372 lines
49 KiB
XML
372 lines
49 KiB
XML
<document id="3C07FCDBFB0962F851264753C5BF4498" ID-CLB-Dataset="28149" ID-DOI="10.11646/zootaxa.4563.2.1" ID-GBIF-Dataset="e6037c5d-a96f-41e5-8e70-71f2659659e2" ID-ISSN="1175-5326" ID-Zenodo-Dep="2601072" ID-ZooBank="CF79B01B-8B5F-4B3A-B642-2CADE4B339AF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553171433896" checkinUser="plazi" docAuthor="Wesener, Thomas" docDate="2019" docId="03EB5506E3168C2BA2EDF9DBFCF0F8F9" docLanguage="en" docName="zootaxa.4563.2.1.pdf" docOrigin="Zootaxa 4563 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Sphaerobelum nigrum Wesener 2019, new species" docType="treatment" docVersion="9" lastPageNumber="232" masterDocId="FFD22D7EE30A8C34A27AFFCFFFE9FFFC" masterDocTitle="First records of giant pill-millipedes from Laos (Diplopoda, Sphaerotheriida, Zephroniidae)" masterLastPageNumber="248" masterPageNumber="201" pageNumber="229" updateTime="1698703176884" updateUser="ExternalLinkService">
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<mods:titleInfo id="0908C89ED52B46EC0551ACD96777BBD2">
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<mods:title id="A4800BD8F2C66998BE300F4DAC5EAD7E">First records of giant pill-millipedes from Laos (Diplopoda, Sphaerotheriida, Zephroniidae)</mods:title>
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<mods:name id="29498746CBE05FCFA6F84AE907C5613F" type="personal">
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<mods:namePart id="92F4D2660910900F4AAB9D9CA37563DB">Wesener, Thomas</mods:namePart>
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<mods:title id="99B969C2E8C15D3268406EF5DBF9DFF3">Zootaxa</mods:title>
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<mods:part id="9BC8022DEB99D93451AE35AA3F420FFA">
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<mods:date id="56F007467D8FA75C767DF712C1E3AFE8">2019</mods:date>
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<mods:number id="3466D4DDEFE7AC248C9ED9F32BFE0B2E">4563</mods:number>
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<treatment id="03EB5506E3168C2BA2EDF9DBFCF0F8F9" ID-DOI="http://doi.org/10.5281/zenodo.5934534" ID-GBIF-Taxon="154474550" ID-Zenodo-Dep="5934534" LSID="urn:lsid:plazi:treatment:03EB5506E3168C2BA2EDF9DBFCF0F8F9" httpUri="http://treatment.plazi.org/id/03EB5506E3168C2BA2EDF9DBFCF0F8F9" lastPageId="31" lastPageNumber="232" pageId="28" pageNumber="229">
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<subSubSection id="C358B79BE3168C28A2EDF9DBFDD6F9D2" box="[151,575,1556,1582]" pageId="28" pageNumber="229" type="nomenclature">
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<paragraph id="8BFDE410E3168C28A2EDF9DBFDD6F9D2" blockId="28.[151,575,1556,1615]" box="[151,575,1556,1582]" pageId="28" pageNumber="229">
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<heading id="D0B5537CE3168C28A2EDF9DBFDD6F9D2" bold="true" box="[151,575,1556,1582]" fontSize="11" level="1" pageId="28" pageNumber="229" reason="1">
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<emphasis id="B9363802E3168C28A2EDF9DBFDD6F9D2" bold="true" box="[151,575,1556,1582]" pageId="28" pageNumber="229">
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<taxonomicName id="4C429F93E3168C28A2EDF9DBFE41F9D2" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[151,424,1556,1582]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="28" pageNumber="229" phylum="Arthropoda" rank="species" species="nigrum" status="sp. nov.">
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<emphasis id="B9363802E3168C28A2EDF9DBFE41F9D2" bold="true" box="[151,424,1556,1582]" italics="true" pageId="28" pageNumber="229">Sphaerobelum nigrum</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2058579E3168C28A3D5F9DBFDD6F9D2" box="[431,575,1556,1582]" pageId="28" pageNumber="229" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C358B79BE3168C28A2EDF9F9FE8CF9B3" box="[151,357,1590,1615]" pageId="28" pageNumber="229" type="description">
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<paragraph id="8BFDE410E3168C28A2EDF9F9FE8CF9B3" blockId="28.[151,575,1556,1615]" box="[151,357,1590,1615]" pageId="28" pageNumber="229">
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<figureCitation id="1379F895E3168C28A2EDF9F9FEFAF9B3" box="[151,275,1590,1615]" captionStart="FIGURE 6" captionStartId="11.[151,250,1916,1938]" captionTargetBox="[150,1423,987,1897]" captionTargetId="figure@11.[150,1436,983,1903]" captionTargetPageId="11" captionText="FIGURE 6. Habitus multi-layer stacked photographs of Sphaerobelum species. A: Sphaerobelum bolavensis n. sp.; B: Sphaerobelum phouloei n. sp.; C: Sphaerobelum denticulatum n. sp.; D: Sphaerobelum spinatum n. sp.; E: Sphaerobelum lachneeis n. sp.; F: Sphaerobelum peterjaegeri n. sp.; G: Sphaerobelum nigrum n. sp.; H: Sphaerobelum splendidum n. sp.; I: Sphaerobelum laoticum n. sp. Not to scale." httpUri="https://zenodo.org/record/2601084/files/figure.png" pageId="28" pageNumber="229">Figures 6G</figureCitation>
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,
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<figureCitation id="1379F895E3168C28A365F9F8FED4F9B3" box="[287,317,1591,1615]" captionStart="FIGURE 20" captionStartId="29.[151,250,1928,1950]" captionTargetBox="[151,1435,337,1906]" captionTargetId="figure@29.[151,1435,337,1906]" captionTargetPageId="29" captionText="FIGURE 20. Sphaerobelum nigrum n. sp., holotype (SMF); A: coxa of first left leg with first stigmatic plate; B: 9th left leg; C: coxa and prefemur of second leg with vulva. Abbreviations: Cx = coxa; O = operculum of vulva; Pre = prefemur; St-Pl = stigmatic plate. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601115/files/figure.png" pageId="28" pageNumber="229">20</figureCitation>
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,
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<figureCitation id="1379F895E3168C28A333F9F8FE8CF9B3" box="[329,357,1591,1615]" captionStart="FIGURE 21" captionStartId="30.[151,250,1950,1972]" captionTargetBox="[174,1418,193,1936]" captionTargetId="figure@30.[174,1418,193,1936]" captionTargetPageId="30" captionText="FIGURE 21. Sphaerobelum nigrum n. sp., holotype (SMF); A: coxa of second left leg with gonopore; B: right anterior telopod, anterior view; C: right anterior telopod, posterior view; D: right anterior telopod, lateral view; E: right posterior telopod, anterior view; F: left posterior telopod, posterior view. Abbreviations: Syn-Cx = syncoxite. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601117/files/figure.png" pageId="28" pageNumber="229">21</figureCitation>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C358B79BE3168C28A2EDF9B1FDA0F8B7" pageId="28" pageNumber="229" type="materials_examined">
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<paragraph id="8BFDE410E3168C28A2EDF9B1FCF2F923" blockId="28.[151,1436,1662,2011]" pageId="28" pageNumber="229">
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<emphasis id="B9363802E3168C28A2EDF9B1FE6DF96B" bold="true" box="[151,388,1662,1687]" pageId="28" pageNumber="229">Material examined:</emphasis>
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Type specimens: 1 M
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<typeStatus id="54F95AB2E3168C28A0F4F9B1FD19F96B" box="[654,752,1662,1687]" pageId="28" pageNumber="229" type="holotype">holotype</typeStatus>
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(SMF) from
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<collectingCountry id="F355A480E3168C28A1F7F9B0FC2FF96A" box="[909,966,1663,1686]" name="Laos" pageId="28" pageNumber="229">Laos</collectingCountry>
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,
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<collectingRegion id="49862AF2E3168C28A1A9F9B1FB2CF96B" box="[979,1221,1662,1687]" country="Laos" name="Champasak" pageId="28" pageNumber="229">Champasak Province</collectingRegion>
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, Muang Bachieng, Ban Lak 35, Tad Etu (
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<geoCoordinate id="EE7682D7E3168C28A3EDF96DFDDAF947" box="[407,563,1698,1723]" direction="north" orientation="latitude" pageId="28" pageNumber="229" precision="1" value="15.193806">N15°11'37.7"</geoCoordinate>
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,
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<geoCoordinate id="EE7682D7E3168C28A03AF96DFD0EF947" box="[576,743,1698,1723]" direction="east" orientation="longitude" pageId="28" pageNumber="229" precision="1" value="106.101746">106°06'06.3"E</geoCoordinate>
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),
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<quantity id="4CBA49F5E3168C28A087F96CFCAEF946" box="[765,839,1699,1723]" metricMagnitude="2" metricUnit="m" metricValue="8.1" pageId="28" pageNumber="229" unit="m" value="810.0">810 m</quantity>
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, coffee plantation with remnant of secondary forest, leg. in vegetation, at night, by hand;
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<date id="FFFCC2D0E3168C28A04BF909FD5DF923" box="[561,692,1734,1759]" pageId="28" pageNumber="229" value="2012-11-16">16.XI.2012</date>
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; P. Jäger.
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</paragraph>
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<paragraph id="8BFDE410E3168C28A2BDF925FDA0F8B7" blockId="28.[151,1436,1662,2011]" pageId="28" pageNumber="229">
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||
<materialsCitation id="3B2AEE4DE3168C28A2BDF925FDACF8B7" ID-GBIF-Occurrence="2013952574" collectingDate="2009-11-25" collectionCode="SMF" collectorName="P. Jager & S. Bayer" country="Laos" county="Muang Bachieng" elevation="810" latitude="15.193806" location="Tad Etu" longLatPrecision="1" longitude="106.101746" municipality="Ban Lak" pageId="28" pageNumber="229" specimenCount="1" stateProvince="Champasak" typeStatus="paratype">
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||
<emphasis id="B9363802E3168C28A2BDF925FED4F8FF" bold="true" box="[199,317,1770,1795]" pageId="28" pageNumber="229">
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||
<typeStatus id="54F95AB2E3168C28A2BDF925FED1F8FF" box="[199,312,1770,1795]" pageId="28" pageNumber="229" type="paratype">Paratype</typeStatus>
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||
:
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</emphasis>
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1 F (
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<collectionCode id="ED537CD5E3168C28A30CF924FE58F8FF" box="[374,433,1771,1795]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="28" pageNumber="229">SMF</collectionCode>
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||
) from
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<collectingCountry id="F355A480E3168C28A386F924FDDCF8FE" box="[508,565,1771,1794]" name="Laos" pageId="28" pageNumber="229">Laos</collectingCountry>
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,
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<collectingRegion id="49862AF2E3168C28A03AF925FCD9F8FF" box="[576,816,1770,1795]" country="Laos" name="Champasak" pageId="28" pageNumber="229">Champasak Province</collectingRegion>
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||
,
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<collectingCounty id="629C9C9CE3168C28A140F924FC14F8FF" box="[826,1021,1770,1795]" pageId="28" pageNumber="229">Muang Bachieng</collectingCounty>
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||
,
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<collectingMunicipality id="6B997E6AE3168C28A67DF924FB8EF8FF" box="[1031,1127,1770,1795]" pageId="28" pageNumber="229">Ban Lak</collectingMunicipality>
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35,
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<location id="8E9DB2CBE3168C28A6EDF925FB07F8FE" LSID="urn:lsid:plazi:treatment:03EB5506E3168C2BA2EDF9DBFCF0F8F9:8E9DB2CBE3168C28A6EDF925FB07F8FE" box="[1175,1262,1770,1795]" country="Laos" county="Muang Bachieng" latitude="15.193806" longLatPrecision="1" longitude="106.101746" municipality="Ban Lak" name="Tad Etu" pageId="28" pageNumber="229" stateProvince="Champasak">Tad Etu</location>
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(
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||
<geoCoordinate id="EE7682D7E3168C28A687F925FA71F8FF" box="[1277,1432,1770,1795]" direction="north" orientation="latitude" pageId="28" pageNumber="229" precision="1" value="15.193806">N15°11'37.7"</geoCoordinate>
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,
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||
<geoCoordinate id="EE7682D7E3168C28A2EDF8C1FED7F8DB" box="[151,318,1806,1831]" direction="east" orientation="longitude" pageId="28" pageNumber="229" precision="1" value="106.101746">106°06'06.3"E</geoCoordinate>
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),
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<quantity id="4CBA49F5E3168C28A329F8C0FE77F8DA" box="[339,414,1807,1831]" metricMagnitude="2" metricUnit="m" metricValue="8.1" pageId="28" pageNumber="229" unit="m" value="810.0">
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<elevation id="006F0323E3168C28A329F8C0FE77F8DA" box="[339,414,1807,1831]" metricMagnitude="2" metricUnit="m" metricValue="8.1" pageId="28" pageNumber="229" unit="m" value="810.0">810 m</elevation>
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||
</quantity>
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, coffee plantation with remnant of secondary forest, close to river, at rock, leg. at day, by hand,
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<date id="FFFCC2D0E3168C28A2A1F8FDFEB7F8B7" box="[219,350,1842,1867]" pageId="28" pageNumber="229" value="2009-11-25">
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<collectingDate id="EFB83B38E3168C28A2A1F8FDFEB7F8B7" box="[219,350,1842,1867]" pageId="28" pageNumber="229" value="2009-11-25">25.XI.2009</collectingDate>
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||
</date>
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||
,
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<collectorName id="26B781C6E3168C28A313F8FCFE57F8B7" box="[361,446,1843,1867]" pageId="28" pageNumber="229">P. Jäger</collectorName>
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||
&
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<collectorName id="26B781C6E3168C28A398F8FCFDACF8B7" box="[482,581,1843,1867]" pageId="28" pageNumber="229">S. Bayer</collectorName>
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||
</materialsCitation>
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||
.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C358B79BE3168C29A2BDF899FBECFF04" lastPageId="29" lastPageNumber="230" pageId="28" pageNumber="229" type="diagnosis">
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||
<paragraph id="8BFDE410E3168C29A2BDF899FBECFF04" blockId="28.[151,1436,1662,2011]" lastBlockId="29.[151,1436,151,285]" lastPageId="29" lastPageNumber="230" pageId="28" pageNumber="229">
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||
<emphasis id="B9363802E3168C28A2BDF899FED3F893" bold="true" box="[199,314,1878,1903]" pageId="28" pageNumber="229">Diagnosis</emphasis>
|
||
:
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||
<taxonomicName id="4C429F93E3168C28A336F898FE55F892" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[332,444,1879,1902]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="28" pageNumber="229" phylum="Arthropoda" rank="species" species="nigrum" status="sp. nov.">
|
||
<emphasis id="B9363802E3168C28A336F898FE55F892" box="[332,444,1879,1902]" italics="true" pageId="28" pageNumber="229">S. nigrum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9363802E3168C28A3BFF899FDEEF893" bold="true" box="[453,519,1878,1903]" pageId="28" pageNumber="229">
|
||
<taxonomicNameLabel id="A2058579E3168C28A3BFF899FDEEF893" box="[453,519,1878,1903]" pageId="28" pageNumber="229" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
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||
belongs to a group of
|
||
<taxonomicName id="4C429F93E3168C28A16FF899FC5FF892" authorityName="Verhoeff" authorityYear="1924" box="[789,950,1878,1902]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="28" pageNumber="229" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9363802E3168C28A16FF899FC5FF892" box="[789,950,1878,1902]" italics="true" pageId="28" pageNumber="229">Sphaerobelum</emphasis>
|
||
</taxonomicName>
|
||
species in which the mesal margin of the femur is modified (
|
||
<figureCitation id="1379F895E3168C28A317F8B5FE3AF86F" box="[365,467,1914,1939]" captionStart="FIGURE 20" captionStartId="29.[151,250,1928,1950]" captionTargetBox="[151,1435,337,1906]" captionTargetId="figure@29.[151,1435,337,1906]" captionTargetPageId="29" captionText="FIGURE 20. Sphaerobelum nigrum n. sp., holotype (SMF); A: coxa of first left leg with first stigmatic plate; B: 9th left leg; C: coxa and prefemur of second leg with vulva. Abbreviations: Cx = coxa; O = operculum of vulva; Pre = prefemur; St-Pl = stigmatic plate. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601115/files/figure.png" pageId="28" pageNumber="229">Fig. 20B</figureCitation>
|
||
).
|
||
<taxonomicName id="4C429F93E3168C28A39CF8B4FDBBF86E" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[486,594,1915,1938]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="28" pageNumber="229" phylum="Arthropoda" rank="species" species="nigrum" status="sp. nov.">
|
||
<emphasis id="B9363802E3168C28A39CF8B4FDBBF86E" box="[486,594,1915,1938]" italics="true" pageId="28" pageNumber="229">S. nigrum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9363802E3168C28A022F8B5FD7FF86F" bold="true" box="[600,662,1914,1939]" pageId="28" pageNumber="229">
|
||
<taxonomicNameLabel id="A2058579E3168C28A022F8B5FD7FF86F" box="[600,662,1914,1939]" pageId="28" pageNumber="229" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
shares only with
|
||
<taxonomicName id="4C429F93E3168C28A123F8B4FC31F86F" authorityName="Verhoeff" authorityYear="1924" box="[857,984,1915,1939]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="28" pageNumber="229" phylum="Arthropoda" rank="species" species="hirsutum">
|
||
<emphasis id="B9363802E3168C28A123F8B4FC31F86F" box="[857,984,1915,1939]" italics="true" pageId="28" pageNumber="229">S. hirsutum</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C429F93E3168C28A19FF8B4FB96F86F" authorityName="Verhoeff" authorityYear="1924" box="[997,1151,1915,1939]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="28" pageNumber="229" phylum="Arthropoda" rank="species" species="clavigerum">
|
||
<emphasis id="B9363802E3168C28A19FF8B4FB96F86F" box="[997,1151,1915,1939]" italics="true" pageId="28" pageNumber="229">S. clavigerum</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C429F93E3168C28A6F6F8B4FAC1F86F" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[1164,1320,1915,1939]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="28" pageNumber="229" phylum="Arthropoda" rank="species" species="splendidum" status="sp. nov.">
|
||
<emphasis id="B9363802E3168C28A6F6F8B4FAC1F86F" box="[1164,1320,1915,1939]" italics="true" pageId="28" pageNumber="229">S. splendidum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9363802E3168C28A755F8B5FA84F86F" bold="true" box="[1327,1389,1914,1939]" pageId="28" pageNumber="229">
|
||
<taxonomicNameLabel id="A2058579E3168C28A755F8B5FA84F86F" box="[1327,1389,1914,1939]" pageId="28" pageNumber="229" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C429F93E3168C28A2EDF850FEB0F84A" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[151,345,1950,1974]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="28" pageNumber="229" phylum="Arthropoda" rank="species" species="schwendingeri" status="sp. nov.">
|
||
<emphasis id="B9363802E3168C28A2EDF850FEB0F84A" box="[151,345,1950,1974]" italics="true" pageId="28" pageNumber="229">S. schwendingeri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9363802E3168C28A319F851FE4DF84B" bold="true" box="[355,420,1950,1975]" pageId="28" pageNumber="229">
|
||
<taxonomicNameLabel id="A2058579E3168C28A319F851FE4DF84B" box="[355,420,1950,1975]" pageId="28" pageNumber="229" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
the presence of a "hook" at telopoditomere 4 of the posterior telopods(
|
||
<figureCitation id="1379F895E3168C28A69FF851FA99F84B" box="[1253,1392,1950,1975]" captionStart="FIGURE 21" captionStartId="30.[151,250,1950,1972]" captionTargetBox="[174,1418,193,1936]" captionTargetId="figure@30.[174,1418,193,1936]" captionTargetPageId="30" captionText="FIGURE 21. Sphaerobelum nigrum n. sp., holotype (SMF); A: coxa of second left leg with gonopore; B: right anterior telopod, anterior view; C: right anterior telopod, posterior view; D: right anterior telopod, lateral view; E: right posterior telopod, anterior view; F: left posterior telopod, posterior view. Abbreviations: Syn-Cx = syncoxite. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601117/files/figure.png" pageId="28" pageNumber="229">Figs 21E, F</figureCitation>
|
||
).
|
||
<taxonomicName id="4C429F93E3168C28A7FDF850FF01F826" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="28" pageNumber="229" phylum="Arthropoda" rank="species" species="nigrum" status="sp. nov.">
|
||
<emphasis id="B9363802E3168C28A7FDF850FF01F826" italics="true" pageId="28" pageNumber="229">S. nigrum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9363802E3168C28A295F80DFEC7F827" bold="true" box="[239,302,1986,2011]" pageId="28" pageNumber="229">
|
||
<taxonomicNameLabel id="A2058579E3168C28A295F80DFEC7F827" box="[239,302,1986,2011]" pageId="28" pageNumber="229" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="4C429F93E3168C28A3BEF80CFDADF827" authorityName="Verhoeff" authorityYear="1924" box="[452,580,1987,2011]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="28" pageNumber="229" phylum="Arthropoda" rank="species" species="hirsutum">
|
||
<emphasis id="B9363802E3168C28A3BEF80CFDADF827" box="[452,580,1987,2011]" italics="true" pageId="28" pageNumber="229">S. hirsutum</emphasis>
|
||
</taxonomicName>
|
||
in the presence of glabrous tergites, from
|
||
<taxonomicName id="4C429F93E3168C28A665F80CFB53F827" authorityName="Verhoeff" authorityYear="1924" box="[1055,1210,1987,2011]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="28" pageNumber="229" phylum="Arthropoda" rank="species" species="clavigerum">
|
||
<emphasis id="B9363802E3168C28A665F80CFB53F827" box="[1055,1210,1987,2011]" italics="true" pageId="28" pageNumber="229">S. clavigerum</emphasis>
|
||
</taxonomicName>
|
||
in the presence of a short locking carina on the anal shield. Differs from
|
||
<taxonomicName id="4C429F93E3178C29A09DFF57FC6CFF4C" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[743,901,152,176]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="29" pageNumber="230" phylum="Arthropoda" rank="species" species="splendidum" status="sp. nov.">
|
||
<emphasis id="B9363802E3178C29A09DFF57FC6CFF4C" box="[743,901,152,176]" italics="true" pageId="29" pageNumber="230">S. splendidum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9363802E3178C29A1F6FF58FC25FF4C" bold="true" box="[908,972,151,176]" pageId="29" pageNumber="230">
|
||
<taxonomicNameLabel id="A2058579E3178C29A1F6FF58FC25FF4C" box="[908,972,151,176]" pageId="29" pageNumber="230" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C429F93E3178C29A67EFF57FB2AFF4C" authorityName="Wesener" authorityYear="2019" box="[1028,1219,152,176]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="29" pageNumber="230" phylum="Arthropoda" rank="species" species="schwendingeri">
|
||
<emphasis id="B9363802E3178C29A67EFF57FB2AFF4C" box="[1028,1219,152,176]" italics="true" pageId="29" pageNumber="230">S. schwendingeri</emphasis>
|
||
</taxonomicName>
|
||
sp. in the presence of an axe-shaped antennomere 6, and the presence of only a short hook at the telopoditomere 4 (
|
||
<figureCitation id="1379F895E3178C29A6B2FF73FAA4FF28" box="[1224,1357,188,213]" captionStart="FIGURE 21" captionStartId="30.[151,250,1950,1972]" captionTargetBox="[174,1418,193,1936]" captionTargetId="figure@30.[174,1418,193,1936]" captionTargetPageId="30" captionText="FIGURE 21. Sphaerobelum nigrum n. sp., holotype (SMF); A: coxa of second left leg with gonopore; B: right anterior telopod, anterior view; C: right anterior telopod, posterior view; D: right anterior telopod, lateral view; E: right posterior telopod, anterior view; F: left posterior telopod, posterior view. Abbreviations: Syn-Cx = syncoxite. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601117/files/figure.png" pageId="29" pageNumber="230">Figs 21E, F</figureCitation>
|
||
) of the posterior telopods (strongly curved telopoditomere
|
||
<quantity id="4CBA49F5E3178C29A0ADFF2FFCEBFF04" box="[727,770,223,248]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="29" pageNumber="230" unit="in" value="4.0">4 in</quantity>
|
||
the other two species).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C358B79BE3178C2BA2BDFECBFB39F941" lastPageId="31" lastPageNumber="232" pageId="29" pageNumber="230" type="description">
|
||
<paragraph id="8BFDE410E3178C29A2BDFECBFCB6FEE1" blockId="29.[151,1436,151,285]" box="[199,863,260,285]" pageId="29" pageNumber="230">
|
||
<emphasis id="B9363802E3178C29A2BDFECBFEB0FEE1" bold="true" box="[199,345,260,285]" pageId="29" pageNumber="230">Description.</emphasis>
|
||
Based on
|
||
<typeStatus id="54F95AB2E3178C29A3B4FECBFDD9FEE1" box="[462,560,260,285]" pageId="29" pageNumber="230" type="holotype">holotype</typeStatus>
|
||
male and
|
||
<typeStatus id="54F95AB2E3178C29A0DEFECAFCEDFEE1" box="[676,772,261,285]" pageId="29" pageNumber="230" type="paratype">paratype</typeStatus>
|
||
female.
|
||
</paragraph>
|
||
<caption id="DF3DB498E3178C29A2EDF847FE15F827" httpUri="https://zenodo.org/record/2601115/files/figure.png" pageId="29" pageNumber="230" startId="29.[151,250,1928,1950]" targetBox="[151,1435,337,1906]" targetPageId="29">
|
||
<paragraph id="8BFDE410E3178C29A2EDF847FE15F827" blockId="29.[151,1436,1928,2011]" pageId="29" pageNumber="230">
|
||
<emphasis id="B9363802E3178C29A2EDF847FEF1F862" bold="true" box="[151,280,1928,1950]" pageId="29" pageNumber="230">FIGURE 20</emphasis>
|
||
.
|
||
<taxonomicName id="4C429F93E3178C29A35EF847FDECF862" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[292,517,1928,1950]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="29" pageNumber="230" phylum="Arthropoda" rank="species" species="nigrum" status="sp. nov.">
|
||
<emphasis id="B9363802E3178C29A35EF847FDECF862" box="[292,517,1928,1950]" italics="true" pageId="29" pageNumber="230">Sphaerobelum nigrum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9363802E3178C29A071F847FDAAF862" bold="true" box="[523,579,1928,1950]" pageId="29" pageNumber="230">
|
||
<taxonomicNameLabel id="A2058579E3178C29A071F847FDAAF862" box="[523,579,1928,1950]" pageId="29" pageNumber="230" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype (SMF);
|
||
<emphasis id="B9363802E3178C29A084F847FCF9F862" bold="true" box="[766,784,1928,1950]" pageId="29" pageNumber="230">A</emphasis>
|
||
: coxa of first left leg with first stigmatic plate;
|
||
<emphasis id="B9363802E3178C29A691F847FB15F862" bold="true" box="[1259,1276,1928,1950]" pageId="29" pageNumber="230">B</emphasis>
|
||
: 9th left leg;
|
||
<emphasis id="B9363802E3178C29A7F9F847FA7CF862" bold="true" box="[1411,1429,1928,1950]" pageId="29" pageNumber="230">C</emphasis>
|
||
: coxa and prefemur of second leg with vulva. Abbreviations:
|
||
<emphasis id="B9363802E3178C29A169F869FCDBF840" bold="true" box="[787,818,1958,1980]" pageId="29" pageNumber="230">Cx</emphasis>
|
||
= coxa;
|
||
<emphasis id="B9363802E3178C29A1EBF869FC4CF840" bold="true" box="[913,933,1958,1980]" pageId="29" pageNumber="230">O</emphasis>
|
||
= operculum of vulva;
|
||
<emphasis id="B9363802E3178C29A6E5F869FB2DF840" bold="true" box="[1183,1220,1958,1980]" pageId="29" pageNumber="230">Pre</emphasis>
|
||
= prefemur;
|
||
<emphasis id="B9363802E3178C29A72BF869FA6DF840" bold="true" box="[1361,1412,1958,1980]" pageId="29" pageNumber="230">St-Pl</emphasis>
|
||
= stigmatic plate. Scale bars = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF3DB498E3148C2AA2EDF851FABEF80E" httpUri="https://zenodo.org/record/2601117/files/figure.png" pageId="30" pageNumber="231" startId="30.[151,250,1950,1972]" targetBox="[174,1418,193,1936]" targetPageId="30">
|
||
<paragraph id="8BFDE410E3148C2AA2EDF851FABEF80E" blockId="30.[151,1436,1950,2034]" pageId="30" pageNumber="231">
|
||
<emphasis id="B9363802E3148C2AA2EDF851FEF4F848" bold="true" box="[151,285,1950,1972]" pageId="30" pageNumber="231">FIGURE 21</emphasis>
|
||
.
|
||
<taxonomicName id="4C429F93E3148C2AA354F851FDFAF849" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[302,531,1950,1973]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="30" pageNumber="231" phylum="Arthropoda" rank="species" species="nigrum" status="sp. nov.">
|
||
<emphasis id="B9363802E3148C2AA354F851FDFAF849" box="[302,531,1950,1973]" italics="true" pageId="30" pageNumber="231">Sphaerobelum nigrum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9363802E3148C2AA067F851FDB3F848" bold="true" box="[541,602,1950,1972]" pageId="30" pageNumber="231">
|
||
<taxonomicNameLabel id="A2058579E3148C2AA067F851FDB3F848" box="[541,602,1950,1972]" pageId="30" pageNumber="231" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype (SMF);
|
||
<emphasis id="B9363802E3148C2AA15EF851FCDFF848" bold="true" box="[804,822,1950,1972]" pageId="30" pageNumber="231">A</emphasis>
|
||
: coxa of second left leg with gonopore;
|
||
<emphasis id="B9363802E3148C2AA68AF851FAE8F848" bold="true" box="[1264,1281,1950,1972]" pageId="30" pageNumber="231">B</emphasis>
|
||
: right anterior telopod, anterior view;
|
||
<emphasis id="B9363802E3148C2AA3F4F872FE40F82F" bold="true" box="[398,425,1981,2003]" pageId="30" pageNumber="231">C:</emphasis>
|
||
right anterior telopod, posterior view;
|
||
<emphasis id="B9363802E3148C2AA132F872FCB3F82F" bold="true" box="[840,858,1981,2003]" pageId="30" pageNumber="231">D</emphasis>
|
||
: right anterior telopod, lateral view;
|
||
<emphasis id="B9363802E3148C2AA69CF872FB1EF82F" bold="true" box="[1254,1271,1981,2003]" pageId="30" pageNumber="231">E</emphasis>
|
||
: right posterior telopod, anterior view;
|
||
<emphasis id="B9363802E3148C2AA3F9F813FE7BF80E" bold="true" box="[387,402,2012,2034]" pageId="30" pageNumber="231">F</emphasis>
|
||
: left posterior telopod, posterior view. Abbreviations:
|
||
<emphasis id="B9363802E3148C2AA1C0F813FBE3F80E" bold="true" box="[954,1034,2012,2034]" pageId="30" pageNumber="231">Syn-Cx</emphasis>
|
||
= syncoxite. Scale bars = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BFDE410E3158C2BA2BDFF58FAE0FEBC" blockId="31.[151,1437,151,1797]" pageId="31" pageNumber="232">
|
||
<emphasis id="B9363802E3158C2BA2BDFF58FE96FF4C" bold="true" box="[199,383,151,176]" pageId="31" pageNumber="232">Measurements:</emphasis>
|
||
Body length:
|
||
<typeStatus id="54F95AB2E3158C2BA053FF58FD62FF4C" box="[553,651,151,176]" pageId="31" pageNumber="232" type="holotype">holotype</typeStatus>
|
||
male: length ca
|
||
<quantity id="4CBA49F5E3158C2BA12BFF57FC52FF53" box="[849,955,152,176]" metricMagnitude="-2" metricUnit="m" metricValue="2.78" pageId="31" pageNumber="232" unit="mm" value="27.8">27.8 mm</quantity>
|
||
. Width, of thoracic shield =
|
||
<quantity id="4CBA49F5E3158C2BA777FF58FA9FFF53" box="[1293,1398,151,176]" metricMagnitude="-2" metricUnit="m" metricValue="1.61" pageId="31" pageNumber="232" unit="mm" value="16.1">16.1 mm</quantity>
|
||
, of tergite 7 =
|
||
<quantity id="4CBA49F5E3158C2BA368FF73FE9FFF28" box="[274,374,188,213]" metricMagnitude="-2" metricUnit="m" metricValue="1.68" pageId="31" pageNumber="232" unit="mm" value="16.8">16.8 mm</quantity>
|
||
(= broadest). Height, of thoracic shield =
|
||
<quantity id="4CBA49F5E3158C2BA12AFF73FC41FF28" box="[848,936,188,212]" metricMagnitude="-3" metricUnit="m" metricValue="9.8" pageId="31" pageNumber="232" unit="mm" value="9.8">9.8 mm</quantity>
|
||
, of tergite 7 =
|
||
<quantity id="4CBA49F5E3158C2BA634FF73FB5BFF28" box="[1102,1202,188,212]" metricMagnitude="-2" metricUnit="m" metricValue="1.01" pageId="31" pageNumber="232" unit="mm" value="10.1">10.1 mm</quantity>
|
||
(= highest). Female: length ca
|
||
<quantity id="4CBA49F5E3158C2BA37CFF2FFE86FF0B" box="[262,367,224,248]" metricMagnitude="-2" metricUnit="m" metricValue="3.47" pageId="31" pageNumber="232" unit="mm" value="34.7">34.7 mm</quantity>
|
||
. Width, of thoracic shield =
|
||
<quantity id="4CBA49F5E3158C2BA0CFFF10FCF5FF0B" box="[693,796,223,248]" metricMagnitude="-2" metricUnit="m" metricValue="1.5699999999999998" pageId="31" pageNumber="232" unit="mm" value="15.7">15.7 mm</quantity>
|
||
, of tergite 7 =
|
||
<quantity id="4CBA49F5E3158C2BA1BDFF2FFBC5FF0B" box="[967,1068,224,248]" metricMagnitude="-2" metricUnit="m" metricValue="1.7399999999999998" pageId="31" pageNumber="232" unit="mm" value="17.4">17.4 mm</quantity>
|
||
(= broadest). Height of thoracic shield =
|
||
<quantity id="4CBA49F5E3158C2BA280FECBFEBDFEE0" box="[250,340,260,284]" metricMagnitude="-3" metricUnit="m" metricValue="9.3" pageId="31" pageNumber="232" unit="mm" value="9.3">9.3 mm</quantity>
|
||
, of tergite 7 =
|
||
<quantity id="4CBA49F5E3158C2BA385FECBFDBFFEE0" box="[511,598,260,284]" metricMagnitude="-3" metricUnit="m" metricValue="9.8" pageId="31" pageNumber="232" unit="mm" value="9.8">9.8 mm</quantity>
|
||
(= highest).
|
||
<emphasis id="B9363802E3158C2BA093FECBFC83FEE1" bold="true" box="[745,874,260,285]" pageId="31" pageNumber="232">Coloration</emphasis>
|
||
: in preserved specimens yellow with dark brown spots or black spots/bands. Head and collum black (
|
||
<figureCitation id="1379F895E3158C2BA0A6FEE8FCDEFEBC" box="[732,823,295,320]" captionStart="FIGURE 6" captionStartId="11.[151,250,1916,1938]" captionTargetBox="[150,1423,987,1897]" captionTargetId="figure@11.[150,1436,983,1903]" captionTargetPageId="11" captionText="FIGURE 6. Habitus multi-layer stacked photographs of Sphaerobelum species. A: Sphaerobelum bolavensis n. sp.; B: Sphaerobelum phouloei n. sp.; C: Sphaerobelum denticulatum n. sp.; D: Sphaerobelum spinatum n. sp.; E: Sphaerobelum lachneeis n. sp.; F: Sphaerobelum peterjaegeri n. sp.; G: Sphaerobelum nigrum n. sp.; H: Sphaerobelum splendidum n. sp.; I: Sphaerobelum laoticum n. sp. Not to scale." httpUri="https://zenodo.org/record/2601084/files/figure.png" pageId="31" pageNumber="232">Fig. 6G</figureCitation>
|
||
). Antennae and legs dark brown to black.
|
||
</paragraph>
|
||
<paragraph id="8BFDE410E3158C2BA2BDFE83FC01FE09" blockId="31.[151,1437,151,1797]" pageId="31" pageNumber="232">
|
||
<emphasis id="B9363802E3158C2BA2BDFE83FEEEFE99" bold="true" box="[199,263,332,357]" pageId="31" pageNumber="232">Head</emphasis>
|
||
: Eyes with ca. 55 ocelli. Antennae short, with rounded joints, extending posteriorly to leg-pair 4. Lengths of antennomeres: 1=2=3=4=5<<6. All antennomeres densely pubescent, sensilla basiconica surrounding apical disc. Last antennomere thickened, apically widened, axe-shaped. Apical disc with 91/89 apical cones (male), 74/78 (female). Organ of Tömösváry located inside antennal groove.
|
||
<emphasis id="B9363802E3158C2BA1CDFE78FB6CFE2C" bold="true" box="[951,1157,439,464]" pageId="31" pageNumber="232">Gnathochilarium</emphasis>
|
||
: structure typical of the order. Palpi sensory cones located in single field.
|
||
<emphasis id="B9363802E3158C2BA0B8FE13FCA9FE09" bold="true" box="[706,832,476,501]" pageId="31" pageNumber="232">Mandibles</emphasis>
|
||
: not dissected.
|
||
</paragraph>
|
||
<paragraph id="8BFDE410E3158C2BA2BDFE30FA75FDE4" blockId="31.[151,1437,151,1797]" box="[199,1436,511,536]" pageId="31" pageNumber="232">
|
||
<emphasis id="B9363802E3158C2BA2BDFE30FE6CFDE4" bold="true" box="[199,389,511,536]" pageId="31" pageNumber="232">Stigmatic plates</emphasis>
|
||
: first stigmatic plate widely rounded, apex well-rounded, curved towards coxa 1 (
|
||
<figureCitation id="1379F895E3158C2BA75CFE30FA79FDE4" box="[1318,1424,511,536]" captionStart="FIGURE 20" captionStartId="29.[151,250,1928,1950]" captionTargetBox="[151,1435,337,1906]" captionTargetId="figure@29.[151,1435,337,1906]" captionTargetPageId="29" captionText="FIGURE 20. Sphaerobelum nigrum n. sp., holotype (SMF); A: coxa of first left leg with first stigmatic plate; B: 9th left leg; C: coxa and prefemur of second leg with vulva. Abbreviations: Cx = coxa; O = operculum of vulva; Pre = prefemur; St-Pl = stigmatic plate. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601115/files/figure.png" pageId="31" pageNumber="232">Fig. 20A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BFDE410E3158C2BA2EDFDEBFA75FBEC" blockId="31.[151,1437,151,1797]" pageId="31" pageNumber="232">
|
||
<emphasis id="B9363802E3158C2BA2EDFDEBFEA9FDC1" bold="true" box="[151,320,548,573]" pageId="31" pageNumber="232">Laterotergites</emphasis>
|
||
: laterotergite 1 and 2 elongated into a long, sharply pointed process.
|
||
<emphasis id="B9363802E3158C2BA644FDEBFB7FFDC1" bold="true" box="[1086,1174,548,573]" pageId="31" pageNumber="232">Collum</emphasis>
|
||
: with glabrous surface, margins with few isolated setae.
|
||
<emphasis id="B9363802E3158C2BA072FD88FD2BFD9C" bold="true" box="[520,706,583,608]" pageId="31" pageNumber="232">Thoracic shield</emphasis>
|
||
: surface glabrous, but covered with small pits like tergites, setae only in grooves. Shallow grooves beset with numerous long setae, slope towards groove with 2 or 3 weak anterior and 2 or 3 posterior keels.
|
||
<emphasis id="B9363802E3158C2BA3B5FD40FDC7FD54" bold="true" box="[463,558,655,680]" pageId="31" pageNumber="232">Tergites</emphasis>
|
||
: glabrous, but entirely covered by tiny pits. Tips of paratergites of midbody tergites projecting posteriorly (
|
||
<figureCitation id="1379F895E3158C2BA388FD7BFDA5FD31" box="[498,588,692,717]" captionStart="FIGURE 6" captionStartId="11.[151,250,1916,1938]" captionTargetBox="[150,1423,987,1897]" captionTargetId="figure@11.[150,1436,983,1903]" captionTargetPageId="11" captionText="FIGURE 6. Habitus multi-layer stacked photographs of Sphaerobelum species. A: Sphaerobelum bolavensis n. sp.; B: Sphaerobelum phouloei n. sp.; C: Sphaerobelum denticulatum n. sp.; D: Sphaerobelum spinatum n. sp.; E: Sphaerobelum lachneeis n. sp.; F: Sphaerobelum peterjaegeri n. sp.; G: Sphaerobelum nigrum n. sp.; H: Sphaerobelum splendidum n. sp.; I: Sphaerobelum laoticum n. sp. Not to scale." httpUri="https://zenodo.org/record/2601084/files/figure.png" pageId="31" pageNumber="232">Fig. 6G</figureCitation>
|
||
).
|
||
<emphasis id="B9363802E3158C2BA01AFD7BFD1EFD31" bold="true" box="[608,759,692,717]" pageId="31" pageNumber="232">Endotergum</emphasis>
|
||
: inner section without any setae or spines. Middle area with a single row of dense, round, cuticular impressions, distance between impressions slightly smaller than their diameter. Apically, 3–4 dense rows of short marginal bristles, the longest just protruding up to half of the distance to tergal margin. Bristles not smooth, but with numerous small spicules.
|
||
<emphasis id="B9363802E3158C2BA67BFCD0FB64FCC4" bold="true" box="[1025,1165,799,824]" pageId="31" pageNumber="232">Anal shield</emphasis>
|
||
: well-rounded, surface completely covered by tiny setae each standing in a tiny pit. Underside with a single, black locking carina, located close to last laterotergite. Locking carinae of similar size to those of tergites.
|
||
<emphasis id="B9363802E3158C2BA673FCA8FBA8FC7C" bold="true" box="[1033,1089,871,896]" pageId="31" pageNumber="232">Legs</emphasis>
|
||
: leg-pair 1 with 2 or 3 ventral spines, leg-pair 2 with 3 or 4, leg-pair 3 with 6. First two leg-pairs without an apical spine. Leg-pairs 4–21 with 9 or 10 ventral spines and one dorso-apical spine. In leg 9 femur 1.6 times, tarsus 3.7 times longer than wide (
|
||
<figureCitation id="1379F895E3158C2BA753FC60FA79FC34" box="[1321,1424,943,968]" captionStart="FIGURE 20" captionStartId="29.[151,250,1928,1950]" captionTargetBox="[151,1435,337,1906]" captionTargetId="figure@29.[151,1435,337,1906]" captionTargetPageId="29" captionText="FIGURE 20. Sphaerobelum nigrum n. sp., holotype (SMF); A: coxa of first left leg with first stigmatic plate; B: 9th left leg; C: coxa and prefemur of second leg with vulva. Abbreviations: Cx = coxa; O = operculum of vulva; Pre = prefemur; St-Pl = stigmatic plate. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601115/files/figure.png" pageId="31" pageNumber="232">Fig. 20B</figureCitation>
|
||
). Femur extended mesally into a dentate margin featuring 12–14 teeth. All podomeres densely setose. Coxa with a well rounded process. Coxa process absent at first leg (
|
||
<figureCitation id="1379F895E3158C2BA081FC38FC88FBEC" box="[763,865,1015,1040]" captionStart="FIGURE 20" captionStartId="29.[151,250,1928,1950]" captionTargetBox="[151,1435,337,1906]" captionTargetId="figure@29.[151,1435,337,1906]" captionTargetPageId="29" captionText="FIGURE 20. Sphaerobelum nigrum n. sp., holotype (SMF); A: coxa of first left leg with first stigmatic plate; B: 9th left leg; C: coxa and prefemur of second leg with vulva. Abbreviations: Cx = coxa; O = operculum of vulva; Pre = prefemur; St-Pl = stigmatic plate. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601115/files/figure.png" pageId="31" pageNumber="232">Fig. 20A</figureCitation>
|
||
) and sharply projecting at second (
|
||
<figureCitation id="1379F895E3158C2BA69CFC38FAB9FBEC" box="[1254,1360,1015,1040]" captionStart="FIGURE 20" captionStartId="29.[151,250,1928,1950]" captionTargetBox="[151,1435,337,1906]" captionTargetId="figure@29.[151,1435,337,1906]" captionTargetPageId="29" captionText="FIGURE 20. Sphaerobelum nigrum n. sp., holotype (SMF); A: coxa of first left leg with first stigmatic plate; B: 9th left leg; C: coxa and prefemur of second leg with vulva. Abbreviations: Cx = coxa; O = operculum of vulva; Pre = prefemur; St-Pl = stigmatic plate. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601115/files/figure.png" pageId="31" pageNumber="232">Figs 20C</figureCitation>
|
||
;
|
||
<figureCitation id="1379F895E3158C2BA721FC37FA66FBEC" box="[1371,1423,1016,1040]" captionStart="FIGURE 21" captionStartId="30.[151,250,1950,1972]" captionTargetBox="[174,1418,193,1936]" captionTargetId="figure@30.[174,1418,193,1936]" captionTargetPageId="30" captionText="FIGURE 21. Sphaerobelum nigrum n. sp., holotype (SMF); A: coxa of second left leg with gonopore; B: right anterior telopod, anterior view; C: right anterior telopod, posterior view; D: right anterior telopod, lateral view; E: right posterior telopod, anterior view; F: left posterior telopod, posterior view. Abbreviations: Syn-Cx = syncoxite. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601117/files/figure.png" pageId="31" pageNumber="232">21A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BFDE410E3158C2BA2BDFBD3FAB0FB81" blockId="31.[151,1437,151,1797]" pageId="31" pageNumber="232">
|
||
<emphasis id="B9363802E3158C2BA2BDFBD3FE10FBC9" bold="true" box="[199,505,1052,1077]" pageId="31" pageNumber="232">Female sexual characters</emphasis>
|
||
: Vulva of gravid female large, covering mesal 1/2 of coxa, extending mesally to anterior half of prefemur (
|
||
<figureCitation id="1379F895E3158C2BA3C7FBF0FDCCFBA4" box="[445,549,1087,1112]" captionStart="FIGURE 20" captionStartId="29.[151,250,1928,1950]" captionTargetBox="[151,1435,337,1906]" captionTargetId="figure@29.[151,1435,337,1906]" captionTargetPageId="29" captionText="FIGURE 20. Sphaerobelum nigrum n. sp., holotype (SMF); A: coxa of first left leg with first stigmatic plate; B: 9th left leg; C: coxa and prefemur of second leg with vulva. Abbreviations: Cx = coxa; O = operculum of vulva; Pre = prefemur; St-Pl = stigmatic plate. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601115/files/figure.png" pageId="31" pageNumber="232">Fig. 20C</figureCitation>
|
||
). Operculum rounded, mesal margin projecting into a well-rounded lobe 1/3 as high as remaining operculum.
|
||
<emphasis id="B9363802E3158C2BA396FBABFD79FB81" bold="true" box="[492,656,1124,1149]" pageId="31" pageNumber="232">Subanal plate</emphasis>
|
||
: large, high, broadly rounded triangular. Apex deeply recessed.
|
||
</paragraph>
|
||
<paragraph id="8BFDE410E3158C2BA2BDFB48FB39F941" blockId="31.[151,1437,151,1797]" pageId="31" pageNumber="232">
|
||
<emphasis id="B9363802E3158C2BA2BDFB48FE08FB5C" bold="true" box="[199,481,1159,1184]" pageId="31" pageNumber="232">Male sexual characters</emphasis>
|
||
: gonopore covered with a single, undivided, circular, sclerotized plate (
|
||
<figureCitation id="1379F895E3158C2BA75EFB48FA66FB5C" box="[1316,1423,1159,1184]" captionStart="FIGURE 21" captionStartId="30.[151,250,1950,1972]" captionTargetBox="[174,1418,193,1936]" captionTargetId="figure@30.[174,1418,193,1936]" captionTargetPageId="30" captionText="FIGURE 21. Sphaerobelum nigrum n. sp., holotype (SMF); A: coxa of second left leg with gonopore; B: right anterior telopod, anterior view; C: right anterior telopod, posterior view; D: right anterior telopod, lateral view; E: right posterior telopod, anterior view; F: left posterior telopod, posterior view. Abbreviations: Syn-Cx = syncoxite. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601117/files/figure.png" pageId="31" pageNumber="232">Fig. 21A</figureCitation>
|
||
).
|
||
<emphasis id="B9363802E3158C2BA2EDFB63FE84FB39" bold="true" box="[151,365,1196,1221]" pageId="31" pageNumber="232">Anterior telopods</emphasis>
|
||
(
|
||
<figureCitation id="1379F895E3158C2BA304FB63FE3FFB38" box="[382,470,1196,1221]" captionStart="FIGURE 21" captionStartId="30.[151,250,1950,1972]" captionTargetBox="[174,1418,193,1936]" captionTargetId="figure@30.[174,1418,193,1936]" captionTargetPageId="30" captionText="FIGURE 21. Sphaerobelum nigrum n. sp., holotype (SMF); A: coxa of second left leg with gonopore; B: right anterior telopod, anterior view; C: right anterior telopod, posterior view; D: right anterior telopod, lateral view; E: right posterior telopod, anterior view; F: left posterior telopod, posterior view. Abbreviations: Syn-Cx = syncoxite. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601117/files/figure.png" pageId="31" pageNumber="232">Figs 21</figureCitation>
|
||
B–D): consisting of 4 telopoditomeres above syncoxite, but telopoditomere 3 and 4 only divided by thin suture. Telopoditomere 1 slightly wider than long. Telopoditomere 2 large, as long as first, almost as long as telopoditomere 3 and 4 combined. Process of telopoditomere 2 located posteriorly, not visible in anterior view. Process of telopoditomere 2 wide, projecting to base of telopoditomere 4, apically with a wellrounded tip. Telopoditomere 3 cylindrical, straight, slightly smaller than telopoditomere 4. Telopoditomere 4 cylindrical, well-rounded, posterior side with two sclerotized spots. Telopoditomeres
|
||
<quantity id="4CBA49F5E3158C2BA622FAAFFB76FA84" box="[1112,1183,1375,1400]" metricMagnitude="-2" metricUnit="m" metricValue="6.35" metricValueMax="10.16" metricValueMin="2.54" pageId="31" pageNumber="232" unit="in" value="2.5" valueMax="4.0" valueMin="1.0">1–4 in</quantity>
|
||
anterior view covered with long setae. In posterior view all telopoditomeres with fewer setae.
|
||
<emphasis id="B9363802E3158C2BA67EFA4BFB0DFA61" bold="true" box="[1028,1252,1412,1437]" pageId="31" pageNumber="232">Posterior telopods</emphasis>
|
||
(
|
||
<figureCitation id="1379F895E3158C2BA680FA4BFA67FA60" box="[1274,1422,1412,1437]" captionStart="FIGURE 22" captionStartId="33.[151,250,1922,1944]" captionTargetBox="[168,1421,193,1901]" captionTargetId="figure@33.[166,1421,193,1901]" captionTargetPageId="33" captionText="FIGURE 22. Sphaerobelum splendidum n. sp., holotype (SMF); A: coxa of first left leg with first stigmatic plate; B: 9th left leg; C: coxa of second left leg with gonopore; D: left anterior telopod, anterior view; E: left anterior telopod, posterior view; F: left anterior telopod, lateral view; G: right posterior telopod, anterior view; H: right posterior telopod, posterior view. Abbreviations: Cx = coxa; St-Pl = stigmatic plate; Syn-Cx = syncoxite. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601119/files/figure.png" pageId="31" pageNumber="232">Figs 22E, F</figureCitation>
|
||
): telopoditomere 1 rectangular, as long as wide. Immovable finger (process of telopoditomere 2) as long as movable finger, consisting of telopoditomeres 3 and 4. Immovable finger with a characteristic, distally swollen apex. Apex expanding more towards movable finger than towards mesal margin, well rounded. Apex almost twice as wide as base. Telopoditomere 3 rectangular, twice as long as wide, towards immoveable finger with a triangular expansion. Telopoditomere 4 reaching 2/3 of length of telopoditomere 3, slenderer, only weakly tapering but tip bent backwards into acute hook, very slightly curved towards immovable finger. Inner margin with two spines towards immovable finger. Telopoditomere 1 at both sides covered by setae. Telopoditomeres 2 and
|
||
<quantity id="4CBA49F5E3158C2BA6B1F94FFB13F964" box="[1227,1274,1663,1688]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="31" pageNumber="232" unit="in" value="3.0">3 in</quantity>
|
||
anterior view covered by long setae, in posterior view mainly with marginal setae. Telopoditomere 4 glabrous.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C358B79BE3158C2BA2BDF908FCF0F8F9" pageId="31" pageNumber="232" type="etymology">
|
||
<paragraph id="8BFDE410E3158C2BA2BDF908FCF0F8F9" blockId="31.[151,1437,151,1797]" pageId="31" pageNumber="232">
|
||
<emphasis id="B9363802E3158C2BA2BDF908FE74F91C" bold="true" box="[199,413,1735,1760]" pageId="31" pageNumber="232">Derivatio nominis</emphasis>
|
||
: nigrum, noun in apposition, after the, for a member of
|
||
<taxonomicName id="4C429F93E3158C2BA675F907FB59F91C" authorityName="Verhoeff" authorityYear="1924" box="[1039,1200,1736,1760]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="31" pageNumber="232" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9363802E3158C2BA675F907FB59F91C" box="[1039,1200,1736,1760]" italics="true" pageId="31" pageNumber="232">Sphaerobelum</emphasis>
|
||
</taxonomicName>
|
||
, unique black colour of the legs, head, collum, antenna and part of the tergites.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |