263 lines
37 KiB
XML
263 lines
37 KiB
XML
<document id="0E867D6BB12A4CE0BF38491690E788D7" ID-CLB-Dataset="28149" ID-DOI="10.11646/zootaxa.4563.2.1" ID-GBIF-Dataset="e6037c5d-a96f-41e5-8e70-71f2659659e2" ID-ISSN="1175-5326" ID-Zenodo-Dep="2601072" ID-ZooBank="CF79B01B-8B5F-4B3A-B642-2CADE4B339AF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553171433896" checkinUser="plazi" docAuthor="Wesener, Thomas" docDate="2019" docId="03EB5506E3048C3BA2EDFDD6FA80FD54" docLanguage="en" docName="zootaxa.4563.2.1.pdf" docOrigin="Zootaxa 4563 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Sphaerobelum phouloei Wesener 2019, new species" docType="treatment" docVersion="9" lastPageNumber="216" masterDocId="FFD22D7EE30A8C34A27AFFCFFFE9FFFC" masterDocTitle="First records of giant pill-millipedes from Laos (Diplopoda, Sphaerotheriida, Zephroniidae)" masterLastPageNumber="248" masterPageNumber="201" pageNumber="215" updateTime="1698703176884" updateUser="ExternalLinkService">
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<mods:title id="EEB9210F963A7DC68028BCFF6C3378E7">First records of giant pill-millipedes from Laos (Diplopoda, Sphaerotheriida, Zephroniidae)</mods:title>
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<mods:name id="0A27DB20B866C605F0E1FED7EBD39070" type="personal">
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<mods:namePart id="1EBBD7DF0DADA4942F49C7F4C13D2460">Wesener, Thomas</mods:namePart>
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<mods:title id="221B58E0190A222FDBF58229FD8C1F62">Zootaxa</mods:title>
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<mods:part id="99526893EED6E3A364700A80CF687CBB">
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<mods:date id="45187D0BC450D6228060AC331C9DA400">2019</mods:date>
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<mods:number id="F46737541C2C88AED36B866000E42DA4">2019-03-01</mods:number>
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<mods:number id="9F438396A2D4A79E632465A834A0374F">4563</mods:number>
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<treatment id="03EB5506E3048C3BA2EDFDD6FA80FD54" ID-DOI="http://doi.org/10.5281/zenodo.5934524" ID-GBIF-Taxon="154474556" ID-Zenodo-Dep="5934524" LSID="urn:lsid:plazi:treatment:03EB5506E3048C3BA2EDFDD6FA80FD54" httpUri="http://treatment.plazi.org/id/03EB5506E3048C3BA2EDFDD6FA80FD54" lastPageId="15" lastPageNumber="216" pageId="14" pageNumber="215">
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<subSubSection id="C358B79BE3048C3AA2EDFDD6FDA6FDCF" box="[151,591,537,563]" pageId="14" pageNumber="215" type="nomenclature">
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<paragraph id="8BFDE410E3048C3AA2EDFDD6FDA6FDCF" blockId="14.[151,591,537,597]" box="[151,591,537,563]" pageId="14" pageNumber="215">
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<heading id="D0B5537CE3048C3AA2EDFDD6FDA6FDCF" bold="true" box="[151,591,537,563]" fontSize="11" level="1" pageId="14" pageNumber="215" reason="1">
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<emphasis id="B9363802E3048C3AA2EDFDD6FDA6FDCF" bold="true" box="[151,591,537,563]" pageId="14" pageNumber="215">
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<taxonomicName id="4C429F93E3048C3AA2EDFDD6FE51FDCF" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[151,440,537,563]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="14" pageNumber="215" phylum="Arthropoda" rank="species" species="phouloei" status="sp. nov.">
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<emphasis id="B9363802E3048C3AA2EDFDD6FE51FDCF" bold="true" box="[151,440,537,563]" italics="true" pageId="14" pageNumber="215">Sphaerobelum phouloei</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2058579E3048C3AA3C5FDD6FDA6FDCF" box="[447,591,537,563]" pageId="14" pageNumber="215" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C358B79BE3048C3AA2EDFDF3FE7AFD66" pageId="14" pageNumber="215" type="description">
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<paragraph id="8BFDE410E3048C3AA2EDFDF3FE72FDA8" blockId="14.[151,591,537,597]" box="[151,411,572,597]" pageId="14" pageNumber="215">
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<figureCitation id="1379F895E3048C3AA2EDFDF3FEFCFDA9" box="[151,277,572,597]" captionStart="FIGURE 6" captionStartId="11.[151,250,1916,1938]" captionTargetBox="[150,1423,987,1897]" captionTargetId="figure@11.[150,1436,983,1903]" captionTargetPageId="11" captionText="FIGURE 6. Habitus multi-layer stacked photographs of Sphaerobelum species. A: Sphaerobelum bolavensis n. sp.; B: Sphaerobelum phouloei n. sp.; C: Sphaerobelum denticulatum n. sp.; D: Sphaerobelum spinatum n. sp.; E: Sphaerobelum lachneeis n. sp.; F: Sphaerobelum peterjaegeri n. sp.; G: Sphaerobelum nigrum n. sp.; H: Sphaerobelum splendidum n. sp.; I: Sphaerobelum laoticum n. sp. Not to scale." httpUri="https://zenodo.org/record/2601084/files/figure.png" pageId="14" pageNumber="215">Figures 6B</figureCitation>
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,
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<figureCitation id="1379F895E3048C3AA35AFDF3FEAAFDA8" box="[288,323,572,596]" captionStart="FIGURE 9" captionStartId="13.[151,250,1423,1445]" captionTargetBox="[152,1434,1021,1400]" captionTargetId="figure@13.[152,1435,1021,1402]" captionTargetPageId="13" captionText="FIGURE 9. Endoterga of the body ring 9 of different Sphaerobelum species. A: Sphaerobelum bolavensis n. sp.; B: Sphaerobelum phouloei n. sp.; C: Sphaerobelum spinatum n. sp.; D: Sphaerobelum splendidum n. sp. Abbreviations: cp = cuticular patterns; iA = inner area." httpUri="https://zenodo.org/record/2601090/files/figure.png" pageId="14" pageNumber="215">9B</figureCitation>
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,
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<figureCitation id="1379F895E3048C3AA335FDF3FE85FDA8" box="[335,364,572,596]" captionStart="FIGURE 11" captionStartId="17.[151,250,1968,1990]" captionTargetBox="[150,1436,250,1894]" captionTargetId="figure@17.[150,1436,250,1894]" captionTargetPageId="17" captionText="FIGURE 11. Sphaerobelum phouloei n. sp., male holotype (ZMUC); A: left first coxa with stigmatic plate; B: 9th left leg. Abbreviations: Cx = coxa; St-Pl = stigmatic plate. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601096/files/figure.png" pageId="14" pageNumber="215">11</figureCitation>
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,
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<figureCitation id="1379F895E3048C3AA302FDF3FE7EFDA8" box="[376,407,572,596]" captionStart="FIGURE 12" captionStartId="18.[151,250,1942,1964]" captionTargetBox="[151,1436,247,1876]" captionTargetId="figure@18.[151,1436,243,1876]" captionTargetPageId="18" captionText="FIGURE 12. Sphaerobelum phouloei n. sp., holotype (ZMUC); A: coxa of second left leg with gonopore; B: left anterior telopod, anterior view; C: left anterior telopod, posterior view; D: left anterior telopod, lateral view; E: right posterior telopod, anterior view; F: right posterior telopod, posterior view. Abbreviations: Cx = coxa; Syn-Cx = syncoxite. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601098/files/figure.png" pageId="14" pageNumber="215">12</figureCitation>
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.
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</paragraph>
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<paragraph id="8BFDE410E3048C3AA2EDFD4BFE7AFD66" blockId="14.[151,403,644,666]" box="[151,403,644,666]" pageId="14" pageNumber="215">
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<taxonomicName id="4C429F93E3048C3AA2EDFD4BFEC0FD66" authorityName="Verhoeff" authorityYear="1924" box="[151,297,644,666]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="14" pageNumber="215" phylum="Arthropoda" rank="genus">Sphaerobelum</taxonomicName>
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sp.
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<date id="FFFCC2D0E3048C3AA329FD4AFE9EFD66" box="[339,375,644,666]" pageId="14" pageNumber="215">II 1</date>
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M
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C358B79BE3048C3AA2EDFD09FC7EFCFE" pageId="14" pageNumber="215" type="materials_examined">
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<paragraph id="8BFDE410E3048C3AA2EDFD09FC7EFCFE" blockId="14.[151,1437,710,2031]" pageId="14" pageNumber="215">
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||
<emphasis id="B9363802E3048C3AA2EDFD09FE90FD23" bold="true" box="[151,377,710,735]" pageId="14" pageNumber="215">Material examined</emphasis>
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: Type specimen. 1 M
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<typeStatus id="54F95AB2E3048C3AA00BFD09FD3AFD23" box="[625,723,710,735]" pageId="14" pageNumber="215" type="holotype">holotype</typeStatus>
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(ZMUC00040257) from
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<collectingCountry id="F355A480E3048C3AA188FD08FBC2FD22" box="[1010,1067,711,734]" name="Laos" pageId="14" pageNumber="215">Laos</collectingCountry>
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,
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<collectingRegion id="49862AF2E3048C3AA64CFD09FAFEFD23" box="[1078,1303,710,735]" country="Laos" name="Houaphan" pageId="14" pageNumber="215">Houaphan Province</collectingRegion>
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, Phou Loei, way from camp 1 to camp 2, ~
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<quantity id="4CBA49F5E3048C3AA38FFD24FDA5FCFE" box="[501,588,747,771]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="14" pageNumber="215" unit="m" value="1200.0">1200 m</quantity>
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, leg.
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<date id="FFFCC2D0E3048C3AA0F2FD24FCE6FCFF" box="[648,783,747,771]" pageId="14" pageNumber="215" value="2008-08-09">9.VIII.2008</date>
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; S. Tarasov.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C358B79BE3048C3AA2BDFCC1FB0EFC27" pageId="14" pageNumber="215" type="diagnosis">
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||
<paragraph id="8BFDE410E3048C3AA2BDFCC1FB0EFC27" blockId="14.[151,1437,710,2031]" pageId="14" pageNumber="215">
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||
<emphasis id="B9363802E3048C3AA2BDFCC1FED3FCDB" bold="true" box="[199,314,782,807]" pageId="14" pageNumber="215">Diagnosis</emphasis>
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:
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<taxonomicName id="4C429F93E3048C3AA331FCC0FE22FCDA" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[331,459,782,806]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="14" pageNumber="215" phylum="Arthropoda" rank="species" species="phouloei" status="sp. nov.">
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||
<emphasis id="B9363802E3048C3AA331FCC0FE22FCDA" box="[331,459,782,806]" italics="true" pageId="14" pageNumber="215">S. phouloei</emphasis>
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</taxonomicName>
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<emphasis id="B9363802E3048C3AA3AEFCC1FDFCFCDB" bold="true" box="[468,533,782,807]" pageId="14" pageNumber="215">
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||
<taxonomicNameLabel id="A2058579E3048C3AA3AEFCC1FDFCFCDB" box="[468,533,782,807]" pageId="14" pageNumber="215" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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belongs to a group of
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<taxonomicName id="4C429F93E3048C3AA166FCC1FC54FCDA" authorityName="Verhoeff" authorityYear="1924" box="[796,957,782,806]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="14" pageNumber="215" phylum="Arthropoda" rank="genus">
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<emphasis id="B9363802E3048C3AA166FCC1FC54FCDA" box="[796,957,782,806]" italics="true" pageId="14" pageNumber="215">Sphaerobelum</emphasis>
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</taxonomicName>
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species in which the mesal margin of the femur is extended in several teeth (
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<figureCitation id="1379F895E3048C3AA053FCFDFD79FCB7" box="[553,656,818,843]" captionStart="FIGURE 11" captionStartId="17.[151,250,1968,1990]" captionTargetBox="[150,1436,250,1894]" captionTargetId="figure@17.[150,1436,250,1894]" captionTargetPageId="17" captionText="FIGURE 11. Sphaerobelum phouloei n. sp., male holotype (ZMUC); A: left first coxa with stigmatic plate; B: 9th left leg. Abbreviations: Cx = coxa; St-Pl = stigmatic plate. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601096/files/figure.png" pageId="14" pageNumber="215">Fig. 11B</figureCitation>
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).
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<taxonomicName id="4C429F93E3048C3AA0DFFCFCFCCCFCB7" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[677,805,819,843]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="14" pageNumber="215" phylum="Arthropoda" rank="species" species="phouloei" status="sp. nov.">
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<emphasis id="B9363802E3048C3AA0DFFCFCFCCCFCB7" box="[677,805,819,843]" italics="true" pageId="14" pageNumber="215">S. phouloei</emphasis>
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||
</taxonomicName>
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||
<emphasis id="B9363802E3048C3AA157FCFDFC84FCB7" bold="true" box="[813,877,818,843]" pageId="14" pageNumber="215">
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||
<taxonomicNameLabel id="A2058579E3048C3AA157FCFDFC84FCB7" box="[813,877,818,843]" pageId="14" pageNumber="215" rank="species">n. sp.</taxonomicNameLabel>
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||
</emphasis>
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shares only with
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<taxonomicName id="4C429F93E3048C3AA643FCFCFB00FCB7" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[1081,1257,819,843]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="14" pageNumber="215" phylum="Arthropoda" rank="species" species="denticulatum" status="sp. nov.">
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||
<emphasis id="B9363802E3048C3AA643FCFCFB00FCB7" box="[1081,1257,819,843]" italics="true" pageId="14" pageNumber="215">S. denticulatum</emphasis>
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||
</taxonomicName>
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||
<emphasis id="B9363802E3048C3AA688FCFDFADBFCB7" bold="true" box="[1266,1330,818,843]" pageId="14" pageNumber="215">
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||
<taxonomicNameLabel id="A2058579E3048C3AA688FCFDFADBFCB7" box="[1266,1330,818,843]" pageId="14" pageNumber="215" rank="species">n. sp.</taxonomicNameLabel>
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||
</emphasis>
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a curved telopoditomere 4 of the posterior telopods which overlaps the immovable finger (
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<figureCitation id="1379F895E3048C3AA64FFC99FB56FC93" box="[1077,1215,854,879]" captionStart="FIGURE 12" captionStartId="18.[151,250,1942,1964]" captionTargetBox="[151,1436,247,1876]" captionTargetId="figure@18.[151,1436,243,1876]" captionTargetPageId="18" captionText="FIGURE 12. Sphaerobelum phouloei n. sp., holotype (ZMUC); A: coxa of second left leg with gonopore; B: left anterior telopod, anterior view; C: left anterior telopod, posterior view; D: left anterior telopod, lateral view; E: right posterior telopod, anterior view; F: right posterior telopod, posterior view. Abbreviations: Cx = coxa; Syn-Cx = syncoxite. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601098/files/figure.png" pageId="14" pageNumber="215">Figs 12E, F</figureCitation>
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||
).
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<taxonomicName id="4C429F93E3048C3AA6AEFC98FABAFC92" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[1236,1363,854,878]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="14" pageNumber="215" phylum="Arthropoda" rank="species" species="phouloei" status="sp. nov.">
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||
<emphasis id="B9363802E3048C3AA6AEFC98FABAFC92" box="[1236,1363,854,878]" italics="true" pageId="14" pageNumber="215">S. phouloei</emphasis>
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||
</taxonomicName>
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||
<emphasis id="B9363802E3048C3AA726FC99FA75FC93" bold="true" box="[1372,1436,854,879]" pageId="14" pageNumber="215">
|
||
<taxonomicNameLabel id="A2058579E3048C3AA726FC99FA75FC93" box="[1372,1436,854,879]" pageId="14" pageNumber="215" rank="species">n. sp.</taxonomicNameLabel>
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||
</emphasis>
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||
differs in several characters from
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<taxonomicName id="4C429F93E3048C3AA067FCB4FD27FC6F" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[541,718,891,915]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="14" pageNumber="215" phylum="Arthropoda" rank="species" species="denticulatum" status="sp. nov.">
|
||
<emphasis id="B9363802E3048C3AA067FCB4FD27FC6F" box="[541,718,891,915]" italics="true" pageId="14" pageNumber="215">S. denticulatum</emphasis>
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||
</taxonomicName>
|
||
<emphasis id="B9363802E3048C3AA0A2FCB5FCF0FC6F" bold="true" box="[728,793,890,915]" pageId="14" pageNumber="215">
|
||
<taxonomicNameLabel id="A2058579E3048C3AA0A2FCB5FCF0FC6F" box="[728,793,890,915]" pageId="14" pageNumber="215" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
: male antenna with only 30–35 apical cones (>
|
||
<quantity id="4CBA49F5E3048C3AA73BFCB5FA97FC6F" box="[1345,1406,890,915]" metricMagnitude="0" metricUnit="m" metricValue="1.27" pageId="14" pageNumber="215" unit="in" value="50.0">50 in</quantity>
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||
<taxonomicName id="4C429F93E3048C3AA7FDFCB4FEC0FC4A" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="14" pageNumber="215" phylum="Arthropoda" rank="species" species="denticulatum" status="sp. nov.">
|
||
<emphasis id="B9363802E3048C3AA7FDFCB4FEC0FC4A" italics="true" pageId="14" pageNumber="215">S. denticulatum</emphasis>
|
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</taxonomicName>
|
||
<emphasis id="B9363802E3048C3AA34BFC51FE98FC4B" bold="true" box="[305,369,926,951]" pageId="14" pageNumber="215">
|
||
<taxonomicNameLabel id="A2058579E3048C3AA34BFC51FE98FC4B" box="[305,369,926,951]" pageId="14" pageNumber="215" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
), prefemur mesal margin well-rounded (indentated in
|
||
<taxonomicName id="4C429F93E3048C3AA1A0FC50FB60FC4A" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[986,1161,926,950]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="14" pageNumber="215" phylum="Arthropoda" rank="species" species="denticulatum" status="sp. nov.">
|
||
<emphasis id="B9363802E3048C3AA1A0FC50FB60FC4A" box="[986,1161,926,950]" italics="true" pageId="14" pageNumber="215">S. denticulatum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9363802E3048C3AA6EBFC51FB39FC4B" bold="true" box="[1169,1232,926,951]" pageId="14" pageNumber="215">
|
||
<taxonomicNameLabel id="A2058579E3048C3AA6EBFC51FB39FC4B" box="[1169,1232,926,951]" pageId="14" pageNumber="215" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
), tergites covered with short setae (glabrous in
|
||
<taxonomicName id="4C429F93E3048C3AA3A1FC0CFD60FC27" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[475,649,963,987]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="14" pageNumber="215" phylum="Arthropoda" rank="species" species="denticulatum" status="sp. nov.">
|
||
<emphasis id="B9363802E3048C3AA3A1FC0CFD60FC27" box="[475,649,963,987]" italics="true" pageId="14" pageNumber="215">S. denticulatum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9363802E3048C3AA0EAFC0DFD26FC27" bold="true" box="[656,719,962,987]" pageId="14" pageNumber="215">
|
||
<taxonomicNameLabel id="A2058579E3048C3AA0EAFC0DFD26FC27" box="[656,719,962,987]" pageId="14" pageNumber="215" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
), legs orange (brown in
|
||
<taxonomicName id="4C429F93E3048C3AA199FC0CFB79FC27" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[995,1168,963,987]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="14" pageNumber="215" phylum="Arthropoda" rank="species" species="denticulatum" status="sp. nov.">
|
||
<emphasis id="B9363802E3048C3AA199FC0CFB79FC27" box="[995,1168,963,987]" italics="true" pageId="14" pageNumber="215">S. denticulatum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9363802E3048C3AA6E2FC0DFB3FFC27" bold="true" box="[1176,1238,962,987]" pageId="14" pageNumber="215">
|
||
<taxonomicNameLabel id="A2058579E3048C3AA6E2FC0DFB3FFC27" box="[1176,1238,962,987]" pageId="14" pageNumber="215" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C358B79BE3048C3BA2BDFC29FA74FD78" lastPageId="15" lastPageNumber="216" pageId="14" pageNumber="215" type="description">
|
||
<paragraph id="8BFDE410E3048C3AA2BDFC29FA74FBDF" blockId="14.[151,1437,710,2031]" pageId="14" pageNumber="215">
|
||
<emphasis id="B9363802E3048C3AA2BDFC29FDF0FC03" bold="true" box="[199,537,998,1023]" pageId="14" pageNumber="215">Description. Measurements:</emphasis>
|
||
Body length:
|
||
<typeStatus id="54F95AB2E3048C3AA0C0FC29FCF5FC03" box="[698,796,998,1023]" pageId="14" pageNumber="215" type="holotype">holotype</typeStatus>
|
||
male: length ca
|
||
<quantity id="4CBA49F5E3048C3AA1A2FC29FBA9FC02" box="[984,1088,998,1023]" metricMagnitude="-2" metricUnit="m" metricValue="2.58" pageId="14" pageNumber="215" unit="mm" value="25.8">25.8 mm</quantity>
|
||
. Width, of thoracic shield
|
||
<quantity id="4CBA49F5E3048C3AA711FC28FF2CFBDE" metricMagnitude="-2" metricUnit="m" metricValue="1.33" pageId="14" pageNumber="215" unit="mm" value="13.3">13.3 mm</quantity>
|
||
, of tergite 8 =
|
||
<quantity id="4CBA49F5E3048C3AA315FBC4FE3DFBDE" box="[367,468,1035,1059]" metricMagnitude="-2" metricUnit="m" metricValue="1.41" pageId="14" pageNumber="215" unit="mm" value="14.1">14.1 mm</quantity>
|
||
(= broadest). Height, of thoracic shield =
|
||
<quantity id="4CBA49F5E3048C3AA1CDFBC4FBF9FBDE" box="[951,1040,1035,1059]" metricMagnitude="-3" metricUnit="m" metricValue="8.9" pageId="14" pageNumber="215" unit="mm" value="8.9">8.9 mm</quantity>
|
||
, of tergite 8 =
|
||
<quantity id="4CBA49F5E3048C3AA6C0FBC4FAFBFBDE" box="[1210,1298,1035,1059]" metricMagnitude="-3" metricUnit="m" metricValue="9.3" pageId="14" pageNumber="215" unit="mm" value="9.3">9.3 mm</quantity>
|
||
(= highest).
|
||
</paragraph>
|
||
<paragraph id="8BFDE410E3048C3AA2EDFBE1FCACFB97" blockId="14.[151,1437,710,2031]" pageId="14" pageNumber="215">
|
||
<emphasis id="B9363802E3048C3AA2EDFBE1FEC8FBBB" bold="true" box="[151,289,1070,1095]" pageId="14" pageNumber="215">Coloration:</emphasis>
|
||
in preserved specimens anterior 2/3 of tergites dark olive green, posterior 1/3 black. Head, ventral side, anal shield orange (
|
||
<figureCitation id="1379F895E3048C3AA3D7FB9DFDECFB97" box="[429,517,1106,1131]" captionStart="FIGURE 6" captionStartId="11.[151,250,1916,1938]" captionTargetBox="[150,1423,987,1897]" captionTargetId="figure@11.[150,1436,983,1903]" captionTargetPageId="11" captionText="FIGURE 6. Habitus multi-layer stacked photographs of Sphaerobelum species. A: Sphaerobelum bolavensis n. sp.; B: Sphaerobelum phouloei n. sp.; C: Sphaerobelum denticulatum n. sp.; D: Sphaerobelum spinatum n. sp.; E: Sphaerobelum lachneeis n. sp.; F: Sphaerobelum peterjaegeri n. sp.; G: Sphaerobelum nigrum n. sp.; H: Sphaerobelum splendidum n. sp.; I: Sphaerobelum laoticum n. sp. Not to scale." httpUri="https://zenodo.org/record/2601084/files/figure.png" pageId="14" pageNumber="215">Fig. 6B</figureCitation>
|
||
). Legs and antennae orange.
|
||
</paragraph>
|
||
<paragraph id="8BFDE410E3048C3AA2BDFBB9FD02FABE" blockId="14.[151,1437,710,2031]" pageId="14" pageNumber="215">
|
||
<emphasis id="B9363802E3048C3AA2BDFBB9FEEEFB73" bold="true" box="[199,263,1142,1167]" pageId="14" pageNumber="215">Head</emphasis>
|
||
: Eyes with 65–70 ocelli. Aberrant ocellus located inside antennal groove. Antennae short, with rounded joints, extending posteriorly to leg-pair 4. First antennomere with cuticular scales. Lengths of antennomeres: 1=2=3<4<5<<6. All antennomeres densely pubescent, sensilla basiconica surrounding apical disc. Last antennomere thickened, apically widened and well rounded. Apical disc with ca 36/33 apical cones. Organ of Tömösváry located inside antennal groove.
|
||
<emphasis id="B9363802E3048C3AA0F3FAC9FCBEFAE3" bold="true" box="[649,855,1286,1311]" pageId="14" pageNumber="215">Gnathochilarium</emphasis>
|
||
: structure typical of the order. Palpi sensory cones located in several clusters.
|
||
<emphasis id="B9363802E3048C3AA3BFFAE5FDAAFABF" bold="true" box="[453,579,1322,1347]" pageId="14" pageNumber="215">Mandibles</emphasis>
|
||
: not dissected.
|
||
</paragraph>
|
||
<paragraph id="8BFDE410E3048C3AA2BDFA81FA75FA9B" blockId="14.[151,1437,710,2031]" box="[199,1436,1358,1383]" pageId="14" pageNumber="215">
|
||
<emphasis id="B9363802E3048C3AA2BDFA81FE6CFA9B" bold="true" box="[199,389,1358,1383]" pageId="14" pageNumber="215">Stigmatic plates</emphasis>
|
||
: first stigmatic plate widely rounded, apex well-rounded, curved towards coxa 1 (
|
||
<figureCitation id="1379F895E3048C3AA75DFA81FA66FA9B" box="[1319,1423,1358,1383]" captionStart="FIGURE 11" captionStartId="17.[151,250,1968,1990]" captionTargetBox="[150,1436,250,1894]" captionTargetId="figure@17.[150,1436,250,1894]" captionTargetPageId="17" captionText="FIGURE 11. Sphaerobelum phouloei n. sp., male holotype (ZMUC); A: left first coxa with stigmatic plate; B: 9th left leg. Abbreviations: Cx = coxa; St-Pl = stigmatic plate. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601096/files/figure.png" pageId="14" pageNumber="215">Fig. 11A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BFDE410E3048C3AA2EDFABDFD09F87F" blockId="14.[151,1437,710,2031]" pageId="14" pageNumber="215">
|
||
<emphasis id="B9363802E3048C3AA2EDFABDFEA9FA77" bold="true" box="[151,320,1394,1419]" pageId="14" pageNumber="215">Laterotergites</emphasis>
|
||
: laterotergite 1 and 2 with a broad, stout projection.
|
||
<emphasis id="B9363802E3048C3AA1E6FABDFC1DFA77" bold="true" box="[924,1012,1394,1419]" pageId="14" pageNumber="215">Collum</emphasis>
|
||
: with glabrous surface, margins with few isolated setae.
|
||
<emphasis id="B9363802E3048C3AA317FA59FDCFFA53" bold="true" box="[365,550,1430,1455]" pageId="14" pageNumber="215">Thoracic shield</emphasis>
|
||
: surface glabrous unlike tergites, setae only in grooves. Shallow grooves beset with numerous long setae, slope towards groove with 3 weak anterior and 2 or 3 posterior keels.
|
||
<emphasis id="B9363802E3048C3AA6ACFA75FADCFA2F" bold="true" box="[1238,1333,1466,1491]" pageId="14" pageNumber="215">Tergites</emphasis>
|
||
: anterior 2/3 of surface densely setose with short setae, posterior 1/3 glabrous. Tips of paratergites of midbody tergites projecting posteriorly (
|
||
<figureCitation id="1379F895E3048C3AA3E4F9CDFE13F9E7" box="[414,506,1538,1563]" captionStart="FIGURE 6" captionStartId="11.[151,250,1916,1938]" captionTargetBox="[150,1423,987,1897]" captionTargetId="figure@11.[150,1436,983,1903]" captionTargetPageId="11" captionText="FIGURE 6. Habitus multi-layer stacked photographs of Sphaerobelum species. A: Sphaerobelum bolavensis n. sp.; B: Sphaerobelum phouloei n. sp.; C: Sphaerobelum denticulatum n. sp.; D: Sphaerobelum spinatum n. sp.; E: Sphaerobelum lachneeis n. sp.; F: Sphaerobelum peterjaegeri n. sp.; G: Sphaerobelum nigrum n. sp.; H: Sphaerobelum splendidum n. sp.; I: Sphaerobelum laoticum n. sp. Not to scale." httpUri="https://zenodo.org/record/2601084/files/figure.png" pageId="14" pageNumber="215">Fig. 6B</figureCitation>
|
||
).
|
||
<emphasis id="B9363802E3048C3AA06AF9CDFD4FF9E7" bold="true" box="[528,678,1538,1563]" pageId="14" pageNumber="215">Endotergum</emphasis>
|
||
(
|
||
<figureCitation id="1379F895E3048C3AA0C3F9CDFCFCF9E7" box="[697,789,1538,1563]" captionStart="FIGURE 9" captionStartId="13.[151,250,1423,1445]" captionTargetBox="[152,1434,1021,1400]" captionTargetId="figure@13.[152,1435,1021,1402]" captionTargetPageId="13" captionText="FIGURE 9. Endoterga of the body ring 9 of different Sphaerobelum species. A: Sphaerobelum bolavensis n. sp.; B: Sphaerobelum phouloei n. sp.; C: Sphaerobelum spinatum n. sp.; D: Sphaerobelum splendidum n. sp. Abbreviations: cp = cuticular patterns; iA = inner area." httpUri="https://zenodo.org/record/2601090/files/figure.png" pageId="14" pageNumber="215">Fig. 9B</figureCitation>
|
||
): inner section lacking any spines or setae. Middle area with a single row of large, sparse, elliptical, cuticular impressions. Distance between impressions as wide as their diameter. Apically, 1–2 sparse rows of long marginal bristles, strongly protruding beyond tergal margin. Bristles not smooth, but with numerous small spinicles.
|
||
<emphasis id="B9363802E3048C3AA0B4F9A1FCB0F97B" bold="true" box="[718,857,1646,1671]" pageId="14" pageNumber="215">Anal shield</emphasis>
|
||
: large, surface completely covered by tiny setae. Underside with a single, long, black locking carina, located close to last laterotergite.
|
||
<emphasis id="B9363802E3048C3AA6F1F95DFB2AF957" bold="true" box="[1163,1219,1682,1707]" pageId="14" pageNumber="215">Legs</emphasis>
|
||
: leg-pair 1 with 2 ventral spines, leg-pair 2 with 5, leg-pair 3 with 6 or 7. First two leg-pairs without an apical spine. Leg pairs 4–21 with 9 or 10 ventral spines and one dorso-apical spine. In leg 9, femur 1.3 times, tarsus 3.7 times longer than wide (
|
||
<figureCitation id="1379F895E3048C3AA2E5F931FEECF8EB" box="[159,261,1790,1815]" captionStart="FIGURE 11" captionStartId="17.[151,250,1968,1990]" captionTargetBox="[150,1436,250,1894]" captionTargetId="figure@17.[150,1436,250,1894]" captionTargetPageId="17" captionText="FIGURE 11. Sphaerobelum phouloei n. sp., male holotype (ZMUC); A: left first coxa with stigmatic plate; B: 9th left leg. Abbreviations: Cx = coxa; St-Pl = stigmatic plate. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601096/files/figure.png" pageId="14" pageNumber="215">Fig. 11B</figureCitation>
|
||
). All podomeres densely setose. Coxa with a large and marginally toothed process. Coxa process absent at first leg (
|
||
<figureCitation id="1379F895E3048C3AA35AF8EDFE6FF8C7" box="[288,390,1826,1851]" captionStart="FIGURE 11" captionStartId="17.[151,250,1968,1990]" captionTargetBox="[150,1436,250,1894]" captionTargetId="figure@17.[150,1436,250,1894]" captionTargetPageId="17" captionText="FIGURE 11. Sphaerobelum phouloei n. sp., male holotype (ZMUC); A: left first coxa with stigmatic plate; B: 9th left leg. Abbreviations: Cx = coxa; St-Pl = stigmatic plate. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601096/files/figure.png" pageId="14" pageNumber="215">Fig. 11B</figureCitation>
|
||
) and sharply projecting at second (
|
||
<figureCitation id="1379F895E3048C3AA15BF8EDFC65F8C7" box="[801,908,1826,1851]" captionStart="FIGURE 12" captionStartId="18.[151,250,1942,1964]" captionTargetBox="[151,1436,247,1876]" captionTargetId="figure@18.[151,1436,243,1876]" captionTargetPageId="18" captionText="FIGURE 12. Sphaerobelum phouloei n. sp., holotype (ZMUC); A: coxa of second left leg with gonopore; B: left anterior telopod, anterior view; C: left anterior telopod, posterior view; D: left anterior telopod, lateral view; E: right posterior telopod, anterior view; F: right posterior telopod, posterior view. Abbreviations: Cx = coxa; Syn-Cx = syncoxite. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601098/files/figure.png" pageId="14" pageNumber="215">Fig. 12A</figureCitation>
|
||
). Prefemur at apical margin with a projection laterally and mesally. Lateral projection triangular and sharply edged, juxtaposed to coxal process. Femur extended mesally into a dentate margin featuring 12–14 teeth.
|
||
</paragraph>
|
||
<paragraph id="8BFDE410E3048C3AA2BDF841FE79F85B" blockId="14.[151,1437,710,2031]" box="[199,400,1934,1959]" pageId="14" pageNumber="215">Female unknown.</paragraph>
|
||
<paragraph id="8BFDE410E3048C3BA2BDF87DFA74FD78" blockId="14.[151,1437,710,2031]" lastBlockId="15.[151,1437,151,680]" lastPageId="15" lastPageNumber="216" pageId="14" pageNumber="215">
|
||
<emphasis id="B9363802E3048C3AA2BDF87DFE08F837" bold="true" box="[199,481,1970,1995]" pageId="14" pageNumber="215">Male sexual characters</emphasis>
|
||
: gonopore covered with a single, undivided, circular, sclerotized plate (
|
||
<figureCitation id="1379F895E3048C3AA75EF87DFA66F837" box="[1316,1423,1970,1995]" captionStart="FIGURE 12" captionStartId="18.[151,250,1942,1964]" captionTargetBox="[151,1436,247,1876]" captionTargetId="figure@18.[151,1436,243,1876]" captionTargetPageId="18" captionText="FIGURE 12. Sphaerobelum phouloei n. sp., holotype (ZMUC); A: coxa of second left leg with gonopore; B: left anterior telopod, anterior view; C: left anterior telopod, posterior view; D: left anterior telopod, lateral view; E: right posterior telopod, anterior view; F: right posterior telopod, posterior view. Abbreviations: Cx = coxa; Syn-Cx = syncoxite. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601098/files/figure.png" pageId="14" pageNumber="215">Fig. 12A</figureCitation>
|
||
).
|
||
<emphasis id="B9363802E3048C3AA2EDF819FE84F813" bold="true" box="[151,365,2006,2031]" pageId="14" pageNumber="215">Anterior telopods</emphasis>
|
||
(
|
||
<figureCitation id="1379F895E3048C3AA304F819FE3FF813" box="[382,470,2006,2031]" captionStart="FIGURE 12" captionStartId="18.[151,250,1942,1964]" captionTargetBox="[151,1436,247,1876]" captionTargetId="figure@18.[151,1436,243,1876]" captionTargetPageId="18" captionText="FIGURE 12. Sphaerobelum phouloei n. sp., holotype (ZMUC); A: coxa of second left leg with gonopore; B: left anterior telopod, anterior view; C: left anterior telopod, posterior view; D: left anterior telopod, lateral view; E: right posterior telopod, anterior view; F: right posterior telopod, posterior view. Abbreviations: Cx = coxa; Syn-Cx = syncoxite. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601098/files/figure.png" pageId="14" pageNumber="215">Figs 12</figureCitation>
|
||
B–D): consisting of 4 telopoditomeres above syncoxite, but telopoditomere 3 and 4 only divided by thin suture. Telopoditomere 1 rectangular, as long as wide. Telopoditomere 2 large, almost as long as telopoditomere 3 and 4 combined. Process of telopoditomere 2 located posteriorly, not visible in anterior view. Process of telopoditomere 2 wide, projecting to half of telopoditomere 4, apically with a well-rounded tip. Telopoditomere 3 cylindrical, straight, as long as telopoditomere 4. Telopoditomere 4 cylindrical, well-rounded, posterior side with a black sclerotized spot and a small, triangular spine. Telopoditomeres
|
||
<quantity id="4CBA49F5E3058C3BA6CAFEE7FB12FEBC" box="[1200,1275,295,320]" metricMagnitude="-2" metricUnit="m" metricValue="6.35" metricValueMax="10.16" metricValueMin="2.54" pageId="15" pageNumber="216" unit="in" value="2.5" valueMax="4.0" valueMin="1.0">1–4 in</quantity>
|
||
anterior view covered with long setae. In posterior view all telopoditomeres with fewer setae.
|
||
<emphasis id="B9363802E3058C3BA664FE83FB11FE99" bold="true" box="[1054,1272,332,357]" pageId="15" pageNumber="216">Posterior telopods</emphasis>
|
||
(
|
||
<figureCitation id="1379F895E3058C3BA77DFE83FA67FE98" box="[1287,1422,332,357]" captionStart="FIGURE 12" captionStartId="18.[151,250,1942,1964]" captionTargetBox="[151,1436,247,1876]" captionTargetId="figure@18.[151,1436,243,1876]" captionTargetPageId="18" captionText="FIGURE 12. Sphaerobelum phouloei n. sp., holotype (ZMUC); A: coxa of second left leg with gonopore; B: left anterior telopod, anterior view; C: left anterior telopod, posterior view; D: left anterior telopod, lateral view; E: right posterior telopod, anterior view; F: right posterior telopod, posterior view. Abbreviations: Cx = coxa; Syn-Cx = syncoxite. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601098/files/figure.png" pageId="15" pageNumber="216">Figs 12E, F</figureCitation>
|
||
): telopoditomere 1 elongated, twice as long as wide. Immovable finger (process of telopoditomere 2) shorter than movable finger, consisting of telopoditomeres 3 and 4. Immovable finger with a characteristic, distally swollen apex, well rounded, apex therefore wider than base. Immovable finger in anterior view with a large spine, at tip opposite to tip of telopoditomere 4 with sclerotized spots. Telopoditomere 3 rectangular, towards immoveable finger with a triangular expansion carrying a spine. Telopoditomere 4 reaching 2/3 of length of telopoditomere 3, apically strongly tapering into acute tip, very slightly curved towards immovable finger. Inner margin with single spine towards immovable finger. Telopoditomere 1 at both sides covered by setae. Telopoditomeres 2 and
|
||
<quantity id="4CBA49F5E3058C3BA714FD87FA75FD9C" box="[1390,1436,583,608]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="15" pageNumber="216" unit="in" value="3.0">3 in</quantity>
|
||
anterior view covered by long setae, in posterior view mostly glabrous. Telopoditomere 4 only with marginal setae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C358B79BE3058C3BA2BDFD40FA80FD54" box="[199,1385,655,680]" pageId="15" pageNumber="216" type="etymology">
|
||
<paragraph id="8BFDE410E3058C3BA2BDFD40FA80FD54" blockId="15.[151,1437,151,680]" box="[199,1385,655,680]" pageId="15" pageNumber="216">
|
||
<emphasis id="B9363802E3058C3BA2BDFD40FE74FD54" bold="true" box="[199,413,655,680]" pageId="15" pageNumber="216">Derivatio nominis</emphasis>
|
||
: phouloei, noun in apposition, after the Phou
|
||
<collectingRegion id="49862AF2E3058C3BA1E4FD40FC3BFD54" box="[926,978,655,680]" country="Thailand" name="Loei" pageId="15" pageNumber="216">Loei</collectingRegion>
|
||
Mountain, the
|
||
<typeStatus id="54F95AB2E3058C3BA606FD5FFB45FD54" box="[1148,1196,656,680]" pageId="15" pageNumber="216">type</typeStatus>
|
||
locality (
|
||
<figureCitation id="1379F895E3058C3BA76DFD40FAB2FD54" box="[1303,1371,655,680]" captionStart="FIGURE 5" captionStartId="9.[151,250,2005,2027]" captionTargetBox="[185,1415,664,1984]" captionTargetId="figure@9.[181,1415,663,1984]" captionTargetPageId="9" captionText="FIGURE 5. Distribution map of Sphaerobelum species, including the unnamed females." httpUri="https://zenodo.org/record/2601082/files/figure.png" pageId="15" pageNumber="216">Fig. 5</figureCitation>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |