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<mods:title id="D55B6266EE37794254DF64FAC784E693">Integrative taxonomic revision of the African taxa of the Racotis Moore, 1887 generic complex (Lepidoptera, Geometridae, Ennominae, Boarmiini)</mods:title>
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<paragraph id="8BFDF53D5D3BFFBA57F5507EFEDDF811" blockId="6.[151,378,1914,1941]" box="[151,378,1914,1941]" pageId="6" pageNumber="7">
<heading id="D0B542515D3BFFBA57F5507EFEDDF811" bold="true" box="[151,378,1914,1941]" fontSize="11" level="2" pageId="6" pageNumber="7" reason="4">
<emphasis id="B936292F5D3BFFBA57F5507EFEDDF811" bold="true" box="[151,378,1914,1941]" pageId="6" pageNumber="7">
<taxonomicName id="4C428EBE5D3BFFBA57F5507EFE87F810" authority="László &amp; Hausmann &amp; Karisch, 2023" authorityName="László &amp; Hausmann &amp; Karisch" authorityYear="2023" box="[151,288,1914,1940]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B936292F5D3BFFBA57F5507EFE87F810" bold="true" box="[151,288,1914,1940]" italics="true" pageId="6" pageNumber="7">Afroracotis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20594545D3BFFBA5645507FFEDDF811" box="[295,378,1915,1941]" pageId="6" pageNumber="7" rank="genus">gen. n.</taxonomicNameLabel>
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<subSubSection id="C358A6B65D3BFFBA57F550BEFC2AF856" box="[151,909,1978,2002]" pageId="6" pageNumber="7" type="type_taxon">
<paragraph id="8BFDF53D5D3BFFBA57F550BEFC2AF856" blockId="6.[151,909,1978,2002]" box="[151,909,1978,2002]" pageId="6" pageNumber="7">
<typeStatus id="54F94B9F5D3BFFBA57F550BEFF6EF856" box="[151,201,1978,2002]" pageId="6" pageNumber="7">Type</typeStatus>
species:
<taxonomicName id="4C428EBE5D3BFFBA564450BEFD3BF856" ID-CoL="97RK5" authority="Butler, 1878" authorityName="Butler" authorityYear="1878" box="[294,668,1978,2002]" class="Insecta" family="Geometridae" genus="Ophthalmodes" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="squalida">
<emphasis id="B936292F5D3BFFBA564450BEFDB0F856" box="[294,535,1978,2002]" italics="true" pageId="6" pageNumber="7">Ophthalmodes squalida</emphasis>
<bibRefCitation id="EFD388CC5D3BFFBA557C50BEFD3BF856" author="Butler, A. G." box="[542,668,1978,2002]" pageId="6" pageNumber="7" pagination="455 - 465" refId="ref82974" refString="Butler, A. G. (1878) Descriptions of some new genera and species of Lepidoptera from Old Calabar and Madagascar. Annals and Magazine of Natural History, Series 5, 2 (12), 455 - 465." type="journal article" year="1878">Butler, 1878</bibRefCitation>
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by present designation.
</paragraph>
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<subSubSection id="C358A6B65D3AFFBB57F55792FE57FA45" pageId="7" pageNumber="8" type="discussion">
<paragraph id="8BFDF53D5D3AFFBB57F55792FDD2FF35" blockId="7.[151,1437,150,2013]" box="[151,629,150,177]" pageId="7" pageNumber="8">
<emphasis id="B936292F5D3AFFBB57F55792FEF9FF35" bold="true" box="[151,350,150,177]" pageId="7" pageNumber="8">Genetic analyses</emphasis>
(
<figureCitation id="1379E9B85D3AFFBB560F5793FE69FF35" box="[365,462,151,177]" captionStart="FIGURE 340" captionStartId="102.[151,250,1920,1944]" captionTargetBox="[169,1417,185,1890]" captionTargetId="figure-21@102.[163,1425,181,1896]" captionTargetPageId="102" captionText="FIGURE 340. ML tree with collapsed branches displaying the relationships of the Racotis s.l. genera based on analysis of COI barcodes. Bootstrap values are indicated above, posterior probability values below the branches." figureDoi="http://doi.org/10.5281/zenodo.8081011" httpUri="https://zenodo.org/record/8081011/files/figure.png" pageId="7" pageNumber="8">Fig. 340</figureCitation>
, Text figs 13)
</paragraph>
<paragraph id="8BFDF53D5D3AFFBB57A557BFFBD5FEC5" blockId="7.[151,1437,150,2013]" pageId="7" pageNumber="8">
Preliminary ML analyses of COI barcode data for
<specimenCount id="9D443EB45D3AFFBB559157BFFC3FFF51" box="[755,920,187,213]" pageId="7" pageNumber="8" type="generic">287 specimens</specimenCount>
of
<taxonomicName id="4C428EBE5D3AFFBB54D857BFFBA9FF50" authorityName="Moore" authorityYear="1887" box="[954,1038,187,212]" class="Insecta" family="Geometridae" genus="Racotis" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus" sensu="lato">
<emphasis id="B936292F5D3AFFBB54D857BFFBA9FF50" box="[954,1038,187,212]" italics="true" pageId="7" pageNumber="8">Racotis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20594545D3AFFBB537657BFFB92FF51" box="[1044,1077,187,213]" pageId="7" pageNumber="8" sensu="lato">s.l.</taxonomicNameLabel>
in BOLD (&gt;500 bp, worldwide) showed a pattern of several well-separated clusters diverging at distances of 57% from each other. All Indomalayan species clustered into two clades, one which included the
<typeStatus id="54F94B9F5D3AFFBB54505607FCC5FE99" box="[818,866,259,285]" pageId="7" pageNumber="8">type</typeStatus>
species of the genus,
<taxonomicName id="4C428EBE5D3AFFBB53005607FAE7FE99" box="[1122,1344,259,285]" class="Insecta" family="Geometridae" genus="Racotis" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="boarmiaria">
<emphasis id="B936292F5D3AFFBB53005607FAE7FE99" box="[1122,1344,259,285]" italics="true" pageId="7" pageNumber="8">Racotis boarmiaria</emphasis>
</taxonomicName>
and the other
<taxonomicName id="4C428EBE5D3AFFBB57BA5623FE0EFEC5" authority="Lucas" authorityName="Lucas" authorityYear="1889" box="[216,425,295,321]" class="Insecta" family="Geometridae" genus="Racotis" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="maculata">
<emphasis id="B936292F5D3AFFBB57BA5623FEF8FEC5" box="[216,351,295,321]" italics="true" pageId="7" pageNumber="8">R. maculata</emphasis>
Lucas
</taxonomicName>
and associated taxa, while all African taxa clustered separately.
</paragraph>
<paragraph id="8BFDF53D5D3AFFBB57A5564FFD84FD9D" blockId="7.[151,1437,150,2013]" pageId="7" pageNumber="8">
In order to obtain a clearer picture of the phylogenetic relationship between the Asian and African lineages of
<taxonomicName id="4C428EBE5D3AFFBB57F5566BFF4CFE0C" authorityName="Moore" authorityYear="1887" box="[151,235,367,392]" class="Insecta" family="Geometridae" genus="Racotis" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus" sensu="lato">
<emphasis id="B936292F5D3AFFBB57F5566BFF4CFE0C" box="[151,235,367,392]" italics="true" pageId="7" pageNumber="8">Racotis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20594545D3AFFBB5791566BFEB2FE0D" box="[243,277,367,393]" pageId="7" pageNumber="8" sensu="lato">s.l.</taxonomicNameLabel>
, as well as within the Afrotropical taxa, 179 morphologically identified specimens were analysed. The phylogenetic inferences obtained from BI and ML analyses recovered similar topologies, although the BI tree was not fully resolved with the Indomalayan
<taxonomicName id="4C428EBE5D3AFFBB553B56B3FD7EFE55" box="[601,729,439,465]" class="Insecta" family="Geometridae" genus="Racotis" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="boarmiaria">
<emphasis id="B936292F5D3AFFBB553B56B3FD7EFE55" box="[601,729,439,465]" italics="true" pageId="7" pageNumber="8">boarmiaria</emphasis>
</taxonomicName>
and the Afrotropical
<taxonomicName id="4C428EBE5D3AFFBB54A656B3FB98FE55" authorityName="László &amp; Hausmann &amp; Karisch" authorityYear="2023" baseAuthorityName="Prout" baseAuthorityYear="1931" box="[964,1087,439,465]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="apodosima">
<emphasis id="B936292F5D3AFFBB54A656B3FB98FE55" box="[964,1087,439,465]" italics="true" pageId="7" pageNumber="8">apodosima</emphasis>
</taxonomicName>
,
<taxonomicName id="4C428EBE5D3AFFBB532E56B3FB19FE55" baseAuthorityName="Warren" baseAuthorityYear="1900" box="[1100,1214,439,465]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="argillacea">
<emphasis id="B936292F5D3AFFBB532E56B3FB19FE55" box="[1100,1214,439,465]" italics="true" pageId="7" pageNumber="8">argillacea</emphasis>
</taxonomicName>
and
<taxonomicName id="4C428EBE5D3AFFBB539156B3FA3AFE55" authority="László &amp; Hausmann &amp; Karisch, 2023" authorityName="László &amp; Hausmann &amp; Karisch" authorityYear="2023" box="[1267,1437,439,465]" pageId="7" pageNumber="8" rank="subGenus" status="subgen. nov." subGenus="Rwandaracotis">
<emphasis id="B936292F5D3AFFBB539156B3FA3AFE55" box="[1267,1437,439,465]" italics="true" pageId="7" pageNumber="8">Rwandaracotis</emphasis>
</taxonomicName>
<emphasis id="B936292F5D3AFFBB57F556DFFEB7FE71" bold="true" box="[151,272,475,501]" pageId="7" pageNumber="8">
<taxonomicNameLabel id="A20594545D3AFFBB57F556DFFEB7FE71" box="[151,272,475,501]" pageId="7" pageNumber="8" rank="subGenus">subgen. n.</taxonomicNameLabel>
</emphasis>
lineages recovered as a polytomy. Due to its better resolution, the ML tree was chosen for illustration in this paper (
<figureCitation id="1379E9B85D3AFFBB567656FBFED1FD9D" box="[276,374,511,537]" captionStart="FIGURE 340" captionStartId="102.[151,250,1920,1944]" captionTargetBox="[169,1417,185,1890]" captionTargetId="figure-21@102.[163,1425,181,1896]" captionTargetPageId="102" captionText="FIGURE 340. ML tree with collapsed branches displaying the relationships of the Racotis s.l. genera based on analysis of COI barcodes. Bootstrap values are indicated above, posterior probability values below the branches." figureDoi="http://doi.org/10.5281/zenodo.8081011" httpUri="https://zenodo.org/record/8081011/files/figure.png" pageId="7" pageNumber="8">Fig. 340</figureCitation>
, Text figs 13).
</paragraph>
<paragraph id="8BFDF53D5D3AFFBB57A55527FDD3FCBD" blockId="7.[151,1437,150,2013]" pageId="7" pageNumber="8">
In the ML analysis, the outgroup taxon
<taxonomicName id="4C428EBE5D3AFFBB55F35527FC4BFDB9" authority="(Warren, 1897)" baseAuthorityName="Warren" baseAuthorityYear="1897" box="[657,1004,547,573]" class="Insecta" family="Geometridae" genus="Eulycia" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="grisea">
<emphasis id="B936292F5D3AFFBB55F35527FC91FDB8" box="[657,822,547,573]" italics="true" pageId="7" pageNumber="8">Eulycia grisea</emphasis>
(
<bibRefCitation id="EFD388CC5D3AFFBB54255527FC43FDB9" author="Warren, W." box="[839,996,547,573]" pageId="7" pageNumber="8" pagination="12 - 130" refId="ref85161" refString="Warren, W. (1897) New genera and species of moths from the Old - World regions in the Tring Museum. Novitates Zoologicae, 4, 12 - 130." type="journal article" year="1897">Warren, 1897</bibRefCitation>
)
</taxonomicName>
+ Asian
<taxonomicName id="4C428EBE5D3AFFBB53355527FB0CFDB8" authorityName="Moore" authorityYear="1887" box="[1111,1195,547,572]" class="Insecta" family="Geometridae" genus="Racotis" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B936292F5D3AFFBB53355527FB0CFDB8" box="[1111,1195,547,572]" italics="true" pageId="7" pageNumber="8">Racotis</emphasis>
</taxonomicName>
clade was recovered as sister to all other African taxa. All African taxa were recovered as monophyletic, although with weak support (BS: 12; PP: -) but considering their genital morphology, the African
<taxonomicName id="4C428EBE5D3AFFBB54D2556FFBA3FD00" authorityName="Moore" authorityYear="1887" box="[944,1028,619,644]" class="Insecta" family="Geometridae" genus="Racotis" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B936292F5D3AFFBB54D2556FFBA3FD00" box="[944,1028,619,644]" italics="true" pageId="7" pageNumber="8">Racotis</emphasis>
</taxonomicName>
are recognised here as belonging to a distinct genus:
<taxonomicName id="4C428EBE5D3AFFBB563E558BFE7CFD2D" authority="László &amp; Hausmann &amp; Karisch, 2023" authorityName="László &amp; Hausmann &amp; Karisch" authorityYear="2023" box="[348,475,655,681]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B936292F5D3AFFBB563E558BFE7CFD2D" box="[348,475,655,681]" italics="true" pageId="7" pageNumber="8">Afroracotis</emphasis>
</taxonomicName>
<emphasis id="B936292F5D3AFFBB5686558BFD92FD2D" bold="true" box="[484,565,655,681]" pageId="7" pageNumber="8">
<taxonomicNameLabel id="A20594545D3AFFBB5686558BFD92FD2D" box="[484,565,655,681]" pageId="7" pageNumber="8" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
Surprisingly, the analyses also suggested that
<taxonomicName id="4C428EBE5D3AFFBB532C558BFF70FD49" authority="(Swinhoe, 1904)" baseAuthorityName="Swinhoe" baseAuthorityYear="1904" class="Insecta" family="Geometridae" genus="Racotis" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="ugandaria">
<emphasis id="B936292F5D3AFFBB532C558BFA86FD2D" box="[1102,1313,655,681]" italics="true" pageId="7" pageNumber="8">Racotis ugandaria</emphasis>
(
<bibRefCitation id="EFD388CC5D3AFFBB5250558BFF68FD49" author="Swinhoe, C." pageId="7" pageNumber="8" pagination="497 - 590" refId="ref84919" refString="Swinhoe, C. (1904) On the Geometridae of tropical Africa in the National Collection. Transactions of the entomological Society of London, 1904 (3), 497 - 590." type="journal article" year="1904">Swinhoe, 1904</bibRefCitation>
)
</taxonomicName>
—a species which has long been treated as the sole Afrotropical member of the genus
<taxonomicName id="4C428EBE5D3AFFBB53B055B7FE72FD75" authority="Walker (Scoble &amp; Hausmann 2007)" authorityName="Walker (Scoble &amp; Hausmann" authorityYear="2007" class="Insecta" family="Geometridae" genus="Chorodna" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B936292F5D3AFFBB53B055B7FAE4FD49" box="[1234,1347,691,717]" italics="true" pageId="7" pageNumber="8">Chorodna</emphasis>
Walker (
<bibRefCitation id="EFD388CC5D3AFFBB57FD55D3FE6AFD75" author="Scoble, M. J. &amp; Hausmann, A." box="[159,461,727,753]" pageId="7" pageNumber="8" refId="ref84778" refString="Scoble, M. J. &amp; Hausmann, A. (2007) Online list of valid and available names of the Geometridae of the world. Available from: http: // www. lepbarcoding. org / geometridae / species _ checklists. php (accessed 31 May 2023)" type="url" year="2007">Scoble &amp; Hausmann 2007</bibRefCitation>
)
</taxonomicName>
and recently transferred to
<taxonomicName id="4C428EBE5D3AFFBB547255D3FB0BFD75" authority="(Murillo-Ramos et al. 2021)" baseAuthorityName="Murillo-Ramos" baseAuthorityYear="2021" box="[784,1196,727,753]" class="Insecta" family="Geometridae" genus="Racotis" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B936292F5D3AFFBB547255D3FCC3FD74" box="[784,868,727,752]" italics="true" pageId="7" pageNumber="8">Racotis</emphasis>
(
<bibRefCitation id="EFD388CC5D3AFFBB541655D3FB03FD75" author="Murillo-Ramos, L. &amp; Chazot, N. &amp; Sihvonen, P. &amp; Ounap, E. &amp; Jiang, N. &amp; Han, H. &amp; Clarke, J. T. &amp; Davis, R. B. &amp; Tammaru, T. &amp; Wahlberg, N." box="[884,1188,727,753]" pageId="7" pageNumber="8" pagination="1 - 10" refId="ref83942" refString="Murillo-Ramos, L., Chazot, N., Sihvonen, P., Ounap, E., Jiang, N., Han, H., Clarke, J. T., Davis, R. B., Tammaru, T. &amp; Wahlberg, N. (2021) Molecular phylogeny, classification, biogeography and diversification patterns of a diverse group of moths (Geometridae, Boarmiini). Molecular Phylogenetics and Evolution, 162 (2021), 107198, 1 - 10." type="journal article" year="2021">
Murillo-Ramos
<emphasis id="B936292F5D3AFFBB534955D3FBC2FD75" box="[1067,1125,727,753]" italics="true" pageId="7" pageNumber="8">et al.</emphasis>
2021
</bibRefCitation>
)
</taxonomicName>
—does not belong to either the Indomalayan
<taxonomicName id="4C428EBE5D3AFFBB56CB55FFFE5AFC90" box="[425,509,763,788]" class="Insecta" family="Geometridae" genus="Racotis" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B936292F5D3AFFBB56CB55FFFE5AFC90" box="[425,509,763,788]" italics="true" pageId="7" pageNumber="8">Racotis</emphasis>
</taxonomicName>
or the Afrotropical
<taxonomicName id="4C428EBE5D3AFFBB558555FFFCC1FC91" authorityName="László &amp; Hausmann &amp; Karisch" authorityYear="2023" box="[743,870,763,789]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B936292F5D3AFFBB558555FFFCC1FC91" box="[743,870,763,789]" italics="true" pageId="7" pageNumber="8">Afroracotis</emphasis>
</taxonomicName>
lineages but represents a distinct genus which is described in this paper:
<taxonomicName id="4C428EBE5D3AFFBB56C1541BFDB9FCBD" authority="László &amp; Hausmann &amp; Karisch, 2023" authorityName="László &amp; Hausmann &amp; Karisch" authorityYear="2023" box="[419,542,799,825]" class="Insecta" family="Geometridae" genus="Chorocotis" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B936292F5D3AFFBB56C1541BFDB9FCBD" box="[419,542,799,825]" italics="true" pageId="7" pageNumber="8">Chorocotis</emphasis>
</taxonomicName>
<emphasis id="B936292F5D3AFFBB5547541BFDD3FCBD" bold="true" box="[549,628,799,825]" pageId="7" pageNumber="8">
<taxonomicNameLabel id="A20594545D3AFFBB5547541BFDD3FCBD" box="[549,628,799,825]" pageId="7" pageNumber="8" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BFDF53D5D3AFFBB57A55447FEEFFC69" blockId="7.[151,1437,150,2013]" pageId="7" pageNumber="8">
A morphologically rather distinct, hitherto undescribed species from
<collectingCountry id="F355B5AD5D3AFFBB54B85447FB91FCD9" box="[986,1078,835,861]" name="Rwanda" pageId="7" pageNumber="8">Rwanda</collectingCountry>
was recovered as a basal sister branch to all other
<taxonomicName id="4C428EBE5D3AFFBB560D5463FE49FC05" authorityName="László &amp; Hausmann &amp; Karisch" authorityYear="2023" box="[367,494,871,897]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B936292F5D3AFFBB560D5463FE49FC05" box="[367,494,871,897]" italics="true" pageId="7" pageNumber="8">Afroracotis</emphasis>
</taxonomicName>
clades (BS: 12; PP: -), and it is recognised here as a representative of a monotypic subgenus
<taxonomicName id="4C428EBE5D3AFFBB5665548FFE16FC21" authority="László &amp; Hausmann &amp; Karisch, 2023" authorityName="László &amp; Hausmann &amp; Karisch" authorityYear="2023" box="[263,433,907,933]" pageId="7" pageNumber="8" rank="subGenus" status="subgen. nov." subGenus="Rwandaracotis">
<emphasis id="B936292F5D3AFFBB5665548FFE16FC21" box="[263,433,907,933]" italics="true" pageId="7" pageNumber="8">Rwandaracotis</emphasis>
</taxonomicName>
<emphasis id="B936292F5D3AFFBB56DA548FFD96FC21" bold="true" box="[440,561,907,933]" pageId="7" pageNumber="8">
<taxonomicNameLabel id="A20594545D3AFFBB56DA548FFD96FC21" box="[440,561,907,933]" pageId="7" pageNumber="8" rank="subGenus">subgen. n.</taxonomicNameLabel>
</emphasis>
Another surprising result was the morphologically very similar
<taxonomicName id="4C428EBE5D3AFFBB5263548FFEB5FC4D" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="argillaceaapodosima">
<emphasis id="B936292F5D3AFFBB5263548FFEB5FC4D" italics="true" pageId="7" pageNumber="8">A. argillaceaapodosima</emphasis>
</taxonomicName>
group (BS: 9; PP: -) and the
<taxonomicName id="4C428EBE5D3AFFBB553454ABFD45FC4D" authorityName="László &amp; Hausmann &amp; Karisch" authorityYear="2023" baseAuthorityName="Herbulot" baseAuthorityYear="1970" box="[598,738,943,969]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="deportata">
<emphasis id="B936292F5D3AFFBB553454ABFD45FC4D" box="[598,738,943,969]" italics="true" pageId="7" pageNumber="8">A. deportata</emphasis>
</taxonomicName>
cluster of undescribed species (BS: 16; PP: -) being recovered as paraphyletic.
</paragraph>
<paragraph id="8BFDF53D5D3AFFBB57A554F3FAB7FAD1" blockId="7.[151,1437,150,2013]" pageId="7" pageNumber="8">
Based on their homologous habitus and genital morphology, both clades are recognised here as representatives of a distinct subgenus,
<taxonomicName id="4C428EBE5D3AFFBB56C7531FFDF7FBB1" authority="László &amp; Hausmann &amp; Karisch, 2023" authorityName="László &amp; Hausmann &amp; Karisch" authorityYear="2023" box="[421,592,1051,1077]" pageId="7" pageNumber="8" rank="subGenus" status="subgen. nov." subGenus="Herbuloracotis">
<emphasis id="B936292F5D3AFFBB56C7531FFDF7FBB1" box="[421,592,1051,1077]" italics="true" pageId="7" pageNumber="8">Herbuloracotis</emphasis>
</taxonomicName>
<emphasis id="B936292F5D3AFFBB5539531FFD70FBB1" bold="true" box="[603,727,1051,1077]" pageId="7" pageNumber="8">
<taxonomicNameLabel id="A20594545D3AFFBB5539531FFD70FBB1" box="[603,727,1051,1077]" pageId="7" pageNumber="8" rank="subGenus">subgen. n.</taxonomicNameLabel>
</emphasis>
(Text fig. 2). Due to the incongruencies between the genetic and morphological analyses, the species-group subdivision of this subgenus is based on morphological features corresponding with the ML topology in the
<taxonomicName id="4C428EBE5D3AFFBB55EC5367FC8DFBF9" authorityName="László &amp; Hausmann &amp; Karisch" authorityYear="2023" baseAuthorityName="Prout" baseAuthorityYear="1931" box="[654,810,1123,1149]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="apodosima">
<emphasis id="B936292F5D3AFFBB55EC5367FC8DFBF9" box="[654,810,1123,1149]" italics="true" pageId="7" pageNumber="8">A. apodosima</emphasis>
</taxonomicName>
and
<taxonomicName id="4C428EBE5D3AFFBB54065367FC55FBF9" authorityName="László &amp; Hausmann &amp; Karisch" authorityYear="2023" baseAuthorityName="Herbulot" baseAuthorityYear="1970" box="[868,1010,1123,1149]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="deportata">
<emphasis id="B936292F5D3AFFBB54065367FC55FBF9" box="[868,1010,1123,1149]" italics="true" pageId="7" pageNumber="8">A. deportata</emphasis>
</taxonomicName>
species-groups, but the
<taxonomicName id="4C428EBE5D3AFFBB526B5367FA3BFBF9" baseAuthorityName="Warren" baseAuthorityYear="1900" box="[1289,1436,1123,1149]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="argillacea">
<emphasis id="B936292F5D3AFFBB526B5367FA3BFBF9" box="[1289,1436,1123,1149]" italics="true" pageId="7" pageNumber="8">A. argillacea</emphasis>
</taxonomicName>
species-group remains paraphyletic. The
<taxonomicName id="4C428EBE5D3AFFBB55075383FD43FB25" authorityName="Butler" authorityYear="1878" box="[613,740,1159,1185]" class="Insecta" family="Geometridae" genus="Ophthalmodes" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="squalida">
<emphasis id="B936292F5D3AFFBB55075383FD43FB25" box="[613,740,1159,1185]" italics="true" pageId="7" pageNumber="8">A. squalida</emphasis>
</taxonomicName>
+
<taxonomicName id="4C428EBE5D3AFFBB54615383FCF0FB25" authorityName="Warren" authorityYear="1899" box="[771,855,1159,1185]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="zebrina">
<emphasis id="B936292F5D3AFFBB54615383FCF0FB25" box="[771,855,1159,1185]" italics="true" pageId="7" pageNumber="8">zebrina</emphasis>
</taxonomicName>
lineage (BS: 55; PP: 97) is clearly divided into two well-supported monophyletic units with clear diagnostic morphological characters representing two subgenera:
<taxonomicName id="4C428EBE5D3AFFBB57F553CBFEB1FB6D" authority="László &amp; Hausmann &amp; Karisch, 2023" authorityName="László &amp; Hausmann &amp; Karisch" authorityYear="2023" box="[151,278,1231,1257]" pageId="7" pageNumber="8" rank="subGenus" status="subgen. nov." subGenus="Afroracotis">
<emphasis id="B936292F5D3AFFBB57F553CBFEB1FB6D" box="[151,278,1231,1257]" italics="true" pageId="7" pageNumber="8">Afroracotis</emphasis>
</taxonomicName>
<emphasis id="B936292F5D3AFFBB564253CBFE3BFB6D" bold="true" box="[288,412,1231,1257]" pageId="7" pageNumber="8">
<taxonomicNameLabel id="A20594545D3AFFBB564253CBFE3BFB6D" box="[288,412,1231,1257]" pageId="7" pageNumber="8" rank="subGenus">subgen. n.</taxonomicNameLabel>
</emphasis>
(Text fig. 1) and
<taxonomicName id="4C428EBE5D3AFFBB550C53CBFD41FB6D" authority="László &amp; Hausmann &amp; Karisch, 2023" authorityName="László &amp; Hausmann &amp; Karisch" authorityYear="2023" box="[622,742,1231,1257]" pageId="7" pageNumber="8" rank="subGenus" status="subgen. nov." subGenus="Zebracotis">
<emphasis id="B936292F5D3AFFBB550C53CBFD41FB6D" box="[622,742,1231,1257]" italics="true" pageId="7" pageNumber="8">Zebracotis</emphasis>
</taxonomicName>
<emphasis id="B936292F5D3AFFBB559253CBFCCBFB6D" bold="true" box="[752,876,1231,1257]" pageId="7" pageNumber="8">
<taxonomicNameLabel id="A20594545D3AFFBB559253CBFCCBFB6D" box="[752,876,1231,1257]" pageId="7" pageNumber="8" rank="subGenus">subgen. n.</taxonomicNameLabel>
</emphasis>
(Text fig. 3).
<taxonomicName id="4C428EBE5D3AFFBB537153CBFB35FB6D" authorityName="László &amp; Hausmann &amp; Karisch" authorityYear="2023" box="[1043,1170,1231,1257]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B936292F5D3AFFBB537153CBFB35FB6D" box="[1043,1170,1231,1257]" italics="true" pageId="7" pageNumber="8">Afroracotis</emphasis>
</taxonomicName>
is subdivided into two species-groups (BS: 96; PP: 100) clearly corresponding with morphology, whereas in
<taxonomicName id="4C428EBE5D3AFFBB53E953F7FAA4FA89" box="[1163,1283,1267,1293]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="78" phylum="Arthropoda" rank="subGenus" subGenus="Zebracotis">
<emphasis id="B936292F5D3AFFBB53E953F7FAA4FA89" box="[1163,1283,1267,1293]" italics="true" pageId="7" pageNumber="8">Zebracotis</emphasis>
</taxonomicName>
both genetic analyses provided a rather heterogenous topology often conflicting with the constant diagnostic morphological traits of the species. The results of the genetic analyses are further discussed in detail under each subgenera.
</paragraph>
<paragraph id="8BFDF53D5D3AFFBB57A5525BFE57FA45" blockId="7.[151,1437,150,2013]" pageId="7" pageNumber="8">
Besides the four subgenera delimited through integrative analyses, a fifth subgenus (
<taxonomicName id="4C428EBE5D3AFFBB5315525BFAB3FAFD" authority="László &amp; Hausmann &amp; Karisch, 2023" authorityName="László &amp; Hausmann &amp; Karisch" authorityYear="2023" box="[1143,1300,1375,1401]" pageId="7" pageNumber="8" rank="subGenus" status="subgen. nov." subGenus="Sokokeracotis">
<emphasis id="B936292F5D3AFFBB5315525BFAB3FAFD" box="[1143,1300,1375,1401]" italics="true" pageId="7" pageNumber="8">Sokokeracotis</emphasis>
</taxonomicName>
<emphasis id="B936292F5D3AFFBB5279525BFA34FAFD" bold="true" box="[1307,1427,1375,1401]" pageId="7" pageNumber="8">
<taxonomicNameLabel id="A20594545D3AFFBB5279525BFA34FAFD" box="[1307,1427,1375,1401]" pageId="7" pageNumber="8" rank="subGenus">subgen. n.</taxonomicNameLabel>
</emphasis>
) is also described in this paper based merely on its unique genital morphology, as the single accessed specimen could not be successfully sequenced.
</paragraph>
</subSubSection>
<subSubSection id="C358A6B65D3AFFBB57A552CEFDDEF859" pageId="7" pageNumber="8" type="diagnosis">
<paragraph id="8BFDF53D5D3AFFBB57A552CEFC6EF939" blockId="7.[151,1437,150,2013]" pageId="7" pageNumber="8">
<emphasis id="B936292F5D3AFFBB57A552CEFE9DFA60" bold="true" box="[199,314,1482,1508]" pageId="7" pageNumber="8">Diagnosis</emphasis>
.
<taxonomicName id="4C428EBE5D3AFFBB562B52CFFE6FFA61" authority="László &amp; Hausmann &amp; Karisch, 2023" authorityName="László &amp; Hausmann &amp; Karisch" authorityYear="2023" box="[329,456,1483,1509]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B936292F5D3AFFBB562B52CFFE6FFA61" box="[329,456,1483,1509]" italics="true" pageId="7" pageNumber="8">Afroracotis</emphasis>
</taxonomicName>
<emphasis id="B936292F5D3AFFBB56B252CFFD87FA61" bold="true" box="[464,544,1483,1509]" pageId="7" pageNumber="8">
<taxonomicNameLabel id="A20594545D3AFFBB56B252CFFD87FA61" box="[464,544,1483,1509]" pageId="7" pageNumber="8" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
shares the general external habitus with the Indomalayan-Australasian
<taxonomicName id="4C428EBE5D3AFFBB522A52CFFE9DF98D" authority="Moore, [1887]" authorityName="Moore" authorityYear="1887" class="Insecta" family="Geometridae" genus="Racotis" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B936292F5D3AFFBB522A52CFFA3BFA60" box="[1352,1436,1483,1508]" italics="true" pageId="7" pageNumber="8">Racotis</emphasis>
Moore, [1887]
</taxonomicName>
(
<typeStatus id="54F94B9F5D3AFFBB562A52EBFEDDF98D" box="[328,378,1519,1545]" pageId="7" pageNumber="8">type</typeStatus>
species:
<taxonomicName id="4C428EBE5D3AFFBB568252EBFC0CF98D" authority="Guenee, [1858]" authorityName="Guenee" authorityYear="1858" box="[480,939,1518,1545]" class="Insecta" family="Geometridae" genus="Hypochroma" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="boarmiaria">
<emphasis id="B936292F5D3AFFBB568252EBFD50F98D" box="[480,759,1519,1545]" italics="true" pageId="7" pageNumber="8">Hypochroma boarmiaria</emphasis>
Guenée, [1858]
</taxonomicName>
described from eastern
<collectingCountry id="F355B5AD5D3AFFBB53D452EBFB55F98D" box="[1206,1266,1519,1545]" name="India" pageId="7" pageNumber="8">India</collectingCountry>
, (
<figureCitation id="1379E9B85D3AFFBB526752EBFAECF98D" box="[1285,1355,1519,1545]" captionStart="TEXT" captionStartId="11.[152,224,1966,1990]" captionTargetBox="[280,1307,888,1933]" captionTargetId="figure-327@11.[274,1313,882,1939]" captionTargetPageId="11" captionText="TEXT FIGURE 1. ML tree of the subgenus Afroracotis based on analysis of COI barcodes. Bootstrap values are indicated above, posterior probability values below the branches." figureDoi="http://doi.org/10.5281/zenodo.8108663" httpUri="https://zenodo.org/record/8108663/files/figure.png" pageId="7" pageNumber="8">Figs 1</figureCitation>
,
<figureCitation id="1379E9B85D3AFFBB523B52EBFA21F98D" box="[1369,1414,1519,1545]" captionStart="FIGURES 191210" captionStartId="81.[151,264,1641,1665]" captionTargetBox="[151,1433,188,1610]" captionTargetId="figure-17@81.[151,1436,181,1617]" captionTargetPageId="81" captionText="FIGURES 191210. Adults, ventral view (all in coll. ANHRT, unless otherwise stated). 191. Afroracotis milesi sp. n., HT, ♁, Gabon; 192. Ibidem, PT, ♁, Cameroon; 193. Ibidem, PT, ♁, Zambia; 194. Ibidem, PT, ♀, Gabon; 195. A. ochsei sp. n., HT, ♁, Gabon; 196. Ibidem, PT, ♁, Gabon; 197. Ibidem, PT, ♀, Gabon; 198. A. helicalis sp. n., HT, ♁, Gabon; 199. Ibidem, PT, ♁, Gabon; 200. Ibidem, PT, ♀, Gabon; 201. A. takanoi sp. n., HT, ♁, Sierra Leone; 202. Ibidem, PT, ♁, Guinea; 203. Ibidem, PT, ♀, Ivory Coast; 204. Ibidem, PT, ♀, Liberia; 205. A. staudei sp. n., HT, ♁, Kenya (RCHS); 206. Racotis boarmiaria, LT, ♁, India (NHMUK); 207. Chorocotis ugandaria, HT of Boarmia ugandaria, ♁, Uganda (NHMUK); 208. Ibidem, ♀, Cameroon (NHMUK); 209. Colocleora breijeri, ♁, RSA (RCDS); 210. C. incauta, ♁, RSA (NHMUK)." figureDoi="http://doi.org/10.5281/zenodo.8080954" httpUri="https://zenodo.org/record/8080954/files/figure.png" pageId="7" pageNumber="8">206</figureCitation>
)), characterised by the dark greenish-brown, often mossy-green colouration usually darkened in the distal third of the wings, the typically speckled wing pattern with the diffuse, interrupted transverse lines often replaced by rows of variably sized patches only and the well-defined, relatively large ovoidal discal spot. The pattern of the underside of the wings is also similar in both genera characterised by the usually sharply divided dark greyish postmedial third and the considerably paler, variably speckled basal and antemedial areas.
</paragraph>
<paragraph id="8BFDF53D5D3AFFBB57A551C3FCEFF8C9" blockId="7.[151,1437,150,2013]" pageId="7" pageNumber="8">
The new Afrotropical genus differs from
<taxonomicName id="4C428EBE5D3AFFBB55E451C3FD7DF964" authorityName="Moore" authorityYear="1887" box="[646,730,1735,1760]" class="Insecta" family="Geometridae" genus="Racotis" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B936292F5D3AFFBB55E451C3FD7DF964" box="[646,730,1735,1760]" italics="true" pageId="7" pageNumber="8">Racotis</emphasis>
</taxonomicName>
by a conspicuous autapomorphic character of the male genitalia, the absence of the saccular process, which is well-developed in species of true
<taxonomicName id="4C428EBE5D3AFFBB536851EFFBF9F880" authorityName="Moore" authorityYear="1887" box="[1034,1118,1771,1796]" class="Insecta" family="Geometridae" genus="Racotis" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B936292F5D3AFFBB536851EFFBF9F880" box="[1034,1118,1771,1796]" italics="true" pageId="7" pageNumber="8">Racotis</emphasis>
</taxonomicName>
(
<figureCitation id="1379E9B85D3AFFBB530F51EFFB68F881" box="[1133,1231,1771,1797]" captionStart="FIGURES 211216" captionStartId="82.[151,264,1807,1831]" captionTargetBox="[182,1403,188,1778]" captionTargetId="figure-21@82.[175,1411,181,1784]" captionTargetPageId="82" captionText="FIGURES 211216. Male genitalia, a: clasping apparatus, b: aedeagus. 211. Racotis boarmiaria, Nepal, slide No. LG 3677 (RCGL); 212. Afroracotis squalida squalida, Togo, slide No. LG 5434 (ANHRT); 213. Ibidem, Uganda, slide No. DS 229-2019 (RCDS); 214. Ibidem, Zambia, slide No. LG 4959 (ANHRT); 215. Ibidem, Ethiopia, slide No. DS 226-2019 (RCDS); 216. A. s. voeltzkowii, Madagascar, slide No. LG 4100 (NHMUK)." figureDoi="http://doi.org/10.5281/zenodo.8080956" httpUri="https://zenodo.org/record/8080956/files/figure.png" pageId="7" pageNumber="8">Fig. 211</figureCitation>
). Additionally, the configuration of the gnathos seems to display constant differences between the species of
<taxonomicName id="4C428EBE5D3AFFBB53E9500BFAADF8AD" authorityName="László &amp; Hausmann &amp; Karisch" authorityYear="2023" box="[1163,1290,1807,1833]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B936292F5D3AFFBB53E9500BFAADF8AD" box="[1163,1290,1807,1833]" italics="true" pageId="7" pageNumber="8">Afroracotis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C428EBE5D3AFFBB5223500BFA32F8AC" authorityName="Moore" authorityYear="1887" box="[1345,1429,1807,1832]" class="Insecta" family="Geometridae" genus="Racotis" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B936292F5D3AFFBB5223500BFA32F8AC" box="[1345,1429,1807,1832]" italics="true" pageId="7" pageNumber="8">Racotis</emphasis>
</taxonomicName>
, being apically rounded in the former and pointed in the latter.
</paragraph>
<paragraph id="8BFDF53D5D3AFFBB57A55053FDDEF859" blockId="7.[151,1437,150,2013]" pageId="7" pageNumber="8">
The gound plan of the female genitalia in
<taxonomicName id="4C428EBE5D3AFFBB55DE5053FC9CF8F5" authorityName="László &amp; Hausmann &amp; Karisch" authorityYear="2023" box="[700,827,1879,1905]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B936292F5D3AFFBB55DE5053FC9CF8F5" box="[700,827,1879,1905]" italics="true" pageId="7" pageNumber="8">Afroracotis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C428EBE5D3AFFBB541B5053FC6AF8F4" authorityName="Moore" authorityYear="1887" box="[889,973,1879,1904]" class="Insecta" family="Geometridae" genus="Racotis" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B936292F5D3AFFBB541B5053FC6AF8F4" box="[889,973,1879,1904]" italics="true" pageId="7" pageNumber="8">Racotis</emphasis>
</taxonomicName>
are rather homologous (cf.
<bibRefCitation id="EFD388CC5D3AFFBB52755053FA37F8F5" author="Sato, R." box="[1303,1424,1879,1905]" pageId="7" pageNumber="8" pagination="130 - 139" refId="ref84694" refString="Sato, R. (2004) Taxonomic notes on Racotis boarmiaria (Guenee) (Geometridae, Ennominae) and its allies from the IndoMalayan region, with descriptions of four new species. Tinea, 18 (2), 130 - 139." type="journal article" year="2004">Sato 2004</bibRefCitation>
), especially the configuration of the fused cervix and corpus bursae suggesting a close affinity between the genera. However, the
<taxonomicName id="4C428EBE5D3AFFBB5657509BFE2EF83C" authorityName="Moore" authorityYear="1887" box="[309,393,1951,1976]" class="Insecta" family="Geometridae" genus="Racotis" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B936292F5D3AFFBB5657509BFE2EF83C" box="[309,393,1951,1976]" italics="true" pageId="7" pageNumber="8">Racotis</emphasis>
</taxonomicName>
females have a uniformly elongate, sack-like bursa copulatrix unlike the remarkably diverse corpus bursae configuration in
<taxonomicName id="4C428EBE5D3AFFBB569650C7FDD4F859" authorityName="László &amp; Hausmann &amp; Karisch" authorityYear="2023" box="[500,627,1987,2013]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B936292F5D3AFFBB569650C7FDD4F859" box="[500,627,1987,2013]" italics="true" pageId="7" pageNumber="8">Afroracotis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C358A6B65D35FFB457F55793FCD9F8C9" pageId="8" pageNumber="9" type="description">
<paragraph id="8BFDF53D5D35FFB457F55793FE8FFF35" blockId="8.[151,296,151,177]" box="[151,296,151,177]" pageId="8" pageNumber="9">
<emphasis id="B936292F5D35FFB457F55793FE8FFF35" bold="true" box="[151,296,151,177]" pageId="8" pageNumber="9">Description</emphasis>
</paragraph>
<paragraph id="8BFDF53D5D35FFB457F557DBFD10FBF9" blockId="8.[151,1437,223,1870]" pageId="8" pageNumber="9">
<emphasis id="B936292F5D35FFB457F557DBFD9EFF7D" bold="true" box="[151,569,223,249]" pageId="8" pageNumber="9">External features of body and wings</emphasis>
(
<figureCitation id="1379E9B85D35FFB4552757DBFD63FF7D" box="[581,708,223,249]" captionStart-0="TEXT" captionStart-1="FIGURES 120" captionStart-2="FIGURES 2140" captionStart-3="FIGURES 4160" captionStart-4="FIGURES 6180" captionStart-5="FIGURES 81100" captionStart-6="FIGURES 101120" captionStartId-0="11.[152,224,1966,1990]" captionStartId-1="17.[151,264,1656,1680]" captionStartId-2="22.[151,264,1647,1671]" captionStartId-3="33.[151,264,1631,1655]" captionStartId-4="41.[151,264,1622,1646]" captionStartId-5="53.[151,264,1614,1638]" captionStartId-6="65.[151,264,1643,1667]" captionTargetBox-0="[280,1307,888,1933]" captionTargetBox-1="[152,1431,183,1623]" captionTargetBox-2="[158,1424,186,1617]" captionTargetBox-3="[163,1428,187,1602]" captionTargetBox-4="[172,1413,190,1591]" captionTargetBox-5="[176,1414,188,1584]" captionTargetBox-6="[185,1402,188,1614]" captionTargetId-0="figure-327@11.[274,1313,882,1939]" captionTargetId-1="figure-17@17.[151,1436,179,1630]" captionTargetId-2="figure-21@22.[157,1430,181,1623]" captionTargetId-3="figure-17@33.[155,1432,181,1608]" captionTargetId-4="figure-17@41.[165,1423,181,1597]" captionTargetId-5="figure-17@53.[167,1421,181,1590]" captionTargetId-6="figure-17@65.[176,1412,181,1620]" captionTargetPageId-0="11" captionTargetPageId-1="17" captionTargetPageId-2="22" captionTargetPageId-3="33" captionTargetPageId-4="41" captionTargetPageId-5="53" captionTargetPageId-6="65" captionText-0="TEXT FIGURE 1. ML tree of the subgenus Afroracotis based on analysis of COI barcodes. Bootstrap values are indicated above, posterior probability values below the branches." captionText-1="FIGURES 120. Adults, dorsal view. 1. Racotis boarmiaria, LT, ♁, India (NHMUK); 2. Afroracotis squalida squalida, HT of Ophthalmodes squalida, ♀, Nigeria (NHMUK); 3. Ibidem, HT of Boarmia uhligi, ♀, Tanzania (MfN); 4. Ibidem, ♁, Cameroon (ANHRT); 5. Ibidem, ♁, Zambia (ANHRT); 6. Ibidem, ♁, Togo (ANHRT); 7. Ibidem, ♁, Gabon (ANHRT); 8. Ibidem, ♁, Ethiopia (RCDS); 9. Ibidem, ♀, Ethiopia (RCDS); 10. Ibidem, ♀, Cameroon (ANHRT); 11. A. s. voeltzkowii, LT of Boarmia voeltzkowii, ♁, Madagascar (MfN); 12. Ibidem, ♁, Comoros (ZSM); 13. Ibidem, ♁, Madagascar (NHMUK); 14. Ibidem, ♀, Madagascar (NHMUK); 15. A. s. thomensis ssp. n., HT, ♁, São Tomé (ANHRT); 16. Ibidem, PT, ♀, São Tomé (ANHRT); 17. Ibidem, ♀, PT, São Tomé (NHMUK); 18. A. incompletaria incompletaria, ♀, Réunion (ZSM); 19. A. i. atrilunaria, HT of Boarmia atrilunaria, ♀, Mauritius (NHMUK); 20. Ibidem, ♀, Mauritius (ANHRT)." captionText-2="FIGURES 2140. Adults, dorsal view. 21. Afroracotis aliena sp. n., HT, ♀, Tanzania (ANHRT); 22. A. angulosa, HT of Racotis angulosa, ♁, Cameroon (ZSM); 23. Ibidem, ♁, Ethiopia (RCDS); 24. Ibidem, ♀, Cameroon (ZSM); 25. Ibidem, ♁, Kenya (RCHS); 26. Ibidem, ♀, Kenya (RCHS); 27. A. canui, HT of Racotis canui, ♁, São Tomé (ZSM); 28. Ibidem, ♁, Principé (ZSM); 29. A. stadiei sp. n., HT, ♁, Tanzania (ZSM); 30. Ibidem, PT, ♀, Tanzania (ZSM); 31. Ibidem, PT, ♁, Rwanda (ZSM); 32. A. violetteae sp. n., HT, ♁, Guinea (ANHRT); 33. Ibidem, PT, ♁, Guinea (ANHRT); 34. Ibidem, PT, ♀, Liberia (ANHRT); 35. A. fiebigi sp. n., HT, ♁, Liberia (ANHRT); 36. Ibidem, PT, ♁, Cameroon (ZSM); 37. Ibidem, PT, ♀, Ethiopia (RCDS); 38. A. albitrigonis, HT of Cleora albitrigonis, ST, ♁, São Tomé (NHMUK); 39. Ibidem, ♁, São Tomé (ANHRT); 40. Ibidem, ♀, São Tomé (ANHRT)." captionText-3="FIGURES 4160. Adults, dorsal view. 41. Afroracotis apodosima, HT of Racotis apodosima, ♁, Mozambique (NHMUK); 42. Ibidem, ♁, Tanzania (ZSM); 43. Ibidem, ♁, Zambia (ANHRT); 44. Ibidem, ♀, Mozambique (ANHRT); 45. A. dargei sp. n., HT, ♁, Cameroon (ZSM); 46. Ibidem, PT, ♁, Cameroon (ZSM); 47. Ibidem, PT, ♀, Cameroon (ZSM); 48. A. deportata, ♁, Madagascar (RCHS); 49. Ibidem, HT of Racotis deportata, ♁, Madagascar (MNHN); 50. Ibidem, ♀, Comoros (NHMUK); 51. A. argillacea argillacea, HT of Alcis argillacea, ♁, DRC (NHMUK); 52. Ibidem, ♁, DRC (NHMUK); 53. Ibidem, ♁, DRC (ZSM); 54. Ibidem, ♁, Cameroon (RCMO); 55. A. argillacea cedrici, HT of Racotis cedrici, ♀, Bioko (ZSM); 56. Ibidem, PT, ♀, Bioko (ZSM); 57. Ibidem, ♀, Bioko (RCTK); 58. A. argillacea morettoi ssp. n., HT, ♁, Ivory Coast (ANHRT); 59. Ibidem, PT, ♀, Ivory Coast (ANHRT); 60. Ibidem, PT, ♀, Ivory Coast (ZSM)." captionText-4="FIGURES 6180. Adults, dorsal view. 61. Afroracotis longicornuta sp. n., HT, ♁, Liberia (ANHRT); 62. Ibidem, PT, ♁, Guinea (ANHRT); 63. A. l. congolana ssp. n., HT, ♁, Cameroon (RCMO); 64. Ibidem, PT, ♁, Cameroon (ZSM); 65. Ibidem, PT, ♁, DRC (NHMUK); 66. Ibidem, PT, ♀, Cameroon (ZSM); 67. A. l. ugandana ssp. n., HT, ♁, Uganda (RCHS); 68. Ibidem, PT, ♀, Uganda (RCHS); 69. A. aristophanousi sp. n., HT, ♁, Liberia (ANHRT); 70. Ibidem, PT, ♁, Cameroon (ZSM); 71. Ibidem, PT, ♁, Uganda (NHMUK); 72. Ibidem, PT, ♀, Cameroon (ZSM); 73. A. muscivirens sp. n., HT, ♁, Liberia (ZSM); 74. Ibidem, PT, ♁, Ivory Coast (ZSM); 75. Ibidem, PT, ♀, Liberia (ANHRT); 76. A. turlini sp. n., HT, ♁, Rwanda (ZSM); 77. Ibidem, PT, ♁, Rwanda (ZSM); 78. Ibidem, PT, ♀, Rwanda (ZSM); 79. A. atriclava, ST of Cleora atriclava, ♁, Madagascar (NHMUK); 80. Ibidem, ♀, Madagascar (ZSM)." captionText-5="FIGURES 81100. Adults, dorsal view. 81. Afroracotis zebrina, LT of Racotis zebrina, ♁, Nigeria (NHMUK); 82. Ibidem, ♁, Guinea (ANHRT); 83. Ibidem, ♀, Ivory Coast (ANHRT); 84. Ibidem, ♀, Liberia (ANHRT); 85. A. chaineyi sp. n., HT, ♁, Gabon (ANHRT); 86. Ibidem, PT, ♁, Cameroon (ZSM); 87. Ibidem, PT, ♁, Zambia (ANHRT); 88. Ibidem, PT, ♀, Guinea (ANHRT); 89. A. lydiae lydiae sp. n., HT, ♁, Liberia (ANHRT); 90. Ibidem, PT, ♁, Cameroon (ZSM); 91. Ibidem, PT, ♀, Guinea (ANHRT); 92. Ibidem, PT, ♀, Bioko (ZSM); 93. A. lydiae orientalis ssp. n., HT, ♁, Tanzania (ANHRT); 94. Ibidem, PT, ♁, Mozambique (ANHRT); 95. Ibidem, PT, ♀, Tanzania (ANHRT); 96. Ibidem, PT, ♀, Mozambique (ANHRT); 97. A. madagascariensis, HT of Racotis madagascariensis, ♁, Madagascar (NHMUK); 98. Ibidem, PT, ♁, Madagascar (NHMUK); 99. Ibidem, PT, ♀, Madagascar (NHMUK); 100. Ibidem, ♀, Madagascar (ANHRT)." captionText-6="FIGURES 101120. Adults, dorsal view (all in coll. ANHRT, unless otherwise stated). 101. A. smithi sp. n., HT, ♁, Zambia; 102. Ibidem, PT, ♁, Gabon; 103. Ibidem, PT, ♀, Zambia; 104. Ibidem, PT, ♀, Cameroon (ZSM); 105. A. milesi sp. n., HT, ♁, Gabon; 106. Ibidem, PT, ♁, Cameroon; 107. Ibidem, PT, ♀, Cameroon; 108. Ibidem, PT, ♁, Zambia; 109. A. ochsei sp. n., HT, ♁, Gabon; 110. Ibidem, PT, ♁, Gabon; 111. Ibidem, PT, ♀, Gabon; 112. Ibidem, PT, ♁, Gabon; 113. A. helicalis sp. n., HT, ♁, Gabon; 114. Ibidem, PT, ♁, Gabon; 115. Ibidem, PT, ♀, Gabon; 116. A. takanoi sp. n., HT, ♁, Sierra Leone; 117. Ibidem, PT, ♁, Guinea; 118. Ibidem, PT, ♁, Liberia; 119. Ibidem, PT, ♀, Liberia; 120. A. staudei sp. n., HT, ♁, Kenya (RCHS)." figureDoi-0="http://doi.org/10.5281/zenodo.8108663" figureDoi-1="http://doi.org/10.5281/zenodo.8080931" figureDoi-2="http://doi.org/10.5281/zenodo.8080933" figureDoi-3="http://doi.org/10.5281/zenodo.8080935" figureDoi-4="http://doi.org/10.5281/zenodo.8080937" figureDoi-5="http://doi.org/10.5281/zenodo.8080939" figureDoi-6="http://doi.org/10.5281/zenodo.8080943" httpUri-0="https://zenodo.org/record/8108663/files/figure.png" httpUri-1="https://zenodo.org/record/8080931/files/figure.png" httpUri-2="https://zenodo.org/record/8080933/files/figure.png" httpUri-3="https://zenodo.org/record/8080935/files/figure.png" httpUri-4="https://zenodo.org/record/8080937/files/figure.png" httpUri-5="https://zenodo.org/record/8080939/files/figure.png" httpUri-6="https://zenodo.org/record/8080943/files/figure.png" pageId="8" pageNumber="9">Figs 1120</figureCitation>
,
<figureCitation id="1379E9B85D35FFB455AF57DBFC94FF7E" box="[717,819,223,250]" captionStart-0="FIGURES 131150" captionStart-1="FIGURES 151170" captionStart-2="FIGURES 171190" captionStart-3="FIGURES 191210" captionStartId-0="78.[151,264,1654,1678]" captionStartId-1="79.[151,264,1648,1672]" captionStartId-2="80.[151,264,1643,1667]" captionStartId-3="81.[151,264,1641,1665]" captionTargetBox-0="[161,1433,189,1625]" captionTargetBox-1="[166,1420,189,1618]" captionTargetBox-2="[182,1405,190,1613]" captionTargetBox-3="[151,1433,188,1610]" captionTargetId-0="figure-21@78.[153,1434,181,1631]" captionTargetId-1="figure-17@79.[159,1428,181,1624]" captionTargetId-2="figure-21@80.[175,1413,181,1619]" captionTargetId-3="figure-17@81.[151,1436,181,1617]" captionTargetPageId-0="78" captionTargetPageId-1="79" captionTargetPageId-2="80" captionTargetPageId-3="81" captionText-0="FIGURES 131150. Adults, ventral view. 131. Afroracotis squalida squalida, ♁, Cameroon (ANHRT); 132. Ibidem, ♀, Cameroon (ANHRT); 133. A. s. voeltzkowii, ♁, Comoros (ZSM); 134. Ibidem, ♀, Madagascar (NHMUK); 135. A. s. thomensis ssp. n., HT, ♁, São Tomé (ANHRT); 136. Ibidem, PT, ♀, São Tomé (ANHRT); 137. A. incompletaria incompletaria, ♀, Réunion (ZSM); 138. A. i. atrilunaria, HT of Boarmia atrilunaria, ♀, Mauritius (NHMUK); 139. A. aliena sp. n., HT, ♀, Tanzania (ANHRT); 140. A. angulosa, HT of Racotis angulosa, ♁, Cameroon (ZSM); 141. Ibidem, ♀, Cameroon (ZSM); 142. A. canui, HT of Racotis canui, ♁, São Tomé (ZSM); 143. Ibidem, ♁, Princípe (ZSM); 144. A. stadiei sp. n., PT, ♀, Tanzania (ZSM); 145. A. violetteae sp. n., HT, ♁, Guinea (ANHRT); 146. Ibidem, PT, ♀, Liberia (ANHRT); 147. A. fiebigi sp. n., HT, ♁, Liberia (ANHRT); 148. Ibidem, PT, ♀, Ethiopia (RCDS); 149. A. albitrigonis, ♁, São Tomé (ANHRT); 150. Ibidem, ♀, São Tomé (ANHRT)." captionText-1="FIGURES 151170. Adults, ventral view. 151. Afroracotis apodosima, ♁, Zambia (ANHRT); 152. Ibidem, ♀, Mozambique (ANHRT); 153. A. dargei sp. n., PT, ♁, Cameroon (ZSM); 154. Ibidem, ♀, Cameroon (ZSM); 155. A. deportata, ♁, Madagascar (RCHS); 156. Ibidem, ♀, Comoros (NHMUK); 157. A. argillacea argillacea, ♁, Cameroon (ZSM); 158. Ibidem, ♁, Cameroon (ZSM); 159. A. argillacea cedrici, HT of Racotis cedrici, ♀, Bioko (ZSM); 160. A. argillacea morettoi ssp. n., HT, ♁, Ivory Coast (ANHRT); 161. Ibidem, PT, ♀, Ivory Coast (ANHRT); 162. A. longicornuta sp. n., HT, ♁, Liberia (ANHRT); 163. Ibidem, PT, ♁, Guinea (ZSM); 164. A. l. congolana ssp. n., PT, ♁, Cameroon (ZSM); 165. Ibidem, PT, ♀, Cameroon (ZSM); 166. A. l. ugandana ssp. n., PT, ♀, Uganda (RCHS); 167. A. aristophanousi sp. n., HT, ♁, Liberia (ANHRT); 168. Ibidem, PT, ♀, Cameroon (ZSM); 169. A. muscivirens sp. n., HT, ♁, Liberia (ZSM); 170. Ibidem, PT, ♀, Liberia (ANHRT)." captionText-2="FIGURES 171190. Adults, ventral view. 171. Afroracotis atriclava, ST of Cleora atriclava, ♁, Madagascar (ZSM); 172. Ibidem, ST, ♀, Madagascar (ZSM); 173. A. turlini sp. n., PT, ♁, Rwanda (ZSM); 174. Ibidem, PT, ♀, Rwanda (ZSM); 175. A. zebrina, LT of Racotis zebrina, ♁, Nigeria (NHMUK); 176. A. zebrina, ♁, Guinea (ANHRT); 177. Ibidem, ♁, Liberia (ANHRT); 178. Ibidem, ♀, Ivory Coast (ANHRT); 179. A. chaineyi sp. n., HT, ♁, Gabon (ANHRT); 180. Ibidem, PT, ♁, Cameroon (ZSM); 181. Ibidem, PT, ♀, Bioko (ZSM); 182. A. lydiae lydiae sp. n., PT, ♁, Liberia (ANHRT); 183. Ibidem, PT, ♀, Guinea (ANHRT); 184. A. lydiae orientalis ssp. n., HT, ♁, Tanzania (ANHRT); 185. Ibidem, PT, ♀, Tanzania (ANHRT); 186. A. madagascariensis, ♁, Madagascar (NHMUK); 187. Ibidem, ♀, Madagascar (ANHRT); 188. A. smithi sp. n., HT, ♁, Zambia (ANHRT); 189. Ibidem, PT, ♀, Zambia (ANHRT); 190. Ibidem, PT, ♀, Cameroon (ZSM)." captionText-3="FIGURES 191210. Adults, ventral view (all in coll. ANHRT, unless otherwise stated). 191. Afroracotis milesi sp. n., HT, ♁, Gabon; 192. Ibidem, PT, ♁, Cameroon; 193. Ibidem, PT, ♁, Zambia; 194. Ibidem, PT, ♀, Gabon; 195. A. ochsei sp. n., HT, ♁, Gabon; 196. Ibidem, PT, ♁, Gabon; 197. Ibidem, PT, ♀, Gabon; 198. A. helicalis sp. n., HT, ♁, Gabon; 199. Ibidem, PT, ♁, Gabon; 200. Ibidem, PT, ♀, Gabon; 201. A. takanoi sp. n., HT, ♁, Sierra Leone; 202. Ibidem, PT, ♁, Guinea; 203. Ibidem, PT, ♀, Ivory Coast; 204. Ibidem, PT, ♀, Liberia; 205. A. staudei sp. n., HT, ♁, Kenya (RCHS); 206. Racotis boarmiaria, LT, ♁, India (NHMUK); 207. Chorocotis ugandaria, HT of Boarmia ugandaria, ♁, Uganda (NHMUK); 208. Ibidem, ♀, Cameroon (NHMUK); 209. Colocleora breijeri, ♁, RSA (RCDS); 210. C. incauta, ♁, RSA (NHMUK)." figureDoi-0="http://doi.org/10.5281/zenodo.8080947" figureDoi-1="http://doi.org/10.5281/zenodo.8080950" figureDoi-2="http://doi.org/10.5281/zenodo.8080952" figureDoi-3="http://doi.org/10.5281/zenodo.8080954" httpUri-0="https://zenodo.org/record/8080947/files/figure.png" httpUri-1="https://zenodo.org/record/8080950/files/figure.png" httpUri-2="https://zenodo.org/record/8080952/files/figure.png" httpUri-3="https://zenodo.org/record/8080954/files/figure.png" pageId="8" pageNumber="9">131205</figureCitation>
). Forewing length
<quantity id="4CBA58D85D35FFB4549C57DBFBD0FF7E" box="[1022,1143,223,250]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" metricValueMax="3.0" metricValueMin="1.6" pageId="8" pageNumber="9" unit="mm" value="23.0" valueMax="30.0" valueMin="16.0">1630 mm</quantity>
. Male antenna bipectinate-fasciculate, rami short or moderately long, distal section filiform, ratio of which varying interspecifically between 1/6 and 1/2 length of entire antenna. Female antenna filiform or bipectinate-fasciculate with short rami in several taxa of
<taxonomicName id="4C428EBE5D35FFB45790564FFE20FEE1" baseAuthorityName="Warren" baseAuthorityYear="1900" box="[242,391,331,357]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="argillacea">
<emphasis id="B936292F5D35FFB45790564FFE20FEE1" box="[242,391,331,357]" italics="true" pageId="8" pageNumber="9">A. argillacea</emphasis>
</taxonomicName>
species-group. Head moderately large, proboscis well-developed, labial palp short, length 1.22.5 times the diameter of eye, porrect, first and second segment dilated, third segment rather short and thin. Compound eyes moderately large. Frons, vertex, collar (patagium), tegula, thorax and abdomen concolourous with wing upperside. Legs rather long, colouration as of wing underside, index of spurs 0-2-4. Forewing broad, triangular, costa straight, apical section slightly arched, termen straight to slightly arcuate, may be moderately scalloped, ventral margin straight. Venation of typical
<taxonomicName id="4C428EBE5D35FFB4546256FBFC27FD9D" box="[768,896,511,537]" class="Insecta" family="Geometridae" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="108" phylum="Arthropoda" rank="subFamily" subFamily="Ennominae">Ennominae</taxonomicName>
ground plan, forewing with four branches of R veins, cell relatively short, wide distally, Sc and R1 running parallely, R1+R2, R3, R4, R5 stalked, M1, M2, M3, CuA1, CuA2 and A well-developed, a longitudinal fold between CuA2 and A present. Hindwing with Sc+R1, Rs, M1, M3, CuA1, CuA2, A2, A3 fully developed, M2 reduced to a fold. Wing venation of all subgenera identical. Ground colour varying interspecifically from greyish dark green usually with a mossy tinge to pale ochreous-yellow, characteristically speckled in most taxa, basal and medial area often paler than terminal one, postmedial area with large dark blotches between M3 and CuA
<quantity id="4CBA58D85D35FFB455CB55D3FD77FD75" box="[681,720,727,753]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="8" pageNumber="9" unit="in" value="1.0">1 in</quantity>
several species. Transverse lines diffuse, interrupted. Basal and antemedial lines detectable, more or less straight, medial line shadow-like, moderately S-curved, postmedial line slightly wavy, subterminal line represented by a row of patches or deleted, terminal line interrupted, consisting of small black dots, short dashes or arches between veins. Fringe (cilia) short, concolourous with forewing. Discal spot well-developed, dot-like, round or ovoidal. Hindwing apex rounded, termen evenly arcuate, or medially angled and moderately scalloped in certain taxa, anal margin straight. Ground colour as of forewing, densely speckled, basal line often diffuse, evenly arched, wavy or sharply angled medially, medial line undulate, postmedial line diffuse, interrupted, terminal line interrupted, consisting of small black dots, short dashes or arches between veins. Discal spot as on forewing. Fringe (cilia) short, colour as of hindwing. Wing underside with broad, dark terminal area with exception for the
<taxonomicName id="4C428EBE5D35FFB45602531FFE54FBB1" baseAuthorityName="Warren" baseAuthorityYear="1900" box="[352,499,1051,1077]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="argillacea">
<emphasis id="B936292F5D35FFB45602531FFE54FBB1" box="[352,499,1051,1077]" italics="true" pageId="8" pageNumber="9">A. argillacea</emphasis>
</taxonomicName>
species-group with limited darker area. Basal and medial areas considerably paler than terminal one, densely suffused with greyish scales. Traces of transverse lines present, often poorly visible, discal spot and postmedial patches well-defined.
</paragraph>
<paragraph id="8BFDF53D5D35FFB457A55383FE3BFA61" blockId="8.[151,1437,223,1870]" pageId="8" pageNumber="9">
<emphasis id="B936292F5D35FFB457A55383FED3FB25" bold="true" box="[199,372,1159,1185]" pageId="8" pageNumber="9">Male genitalia</emphasis>
(
<figureCitation id="1379E9B85D35FFB456E75383FD80FB25" box="[389,551,1159,1185]" captionStart-0="FIGURES 211216" captionStart-1="FIGURES 217224" captionStart-10="FIGURES 274279" captionStart-11="FIGURES 280284" captionStart-2="FIGURES 225232" captionStart-3="FIGURES 233238" captionStart-4="FIGURES 239244" captionStart-5="FIGURES 245250" captionStart-6="FIGURES 251255" captionStart-7="FIGURES 256261" captionStart-8="FIGURES 262267" captionStart-9="FIGURES 268273" captionStartId-0="82.[151,264,1807,1831]" captionStartId-1="83.[151,264,1832,1856]" captionStartId-10="92.[151,264,1818,1842]" captionStartId-11="93.[151,264,1824,1848]" captionStartId-2="84.[151,264,1829,1853]" captionStartId-3="85.[151,264,1828,1852]" captionStartId-4="86.[151,264,1818,1842]" captionStartId-5="87.[151,264,1775,1799]" captionStartId-6="88.[151,264,1832,1856]" captionStartId-7="89.[151,264,1832,1856]" captionStartId-8="90.[151,264,1842,1866]" captionStartId-9="91.[151,264,1844,1868]" captionTargetBox-0="[182,1403,188,1778]" captionTargetBox-1="[155,1436,189,1802]" captionTargetBox-10="[151,1433,187,1788]" captionTargetBox-11="[155,1431,189,1794]" captionTargetBox-2="[151,1434,185,1799]" captionTargetBox-3="[152,1429,181,1798]" captionTargetBox-4="[154,1423,187,1788]" captionTargetBox-5="[162,1425,184,1744]" captionTargetBox-6="[158,1431,190,1802]" captionTargetBox-7="[151,1433,187,1802]" captionTargetBox-8="[151,1436,187,1812]" captionTargetBox-9="[151,1436,182,1815]" captionTargetId-0="figure-21@82.[175,1411,181,1784]" captionTargetId-1="figure-17@83.[151,1436,181,1808]" captionTargetId-10="figure-21@92.[151,1436,181,1794]" captionTargetId-11="figure-17@93.[151,1436,181,1800]" captionTargetId-2="figure-21@84.[151,1436,181,1805]" captionTargetId-3="figure-17@85.[151,1436,181,1804]" captionTargetId-4="figure-21@86.[153,1434,181,1794]" captionTargetId-5="figure-17@87.[156,1431,181,1750]" captionTargetId-6="figure-21@88.[152,1435,181,1808]" captionTargetId-7="figure-17@89.[151,1436,181,1808]" captionTargetId-8="figure-21@90.[151,1436,181,1818]" captionTargetId-9="figure-17@91.[151,1436,181,1821]" captionTargetPageId-0="82" captionTargetPageId-1="83" captionTargetPageId-10="92" captionTargetPageId-11="93" captionTargetPageId-2="84" captionTargetPageId-3="85" captionTargetPageId-4="86" captionTargetPageId-5="87" captionTargetPageId-6="88" captionTargetPageId-7="89" captionTargetPageId-8="90" captionTargetPageId-9="91" captionText-0="FIGURES 211216. Male genitalia, a: clasping apparatus, b: aedeagus. 211. Racotis boarmiaria, Nepal, slide No. LG 3677 (RCGL); 212. Afroracotis squalida squalida, Togo, slide No. LG 5434 (ANHRT); 213. Ibidem, Uganda, slide No. DS 229-2019 (RCDS); 214. Ibidem, Zambia, slide No. LG 4959 (ANHRT); 215. Ibidem, Ethiopia, slide No. DS 226-2019 (RCDS); 216. A. s. voeltzkowii, Madagascar, slide No. LG 4100 (NHMUK)." captionText-1="FIGURES 217224. Male genitalia, a: clasping apparatus, b: aedeagus. 217. Afroracotis squalida thomensis ssp. n., HT, São Tomé, slide No. LG 4931 (ANHRT); 218. A. angulosa, HT, Cameroon, slide No. LG 4612 (ZSM); 219. Ibidem, DRC, slide No. LG 4091 (NHMUK); 220. Ibidem,Ethiopia, slide No. DS 222-2019 (RCDS); 221. Ibidem, Uganda, slide No. DS 227-2019 (RCDS); 222. Ibidem, Kenya, slide No. LG 5552 (RCHS); 223. A. canui, Princípe, slide No. LG 5037 (ZSM); 224. A. stadiei sp. n., HT, Tanzania, slide No. LG 4763 (ZSM)." captionText-10="FIGURES 274279. Male genitalia, a: clasping apparatus, b: aedeagus. 274. Afroracotis milesi sp. n., PT, Zambia, slide No. LG 4061 (ANHRT); 275. A. ochsei sp. n., HT, Gabon, slide No. LG 5406 (ANHRT); 276. Ibidem, PT, Cameroon, slide No. LG 4413 (ZSM); 277. Ibidem, PT, Gabon, slide No. LG 5414 (ANHRT); 278. A. helicalis sp. n., HT, Gabon, slide No. LG 5621 (ANHRT); 279. Ibidem, PT, Gabon, slide No. LG 5412 (ANHRT)." captionText-11="FIGURES 280284. Male genitalia, a: clasping apparatus, b: aedeagus. 280. Afroracotis helicalis sp. n., PT, Gabon, slide No. LG 5399 (ANHRT); 281. A. takanoi sp. n., HT, Sierra Leone, slide No. LG 4810 (ANHRT); 282. Ibidem, PT, Guinea, slide No. LG 5445 (ANHRT); 283. Ibidem, PT, Liberia, slide No. LG 4982 (ANHRT); 284. A. staudei sp. n., HT, Kenya, slide No. LG 5540 (RCHS)." captionText-2="FIGURES 225232. Male genitalia, a: clasping apparatus, b: aedeagus. 225. Afroracotis stadiei sp. n., PT, Rwanda, slide No. LG 4789 (ZSM); 226. A. violetteae sp. n., HT, Guinea, slide No. LG 5433 (ANHRT); 227. Ibidem, PT, Liberia, slide No. LG 4999 (ANHRT); 228. A. fiebigi sp. n., HT, Liberia, slide No. LG 5000 (ANHRT); 229. Ibidem, PT, Cameroon, slide No. LG 5070 (ZSM); 230. Ibidem, PT, Uganda, slide No. LG 4106 (NHMUK); 231. A. albitrigonis, São Tomé, slide No. LG 4800 (ANHRT); 232. A. apodosima, Mozambique, slide No. LG 4827 (ANHRT)." captionText-3="FIGURES 233238. Male genitalia, a: clasping apparatus, b: aedeagus.; 233. Afroracotis apodosima, Zambia, slide No. LG 4930 (ANHRT); 234. Ibidem, Gabon, slide No. LG 4770 (ZSM); 235. A. dargei sp. n., HT, Cameroon, slide No. LG 4615 (ZSM); 236. Ibidem, PT, Cameroon, slide No. LG 5041 (ZSM); 237. A. deportata, Madagascar, slide No. LG 4067 (ANHRT); 238. A. argillacea argillacea, DRC, slide No. NHMUK 010315180 (NHMUK)." captionText-4="FIGURES 239244. Male genitalia, a: clasping apparatus, b: aedeagus. 239. Afroracotis argillacea argillacea, Cameroon, slide No. LG 4405 (ZSM); 240. A. a. morettoi ssp. n., HT, Ivory Coast, slide No. LG 5416 (ANHRT); 241. A. longicornuta longicornuta sp. n., HT, Liberia, slide No. LG 5064 (ANHRT); 242. Ibidem, Guinea, slide No. LG 4424 (ZSM); 243. A. l. congolana ssp. n., HT Cameroon, slide No. LG 4088 (RCMO); 244. Ibidem, PT, Cameroon, slide No. LG 5043 (ZSM)." captionText-5="FIGURES 245250. Male genitalia, a: clasping apparatus, b: aedeagus. 245. Afroracotis longicornuta congolana ssp. n., PT, DRC, slide No. NHMUK 010315179 (NHMUK); 246. A. l. ugandana ssp. n., HT, Uganda, slide No. LG 5529 (RCHS). 247. Ibidem, PT, Uganda, slide No. LG 5530 (RCHS); 248. A. aristophanousi sp. n., HT, Liberia, slide No. LG 5418 (ANHRT); 249. Ibidem, PT, Uganda, slide No. NHMUK 010315182 (NHMUK); 250. A. muscivirens sp. n., HT, Liberia, slide No. LG 4758 (ZSM)." captionText-6="FIGURES 251255. Male genitalia, a: clasping apparatus, b: aedeagus. 251. Afroracotis muscivirens sp. n., PT, Ivory Coast, slide No. LG 4617 (ZSM); 252. A. atriclava, ST of Cleora atriclava, Madagascar, slide No. LG 5039 (ZSM); 253. A. turlini sp. n., HT, Rwanda, slide No. LG 4619 (ZSM). 254. Afroracotis zebrina, Ivory Coast, slide No. LG 4176 (ANHRT); 255. Ibidem, Liberia, slide No. LG 4965 (ANHRT)." captionText-7="FIGURES 256261. Male genitalia, a: clasping apparatus, b: aedeagus. 256. Afroracotis zebrina, Ivory Coast, slide No. LG 4785 (ZSM); 257. A. chaineyi sp. n., HT, Gabon, slide No. LG 5405 (ANHRT); 258. Ibidem, PT, Nigeria, slide No. LG 4092 (NHMUK); 259. Ibidem, PT, Zambia, slide No. LG 4971 (ANHRT); 260. A. lydiae lydiae sp. n., HT, Liberia, slide No. LG 5788 (ANHRT); 261. Ibidem, PT, Gabon, slide No. LG 4824 (ANHRT)." captionText-8="FIGURES 262267. Male genitalia, a: clasping apparatus, b: aedeagus. 262. Afroracotis lydiae lydiae sp. n., PT, Princípe, slide No. LG 5789 (ANHRT); 263. A. l. orientalis ssp. n., HT, Tanzania, slide No.LG 4172 (ANHRT); 264. Ibidem, Mozambique, slide No. LG 4932 (ANHRT); 265. Ibidem, Rwanda, slide No. LG 5536 (RCHS); 266. A. madagascariensis, Madagascar, slide No. LG 4102 (NHMUK); 267. Ibidem, Madagascar, slide No. LG 4787 (ZSM)." captionText-9="FIGURES 268273. Male genitalia, a: clasping apparatus, b: aedeagus. 268. Afroracotis madagascariensis, Madagascar, slide No. LG 4078 (ANHRT); 269. A. smithi sp. n., HT, Zambia, slide No. LG 4972 (ANHRT); 270. Ibidem, PT, Gabon, slide No. LG 5398 (ANHRT); 271. Ibidem, PT, Kenya, slide No. LG 5549 (RCHS); 272. Afroracotis milesi sp. n., HT, Gabon, slide No. LG 5619 (ANHRT); 273. Ibidem, PT, Uganda, slide No. DS 228-2019 (RCDS)." figureDoi-0="http://doi.org/10.5281/zenodo.8080956" figureDoi-1="http://doi.org/10.5281/zenodo.8080960" figureDoi-10="http://doi.org/10.5281/zenodo.8080985" figureDoi-11="http://doi.org/10.5281/zenodo.8080987" figureDoi-2="http://doi.org/10.5281/zenodo.8080964" figureDoi-3="http://doi.org/10.5281/zenodo.8080966" figureDoi-4="http://doi.org/10.5281/zenodo.8080968" figureDoi-5="http://doi.org/10.5281/zenodo.8080970" figureDoi-6="http://doi.org/10.5281/zenodo.8080974" figureDoi-7="http://doi.org/10.5281/zenodo.8080976" figureDoi-8="http://doi.org/10.5281/zenodo.8080978" figureDoi-9="http://doi.org/10.5281/zenodo.8080983" httpUri-0="https://zenodo.org/record/8080956/files/figure.png" httpUri-1="https://zenodo.org/record/8080960/files/figure.png" httpUri-10="https://zenodo.org/record/8080985/files/figure.png" httpUri-11="https://zenodo.org/record/8080987/files/figure.png" httpUri-2="https://zenodo.org/record/8080964/files/figure.png" httpUri-3="https://zenodo.org/record/8080966/files/figure.png" httpUri-4="https://zenodo.org/record/8080968/files/figure.png" httpUri-5="https://zenodo.org/record/8080970/files/figure.png" httpUri-6="https://zenodo.org/record/8080974/files/figure.png" httpUri-7="https://zenodo.org/record/8080976/files/figure.png" httpUri-8="https://zenodo.org/record/8080978/files/figure.png" httpUri-9="https://zenodo.org/record/8080983/files/figure.png" pageId="8" pageNumber="9">Figs 212284</figureCitation>
). Uncus short, bifid with short apical projections or simple, moderately long, distally tapered. Tegumen short and broad, gnathos well-developed, apically rounded. Valva short, broad at base, distally tapered; costal margin sclerotised, often setose, may have a sclerotised apical projection outreaching the membranous ventro-distal section of valva. Ventral margin of valva moderately arched, medially often concave, slightly varying intra- and interspecifically; sacculus short and broad, weakly sclerotised, without process. Juxta weakly sclerotised, shield-like without arms. Vinculum short or moderately long and broad, V-shaped or medially tapered, apically rounded. Aedeagus rather thick, short to medium long, straight or slightly curved, without carina. Vesica thick, moderately long, in most species armed with robust cornutus, may bear additional cornuti field and well-developed diverticulum. Configuration of vesica-cornuti-diverticulum complex is an important diagnostic species-level character.
</paragraph>
<paragraph id="8BFDF53D5D35FFB457A552EBFCD9F8C9" blockId="8.[151,1437,223,1870]" pageId="8" pageNumber="9">
<emphasis id="B936292F5D35FFB457A552EBFE2BF98D" bold="true" box="[199,396,1519,1545]" pageId="8" pageNumber="9">Female genitalia</emphasis>
(
<figureCitation id="1379E9B85D35FFB456F952EBFD9DF98D" box="[411,570,1519,1545]" captionStart-0="FIGURES 290297" captionStart-1="FIGURES 298305" captionStart-2="FIGURES 306313" captionStart-3="FIGURES 314319" captionStart-4="FIGURES 320327" captionStart-5="FIGURES 328335" captionStart-6="FIGURES 336339" captionStartId-0="95.[151,264,1782,1806]" captionStartId-1="96.[151,264,1790,1814]" captionStartId-2="97.[151,264,1804,1828]" captionStartId-3="98.[151,264,1722,1746]" captionStartId-4="99.[151,264,1778,1802]" captionStartId-5="100.[151,264,1757,1781]" captionStartId-6="101.[151,264,1013,1037]" captionTargetBox-0="[259,1392,188,1752]" captionTargetBox-1="[193,1395,182,1760]" captionTargetBox-2="[205,1372,188,1774]" captionTargetBox-3="[223,1364,187,1692]" captionTargetBox-4="[208,1389,194,1748]" captionTargetBox-5="[197,1392,189,1727]" captionTargetBox-6="[191,1397,187,983]" captionTargetId-0="figure-17@95.[185,1402,181,1758]" captionTargetId-1="figure-21@96.[185,1402,181,1766]" captionTargetId-2="figure-17@97.[199,1387,181,1780]" captionTargetId-3="figure-21@98.[216,1371,181,1698]" captionTargetId-4="figure-17@99.[186,1401,181,1754]" captionTargetId-5="figure-21@100.[190,1398,181,1733]" captionTargetId-6="figure-17@101.[184,1404,181,989]" captionTargetPageId-0="95" captionTargetPageId-1="96" captionTargetPageId-2="97" captionTargetPageId-3="98" captionTargetPageId-4="99" captionTargetPageId-5="100" captionTargetPageId-6="101" captionText-0="FIGURES 290297. Female genitalia. 290. Afroracotis squalida squalida, Cameroon, slide No.: LG 4088 (NHMUK); 291. Ibidem, Tanzania, slide No.: LG 4068 (ANHRT); 292. A. s. voeltzkowii, Madagascar, slide No.: LG 4101 (NHMUK); 293. A. s. thomensis ssp. n., PT, São Tomé, slide No.: LG 5447 (ANHRT); 294. Ibidem, PT, São Tomé, slide No.: NHMUK 010316166 (NHMUK); 295. A. incompletaria incompletaria, Réunion, slide No.: LG 5048 (ZSM); 296. A. i. atrilunaria, Mauritius, slide No.: LG 4105 (NHMUK); 297. A. aliena sp. n., HT, Tanzania, slide No.: LG 4084 (ANHRT)." captionText-1="FIGURES 298305. Female genitalia. 298. Afroracotis angulosa, Cameroon, slide No.: ZSM G 20982 (ZSM); 299. Ibidem, Kenya, slide No.: LG 5558 (RCHS); 300. A. stadiei sp. n., PT, Tanzania, slide No.: LG 4760 (ZSM); 301. Ibidem, PT, Tanzania, slide No.: LG 5050 (ZSM); 302. A. violetteae sp. n., PT, Liberia, slide No.: LG 4977 (ANHRT); 303. A. fiebigi sp. n., PT, Liberia, slide No.: LG 4985 (ANHRT); 304. Ibidem, PT, Uganda, slide No.: LG 4107 (NHMUK); 305. A. albitrigonis, São Tomé, slide No.: LG 4446 (ANHRT)." captionText-2="FIGURES 306313. Female genitalia. 306. Afroracotis apodosima, Tanzania, slide No.: LG 4767 (ZSM); 307. Ibidem, Mozambique, slide No.: LG 4826 (ANHRT); 308. A. dargei sp. n., PT, Cameroon, slide No.: LG 4771 (ZSM); 309. A. deportata, Madagascar, slide No.: LG 4082 (ANHRT); 310. A. argillacea cedrici, PT, Bioko, slide No.: HERB 7255 (ZSM); 311. A. a. morettoi ssp. n., PT, Ivory Coast, slide No.: LG 5417 (ANHRT); 312. A. longicornuta congolana ssp. n., PT, Cameroon, slide No.: LG 5044 (ZSM); 313. A. l. ugandana ssp. n., PT, Uganda, gen. slide No.: LG 5531 (RCHS)." captionText-3="FIGURES 314319. Female genitalia. 314. Afroracotis aristophanousi sp. n., PT, Ghana, slide No.: NHMUK 010316165 (NHMUK); 315. Ibidem, PT, Cameroon, slide No.: LG 5045 (ZSM); 316. Ibidem, PT, Cameroon, slide No.: LG 5849 (RMCA); 317. A. atriclava, ST, Madagascar, slide No.: LG 5040 (ZSM); 318. A. muscivirens sp. n., PT, Liberia, slide No.: LG 5430 (ANHRT); 319. A. turlini sp. n., PT, Rwanda, slide No.: LG 4773 (ZSM)." captionText-4="FIGURES 320327. Female genitalia. 320. Afroracotis zebrina, Liberia, slide No.: LG 5436 (ANHRT); 321. Ibidem, Ivory Coast, slide No.: LG 4743 (ZSM); 322. A. chaineyi sp. n., PT, Guinea, slide No.: LG 5446 (ANHRT); 323. Ibidem, PT, Bioko, slide No.: LG 5073 (ZSM); 324. Ibidem, PT, Zambia, slide No.: LG 5432 (ANHRT); 325. Ibidem, PT, DRC, slide No.: LG 5054 (ZSM); 326. A. lydiae lydiae sp. n., PT, Guinea, slide No.: LG 5439 (ANHRT); 327 Ibidem, PT, Gabon, gen. slide No.: LG 5404 (ANHRT)." captionText-5="FIGURES 328335. Female genitalia. 328. Afroracotis lydiae orientalis ssp. n., PT, Tanzania, slide No.: LG 4173 (ANHRT); 329. Ibidem, PT, Mozambique, slide No.: LG 4955 (ANHRT); 330. A. madagascariensis, Madagascar, slide No.: LG 4750 (ZSM); 331. A. smithi sp. n., PT, Zambia, slide No.: LG 5400 (ANHRT); 332. Ibidem, PT, Cameroon, slide No.: LG 5074 (ZSM); 333. A. milesi sp. n., PT, Gabon, slide No.: LG 5411 (ANHRT); 334. Ibidem, PT, Cameroon, slide No.: LG 5437 (ANHRT); 335. A. ochsei sp. n., PT, Gabon, gen. slide No.: LG 5623 (ANHRT)." captionText-6="FIGURES 336339. Female genitalia. 336. Afroracotis helicalis sp. n., PT, Gabon, slide No.: LG 5624 (ANHRT); 337. A. takanoi sp. n., PT, Liberia, slide No.: LG 5438 (ANHRT); 338. Ibidem, PT, Ivory Coast, slide No.: LG 4809 (ANHRT); 339. Chorocotis ugandaria, Cameroon, slide No.: NHMUK 010317818 (NHMUK)." figureDoi-0="http://doi.org/10.5281/zenodo.8080993" figureDoi-1="http://doi.org/10.5281/zenodo.8080995" figureDoi-2="http://doi.org/10.5281/zenodo.8080997" figureDoi-3="http://doi.org/10.5281/zenodo.8080999" figureDoi-4="http://doi.org/10.5281/zenodo.8081003" figureDoi-5="http://doi.org/10.5281/zenodo.8081005" figureDoi-6="http://doi.org/10.5281/zenodo.8081007" httpUri-0="https://zenodo.org/record/8080993/files/figure.png" httpUri-1="https://zenodo.org/record/8080995/files/figure.png" httpUri-2="https://zenodo.org/record/8080997/files/figure.png" httpUri-3="https://zenodo.org/record/8080999/files/figure.png" httpUri-4="https://zenodo.org/record/8081003/files/figure.png" httpUri-5="https://zenodo.org/record/8081005/files/figure.png" httpUri-6="https://zenodo.org/record/8081007/files/figure.png" pageId="8" pageNumber="9">Figs 290339</figureCitation>
). Ovipositor medium long, papilla analis narrow triangular, apically rounded, sparsely setose, apophysis posterioris moderately long. Eighth tergite medium long, quadrangular or trapezoidal, distal margin more or less straight, proximal margin often undulate, apophysis anterioris shorter than apophysis posterioris. Ostium bursae broad, membranous, antrum moderately sclerotised, short, funnel-, cup- or goblet-shaped. Ductus bursae membranous, short and narrow, may be fully reduced, substituted by tubular distal part of cervix-corpus bursae complex. Cervix bursae well-developed, heavily sclerotised, often present as a rounded bulge or may be fully merged with corpus bursae forming a sclerotised, distal section of bursa copulatrix. Corpus bursae varying in size interspecifically, may be highly reduced and fully fused with sclerotised, often rugose cervix bursae or well-developed and separated from cervix; may bear a strongly sclerotised disc-shaped plate supposedly serving as receptive surface of cornutus during copulation; signum absent.
</paragraph>
</subSubSection>
<subSubSection id="C358A6B65D34FFB657F55793FB22FCA7" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10" type="multiple">
<paragraph id="8BFDF53D5D34FFB557F55793FD61FF35" blockId="9.[151,710,151,177]" box="[151,710,151,177]" pageId="9" pageNumber="10">
<emphasis id="B936292F5D34FFB557F55793FD61FF35" bold="true" box="[151,710,151,177]" pageId="9" pageNumber="10">
Taxonomic overview of the genus
<taxonomicName id="4C428EBE5D34FFB5555F5793FD61FF35" authorityName="László &amp; Hausmann &amp; Karisch" authorityYear="2023" box="[573,710,151,177]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B936292F5D34FFB5555F5793FD61FF35" bold="true" box="[573,710,151,177]" italics="true" pageId="9" pageNumber="10">Afroracotis</emphasis>
</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="8BFDF53D5D34FFB557F557DBFE01FF7D" blockId="9.[151,422,223,249]" box="[151,422,223,249]" pageId="9" pageNumber="10">
<heading id="D0B542515D34FFB557F557DBFE01FF7D" bold="true" box="[151,422,223,249]" fontSize="11" level="2" pageId="9" pageNumber="10" reason="4">
<emphasis id="B936292F5D34FFB557F557DBFE01FF7D" bold="true" box="[151,422,223,249]" pageId="9" pageNumber="10">
<taxonomicName id="4C428EBE5D34FFB557F557DBFE87FF7D" authority="László &amp; Hausmann &amp; Karisch, 2023" authorityName="László &amp; Hausmann &amp; Karisch" authorityYear="2023" box="[151,288,223,249]" pageId="9" pageNumber="10" rank="subGenus" status="subgen. nov." subGenus="Afroracotis">
<emphasis id="B936292F5D34FFB557F557DBFE87FF7D" bold="true" box="[151,288,223,249]" italics="true" pageId="9" pageNumber="10">Afroracotis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20594545D34FFB5564557DBFE01FF7D" box="[295,422,223,249]" pageId="9" pageNumber="10" rank="subGenus">subgen. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BFDF53D5D34FFB557F55623FDDEFEC5" blockId="9.[151,633,295,322]" box="[151,633,295,322]" pageId="9" pageNumber="10">
<emphasis id="B936292F5D34FFB557F55623FDDEFEC5" bold="true" box="[151,633,295,322]" pageId="9" pageNumber="10">
The
<taxonomicName id="4C428EBE5D34FFB557AD5623FE60FEC6" box="[207,455,295,322]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="squalida">
<emphasis id="B936292F5D34FFB557AD5623FE60FEC6" bold="true" box="[207,455,295,322]" italics="true" pageId="9" pageNumber="10">Afroracotis squalida</emphasis>
</taxonomicName>
species-group
</emphasis>
</paragraph>
<paragraph id="8BFDF53D5D34FFB557F55668FCE1FE00" blockId="9.[151,1018,364,541]" box="[151,838,364,388]" pageId="9" pageNumber="10">
<taxonomicName id="4C428EBE5D34FFB557F55668FD44FE00" authority="(Butler, 1878)" authorityName="László &amp; Hausmann &amp; Karisch" authorityYear="2023" baseAuthorityName="Butler" baseAuthorityYear="1878" box="[151,739,364,388]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="subSpecies" species="squalida" status="comb. nov." subGenus="Afroracotis" subSpecies="squalida">
<emphasis id="B936292F5D34FFB557F55668FDE9FE00" box="[151,590,364,388]" italics="true" pageId="9" pageNumber="10">Afroracotis (Afroracotis) squalida squalida</emphasis>
(
<bibRefCitation id="EFD388CC5D34FFB5553E5668FD7BFE00" author="Butler, A. G." box="[604,732,364,388]" pageId="9" pageNumber="10" pagination="455 - 465" refId="ref82974" refString="Butler, A. G. (1878) Descriptions of some new genera and species of Lepidoptera from Old Calabar and Madagascar. Annals and Magazine of Natural History, Series 5, 2 (12), 455 - 465." type="journal article" year="1878">Butler, 1878</bibRefCitation>
)
</taxonomicName>
<emphasis id="B936292F5D34FFB555885668FCE1FE00" bold="true" box="[746,838,364,388]" pageId="9" pageNumber="10">
<taxonomicNameLabel id="A20594545D34FFB555885668FCE1FE00" box="[746,838,364,388]" pageId="9" pageNumber="10" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BFDF53D5D34FFB557F5568EFC00FE26" blockId="9.[151,1018,364,541]" box="[151,935,394,418]" pageId="9" pageNumber="10">
<taxonomicName id="4C428EBE5D34FFB557F5568EFCE3FE26" authority="(Pagenstecher, 1907)" authorityName="László &amp; Hausmann &amp; Karisch" authorityYear="2023" baseAuthorityName="Pagenstecher" baseAuthorityYear="1907" box="[151,836,394,418]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="subSpecies" species="squalida" status="comb. nov." subGenus="Afroracotis" subSpecies="voeltzkowii">
<emphasis id="B936292F5D34FFB557F5568EFDCEFE26" box="[151,617,394,418]" italics="true" pageId="9" pageNumber="10">Afroracotis (Afroracotis) squalida voeltzkowii</emphasis>
(
<bibRefCitation id="EFD388CC5D34FFB55514568EFC9AFE26" author="Pagenstecher, A." box="[630,829,394,418]" pageId="9" pageNumber="10" pagination="93 - 146" refId="ref84035" refString="Pagenstecher, A. (1907) Lepidoptera - Heterocera (Uraniidae, Geometridae, Noctuidae, Pyralidae, Thyrididae, Tortricidae, Tineidae, Pterophoridae) von Madagaskar, den Comoren und Ostafrika. In: Voeltzkow, A., Reise in Ostafrika in den Jahren 1903 - 05. Wissenschaftliche Ergebnisse 2. Systematische Arbeiten, 2, pp. 93 - 146." type="journal article" year="1907">Pagenstecher, 1907</bibRefCitation>
)
</taxonomicName>
<emphasis id="B936292F5D34FFB55429568EFC00FE26" bold="true" box="[843,935,394,418]" pageId="9" pageNumber="10">
<taxonomicNameLabel id="A20594545D34FFB55429568EFC00FE26" box="[843,935,394,418]" pageId="9" pageNumber="10" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BFDF53D5D34FFB557F556ADFD0DFE45" blockId="9.[151,1018,364,541]" box="[151,682,424,449]" pageId="9" pageNumber="10">
<taxonomicName id="4C428EBE5D34FFB557F556ADFDC6FE44" authority="László &amp; Hausmann &amp; Karisch, 2023" authorityName="László &amp; Hausmann &amp; Karisch" authorityYear="2023" box="[151,609,424,449]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="19" phylum="Arthropoda" rank="subSpecies" species="squalida" status="subsp. nov." subGenus="Afroracotis" subSpecies="thomensis">
<emphasis id="B936292F5D34FFB557F556ADFDC6FE44" box="[151,609,424,449]" italics="true" pageId="9" pageNumber="10">
Afroracotis (Afroracotis) squalida
<emphasis id="B936292F5D34FFB5569556ACFDC6FE44" bold="true" box="[503,609,424,448]" pageId="9" pageNumber="10">thomensis</emphasis>
</emphasis>
</taxonomicName>
ssp. n.
</paragraph>
<paragraph id="8BFDF53D5D34FFB557F556CCFC64FE64" blockId="9.[151,1018,364,541]" box="[151,963,455,480]" pageId="9" pageNumber="10">
<taxonomicName id="4C428EBE5D34FFB557F556CCFCC7FE5B" authority="(Guenee, 1862)" authorityName="László &amp; Hausmann &amp; Karisch" authorityYear="2023" baseAuthorityName="Guenee" baseAuthorityYear="1862" box="[151,864,455,480]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="subSpecies" species="incompletaria" status="comb. nov." subGenus="Afroracotis" subSpecies="incompletaria">
<emphasis id="B936292F5D34FFB557F556CCFD1BFE64" box="[151,700,456,480]" italics="true" pageId="9" pageNumber="10">Afroracotis (Afroracotis) incompletaria incompletaria</emphasis>
(
<bibRefCitation id="EFD388CC5D34FFB555A856CCFCFEFE5B" author="Guenee, A. M." box="[714,857,455,480]" pageId="9" pageNumber="10" pagination="1 - 72" refId="ref83173" refString="Guenee, A. M. (1862) Lepidopteres. In: Maillard, L., Notes sur l'ile de la Reunion (Bourbon), Annexe G, pp. 1 - 72, pls. 12 - 13." type="book chapter" year="1862">Guenée, 1862</bibRefCitation>
)
</taxonomicName>
<emphasis id="B936292F5D34FFB5540456CCFC64FE64" bold="true" box="[870,963,456,480]" pageId="9" pageNumber="10">
<taxonomicNameLabel id="A20594545D34FFB5540456CCFC64FE64" box="[870,963,456,480]" pageId="9" pageNumber="10" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BFDF53D5D34FFB557F556E2FC5DFE7A" blockId="9.[151,1018,364,541]" box="[151,1018,486,510]" pageId="9" pageNumber="10">
<taxonomicName id="4C428EBE5D34FFB557F556E2FCE5FE7A" authority="(Mabille, 1893)" authorityName="László &amp; Hausmann &amp; Karisch" authorityYear="2023" baseAuthorityName="Mabille" baseAuthorityYear="1893" box="[151,834,486,510]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="subSpecies" species="incompletaria" status="comb. nov." subGenus="Afroracotis" subSpecies="atrilunaria">
<emphasis id="B936292F5D34FFB557F556E2FD3AFE7A" box="[151,669,486,510]" italics="true" pageId="9" pageNumber="10">Afroracotis (Afroracotis) incompletaria atrilunaria</emphasis>
(
<bibRefCitation id="EFD388CC5D34FFB555C956E2FC9BFE7A" author="Mabille, P." box="[683,828,486,510]" pageId="9" pageNumber="10" pagination="50 - 65" refId="ref83882" refString="Mabille, P. (1893) Descriptions de Lepidopteres nouveaux. Annales de la Societe entomologique de Belgique, 37, 50 - 65." type="journal article" year="1893">Mabille, 1893</bibRefCitation>
)
</taxonomicName>
<emphasis id="B936292F5D34FFB5542B56E2FC02FE7A" bold="true" box="[841,933,486,510]" pageId="9" pageNumber="10">
<taxonomicNameLabel id="A20594545D34FFB5542B56E2FC02FE7A" box="[841,933,486,510]" pageId="9" pageNumber="10" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
,
<emphasis id="B936292F5D34FFB554D056E2FC5DFE7A" bold="true" box="[946,1018,486,510]" pageId="9" pageNumber="10">stat. n.</emphasis>
</paragraph>
<paragraph id="8BFDF53D5D34FFB557F55501FDBDFD99" blockId="9.[151,1018,364,541]" box="[151,538,516,541]" pageId="9" pageNumber="10">
<taxonomicName id="4C428EBE5D34FFB557F55501FE7CFD98" authority="László &amp; Hausmann &amp; Karisch, 2023" authorityName="László &amp; Hausmann &amp; Karisch" authorityYear="2023" box="[151,475,516,541]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="aliena" status="sp. nov." subGenus="Afroracotis">
<emphasis id="B936292F5D34FFB557F55501FE7CFD98" box="[151,475,516,541]" italics="true" pageId="9" pageNumber="10">
Afroracotis (Afroracotis)
<emphasis id="B936292F5D34FFB556F85500FE7CFD98" bold="true" box="[410,475,516,540]" pageId="9" pageNumber="10">aliena</emphasis>
</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20594545D34FFB556835501FDBDFD99" box="[481,538,517,541]" pageId="9" pageNumber="10" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BFDF53D5D34FFB557F5556BFD26FD0D" blockId="9.[151,641,623,650]" box="[151,641,623,650]" pageId="9" pageNumber="10">
<emphasis id="B936292F5D34FFB557F5556BFD26FD0D" bold="true" box="[151,641,623,650]" pageId="9" pageNumber="10">
The
<taxonomicName id="4C428EBE5D34FFB557AD556BFE68FD0E" authorityName="Herbulot" authorityYear="1973" box="[207,463,623,650]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="angulosa">
<emphasis id="B936292F5D34FFB557AD556BFE68FD0E" bold="true" box="[207,463,623,650]" italics="true" pageId="9" pageNumber="10">Afroracotis angulosa</emphasis>
</taxonomicName>
species-group
</emphasis>
</paragraph>
<paragraph id="8BFDF53D5D34FFB557F555AAFCADFD42" blockId="9.[151,778,686,864]" box="[151,778,686,710]" pageId="9" pageNumber="10">
<taxonomicName id="4C428EBE5D34FFB557F555AAFD00FD42" authority="(Herbulot, 1973)" authorityName="László &amp; Hausmann &amp; Karisch" authorityYear="2023" baseAuthorityName="Herbulot" baseAuthorityYear="1973" box="[151,679,686,710]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="angulosa" status="comb. nov." subGenus="Afroracotis">
<emphasis id="B936292F5D34FFB557F555AAFE50FD42" box="[151,503,686,710]" italics="true" pageId="9" pageNumber="10">Afroracotis (Afroracotis) angulosa</emphasis>
(
<bibRefCitation id="EFD388CC5D34FFB5556655AAFD07FD42" author="Herbulot, C." box="[516,672,686,710]" pageId="9" pageNumber="10" pagination="141 - 148" refId="ref83407" refString="Herbulot, C. (1973) Nouveaux Ennominae du Cameroun (Lepidoptera, Geometridae). Annales de la Faculte des Sciences du Cameroun, 1973 (14), 141 - 148." type="journal article" year="1973">Herbulot, 1973</bibRefCitation>
)
</taxonomicName>
<emphasis id="B936292F5D34FFB555CF55AAFCADFD42" bold="true" box="[685,778,686,710]" pageId="9" pageNumber="10">
<taxonomicNameLabel id="A20594545D34FFB555CF55AAFCADFD42" box="[685,778,686,710]" pageId="9" pageNumber="10" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BFDF53D5D34FFB557F555C9FD42FD61" blockId="9.[151,778,686,864]" box="[151,741,717,741]" pageId="9" pageNumber="10">
<taxonomicName id="4C428EBE5D34FFB557F555C9FD25FD61" authority="(Herbulot, 1991)" authorityName="László &amp; Hausmann &amp; Karisch" authorityYear="2023" baseAuthorityName="Herbulot" baseAuthorityYear="1991" box="[151,642,717,741]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="canui" status="comb. nov." subGenus="Afroracotis">
<emphasis id="B936292F5D34FFB557F555C9FE75FD61" box="[151,466,717,741]" italics="true" pageId="9" pageNumber="10">Afroracotis (Afroracotis) canui</emphasis>
(
<bibRefCitation id="EFD388CC5D34FFB556BD55C9FDDCFD61" author="Herbulot, C." box="[479,635,717,741]" pageId="9" pageNumber="10" pagination="16" refId="ref83442" refString="Herbulot, C. (1991) Un nouveau Racotis africain (Lepidoptera Geometridae). Bulletin de la Societe entomologique de Mulhouse, 21 (1), 16." type="journal article" year="1991">Herbulot, 1991</bibRefCitation>
)
</taxonomicName>
<emphasis id="B936292F5D34FFB555EB55C9FD42FD61" bold="true" box="[649,741,717,741]" pageId="9" pageNumber="10">
<taxonomicNameLabel id="A20594545D34FFB555EB55C9FD42FD61" box="[649,741,717,741]" pageId="9" pageNumber="10" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BFDF53D5D34FFB557F555E8FDBAFC80" blockId="9.[151,778,686,864]" box="[151,541,747,772]" pageId="9" pageNumber="10">
<taxonomicName id="4C428EBE5D34FFB557F555E8FE79FC87" authority="László &amp; Hausmann &amp; Karisch, 2023" authorityName="László &amp; Hausmann &amp; Karisch" authorityYear="2023" box="[151,478,747,772]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="stadiei" status="sp. nov." subGenus="Afroracotis">
<emphasis id="B936292F5D34FFB557F555E8FE79FC87" box="[151,478,747,772]" italics="true" pageId="9" pageNumber="10">
Afroracotis (Afroracotis)
<emphasis id="B936292F5D34FFB556F855EFFE79FC87" bold="true" box="[410,478,747,771]" pageId="9" pageNumber="10">stadiei</emphasis>
</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20594545D34FFB5568655E8FDBAFC80" box="[484,541,748,772]" pageId="9" pageNumber="10" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BFDF53D5D34FFB557F5540EFD9CFCA6" blockId="9.[151,778,686,864]" box="[151,571,778,802]" pageId="9" pageNumber="10">
<taxonomicName id="4C428EBE5D34FFB557F5540EFE5BFCA6" authority="László &amp; Hausmann &amp; Karisch, 2023" authorityName="László &amp; Hausmann &amp; Karisch" authorityYear="2023" box="[151,508,778,802]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="violetteae" status="sp. nov." subGenus="Afroracotis">
<emphasis id="B936292F5D34FFB557F5540EFE5BFCA6" box="[151,508,778,802]" italics="true" pageId="9" pageNumber="10">
Afroracotis (Afroracotis)
<emphasis id="B936292F5D34FFB556F8540EFE5BFCA6" bold="true" box="[410,508,778,802]" pageId="9" pageNumber="10">violetteae</emphasis>
</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20594545D34FFB55560540EFD9CFCA6" box="[514,571,778,802]" pageId="9" pageNumber="10" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BFDF53D5D34FFB557F5542DFDBCFCC5" blockId="9.[151,778,686,864]" box="[151,539,808,833]" pageId="9" pageNumber="10">
<taxonomicName id="4C428EBE5D34FFB557F5542DFE7BFCC4" authority="László &amp; Hausmann &amp; Karisch, 2023" authorityName="László &amp; Hausmann &amp; Karisch" authorityYear="2023" box="[151,476,808,833]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="fiebigi" status="sp. nov." subGenus="Afroracotis">
<emphasis id="B936292F5D34FFB557F5542DFE7BFCC4" box="[151,476,808,833]" italics="true" pageId="9" pageNumber="10">
Afroracotis (Afroracotis)
<emphasis id="B936292F5D34FFB556F8542CFE7BFCC4" bold="true" box="[410,476,808,832]" pageId="9" pageNumber="10">fiebigi</emphasis>
</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20594545D34FFB55680542DFDBCFCC5" box="[482,539,809,833]" pageId="9" pageNumber="10" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BFDF53D5D34FFB557F5544CFCA7FCE4" blockId="9.[151,778,686,864]" box="[151,768,840,864]" pageId="9" pageNumber="10">
<taxonomicName id="4C428EBE5D34FFB557F5544CFD3AFCE4" authority="(Prout, 1927)" authorityName="László &amp; Hausmann &amp; Karisch" authorityYear="2023" baseAuthorityName="Prout" baseAuthorityYear="1927" box="[151,669,840,864]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="albitrigonis" status="comb. nov." subGenus="Afroracotis">
<emphasis id="B936292F5D34FFB557F5544CFDB7FCE4" box="[151,528,840,864]" italics="true" pageId="9" pageNumber="10">Afroracotis (Afroracotis) albitrigonis</emphasis>
(
<bibRefCitation id="EFD388CC5D34FFB5557F544CFD31FCE4" author="Prout, L. B." box="[541,662,840,864]" pageId="9" pageNumber="10" pagination="187 - 200" refId="ref84205" refString="Prout, L. B. (1927) A list of the Geometridae (Lep. Het.) known to occur in the Island of Sai Thome, with descriptions of some new species collected by Mr. T. A. Barns. Transactions of the entomological Society of London, 75 (1), 187 - 200." type="journal article" year="1927">Prout, 1927</bibRefCitation>
)
</taxonomicName>
<emphasis id="B936292F5D34FFB555C6544CFCA7FCE4" bold="true" box="[676,768,840,864]" pageId="9" pageNumber="10">
<taxonomicNameLabel id="A20594545D34FFB555C6544CFCA7FCE4" box="[676,768,840,864]" pageId="9" pageNumber="10" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BFDF53D5D34FFB557F554B5FE75FC48" blockId="9.[151,466,945,972]" box="[151,466,945,972]" pageId="9" pageNumber="10">
<heading id="D0B542515D34FFB557F554B5FE75FC48" bold="true" box="[151,466,945,972]" fontSize="11" level="2" pageId="9" pageNumber="10" reason="4">
<emphasis id="B936292F5D34FFB557F554B5FE75FC48" bold="true" box="[151,466,945,972]" pageId="9" pageNumber="10">
<taxonomicName id="4C428EBE5D34FFB557F554B5FEEBFC48" authority="László &amp; Hausmann &amp; Karisch, 2023" authorityName="László &amp; Hausmann &amp; Karisch" authorityYear="2023" box="[151,332,945,972]" pageId="9" pageNumber="10" rank="subGenus" status="subgen. nov." subGenus="Rwandaracotis">
<emphasis id="B936292F5D34FFB557F554B5FEEBFC48" bold="true" box="[151,332,945,972]" italics="true" pageId="9" pageNumber="10">Rwandaracotis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20594545D34FFB5563154B6FE75FC48" box="[339,466,946,972]" pageId="9" pageNumber="10" rank="subGenus">subgen. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BFDF53D5D34FFB557F554F2FDE5FB8A" blockId="9.[151,578,1014,1038]" box="[151,578,1014,1038]" pageId="9" pageNumber="10">
<taxonomicName id="4C428EBE5D34FFB557F554F2FDA4FB8A" authority="László &amp; Hausmann &amp; Karisch, 2023" authorityName="László &amp; Hausmann &amp; Karisch" authorityYear="2023" box="[151,515,1014,1038]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="49" phylum="Arthropoda" rank="species" species="turlini" status="sp. nov." subGenus="Rwandaracotis">
<emphasis id="B936292F5D34FFB557F554F2FDA4FB8A" box="[151,515,1014,1038]" italics="true" pageId="9" pageNumber="10">
Afroracotis (Rwandaracotis)
<emphasis id="B936292F5D34FFB556A354F2FDA4FB8A" bold="true" box="[449,515,1014,1038]" pageId="9" pageNumber="10">turlini</emphasis>
</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20594545D34FFB5556B54F2FDE5FB8A" box="[521,578,1014,1038]" pageId="9" pageNumber="10" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BFDF53D5D34FFB557F55364FE73FBFF" blockId="9.[151,468,1120,1147]" box="[151,468,1120,1147]" pageId="9" pageNumber="10">
<heading id="D0B542515D34FFB557F55364FE73FBFF" bold="true" box="[151,468,1120,1147]" fontSize="11" level="2" pageId="9" pageNumber="10" reason="4">
<emphasis id="B936292F5D34FFB557F55364FE73FBFF" bold="true" box="[151,468,1120,1147]" pageId="9" pageNumber="10">
<taxonomicName id="4C428EBE5D34FFB557F55364FEE9FBFF" authority="László &amp; Hausmann &amp; Karisch, 2023" authorityName="László &amp; Hausmann &amp; Karisch" authorityYear="2023" box="[151,334,1120,1147]" pageId="9" pageNumber="10" rank="subGenus" status="subgen. nov." subGenus="Herbuloracotis">
<emphasis id="B936292F5D34FFB557F55364FEE9FBFF" bold="true" box="[151,334,1120,1147]" italics="true" pageId="9" pageNumber="10">Herbuloracotis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20594545D34FFB556375365FE73FBFF" box="[341,468,1121,1147]" pageId="9" pageNumber="10" rank="subGenus">subgen. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BFDF53D5D34FFB557F553ADFD33FB46" blockId="9.[151,660,1192,1219]" box="[151,660,1192,1219]" pageId="9" pageNumber="10">
<emphasis id="B936292F5D34FFB557F553ADFD33FB46" bold="true" box="[151,660,1192,1219]" pageId="9" pageNumber="10">
The
<taxonomicName id="4C428EBE5D34FFB557AD53ACFE45FB47" authorityName="László &amp; Hausmann &amp; Karisch" authorityYear="2023" baseAuthorityName="Prout" baseAuthorityYear="1931" box="[207,482,1192,1219]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="apodosima">
<emphasis id="B936292F5D34FFB557AD53ACFE45FB47" bold="true" box="[207,482,1192,1219]" italics="true" pageId="9" pageNumber="10">Afroracotis apodosima</emphasis>
</taxonomicName>
species-group
</emphasis>
</paragraph>
<paragraph id="8BFDF53D5D34FFB557F553E9FC86FA81" blockId="9.[151,801,1261,1316]" box="[151,801,1261,1285]" pageId="9" pageNumber="10">
<taxonomicName id="4C428EBE5D34FFB557F553E9FD19FA81" authority="(Prout, 1931)" authorityName="László &amp; Hausmann &amp; Karisch" authorityYear="2023" baseAuthorityName="Prout" baseAuthorityYear="1931" box="[151,702,1261,1285]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="apodosima" status="comb. nov." subGenus="Herbuloracotis">
<emphasis id="B936292F5D34FFB557F553E9FD96FA81" box="[151,561,1261,1285]" italics="true" pageId="9" pageNumber="10">Afroracotis (Herbuloracotis) apodosima</emphasis>
(
<bibRefCitation id="EFD388CC5D34FFB5555C53E9FD1FFA81" author="Prout, L. B." box="[574,696,1261,1285]" pageId="9" pageNumber="10" pagination="151 - 180" refId="ref84268" refString="Prout, L. B. (1931) Geometrid descriptions and notes. Novitates Zoologicae, 36 (2), 151 - 180." type="journal article" year="1931">Prout, 1931</bibRefCitation>
)
</taxonomicName>
<emphasis id="B936292F5D34FFB555A753E9FC86FA81" bold="true" box="[709,801,1261,1285]" pageId="9" pageNumber="10">
<taxonomicNameLabel id="A20594545D34FFB555A753E9FC86FA81" box="[709,801,1261,1285]" pageId="9" pageNumber="10" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BFDF53D5D34FFB557F55208FDE4FAA0" blockId="9.[151,801,1261,1316]" box="[151,579,1291,1316]" pageId="9" pageNumber="10">
<taxonomicName id="4C428EBE5D34FFB557F55208FDA3FAA7" authority="László &amp; Hausmann &amp; Karisch, 2023" authorityName="László &amp; Hausmann &amp; Karisch" authorityYear="2023" box="[151,516,1291,1316]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="35" phylum="Arthropoda" rank="species" species="dargei" status="sp. nov." subGenus="Herbuloracotis">
<emphasis id="B936292F5D34FFB557F55208FDA3FAA7" box="[151,516,1291,1316]" italics="true" pageId="9" pageNumber="10">
Afroracotis (Herbuloracotis)
<emphasis id="B936292F5D34FFB556A0520FFDA3FAA7" bold="true" box="[450,516,1291,1315]" pageId="9" pageNumber="10">dargei</emphasis>
</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20594545D34FFB555685208FDE4FAA0" box="[522,579,1292,1316]" pageId="9" pageNumber="10" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BFDF53D5D34FFB557F55272FD23FA0B" blockId="9.[151,644,1397,1424]" box="[151,644,1397,1424]" pageId="9" pageNumber="10">
<emphasis id="B936292F5D34FFB557F55272FD23FA0B" bold="true" box="[151,644,1397,1424]" pageId="9" pageNumber="10">
The
<taxonomicName id="4C428EBE5D34FFB557AD5271FE75FA14" authorityName="László &amp; Hausmann &amp; Karisch" authorityYear="2023" baseAuthorityName="Herbulot" baseAuthorityYear="1970" box="[207,466,1397,1424]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="deportata">
<emphasis id="B936292F5D34FFB557AD5271FE75FA14" bold="true" box="[207,466,1397,1424]" italics="true" pageId="9" pageNumber="10">Afroracotis deportata</emphasis>
</taxonomicName>
species-group
</emphasis>
</paragraph>
<paragraph id="8BFDF53D5D34FFB557F552BEFC9FFA56" blockId="9.[151,824,1466,1490]" box="[151,824,1466,1490]" pageId="9" pageNumber="10">
<taxonomicName id="4C428EBE5D34FFB557F552BEFD72FA56" authority="(Herbulot, 1970)" authorityName="László &amp; Hausmann &amp; Karisch" authorityYear="2023" baseAuthorityName="Herbulot" baseAuthorityYear="1970" box="[151,725,1466,1490]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="deportata" status="comb. nov." subGenus="Herbuloracotis">
<emphasis id="B936292F5D34FFB557F552BEFD82FA56" box="[151,549,1466,1490]" italics="true" pageId="9" pageNumber="10">Afroracotis (Herbuloracotis) deportata</emphasis>
(
<bibRefCitation id="EFD388CC5D34FFB5555052BEFD69FA56" author="Herbulot, C." box="[562,718,1466,1490]" pageId="9" pageNumber="10" pagination="157 - 166" refId="ref83371" refString="Herbulot, C. (1970) Lepidoptera Geometridae du Tsaratanana (Madagascar Nord). Memoires O. R. S. T. O. M., 37, 157 - 166." type="journal article" year="1970">Herbulot, 1970</bibRefCitation>
)
</taxonomicName>
<emphasis id="B936292F5D34FFB555B952BEFC9FFA56" bold="true" box="[731,824,1466,1490]" pageId="9" pageNumber="10">
<taxonomicNameLabel id="A20594545D34FFB555B952BEFC9FFA56" box="[731,824,1466,1490]" pageId="9" pageNumber="10" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BFDF53D5D34FFB557F55121FD2CF9BA" blockId="9.[151,651,1572,1599]" box="[151,651,1572,1599]" pageId="9" pageNumber="10">
<emphasis id="B936292F5D34FFB557F55121FD2CF9BA" bold="true" box="[151,651,1572,1599]" pageId="9" pageNumber="10">
The
<taxonomicName id="4C428EBE5D34FFB557AD5120FE7EF9BB" baseAuthorityName="Warren" baseAuthorityYear="1900" box="[207,473,1572,1599]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="argillacea">
<emphasis id="B936292F5D34FFB557AD5120FE7EF9BB" bold="true" box="[207,473,1572,1599]" italics="true" pageId="9" pageNumber="10">Afroracotis argillacea</emphasis>
</taxonomicName>
species-group
</emphasis>
</paragraph>
<paragraph id="8BFDF53D5D34FFB557F5516DFC3DF905" blockId="9.[151,922,1641,1910]" box="[151,922,1641,1665]" pageId="9" pageNumber="10">
<taxonomicName id="4C428EBE5D34FFB557F5516DFC90F905" authority="(Warren, 1900)" authorityName="László &amp; Hausmann &amp; Karisch" authorityYear="2023" baseAuthorityName="Warren" baseAuthorityYear="1900" box="[151,823,1641,1665]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="subSpecies" species="argillacea" status="comb. nov." subGenus="Herbuloracotis" subSpecies="argillacea">
<emphasis id="B936292F5D34FFB557F5516DFD30F905" box="[151,663,1641,1665]" italics="true" pageId="9" pageNumber="10">Afroracotis (Herbuloracotis) argillacea argillacea</emphasis>
(
<bibRefCitation id="EFD388CC5D34FFB555C6516DFC97F905" author="Warren, W." box="[676,816,1641,1665]" pageId="9" pageNumber="10" pagination="90 - 98" refId="ref85235" refString="Warren, W. (1900) New genera and species of Thyrididae and Geometridae from Africa. Novitates Zoologicae, 7, 90 - 98." type="journal article" year="1900">Warren, 1900</bibRefCitation>
)
</taxonomicName>
<emphasis id="B936292F5D34FFB5545F516DFC3DF905" bold="true" box="[829,922,1641,1665]" pageId="9" pageNumber="10">
<taxonomicNameLabel id="A20594545D34FFB5545F516DFC3DF905" box="[829,922,1641,1665]" pageId="9" pageNumber="10" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BFDF53D5D34FFB557F5518CFD77F924" blockId="9.[151,922,1641,1910]" box="[151,720,1671,1696]" pageId="9" pageNumber="10">
<taxonomicName id="4C428EBE5D34FFB557F5518CFD20F91B" authority="László &amp; Hausmann &amp; Karisch, 2023" authorityName="László &amp; Hausmann &amp; Karisch" authorityYear="2023" box="[151,647,1671,1696]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="41" phylum="Arthropoda" rank="subSpecies" species="argillacea" status="subsp. nov." subGenus="Herbuloracotis" subSpecies="morettoi">
<emphasis id="B936292F5D34FFB557F5518CFD20F91B" box="[151,647,1671,1696]" italics="true" pageId="9" pageNumber="10">
Afroracotis (Herbuloracotis) argillacea
<emphasis id="B936292F5D34FFB555525183FD20F91B" bold="true" box="[560,647,1671,1695]" pageId="9" pageNumber="10">morettoi</emphasis>
</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20594545D34FFB555EF518CFD77F924" box="[653,720,1672,1696]" pageId="9" pageNumber="10" rank="subSpecies">ssp. n.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BFDF53D5D34FFB557F551A2FCD2F93A" blockId="9.[151,922,1641,1910]" box="[151,885,1702,1726]" pageId="9" pageNumber="10">
<taxonomicName id="4C428EBE5D34FFB557F551A2FC81F93A" authority="(Herbulot, 1998)" authorityName="László &amp; Hausmann &amp; Karisch" authorityYear="2023" baseAuthorityName="Herbulot" baseAuthorityYear="1998" box="[151,806,1702,1726]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="subSpecies" species="argillacea" subGenus="Herbuloracotis" subSpecies="cedrici">
<emphasis id="B936292F5D34FFB557F551A2FDD1F93A" box="[151,630,1702,1726]" italics="true" pageId="9" pageNumber="10">Afroracotis (Herbuloracotis) argillacea cedrici</emphasis>
(
<bibRefCitation id="EFD388CC5D34FFB555E151A2FCB8F93A" author="Herbulot, C." box="[643,799,1702,1726]" pageId="9" pageNumber="10" pagination="287 - 292" refId="ref83473" refString="Herbulot, C. (1998) Lepidopteres recoltes par le Dr. J. - G. Canu dans l'ile de Bioko (ex Fernando Poo) (Lepidoptera). Lambillionea, 98 (2), 287 - 292." type="journal article" year="1998">Herbulot, 1998</bibRefCitation>
)
</taxonomicName>
<emphasis id="B936292F5D34FFB5544F51A2FCD2F93A" bold="true" box="[813,885,1702,1726]" pageId="9" pageNumber="10">stat. n.</emphasis>
</paragraph>
<paragraph id="8BFDF53D5D34FFB557F551C1FCB1F959" blockId="9.[151,922,1641,1910]" box="[151,790,1732,1757]" pageId="9" pageNumber="10">
<emphasis id="B936292F5D34FFB557F551C1FD71F958" box="[151,726,1732,1757]" italics="true" pageId="9" pageNumber="10">
<taxonomicName id="4C428EBE5D34FFB557F551C1FDEEF958" authorityName="László &amp; Hausmann &amp; Karisch" authorityYear="2023" box="[151,585,1732,1757]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="longicornuta" subGenus="Herbuloracotis">
Afroracotis (Herbuloracotis)
<emphasis id="B936292F5D34FFB556A051C0FDEEF958" bold="true" box="[450,585,1732,1756]" pageId="9" pageNumber="10">longicornuta</emphasis>
</taxonomicName>
<taxonomicName id="4C428EBE5D34FFB5552D51C0FD71F958" authority="László &amp; Hausmann &amp; Karisch, 2023" authorityName="László &amp; Hausmann &amp; Karisch" authorityYear="2023" box="[591,726,1732,1756]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="longicornuta" status="sp. nov.">
<emphasis id="B936292F5D34FFB5552D51C0FD71F958" bold="true" box="[591,726,1732,1756]" pageId="9" pageNumber="10">longicornuta</emphasis>
</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A20594545D34FFB555BF51C1FCB1F959" box="[733,790,1733,1757]" pageId="9" pageNumber="10" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BFDF53D5D34FFB557F551E0FCA1F978" blockId="9.[151,922,1641,1910]" box="[151,774,1763,1788]" pageId="9" pageNumber="10">
<taxonomicName id="4C428EBE5D34FFB557F551E0FD1AF97F" authority="László &amp; Hausmann &amp; Karisch, 2023" authorityName="László &amp; Hausmann &amp; Karisch" authorityYear="2023" box="[151,701,1763,1788]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="43" phylum="Arthropoda" rank="subSpecies" species="longicornuta" status="subsp. nov." subGenus="Herbuloracotis" subSpecies="congolana">
<emphasis id="B936292F5D34FFB557F551E0FD1AF97F" box="[151,701,1763,1788]" italics="true" pageId="9" pageNumber="10">
Afroracotis (Herbuloracotis)
<emphasis id="B936292F5D34FFB556A051E7FD1AF97F" bold="true" box="[450,701,1763,1787]" pageId="9" pageNumber="10">longicornuta congolana</emphasis>
</emphasis>
</taxonomicName>
ssp. n.
</paragraph>
<paragraph id="8BFDF53D5D34FFB557F55006FCA5F89E" blockId="9.[151,922,1641,1910]" box="[151,770,1794,1818]" pageId="9" pageNumber="10">
<taxonomicName id="4C428EBE5D34FFB557F55006FD1EF89E" authority="László &amp; Hausmann &amp; Karisch, 2023" authorityName="László &amp; Hausmann &amp; Karisch" authorityYear="2023" box="[151,697,1794,1818]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="44" phylum="Arthropoda" rank="subSpecies" species="longicornuta" status="subsp. nov." subGenus="Herbuloracotis" subSpecies="ugandana">
<emphasis id="B936292F5D34FFB557F55006FD1EF89E" box="[151,697,1794,1818]" italics="true" pageId="9" pageNumber="10">
Afroracotis (Herbuloracotis)
<emphasis id="B936292F5D34FFB556A05006FD1EF89E" bold="true" box="[450,697,1794,1818]" pageId="9" pageNumber="10">longicornuta ugandana</emphasis>
</emphasis>
</taxonomicName>
ssp. n.
</paragraph>
<paragraph id="8BFDF53D5D34FFB557F55025FD39F8BD" blockId="9.[151,922,1641,1910]" box="[151,670,1824,1849]" pageId="9" pageNumber="10">
<taxonomicName id="4C428EBE5D34FFB557F55025FDF9F8BC" authority="László &amp; Hausmann &amp; Karisch, 2023" authorityName="László &amp; Hausmann &amp; Karisch" authorityYear="2023" box="[151,606,1824,1849]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="aristophanousi" status="sp. nov." subGenus="Herbuloracotis">
<emphasis id="B936292F5D34FFB557F55025FDF9F8BC" box="[151,606,1824,1849]" italics="true" pageId="9" pageNumber="10">
Afroracotis (Herbuloracotis)
<emphasis id="B936292F5D34FFB556A05024FDF9F8BC" bold="true" box="[450,606,1824,1848]" pageId="9" pageNumber="10">aristophanousi</emphasis>
</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20594545D34FFB555075025FD39F8BD" box="[613,670,1825,1849]" pageId="9" pageNumber="10" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BFDF53D5D34FFB557F55044FDD8F8DC" blockId="9.[151,922,1641,1910]" box="[151,639,1855,1880]" pageId="9" pageNumber="10">
<taxonomicName id="4C428EBE5D34FFB557F55044FD98F8D3" authority="László &amp; Hausmann &amp; Karisch, 2023" authorityName="László &amp; Hausmann &amp; Karisch" authorityYear="2023" box="[151,575,1855,1880]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="muscivirens" status="sp. nov." subGenus="Herbuloracotis">
<emphasis id="B936292F5D34FFB557F55044FD98F8D3" box="[151,575,1855,1880]" italics="true" pageId="9" pageNumber="10">
Afroracotis (Herbuloracotis)
<emphasis id="B936292F5D34FFB556A0503BFD98F8D3" bold="true" box="[450,575,1855,1879]" pageId="9" pageNumber="10">muscivirens</emphasis>
</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20594545D34FFB555245044FDD8F8DC" box="[582,639,1856,1880]" pageId="9" pageNumber="10" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BFDF53D5D34FFB557F5505AFCAAF8F2" blockId="9.[151,922,1641,1910]" box="[151,781,1886,1910]" pageId="9" pageNumber="10">
<taxonomicName id="4C428EBE5D34FFB557F5505AFD0CF8F2" authority="(Prout, 1926)" authorityName="László &amp; Hausmann &amp; Karisch" authorityYear="2023" baseAuthorityName="Prout" baseAuthorityYear="1926" box="[151,683,1886,1910]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="atriclava" status="comb. nov." subGenus="Herbuloracotis">
<emphasis id="B936292F5D34FFB557F5505AFDBAF8F2" box="[151,541,1886,1910]" italics="true" pageId="9" pageNumber="10">Afroracotis (Herbuloracotis) atriclava</emphasis>
(
<bibRefCitation id="EFD388CC5D34FFB55549505AFD03F8F2" author="Prout, L. B." box="[555,676,1886,1910]" pageId="9" pageNumber="10" pagination="179 - 188" refId="ref84180" refString="Prout, L. B. (1926) New Geometridae in the Tring Museum. Novitates Zoologicae, 33, 179 - 188." type="journal article" year="1926">Prout, 1926</bibRefCitation>
)
</taxonomicName>
<emphasis id="B936292F5D34FFB555D3505AFCAAF8F2" bold="true" box="[689,781,1886,1910]" pageId="9" pageNumber="10">
<taxonomicNameLabel id="A20594545D34FFB555D3505AFCAAF8F2" box="[689,781,1886,1910]" pageId="9" pageNumber="10" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BFDF53D5D37FFB657F55793FE3BFF35" blockId="10.[151,412,151,178]" box="[151,412,151,178]" pageId="10" pageNumber="11">
<heading id="D0B542515D37FFB657F55793FE3BFF35" bold="true" box="[151,412,151,178]" fontSize="11" level="2" pageId="10" pageNumber="11" reason="4">
<emphasis id="B936292F5D37FFB657F55793FE3BFF35" bold="true" box="[151,412,151,178]" pageId="10" pageNumber="11">
<taxonomicName id="4C428EBE5D37FFB657F55793FEB1FF36" authority="László &amp; Hausmann &amp; Karisch, 2023" authorityName="László &amp; Hausmann &amp; Karisch" authorityYear="2023" box="[151,278,151,178]" pageId="10" pageNumber="11" rank="subGenus" status="subgen. nov." subGenus="Zebracotis">
<emphasis id="B936292F5D37FFB657F55793FEB1FF36" bold="true" box="[151,278,151,178]" italics="true" pageId="10" pageNumber="11">Zebracotis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20594545D37FFB6567C5793FE3BFF35" box="[286,412,151,177]" pageId="10" pageNumber="11" rank="subGenus">subgen. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BFDF53D5D37FFB657F557D8FD44FF70" blockId="10.[151,967,220,520]" box="[151,739,220,244]" pageId="10" pageNumber="11">
<taxonomicName id="4C428EBE5D37FFB657F557D8FD27FF70" authority="(Warren, 1899)" authorityName="László &amp; Hausmann &amp; Karisch" authorityYear="2023" baseAuthorityName="Warren" baseAuthorityYear="1899" box="[151,640,220,244]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="zebrina" status="comb. nov." subGenus="Zebracotis">
<emphasis id="B936292F5D37FFB657F557D8FE47FF70" box="[151,480,220,244]" italics="true" pageId="10" pageNumber="11">Afroracotis (Zebracotis) zebrina</emphasis>
(
<bibRefCitation id="EFD388CC5D37FFB6568F57D8FDDEFF70" author="Warren, W." box="[493,633,220,244]" pageId="10" pageNumber="11" pagination="1 - 66" refId="ref85194" refString="Warren, W. (1899) New species and genera of the families Drepanulidae, Thyrididae, Uraniidae, Epiplemidae, and Geometridae from the Old-World regions. Novitates Zoologicae, 6 (1), 1 - 66." type="journal article" year="1899">Warren, 1899</bibRefCitation>
)
</taxonomicName>
<emphasis id="B936292F5D37FFB655E457D8FD44FF70" bold="true" box="[646,739,220,244]" pageId="10" pageNumber="11">
<taxonomicNameLabel id="A20594545D37FFB655E457D8FD44FF70" box="[646,739,220,244]" pageId="10" pageNumber="11" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BFDF53D5D37FFB657F557FEFD8BFE96" blockId="10.[151,967,220,520]" box="[151,556,250,274]" pageId="10" pageNumber="11">
<taxonomicName id="4C428EBE5D37FFB657F557FEFE4AFE96" authority="László &amp; Hausmann &amp; Karisch, 2023" authorityName="László &amp; Hausmann &amp; Karisch" authorityYear="2023" box="[151,493,250,274]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="chaineyi" status="sp. nov." subGenus="Zebracotis">
<emphasis id="B936292F5D37FFB657F557FEFE4AFE96" box="[151,493,250,274]" italics="true" pageId="10" pageNumber="11">
Afroracotis (Zebracotis)
<emphasis id="B936292F5D37FFB656F657FEFE4AFE96" bold="true" box="[404,493,250,274]" pageId="10" pageNumber="11">chaineyi</emphasis>
</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20594545D37FFB6569157FEFD8BFE96" box="[499,556,250,274]" pageId="10" pageNumber="11" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BFDF53D5D37FFB657F5561DFC60FEB5" blockId="10.[151,967,220,520]" box="[151,967,281,305]" pageId="10" pageNumber="11">
<taxonomicName id="4C428EBE5D37FFB657F5561DFCC2FEB5" authority="(Chainey &amp; Karisch, 2017)" authorityName="László &amp; Hausmann &amp; Karisch" authorityYear="2023" baseAuthorityName="Chainey &amp; Karisch" baseAuthorityYear="2017" box="[151,869,281,305]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="madagascariensis" status="comb. nov." subGenus="Zebracotis">
<emphasis id="B936292F5D37FFB657F5561DFDECFEB5" box="[151,587,281,305]" italics="true" pageId="10" pageNumber="11">Afroracotis (Zebracotis) madagascariensis</emphasis>
(
<bibRefCitation id="EFD388CC5D37FFB6553A561DFCF9FEB5" author="Chainey, J. &amp; Karisch, T." box="[600,862,281,305]" pageId="10" pageNumber="11" pagination="49 - 53" refId="ref83017" refString="Chainey, J. &amp; Karisch, T. (2017) A new species of Racotis Moore, [1887] (Lepidoptera: Geometridae: Ennominae) from Madagascar. Metamorphosis, 28, 49 - 53." type="journal article" year="2017">Chainey &amp; Karisch, 2017</bibRefCitation>
)
</taxonomicName>
<emphasis id="B936292F5D37FFB65409561DFC60FEB5" bold="true" box="[875,967,281,305]" pageId="10" pageNumber="11">
<taxonomicNameLabel id="A20594545D37FFB65409561DFC60FEB5" box="[875,967,281,305]" pageId="10" pageNumber="11" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BFDF53D5D37FFB657F5563CFDF2FED4" blockId="10.[151,967,220,520]" box="[151,597,311,336]" pageId="10" pageNumber="11">
<taxonomicName id="4C428EBE5D37FFB657F5563CFDB1FECB" authority="László &amp; Hausmann &amp; Karisch, 2023" authorityName="László &amp; Hausmann &amp; Karisch" authorityYear="2023" box="[151,534,311,336]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="59" phylum="Arthropoda" rank="subSpecies" species="lydiae" status="sp. nov." subGenus="Zebracotis" subSpecies="lydiae">
<emphasis id="B936292F5D37FFB657F5563CFDB1FECB" box="[151,534,311,336]" italics="true" pageId="10" pageNumber="11">
Afroracotis (Zebracotis)
<emphasis id="B936292F5D37FFB656F65633FDB1FECB" bold="true" box="[404,534,311,335]" pageId="10" pageNumber="11">lydiae lydiae</emphasis>
</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20594545D37FFB6557E563CFDF2FED4" box="[540,597,312,336]" pageId="10" pageNumber="11" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BFDF53D5D37FFB657F55652FD23FEEA" blockId="10.[151,967,220,520]" box="[151,644,342,366]" pageId="10" pageNumber="11">
<taxonomicName id="4C428EBE5D37FFB657F55652FD9CFEEA" authority="László &amp; Hausmann &amp; Karisch, 2023" authorityName="László &amp; Hausmann &amp; Karisch" authorityYear="2023" box="[151,571,342,366]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="62" phylum="Arthropoda" rank="subSpecies" species="lydiae" status="subsp. nov." subGenus="Zebracotis" subSpecies="orientalis">
<emphasis id="B936292F5D37FFB657F55652FD9CFEEA" box="[151,571,342,366]" italics="true" pageId="10" pageNumber="11">
Afroracotis (Zebracotis)
<emphasis id="B936292F5D37FFB656F65652FD9CFEEA" bold="true" box="[404,571,342,366]" pageId="10" pageNumber="11">lydiae orientalis</emphasis>
</emphasis>
</taxonomicName>
ssp. n.
</paragraph>
<paragraph id="8BFDF53D5D37FFB657F55671FDB3FE09" blockId="10.[151,967,220,520]" box="[151,532,372,397]" pageId="10" pageNumber="11">
<taxonomicName id="4C428EBE5D37FFB657F55671FE72FE08" authority="László &amp; Hausmann &amp; Karisch, 2023" authorityName="László &amp; Hausmann &amp; Karisch" authorityYear="2023" box="[151,469,372,397]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="63" phylum="Arthropoda" rank="species" species="smithi" status="sp. nov." subGenus="Zebracotis">
<emphasis id="B936292F5D37FFB657F55671FE72FE08" box="[151,469,372,397]" italics="true" pageId="10" pageNumber="11">
Afroracotis (Zebracotis)
<emphasis id="B936292F5D37FFB656F65670FE72FE08" bold="true" box="[404,469,372,396]" pageId="10" pageNumber="11">smithi</emphasis>
</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20594545D37FFB656B95671FDB3FE09" box="[475,532,373,397]" pageId="10" pageNumber="11" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BFDF53D5D37FFB657F55690FDB6FE28" blockId="10.[151,967,220,520]" box="[151,529,403,428]" pageId="10" pageNumber="11">
<taxonomicName id="4C428EBE5D37FFB657F55690FE75FE2F" authority="László &amp; Hausmann &amp; Karisch, 2023" authorityName="László &amp; Hausmann &amp; Karisch" authorityYear="2023" box="[151,466,403,428]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="milesi" status="sp. nov." subGenus="Zebracotis">
<emphasis id="B936292F5D37FFB657F55690FE75FE2F" box="[151,466,403,428]" italics="true" pageId="10" pageNumber="11">
Afroracotis (Zebracotis)
<emphasis id="B936292F5D37FFB656F65697FE75FE2F" bold="true" box="[404,466,403,427]" pageId="10" pageNumber="11">milesi</emphasis>
</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20594545D37FFB656BA5690FDB6FE28" box="[472,529,404,428]" pageId="10" pageNumber="11" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BFDF53D5D37FFB657F556B6FDB2FE4E" blockId="10.[151,967,220,520]" box="[151,533,434,458]" pageId="10" pageNumber="11">
<taxonomicName id="4C428EBE5D37FFB657F556B6FE71FE4E" authority="László &amp; Hausmann &amp; Karisch, 2023" authorityName="László &amp; Hausmann &amp; Karisch" authorityYear="2023" box="[151,470,434,458]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="ochsei" status="sp. nov." subGenus="Zebracotis">
<emphasis id="B936292F5D37FFB657F556B6FE71FE4E" box="[151,470,434,458]" italics="true" pageId="10" pageNumber="11">
Afroracotis (Zebracotis)
<emphasis id="B936292F5D37FFB656F656B6FE71FE4E" bold="true" box="[404,470,434,458]" pageId="10" pageNumber="11">ochsei</emphasis>
</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20594545D37FFB656BE56B6FDB2FE4E" box="[476,533,434,458]" pageId="10" pageNumber="11" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BFDF53D5D37FFB657F556D5FD8DFE6D" blockId="10.[151,967,220,520]" box="[151,554,464,489]" pageId="10" pageNumber="11">
<taxonomicName id="4C428EBE5D37FFB657F556D5FE4CFE6C" authority="László &amp; Hausmann &amp; Karisch, 2023" authorityName="László &amp; Hausmann &amp; Karisch" authorityYear="2023" box="[151,491,464,489]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="helicalis" status="sp. nov." subGenus="Zebracotis">
<emphasis id="B936292F5D37FFB657F556D5FE4CFE6C" box="[151,491,464,489]" italics="true" pageId="10" pageNumber="11">
Afroracotis (Zebracotis)
<emphasis id="B936292F5D37FFB656F656D4FE4CFE6C" bold="true" box="[404,491,464,488]" pageId="10" pageNumber="11">helicalis</emphasis>
</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20594545D37FFB6569356D5FD8DFE6D" box="[497,554,465,489]" pageId="10" pageNumber="11" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BFDF53D5D37FFB657F556F4FD85FD8C" blockId="10.[151,967,220,520]" box="[151,546,495,520]" pageId="10" pageNumber="11">
<taxonomicName id="4C428EBE5D37FFB657F556F4FE44FD83" authority="László &amp; Hausmann &amp; Karisch, 2023" authorityName="László &amp; Hausmann &amp; Karisch" authorityYear="2023" box="[151,483,495,520]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="69" phylum="Arthropoda" rank="species" species="takanoi" status="sp. nov." subGenus="Zebracotis">
<emphasis id="B936292F5D37FFB657F556F4FE44FD83" box="[151,483,495,520]" italics="true" pageId="10" pageNumber="11">
Afroracotis (Zebracotis)
<emphasis id="B936292F5D37FFB656F656EBFE44FD83" bold="true" box="[404,483,495,519]" pageId="10" pageNumber="11">takanoi</emphasis>
</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20594545D37FFB6568B56F4FD85FD8C" box="[489,546,496,520]" pageId="10" pageNumber="11" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BFDF53D5D37FFB657F5555DFE60FDF0" blockId="10.[151,455,601,628]" box="[151,455,601,628]" pageId="10" pageNumber="11">
<heading id="D0B542515D37FFB657F5555DFE60FDF0" bold="true" box="[151,455,601,628]" fontSize="11" level="2" pageId="10" pageNumber="11" reason="4">
<emphasis id="B936292F5D37FFB657F5555DFE60FDF0" bold="true" box="[151,455,601,628]" pageId="10" pageNumber="11">
<taxonomicName id="4C428EBE5D37FFB657F5555DFEE7FDF0" authority="László &amp; Hausmann &amp; Karisch, 2023" authorityName="László &amp; Hausmann &amp; Karisch" authorityYear="2023" box="[151,320,601,628]" pageId="10" pageNumber="11" rank="subGenus" status="subgen. nov." subGenus="Sokokeracotis">
<emphasis id="B936292F5D37FFB657F5555DFEE7FDF0" bold="true" box="[151,320,601,628]" italics="true" pageId="10" pageNumber="11">Sokokeracotis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20594545D37FFB6562A555EFE60FDF0" box="[328,455,602,628]" pageId="10" pageNumber="11" rank="subGenus">subgen. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BFDF53D5D37FFB657F5559AFD98FD32" blockId="10.[151,575,670,694]" box="[151,575,670,694]" pageId="10" pageNumber="11">
<taxonomicName id="4C428EBE5D37FFB657F5559AFDA7FD32" authority="László &amp; Hausmann &amp; Karisch, 2023" authorityName="László &amp; Hausmann &amp; Karisch" authorityYear="2023" box="[151,512,670,694]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="staudei" status="sp. nov." subGenus="Sokokeracotis">
<emphasis id="B936292F5D37FFB657F5559AFDA7FD32" box="[151,512,670,694]" italics="true" pageId="10" pageNumber="11">
Afroracotis (Sokokeracotis)
<emphasis id="B936292F5D37FFB656D7559AFDA7FD32" bold="true" box="[437,512,670,694]" pageId="10" pageNumber="11">staudei</emphasis>
</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20594545D37FFB65564559AFD98FD32" box="[518,575,670,694]" pageId="10" pageNumber="11" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BFDF53D5D37FFB657F5540DFB22FCA7" blockId="10.[151,1157,776,803]" box="[151,1157,776,803]" pageId="10" pageNumber="11">
<emphasis id="B936292F5D37FFB657F5540DFB22FCA7" bold="true" box="[151,1157,776,803]" pageId="10" pageNumber="11">
Key to the subgenera of the genus
<taxonomicName id="4C428EBE5D37FFB65527540CFD69FCA6" authorityName="László &amp; Hausmann &amp; Karisch" authorityYear="2023" box="[581,718,776,802]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B936292F5D37FFB65527540CFD69FCA6" bold="true" box="[581,718,776,802]" italics="true" pageId="10" pageNumber="11">Afroracotis</emphasis>
</taxonomicName>
based on male genitalia characters
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C358A6B65D37FFB657F5544AFA3AFBB4" pageId="10" pageNumber="11" type="key">
<key id="FFD3A20C5D37FFB657F5544AFA3AFBB4" pageId="10" pageNumber="11">
<keyStep id="30B6ED985D37FFB657F5544AFA3BFC04" pageId="10" pageNumber="11">
<paragraph id="8BFDF53D5D37FFB657F5544AFA3BFCE7" blockId="10.[151,1437,845,1073]" box="[151,1436,845,867]" pageId="10" pageNumber="11">
<keyLead id="30B356085D37FFB657F5544AFA3BFCE7" box="[151,1436,845,867]" pageId="10" pageNumber="11">1. Uncus short, bifid.................................................................................... 2</keyLead>
</paragraph>
<paragraph id="8BFDF53D5D37FFB657F5546FFA3BFC04" blockId="10.[151,1437,845,1073]" box="[151,1436,875,897]" pageId="10" pageNumber="11">
<keyLead id="30B356085D37FFB657F5546FFA3BFC04" box="[151,1436,875,897]" pageId="10" pageNumber="11">-. Uncus moderately long, simple, pointed................................................................... 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B6ED985D37FFB657F5548CFA3AFC3E" pageId="10" pageNumber="11">
<paragraph id="8BFDF53D5D37FFB657F5548CFA3BFC19" blockId="10.[151,1437,845,1073]" box="[151,1436,904,926]" pageId="10" pageNumber="11">
<keyLead id="30B356085D37FFB657F5548CFA3BFC19" box="[151,1436,904,926]" pageId="10" pageNumber="11">
2. Valva tip straight, valva without medial crest.......................................................
<taxonomicName id="4C428EBE5D37FFB65252548CFA3BFC19" authorityName="László &amp; Hausmann &amp; Karisch" authorityYear="2023" box="[1328,1436,904,925]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B936292F5D37FFB65252548CFA3BFC19" box="[1328,1436,904,925]" italics="true" pageId="10" pageNumber="11">Afroracotis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BFDF53D5D37FFB657F554A2FA3AFC3E" blockId="10.[151,1437,845,1073]" box="[151,1437,933,955]" pageId="10" pageNumber="11">
<keyLead id="30B356085D37FFB657F554A2FA3AFC3E" box="[151,1437,933,955]" pageId="10" pageNumber="11">
-. Valva tip curved medio-caudad, valva with medial crest.............................................
<taxonomicName id="4C428EBE5D37FFB6527454A1FA3AFC3E" box="[1302,1437,933,954]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="104" phylum="Arthropoda" rank="subGenus" subGenus="Sokokeracotis">
<emphasis id="B936292F5D37FFB6527454A1FA3AFC3E" box="[1302,1437,933,954]" italics="true" pageId="10" pageNumber="11">Sokokeracotis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B6ED985D37FFB657F554C7FA3AFC71" pageId="10" pageNumber="11">
<paragraph id="8BFDF53D5D37FFB657F554C7FA3BFC5C" blockId="10.[151,1437,845,1073]" box="[151,1436,963,985]" pageId="10" pageNumber="11">
<keyLead id="30B356085D37FFB657F554C7FA3BFC5C" box="[151,1436,963,985]" pageId="10" pageNumber="11">
3. Vesica with a single, usually large (reduced in
<taxonomicName id="4C428EBE5D37FFB655C354C7FCADFC5C" authorityName="László &amp; Hausmann &amp; Karisch" authorityYear="2023" baseAuthorityName="Prout" baseAuthorityYear="1931" box="[673,778,963,984]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="apodosima">
<emphasis id="B936292F5D37FFB655C354C7FCADFC5C" box="[673,778,963,984]" italics="true" pageId="10" pageNumber="11">apodosima</emphasis>
</taxonomicName>
group), acute distal cornutus.............................. 4
</keyLead>
</paragraph>
<paragraph id="8BFDF53D5D37FFB657F554E5FA3AFC71" blockId="10.[151,1437,845,1073]" box="[151,1437,992,1014]" pageId="10" pageNumber="11">
<keyLead id="30B356085D37FFB657F554E5FA3AFC71" box="[151,1437,992,1014]" pageId="10" pageNumber="11">
-. Vesica with a diverticular cornutus, or a distal cluster of cornuti or both...................................
<taxonomicName id="4C428EBE5D37FFB6525454E4FA3AFC71" box="[1334,1437,992,1013]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="78" phylum="Arthropoda" rank="subGenus" subGenus="Zebracotis">
<emphasis id="B936292F5D37FFB6525454E4FA3AFC71" box="[1334,1437,992,1013]" italics="true" pageId="10" pageNumber="11">Zebracotis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B6ED985D37FFB657F554FAFA3AFBB4" pageId="10" pageNumber="11">
<paragraph id="8BFDF53D5D37FFB657F554FAFA3BFB96" blockId="10.[151,1437,845,1073]" box="[151,1436,1021,1043]" pageId="10" pageNumber="11">
<keyLead id="30B356085D37FFB657F554FAFA3BFB96" box="[151,1436,1021,1043]" pageId="10" pageNumber="11">
4. Vesica extremely long and thin................................................................
<taxonomicName id="4C428EBE5D37FFB6526954F9FA3BFB96" box="[1291,1436,1021,1042]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="50" phylum="Arthropoda" rank="subGenus" subGenus="Rwandaracotis">
<emphasis id="B936292F5D37FFB6526954F9FA3BFB96" box="[1291,1436,1021,1042]" italics="true" pageId="10" pageNumber="11">Rwandaracotis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BFDF53D5D37FFB657F5531FFA3AFBB4" blockId="10.[151,1437,845,1073]" box="[151,1437,1051,1073]" pageId="10" pageNumber="11">
<keyLead id="30B356085D37FFB657F5531FFA3AFBB4" box="[151,1437,1051,1073]" pageId="10" pageNumber="11">
-. Vesica short and thick.......................................................................
<taxonomicName id="4C428EBE5D37FFB65268531FFA3AFBB4" box="[1290,1437,1051,1072]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="105" phylum="Arthropoda" rank="subGenus" subGenus="Herbuloracotis">
<emphasis id="B936292F5D37FFB65268531FFA3AFBB4" box="[1290,1437,1051,1072]" italics="true" pageId="10" pageNumber="11">Herbuloracotis</emphasis>
</taxonomicName>
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</paragraph>
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</key>
</subSubSection>
</treatment>
</document>