendidum" status="sp. nov.">
|
||
<emphasis id="B9363802E32E8C10A7FDFE86FEF1FE79" italics="true" pageId="36" pageNumber="237">S. splendidum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9363802E32E8C10A35EFEA3FE8EFE79" bold="true" box="[292,359,364,389]" pageId="36" pageNumber="237">
|
||
<taxonomicNameLabel id="A2058579E32E8C10A35EFEA3FE8EFE79" box="[292,359,364,389]" pageId="36" pageNumber="237" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C429F93E32E8C10A3DDFEA2FD85FE79" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[423,620,365,389]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="36" pageNumber="237" phylum="Arthropoda" rank="species" species="schwendingeri" status="sp. nov.">
|
||
<emphasis id="B9363802E32E8C10A3DDFEA2FD85FE79" box="[423,620,365,389]" italics="true" pageId="36" pageNumber="237">S. schwendingeri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9363802E32E8C10A00DFEA3FD52FE79" bold="true" box="[631,699,364,389]" pageId="36" pageNumber="237">
|
||
<taxonomicNameLabel id="A2058579E32E8C10A00DFEA3FD52FE79" box="[631,699,364,389]" pageId="36" pageNumber="237" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
the presence of a "hook" at telopoditomere 4 of the posterior telopods (
|
||
<figureCitation id="1379F895E32E8C10A372FE5FFE7DFE54" box="[264,404,400,425]" captionStart="FIGURE 24" captionStartId="35.[151,250,1945,1967]" captionTargetBox="[161,1430,193,1928]" captionTargetId="figure@35.[161,1430,193,1928]" captionTargetPageId="35" captionText="FIGURE 24. Sphaerobelum laoticum n. sp., holotype (SMF); A: coxa of second left leg with gonopore; B: left anterior telopod, anterior view; C: left anterior telopod, posterior view; D: left anterior telopod, lateral view; E: left posterior telopod, anterior view; F: left posterior telopod, posterior view. Abbreviations: Cx = coxa; Syn-Cx = syncoxite. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601123/files/figure.png" pageId="36" pageNumber="237">Figs 24E, F</figureCitation>
|
||
).
|
||
<taxonomicName id="4C429F93E32E8C10A3D1FE5FFDC5FE54" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[427,556,400,424]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="36" pageNumber="237" phylum="Arthropoda" rank="species" species="laoticum" status="sp. nov.">
|
||
<emphasis id="B9363802E32E8C10A3D1FE5FFDC5FE54" box="[427,556,400,424]" italics="true" pageId="36" pageNumber="237">S. laoticum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9363802E32E8C10A04FFE5FFD9EFE55" bold="true" box="[565,631,400,425]" pageId="36" pageNumber="237">
|
||
<taxonomicNameLabel id="A2058579E32E8C10A04FFE5FFD9EFE55" box="[565,631,400,425]" pageId="36" pageNumber="237" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="4C429F93E32E8C10A16EFE5FFC7EFE54" authorityName="Verhoeff" authorityYear="1924" box="[788,919,400,424]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="36" pageNumber="237" phylum="Arthropoda" rank="species" species="hirsutum">
|
||
<emphasis id="B9363802E32E8C10A16EFE5FFC7EFE54" box="[788,919,400,424]" italics="true" pageId="36" pageNumber="237">S. hirsutum</emphasis>
|
||
</taxonomicName>
|
||
in the presence of glabrous tergites, from
|
||
<taxonomicName id="4C429F93E32E8C10A7FDFE5FFEFCFE30" authorityName="Verhoeff" authorityYear="1924" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="36" pageNumber="237" phylum="Arthropoda" rank="species" species="clavigerum">
|
||
<emphasis id="B9363802E32E8C10A7FDFE5FFEFCFE30" italics="true" pageId="36" pageNumber="237">S. clavigerum</emphasis>
|
||
</taxonomicName>
|
||
in the presence of a short locking carina on the anal shield. It differs from
|
||
<taxonomicName id="4C429F93E32E8C10A6E8FE7BFAEBFE37" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[1170,1282,436,459]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="36" pageNumber="237" phylum="Arthropoda" rank="species" species="nigrum" status="sp. nov.">
|
||
<emphasis id="B9363802E32E8C10A6E8FE7BFAEBFE37" box="[1170,1282,436,459]" italics="true" pageId="36" pageNumber="237">S. nigrum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9363802E32E8C10A776FE7CFAA6FE30" bold="true" box="[1292,1359,435,460]" pageId="36" pageNumber="237">
|
||
<taxonomicNameLabel id="A2058579E32E8C10A776FE7CFAA6FE30" box="[1292,1359,435,460]" pageId="36" pageNumber="237" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in the presence of a cylindrical antennomere 6, the presence of only 5–7 ventral spines on the tarsi (
|
||
<quantity id="4CBA49F5E32E8C10A6AFFE18FAC7FE0C" box="[1237,1326,471,496]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" metricValueMax="2.54" metricValueMin="2.032" pageId="36" pageNumber="237" unit="in" value="9.0" valueMax="10.0" valueMin="8.0">8–10 in</quantity>
|
||
the other species) (
|
||
<figureCitation id="1379F895E32E8C10A37BFE35FE8EFDEF" box="[257,359,506,531]" captionStart="FIGURE 23" captionStartId="34.[151,250,1968,1990]" captionTargetBox="[210,1381,193,1950]" captionTargetId="figure@34.[210,1381,193,1950]" captionTargetPageId="34" captionText="FIGURE 23. Sphaerobelum laoticum n. sp., holotype (SMF); A: coxa of first left leg with first stigmatic plate; B: 9th left leg. Abbreviations: Cx = coxa; St-Pl = stigmatic plate. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601121/files/figure.png" pageId="36" pageNumber="237">Fig. 23B</figureCitation>
|
||
), and the presence of a strongly curved telopoditomere 4 of the posterior telopod, all characters only shared with
|
||
<taxonomicName id="4C429F93E32E8C10A322FDD1FE1CFDCA" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[344,501,542,566]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="36" pageNumber="237" phylum="Arthropoda" rank="species" species="splendidum" status="sp. nov.">
|
||
<emphasis id="B9363802E32E8C10A322FDD1FE1CFDCA" box="[344,501,542,566]" italics="true" pageId="36" pageNumber="237">S. splendidum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9363802E32E8C10A386FDD2FDD2FDCA" bold="true" box="[508,571,541,566]" pageId="36" pageNumber="237">
|
||
<taxonomicNameLabel id="A2058579E32E8C10A386FDD2FDD2FDCA" box="[508,571,541,566]" pageId="36" pageNumber="237" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C429F93E32E8C10A00BFDD1FCD9FDCA" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[625,816,542,566]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="36" pageNumber="237" phylum="Arthropoda" rank="species" species="schwendingeri" status="sp. nov.">
|
||
<emphasis id="B9363802E32E8C10A00BFDD1FCD9FDCA" box="[625,816,542,566]" italics="true" pageId="36" pageNumber="237">S. schwendingeri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9363802E32E8C10A14DFDD2FC9FFDCA" bold="true" box="[823,886,541,566]" pageId="36" pageNumber="237">
|
||
<taxonomicNameLabel id="A2058579E32E8C10A14DFDD2FC9FFDCA" box="[823,886,541,566]" pageId="36" pageNumber="237" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
For a differentiation of
|
||
<taxonomicName id="4C429F93E32E8C10A6FEFDD1FAE8FDCA" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[1156,1281,542,566]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="36" pageNumber="237" phylum="Arthropoda" rank="species" species="laoticum" status="sp. nov.">
|
||
<emphasis id="B9363802E32E8C10A6FEFDD1FAE8FDCA" box="[1156,1281,542,566]" italics="true" pageId="36" pageNumber="237">S. laoticum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9363802E32E8C10A772FDD2FAAEFDCA" bold="true" box="[1288,1351,541,566]" pageId="36" pageNumber="237">
|
||
<taxonomicNameLabel id="A2058579E32E8C10A772FDD2FAAEFDCA" box="[1288,1351,541,566]" pageId="36" pageNumber="237" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
with
|
||
<taxonomicName id="4C429F93E32E8C10A7FDFDD1FEF1FDA5" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="36" pageNumber="237" phylum="Arthropoda" rank="species" species="splendidum" status="sp. nov.">
|
||
<emphasis id="B9363802E32E8C10A7FDFDD1FEF1FDA5" italics="true" pageId="36" pageNumber="237">S. splendidum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9363802E32E8C10A359FD8EFE8FFDA6" bold="true" box="[291,358,577,602]" pageId="36" pageNumber="237">
|
||
<taxonomicNameLabel id="A2058579E32E8C10A359FD8EFE8FFDA6" box="[291,358,577,602]" pageId="36" pageNumber="237" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
see above.
|
||
<taxonomicName id="4C429F93E32E8C10A38CFD8DFD91FDA5" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[502,632,577,601]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="36" pageNumber="237" phylum="Arthropoda" rank="species" species="laoticum" status="sp. nov.">
|
||
<emphasis id="B9363802E32E8C10A38CFD8DFD91FDA5" box="[502,632,577,601]" italics="true" pageId="36" pageNumber="237">S. laoticum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9363802E32E8C10A0FEFD8EFD2EFDA6" bold="true" box="[644,711,577,602]" pageId="36" pageNumber="237">
|
||
<taxonomicNameLabel id="A2058579E32E8C10A0FEFD8EFD2EFDA6" box="[644,711,577,602]" pageId="36" pageNumber="237" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="4C429F93E32E8C10A112FD8DFBC2FDA5" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[872,1067,577,601]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="36" pageNumber="237" phylum="Arthropoda" rank="species" species="schwendingeri" status="sp. nov.">
|
||
<emphasis id="B9363802E32E8C10A112FD8DFBC2FDA5" box="[872,1067,577,601]" italics="true" pageId="36" pageNumber="237">S. schwendingeri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9363802E32E8C10A64CFD8EFB90FDA6" bold="true" box="[1078,1145,577,602]" pageId="36" pageNumber="237">
|
||
<taxonomicNameLabel id="A2058579E32E8C10A64CFD8EFB90FDA6" box="[1078,1145,577,602]" pageId="36" pageNumber="237" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, which occurs almost in sympatry, in a genetic distance of 13.6% in the COI gene, the process of telopoditomere 2 of the anterior telopods projecting as high as basal 1/3 (3/
|
||
<quantity id="4CBA49F5E32E8C10A074FD46FDD3FD5D" box="[526,570,648,673]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="36" pageNumber="237" unit="in" value="4.0">4 in</quantity>
|
||
<taxonomicName id="4C429F93E32E8C10A03AFD46FD16FD5C" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[576,767,648,672]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="36" pageNumber="237" phylum="Arthropoda" rank="species" species="schwendingeri" status="sp. nov.">
|
||
<emphasis id="B9363802E32E8C10A03AFD46FD16FD5C" box="[576,767,648,672]" italics="true" pageId="36" pageNumber="237">S. schwendingeri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9363802E32E8C10A17CFD47FCACFD5D" bold="true" box="[774,837,648,673]" pageId="36" pageNumber="237">
|
||
<taxonomicNameLabel id="A2058579E32E8C10A17CFD47FCACFD5D" box="[774,837,648,673]" pageId="36" pageNumber="237" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
) of telopoditomere 3+4 (
|
||
<figureCitation id="1379F895E32E8C10A625FD47FB2AFD5D" box="[1119,1219,648,673]" captionStart="FIGURE 22" captionStartId="33.[151,250,1922,1944]" captionTargetBox="[168,1421,193,1901]" captionTargetId="figure@33.[166,1421,193,1901]" captionTargetPageId="33" captionText="FIGURE 22. Sphaerobelum splendidum n. sp., holotype (SMF); A: coxa of first left leg with first stigmatic plate; B: 9th left leg; C: coxa of second left leg with gonopore; D: left anterior telopod, anterior view; E: left anterior telopod, posterior view; F: left anterior telopod, lateral view; G: right posterior telopod, anterior view; H: right posterior telopod, posterior view. Abbreviations: Cx = coxa; St-Pl = stigmatic plate; Syn-Cx = syncoxite. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601119/files/figure.png" pageId="36" pageNumber="237">Fig. 22F</figureCitation>
|
||
); anal shield with a single short locking carinae (absent in
|
||
<taxonomicName id="4C429F93E32E8C10A032FD63FCE1FD38" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[584,776,684,708]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="36" pageNumber="237" phylum="Arthropoda" rank="species" species="schwendingeri" status="sp. nov.">
|
||
<emphasis id="B9363802E32E8C10A032FD63FCE1FD38" box="[584,776,684,708]" italics="true" pageId="36" pageNumber="237">S. schwendingeri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9363802E32E8C10A175FD63FCA7FD39" bold="true" box="[783,846,684,709]" pageId="36" pageNumber="237">
|
||
<taxonomicNameLabel id="A2058579E32E8C10A175FD63FCA7FD39" box="[783,846,684,709]" pageId="36" pageNumber="237" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
); palpi of gnathochilarium, sensory cones located in single field (in several clusters in
|
||
<taxonomicName id="4C429F93E32E8C10A06BFD1FFD39FD1B" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[529,720,719,743]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="36" pageNumber="237" phylum="Arthropoda" rank="species" species="schwendingeri" status="sp. nov.">
|
||
<emphasis id="B9363802E32E8C10A06BFD1FFD39FD1B" box="[529,720,719,743]" italics="true" pageId="36" pageNumber="237">S. schwendingeri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9363802E32E8C10A0ADFD00FCFFFD14" bold="true" box="[727,790,719,744]" pageId="36" pageNumber="237">
|
||
<taxonomicNameLabel id="A2058579E32E8C10A0ADFD00FCFFFD14" box="[727,790,719,744]" pageId="36" pageNumber="237" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C358B79BE32E8C11A2BDFD3CFF13FE99" lastPageId="37" lastPageNumber="238" pageId="36" pageNumber="237" type="description">
|
||
<paragraph id="8BFDE410E32E8C10A2BDFD3CFD86FCF0" blockId="36.[151,1437,151,2021]" box="[199,623,755,780]" pageId="36" pageNumber="237">
|
||
<emphasis id="B9363802E32E8C10A2BDFD3CFEB2FCF0" bold="true" box="[199,347,755,780]" pageId="36" pageNumber="237">Description:</emphasis>
|
||
Based on male
|
||
<typeStatus id="54F95AB2E32E8C10A077FD3CFD86FCF0" box="[525,623,755,780]" pageId="36" pageNumber="237" type="holotype">holotype</typeStatus>
|
||
</paragraph>
|
||
<paragraph id="8BFDE410E32E8C10A2BDFCD9FB66FC8A" blockId="36.[151,1437,151,2021]" pageId="36" pageNumber="237">
|
||
<emphasis id="B9363802E32E8C10A2BDFCD9FE9FFCD3" bold="true" box="[199,374,790,815]" pageId="36" pageNumber="237">Measurements</emphasis>
|
||
: body length
|
||
<quantity id="4CBA49F5E32E8C10A062FCD8FD6BFCD2" box="[536,642,791,815]" metricMagnitude="-2" metricUnit="m" metricValue="2.44" pageId="36" pageNumber="237" unit="mm" value="24.4">24.4 mm</quantity>
|
||
. Width, of thoracic shield =
|
||
<quantity id="4CBA49F5E32E8C10A1B5FCD8FBDCFCD2" box="[975,1077,791,815]" metricMagnitude="-2" metricUnit="m" metricValue="1.27" pageId="36" pageNumber="237" unit="mm" value="12.7">12.7 mm</quantity>
|
||
(= widest). Height, of thoracic shield =
|
||
<quantity id="4CBA49F5E32E8C10A37BFCF5FEB5FCAD" box="[257,348,826,850]" metricMagnitude="-3" metricUnit="m" metricValue="7.4" pageId="36" pageNumber="237" unit="mm" value="7.4">7.4 mm</quantity>
|
||
(= highest).
|
||
<emphasis id="B9363802E32E8C10A380FCF6FD92FCAE" bold="true" box="[506,635,825,850]" pageId="36" pageNumber="237">Coloration</emphasis>
|
||
: Head and collum black. Tergites black (fading to brown in longer preserved specimens), paratergite depressions olive green (
|
||
<figureCitation id="1379F895E32E8C10A151FC92FC93FC8A" box="[811,890,861,886]" captionStart="FIGURE 6" captionStartId="11.[151,250,1916,1938]" captionTargetBox="[150,1423,987,1897]" captionTargetId="figure@11.[150,1436,983,1903]" captionTargetPageId="11" captionText="FIGURE 6. Habitus multi-layer stacked photographs of Sphaerobelum species. A: Sphaerobelum bolavensis n. sp.; B: Sphaerobelum phouloei n. sp.; C: Sphaerobelum denticulatum n. sp.; D: Sphaerobelum spinatum n. sp.; E: Sphaerobelum lachneeis n. sp.; F: Sphaerobelum peterjaegeri n. sp.; G: Sphaerobelum nigrum n. sp.; H: Sphaerobelum splendidum n. sp.; I: Sphaerobelum laoticum n. sp. Not to scale." httpUri="https://zenodo.org/record/2601084/files/figure.png" pageId="36" pageNumber="237">
|
||
Fig.
|
||
<date id="FFFCC2D0E32E8C10A125FC92FC93FC8A" box="[863,890,861,886]" pageId="36" pageNumber="237">6I</date>
|
||
</figureCitation>
|
||
). Antennae and legs red.
|
||
</paragraph>
|
||
<paragraph id="8BFDE410E32E8C10A2BDFC4FFD37FBF8" blockId="36.[151,1437,151,2021]" pageId="36" pageNumber="237">
|
||
<emphasis id="B9363802E32E8C10A2BDFC4FFEEEFC65" bold="true" box="[199,263,896,921]" pageId="36" pageNumber="237">Head</emphasis>
|
||
: with ca. 65 ocelli. Antennae extending posteriorly to leg 5. Size of antennomeres 1=2=3=4=5<<6, 6th apically widened. Apical disc carrying 81/86 apical cones. Antennomere 1–5 with only few setae. Organ of Tömösváry located in antennal groove.
|
||
<emphasis id="B9363802E32E8C10A014FC08FCD4FC1C" bold="true" box="[622,829,967,992]" pageId="36" pageNumber="237">Gnathochilarium</emphasis>
|
||
: with the usual shape for the order, sensory cones located in single field.
|
||
<emphasis id="B9363802E32E8C10A3EDFC24FDFCFBF8" bold="true" box="[407,533,1003,1028]" pageId="36" pageNumber="237">Mandibles</emphasis>
|
||
: not investigated.
|
||
</paragraph>
|
||
<paragraph id="8BFDE410E32E8C10A2BDFBC1FA75FBDB" blockId="36.[151,1437,151,2021]" box="[199,1436,1038,1063]" pageId="36" pageNumber="237">
|
||
<emphasis id="B9363802E32E8C10A2BDFBC1FE22FBDB" bold="true" box="[199,459,1038,1063]" pageId="36" pageNumber="237">First stigmatic plate</emphasis>
|
||
: small, well-rounded, inconspicuous, slightly curved towards coxa (
|
||
<figureCitation id="1379F895E32E8C10A767FBC1FA79FBDB" box="[1309,1424,1038,1063]" captionStart="FIGURE 23" captionStartId="34.[151,250,1968,1990]" captionTargetBox="[210,1381,193,1950]" captionTargetId="figure@34.[210,1381,193,1950]" captionTargetPageId="34" captionText="FIGURE 23. Sphaerobelum laoticum n. sp., holotype (SMF); A: coxa of first left leg with first stigmatic plate; B: 9th left leg. Abbreviations: Cx = coxa; St-Pl = stigmatic plate. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601121/files/figure.png" pageId="36" pageNumber="237">Fig. 23A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BFDE410E32E8C10A2EDFBFDFB96FA2D" blockId="36.[151,1437,151,2021]" pageId="36" pageNumber="237">
|
||
<emphasis id="B9363802E32E8C10A2EDFBFDFEA9FBB7" bold="true" box="[151,320,1074,1099]" pageId="36" pageNumber="237">Laterotergites</emphasis>
|
||
: 1 and 2 with a short, well-rounded projection.
|
||
<emphasis id="B9363802E32E8C10A1FFFBFDFC34FBB7" bold="true" box="[901,989,1074,1099]" pageId="36" pageNumber="237">Collum</emphasis>
|
||
: glabrous only short setae at margin.
|
||
<emphasis id="B9363802E32E8C10A2EDFB9AFEBBFB92" bold="true" box="[151,338,1109,1134]" pageId="36" pageNumber="237">Thoracic shield</emphasis>
|
||
: Shallow grooves with few setae, surface glabrous. Margin towards grooves with three or four weak anterior, and 2 or 3 posterior crests.
|
||
<emphasis id="B9363802E32E8C10A0EBFBB6FD19FB6E" bold="true" box="[657,752,1145,1170]" pageId="36" pageNumber="237">Tergites</emphasis>
|
||
: shiny, polished, no pits or setae present in posterior 2/3. Anterior 1/3, which is mainly visible when the specimen is rolled up, with longer setae providing a golden sheen. Paratergite tips weakly projecting posteriorly (
|
||
<figureCitation id="1379F895E32E8C10A0D7FB70FD16FB24" box="[685,767,1215,1240]" captionStart="FIGURE 6" captionStartId="11.[151,250,1916,1938]" captionTargetBox="[150,1423,987,1897]" captionTargetId="figure@11.[150,1436,983,1903]" captionTargetPageId="11" captionText="FIGURE 6. Habitus multi-layer stacked photographs of Sphaerobelum species. A: Sphaerobelum bolavensis n. sp.; B: Sphaerobelum phouloei n. sp.; C: Sphaerobelum denticulatum n. sp.; D: Sphaerobelum spinatum n. sp.; E: Sphaerobelum lachneeis n. sp.; F: Sphaerobelum peterjaegeri n. sp.; G: Sphaerobelum nigrum n. sp.; H: Sphaerobelum splendidum n. sp.; I: Sphaerobelum laoticum n. sp. Not to scale." httpUri="https://zenodo.org/record/2601084/files/figure.png" pageId="36" pageNumber="237">
|
||
Fig.
|
||
<date id="FFFCC2D0E32E8C10A09EFB70FD16FB24" box="[740,767,1215,1240]" pageId="36" pageNumber="237">6I</date>
|
||
</figureCitation>
|
||
).
|
||
<emphasis id="B9363802E32E8C10A16FFB70FC45FB24" bold="true" box="[789,940,1215,1240]" pageId="36" pageNumber="237">Endotergum</emphasis>
|
||
: inner section without any setae or spines. Middle area with a single row of sparse elliptical, cuticular impressions, distance between impressions larger than their diameter. Apically 3 dense rows of short marginal bristles, the longest just protruding up to tergal margin. Bristles not smooth, but with numerous small spicules.
|
||
<emphasis id="B9363802E32E8C10A163FAE5FC4AFABF" bold="true" box="[793,931,1322,1347]" pageId="36" pageNumber="237">Anal shield</emphasis>
|
||
: well-rounded, covered with numerous tiny setae, underside with a single very small, almost dot-like, locking carina.
|
||
<emphasis id="B9363802E32E8C10A19CFA82FBF7FA9A" bold="true" box="[998,1054,1357,1382]" pageId="36" pageNumber="237">Legs</emphasis>
|
||
: leg-pair 1 with 2 ventral spines, leg-pair 2 with 4, leg-pair 3 with 6, leg-pairs 4–21 with only 4–7. Apical spines at present tarsi 3–21. In leg 9, femur 1.6 times, tarsus 3.9 times longer than wide (
|
||
<figureCitation id="1379F895E32E8C10A0ADFA5BFCD7FA51" box="[727,830,1428,1453]" captionStart="FIGURE 23" captionStartId="34.[151,250,1968,1990]" captionTargetBox="[210,1381,193,1950]" captionTargetId="figure@34.[210,1381,193,1950]" captionTargetPageId="34" captionText="FIGURE 23. Sphaerobelum laoticum n. sp., holotype (SMF); A: coxa of first left leg with first stigmatic plate; B: 9th left leg. Abbreviations: Cx = coxa; St-Pl = stigmatic plate. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601121/files/figure.png" pageId="36" pageNumber="237">Fig. 23B</figureCitation>
|
||
). Leg 1 without a coxal process (
|
||
<figureCitation id="1379F895E32E8C10A6CAFA5BFAF1FA51" box="[1200,1304,1428,1453]" captionStart="FIGURE 23" captionStartId="34.[151,250,1968,1990]" captionTargetBox="[210,1381,193,1950]" captionTargetId="figure@34.[210,1381,193,1950]" captionTargetPageId="34" captionText="FIGURE 23. Sphaerobelum laoticum n. sp., holotype (SMF); A: coxa of first left leg with first stigmatic plate; B: 9th left leg. Abbreviations: Cx = coxa; St-Pl = stigmatic plate. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601121/files/figure.png" pageId="36" pageNumber="237">Fig. 23A</figureCitation>
|
||
), leg 2 with a sharp coxal process (
|
||
<figureCitation id="1379F895E32E8C10A3EFFA77FE14FA2C" box="[405,509,1464,1489]" captionStart="FIGURE 24" captionStartId="35.[151,250,1945,1967]" captionTargetBox="[161,1430,193,1928]" captionTargetId="figure@35.[161,1430,193,1928]" captionTargetPageId="35" captionText="FIGURE 24. Sphaerobelum laoticum n. sp., holotype (SMF); A: coxa of second left leg with gonopore; B: left anterior telopod, anterior view; C: left anterior telopod, posterior view; D: left anterior telopod, lateral view; E: left posterior telopod, anterior view; F: left posterior telopod, posterior view. Abbreviations: Cx = coxa; Syn-Cx = syncoxite. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601123/files/figure.png" pageId="36" pageNumber="237">Fig. 24A</figureCitation>
|
||
), while the process is well-rounded in the following legs.
|
||
</paragraph>
|
||
<paragraph id="8BFDE410E32E8C10A2BDFA14FC49F819" blockId="36.[151,1437,151,2021]" pageId="36" pageNumber="237">
|
||
<emphasis id="B9363802E32E8C10A2BDFA14FE36FA08" bold="true" box="[199,479,1499,1524]" pageId="36" pageNumber="237">Male sexual characters</emphasis>
|
||
: male gonopore large, covered by small, inconspicuous plate (
|
||
<figureCitation id="1379F895E32E8C10A6C9FA14FAF7FA08" box="[1203,1310,1499,1524]" captionStart="FIGURE 24" captionStartId="35.[151,250,1945,1967]" captionTargetBox="[161,1430,193,1928]" captionTargetId="figure@35.[161,1430,193,1928]" captionTargetPageId="35" captionText="FIGURE 24. Sphaerobelum laoticum n. sp., holotype (SMF); A: coxa of second left leg with gonopore; B: left anterior telopod, anterior view; C: left anterior telopod, posterior view; D: left anterior telopod, lateral view; E: left posterior telopod, anterior view; F: left posterior telopod, posterior view. Abbreviations: Cx = coxa; Syn-Cx = syncoxite. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601123/files/figure.png" pageId="36" pageNumber="237">Fig. 24A</figureCitation>
|
||
).
|
||
<emphasis id="B9363802E32E8C10A74EFA14FF13F9E4" bold="true" pageId="36" pageNumber="237">Anterior telopods</emphasis>
|
||
(
|
||
<figureCitation id="1379F895E32E8C10A377FA30FE81F9EB" box="[269,360,1535,1560]" captionStart="FIGURE 24" captionStartId="35.[151,250,1945,1967]" captionTargetBox="[161,1430,193,1928]" captionTargetId="figure@35.[161,1430,193,1928]" captionTargetPageId="35" captionText="FIGURE 24. Sphaerobelum laoticum n. sp., holotype (SMF); A: coxa of second left leg with gonopore; B: left anterior telopod, anterior view; C: left anterior telopod, posterior view; D: left anterior telopod, lateral view; E: left posterior telopod, anterior view; F: left posterior telopod, posterior view. Abbreviations: Cx = coxa; Syn-Cx = syncoxite. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601123/files/figure.png" pageId="36" pageNumber="237">Figs 24</figureCitation>
|
||
B–D): consisting of three telopoditomere above syncoxite. Last telopoditomere divided by a suture still visible in lateral view. Telopoditomere 1 rectangular, slightly longer than wide. Telopoditomere 2 large, as long as telopoditomere 3. Process of telopoditomere 2 located posteriorly, not visible in anterior view. Process of telopoditomere 2 wide, projecting to half of telopoditomere 3, slightly curved, apically with a sharp tip. Telopoditomere 3 massive, cylindrical, straight, apically slightly tapering. Posterior side with two large, triangular spines. In anterior view all telopoditomeres covered by long setae, in posterior view telopoditomeres 2 and 3 only with marginal setae.
|
||
<emphasis id="B9363802E32E8C10A3F3F91BFD8CF911" bold="true" box="[393,613,1748,1773]" pageId="36" pageNumber="237">Posterior telopods</emphasis>
|
||
(
|
||
<figureCitation id="1379F895E32E8C10A00CF91BFCEAF910" box="[630,771,1748,1773]" captionStart="FIGURE 25" captionStartId="41.[151,250,1885,1907]" captionTargetBox="[207,1378,195,1868]" captionTargetId="figure@41.[207,1378,195,1868]" captionTargetPageId="41" captionText="FIGURE 25. Sphaerobelum schwendingeri n. sp. A–C female paratype (ZMUC), D–G male holotype (MHNG). A: coxa of first right leg with first stigmatic plate; B: 9th left leg; C: coxa and prefemur of second leg with vulva; D: left anterior telopod, anterior view; E: left anterior telopod, posterior view; F: right posterior telopod, anterior view; G: right posterior telopod, posterior view. Abbreviations: Cx = coxa; O = operculum of vulva; Pre = prefemur; St-Pl = stigmatic plate; Syn-Cx = syncoxite. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601125/files/figure.png" pageId="36" pageNumber="237">Figs 25E, F</figureCitation>
|
||
): telopoditomere 1 rectangular, slightly longer than wide. Immovable finger (process of telopoditomere 2) slightly shorter than movable finger, consisting of telopoditomeres 3 and 4. Immovable finger with a characteristic, distally swollen apex, slightly swollen. Apically broader than base, at tip in anterior view with a few large, crenulated teeth. Telopoditomere 3 elongated, towards immovable finger at midpoint with a triangular extension carrying a spines. Telopoditomere 4 tapering, anterior half strongly curved 90° towards immovable finger, which it overlaps. At inner margin with two larger spines. Entire telopoditomeres 1 and 2 on both sides covered with long setae, except for the immovable finger. Telopoditomere 3 only basally in anterior view with setae, remaining part as well as telopoditomere 4 glabrous.
|
||
</paragraph>
|
||
<paragraph id="8BFDE410E32F8C11A2BDFF58FF13FE99" blockId="37.[151,1436,151,573]" pageId="37" pageNumber="238">
|
||
<emphasis id="B9363802E32F8C11A2BDFF58FE3BFF4C" bold="true" box="[199,466,151,176]" pageId="37" pageNumber="238">Intraspecific variation</emphasis>
|
||
:
|
||
<taxonomicName id="4C429F93E32F8C11A398FF57FD88FF4C" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[482,609,152,176]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="37" pageNumber="238" phylum="Arthropoda" rank="species" species="laoticum" status="sp. nov.">
|
||
<emphasis id="B9363802E32F8C11A398FF57FD88FF4C" box="[482,609,152,176]" italics="true" pageId="37" pageNumber="238">S. laoticum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9363802E32F8C11A010FF58FD40FF4C" bold="true" box="[618,681,151,176]" pageId="37" pageNumber="238">
|
||
<taxonomicNameLabel id="A2058579E32F8C11A010FF58FD40FF4C" box="[618,681,151,176]" pageId="37" pageNumber="238" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is the giant pill-millipede species from
|
||
<collectingCountry id="F355A480E32F8C11A615FF57FB4CFF53" box="[1135,1189,152,175]" name="Laos" pageId="37" pageNumber="238">Laos</collectingCountry>
|
||
with the most known localities (3, all in a
|
||
<quantity id="4CBA49F5E32F8C11A3EEFF73FE09FF29" box="[404,480,188,213]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="37" pageNumber="238" unit="km" value="15.0">15 km</quantity>
|
||
radius,
|
||
<figureCitation id="1379F895E32F8C11A03EFF73FD63FF29" box="[580,650,188,213]" captionStart="FIGURE 5" captionStartId="9.[151,250,2005,2027]" captionTargetBox="[185,1415,664,1984]" captionTargetId="figure@9.[181,1415,663,1984]" captionTargetPageId="9" captionText="FIGURE 5. Distribution map of Sphaerobelum species, including the unnamed females." httpUri="https://zenodo.org/record/2601082/files/figure.png" pageId="37" pageNumber="238">Fig. 5</figureCitation>
|
||
) and specimens. The specimens from the different localities show considerable intraspecific variation. The smaller, not fully mature specimens from Ban Sok Say and Tham Loup display one, sometimes two weak short locking carinae. These samples were collected recently and might require Nagoya permissions to be barcoded, so an analysis of the intraspecific variation between these populations was not possible.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C358B79BE32F8C11A2BDFEA0FABAFE09" pageId="37" pageNumber="238" type="discussion">
|
||
<paragraph id="8BFDE410E32F8C11A2BDFEA0FABAFE09" blockId="37.[151,1436,151,573]" pageId="37" pageNumber="238">
|
||
<emphasis id="B9363802E32F8C11A2BDFEA0FEDDFE74" bold="true" box="[199,308,367,392]" pageId="37" pageNumber="238">Remarks</emphasis>
|
||
: This species, only known from males, lives syntopically with two other
|
||
<taxonomicName id="4C429F93E32F8C11A616FEBFFAE4FE74" authorityName="Verhoeff" authorityYear="1924" box="[1132,1293,368,392]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="37" pageNumber="238" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9363802E32F8C11A616FEBFFAE4FE74" box="[1132,1293,368,392]" italics="true" pageId="37" pageNumber="238">Sphaerobelum</emphasis>
|
||
</taxonomicName>
|
||
species, one only known from females.
|
||
<taxonomicName id="4C429F93E32F8C11A3AAFE5BFD98FE50" authorityName="Verhoeff" authorityYear="1924" box="[464,625,404,428]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="37" pageNumber="238" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9363802E32F8C11A3AAFE5BFD98FE50" box="[464,625,404,428]" italics="true" pageId="37" pageNumber="238">Sphaerobelum</emphasis>
|
||
</taxonomicName>
|
||
sp. L10, and
|
||
<taxonomicName id="4C429F93E32F8C11A16FFE5BFC3EFE50" authorityName="Wesener" authorityYear="2019" box="[789,983,404,428]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="37" pageNumber="238" phylum="Arthropoda" rank="species" species="schwendingeri">
|
||
<emphasis id="B9363802E32F8C11A16FFE5BFC3EFE50" box="[789,983,404,428]" italics="true" pageId="37" pageNumber="238">S. schwendingeri</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="1379F895E32F8C11A193FE5BFBD9FE51" box="[1001,1072,404,429]" captionStart="FIGURE 5" captionStartId="9.[151,250,2005,2027]" captionTargetBox="[185,1415,664,1984]" captionTargetId="figure@9.[181,1415,663,1984]" captionTargetPageId="9" captionText="FIGURE 5. Distribution map of Sphaerobelum species, including the unnamed females." httpUri="https://zenodo.org/record/2601082/files/figure.png" pageId="37" pageNumber="238">Fig. 5</figureCitation>
|
||
, see below). Genetic barcoding data clearly reveals that the specimens, collected from a close-by location, are distinct species, differing in 15.9% (to S. sp. L10) or 13.6 % p-distance (to
|
||
<taxonomicName id="4C429F93E32F8C11A02EFE13FCFAFE08" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[596,787,476,500]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="37" pageNumber="238" phylum="Arthropoda" rank="species" species="schwendingeri" status="sp. nov.">
|
||
<emphasis id="B9363802E32F8C11A02EFE13FCFAFE08" box="[596,787,476,500]" italics="true" pageId="37" pageNumber="238">S. schwendingeri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9363802E32F8C11A160FE13FCB0FE09" bold="true" box="[794,857,476,501]" pageId="37" pageNumber="238">
|
||
<taxonomicNameLabel id="A2058579E32F8C11A160FE13FCB0FE09" box="[794,857,476,501]" pageId="37" pageNumber="238" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
) in the COI gene in the barcoding COI gene.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C358B79BE32F8C11A2BDFE30FF1BFDC1" pageId="37" pageNumber="238" type="etymology">
|
||
<paragraph id="8BFDE410E32F8C11A2BDFE30FF1BFDC1" blockId="37.[151,1436,151,573]" pageId="37" pageNumber="238">
|
||
<emphasis id="B9363802E32F8C11A2BDFE30FE74FDE4" bold="true" box="[199,413,511,536]" pageId="37" pageNumber="238">Derivatio nominis</emphasis>
|
||
: laoticum, noun in apposition, after
|
||
<collectingCountry id="F355A480E32F8C11A149FDCFFC85FDEB" box="[819,876,512,535]" name="Laos" pageId="37" pageNumber="238">Laos</collectingCountry>
|
||
, as this is the first
|
||
<taxonomicName id="4C429F93E32F8C11A646FDCFFB34FDE4" authorityName="Verhoeff" authorityYear="1924" box="[1084,1245,512,536]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="37" pageNumber="238" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9363802E32F8C11A646FDCFFB34FDE4" box="[1084,1245,512,536]" italics="true" pageId="37" pageNumber="238">Sphaerobelum</emphasis>
|
||
</taxonomicName>
|
||
known from this country.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |