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<mods:title id="E28AAB6D20F6030EFD7D9168FF4AD790">A new species of the southern Central American genus Linoderus Sharp (Coleoptera: Staphylinidae: Staphylininae) with a revised key and an updated phylogeny</mods:title>
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<mods:namePart id="BB46156791E7AC97D8EC8256B4EAD82C">Chani-Posse, Mariana Raquel</mods:namePart>
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<mods:affiliation id="4E6DB20A30B32848D8A2D417FDEFD3A3">Laboratorio de Entomología, Instituto Argentino de Investigaciones de las Zonas Aridas (IADIZA, CCT CONICET, Mendoza), Casilla de Correo 507, 5500 Mendoza, Argentina &amp; Facultad de Ciencias Exactas y Naturales, Universidad Nacional de Cuyo, Mendoza, Argentina mchani @ mendoza-conicet. gob. ar; https: // orcid. org / 0000 - 0002 - 3466 - 7669</mods:affiliation>
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<treatment id="03E6D042FFF2310A35B6F361BE90F891" ID-DOI="http://doi.org/10.5281/zenodo.5636644" ID-GBIF-Taxon="189798538" ID-Zenodo-Dep="5636644" LSID="urn:lsid:plazi:treatment:03E6D042FFF2310A35B6F361BE90F891" httpUri="http://treatment.plazi.org/id/03E6D042FFF2310A35B6F361BE90F891" lastPageId="2" lastPageNumber="441" pageId="1" pageNumber="440">
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<paragraph id="8BF06154FFF2310935B6F361BE9EF8D9" blockId="1.[151,953,1843,1941]" box="[151,465,1843,1870]" pageId="1" pageNumber="440">
<heading id="D0B8D638FFF2310935B6F361BE9EF8D9" bold="true" box="[151,465,1843,1870]" fontSize="11" level="1" pageId="1" pageNumber="440" reason="1">
<emphasis id="B93BBD46FFF2310935B6F361BE9EF8D9" bold="true" box="[151,465,1843,1870]" pageId="1" pageNumber="440">
<taxonomicName id="4C4F1AD7FFF2310935B6F361BEC8F8D9" ID-CoL="9GCR7" authority="Chani-Posse, 2021" authorityName="Chani-Posse" authorityYear="2021" box="[151,391,1843,1870]" class="Insecta" family="Staphylinidae" genus="Linoderus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="440" phylum="Arthropoda" rank="species" species="smetanai" status="sp. nov.">
<emphasis id="B93BBD46FFF2310935B6F361BEC8F8D9" bold="true" box="[151,391,1843,1870]" italics="true" pageId="1" pageNumber="440">Linoderus smetanai</emphasis>
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<taxonomicNameLabel id="A208003DFFF2310934AEF361BE9EF8D9" box="[399,465,1843,1869]" pageId="1" pageNumber="440" rank="species">sp. n.</taxonomicNameLabel>
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</paragraph>
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<subSubSection id="C35532DFFFF2310935B6F329BE6FF801" box="[151,288,1915,1941]" pageId="1" pageNumber="440" type="description">
<paragraph id="8BF06154FFF2310935B6F329BE6FF801" blockId="1.[151,953,1843,1941]" box="[151,288,1915,1941]" pageId="1" pageNumber="440">
(
<figureCitation id="13747DD1FFF2310935BEF329BE57F801" box="[159,280,1915,1941]" captionStart="FIGURES 112" captionStartId="4.[151,264,1864,1888]" captionTargetBox="[151,1413,179,1837]" captionTargetId="figure-23@4.[151,1436,179,1838]" captionTargetPageId="4" captionText="FIGURES 112. Habitus of Linoderus smetanai sp. n.: (1) male holotype and (2) female paratype. Linoderus smetanai, male and female genitalia: (3) male sternite 8; (4) sternite 9; (5) male tergite 10; (6) aedeagus in ventral view; (7) aedeagus in lateral view; (8) male lateral tergal sclerite 9; (9) female tergite 10; (10) female sternite 8; (11) gonocoxites of female genital segment; (12) female lateral tergal sclerites 9." figureDoi="http://doi.org/10.5281/zenodo.5635986" httpUri="https://zenodo.org/record/5635986/files/figure.png" pageId="1" pageNumber="440">Figs. 112</figureCitation>
)
</paragraph>
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<subSubSection id="C35532DFFFF2310A35B6F391BEC9FE61" lastPageId="2" lastPageNumber="441" pageId="1" pageNumber="440" type="diagnosis">
<paragraph id="8BF06154FFF2310A35B6F391BE8CFE1D" blockId="1.[151,1436,1987,2014]" lastBlockId="2.[151,1436,151,393]" lastPageId="2" lastPageNumber="441" pageId="1" pageNumber="440">
<emphasis id="B93BBD46FFF2310935B6F391BE5DF849" bold="true" box="[151,274,1987,2013]" pageId="1" pageNumber="440">Diagnosis.</emphasis>
<taxonomicName id="4C4F1AD7FFF23109343AF391BEB4F849" authorityName="Chani-Posse" authorityYear="2021" box="[283,507,1987,2013]" class="Insecta" family="Staphylinidae" genus="Linoderus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="440" phylum="Arthropoda" rank="species" species="smetanai">
<emphasis id="B93BBD46FFF23109343AF391BEB4F849" box="[283,507,1987,2013]" italics="true" pageId="1" pageNumber="440">Linoderus smetanai</emphasis>
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differs from
<taxonomicName id="4C4F1AD7FFF2310937B7F396BC73F849" authorityName="Chani-Posse &amp; Solodovnikov" authorityYear="2017" box="[662,828,1987,2013]" class="Insecta" family="Staphylinidae" genus="Linoderus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="440" phylum="Arthropoda" rank="species" species="alajuelensis">
<emphasis id="B93BBD46FFF2310937B7F396BC73F849" box="[662,828,1987,2013]" italics="true" pageId="1" pageNumber="440">L. alajuelensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4F1AD7FFF231093656F396BB48F849" box="[887,1031,1987,2013]" class="Insecta" family="Staphylinidae" genus="Linoderus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="440" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="B93BBD46FFF231093656F396BB48F849" box="[887,1031,1987,2013]" italics="true" pageId="1" pageNumber="440">L. gracilipes</emphasis>
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in the distinct colour pattern: head and prothorax dark metallic blue, elytra metallic green, abdomen with first five visible abdominal segments reddish brown and apical abdominal segments dark brown to black; antennae and palpi mostly reddish brown to brown, with legs and styli dark brown to black as well as the first antennomere moderately longer than half of head length*, labial palpus with second palpomere about as long as third*, the first metatarsomere not sexually dimorphic in length*; the abdominal tergites 35 with posterior basal transverse carina (PBTC) complete and straight, space behind PBTC not distinctly punctate* and the sternite 9 with several apical setae at each side of emargination are also unique to this species.
</paragraph>
<paragraph id="8BF06154FFF1310A35B6F5E5BEC9FE61" blockId="2.[151,1436,439,501]" pageId="2" pageNumber="441">
* Not shown as characters with interspecific variation in the redescription of the genus by
<bibRefCitation id="EFDE1CA5FFF1310A31CCF5E5BEC9FE61" author="Chani-Posse, M. &amp; Solodovnikov, A." pageId="2" pageNumber="441" pagination="83 - 97" refId="ref4307" refString="Chani-Posse, M. &amp; Solodovnikov, A. (2017) Tackling generic limits for the Neotropical Philonthina with a phylogenetic approach: revision of the genera Linoderus Sharp and Odontolinus Sharp (Coleoptera: Staphylinidae). Arthropod Systematics &amp; Phylogeny, 75 (1), 83 - 97." type="journal article" year="2017">Chani-Posse &amp; Solodovnikov (2017)</bibRefCitation>
</paragraph>
</subSubSection>
<subSubSection id="C35532DFFFF1310A35E6F670BC4BFAA5" pageId="2" pageNumber="441" type="description">
<paragraph id="8BF06154FFF1310A35E6F670BD81FB7D" blockId="2.[151,1437,546,1797]" pageId="2" pageNumber="441">
<emphasis id="B93BBD46FFF1310A35E6F670BE16FDA8" bold="true" box="[199,345,546,572]" pageId="2" pageNumber="441">Description.</emphasis>
Body length
<quantity id="4CB7CCB1FFF1310A34DBF670BDC7FDA9" box="[506,648,546,573]" metricMagnitude="-3" metricUnit="m" metricValue="8.1" metricValueMax="8.5" metricValueMin="7.7" pageId="2" pageNumber="441" unit="mm" value="8.1" valueMax="8.5" valueMin="7.7">7.78.5 mm</quantity>
. Colouration: as in diagnosis (
<figureCitation id="13747DD1FFF1310A36CDF671BB10FDA9" box="[1004,1119,547,573]" captionStart="FIGURES 112" captionStartId="4.[151,264,1864,1888]" captionTargetBox="[151,1413,179,1837]" captionTargetId="figure-23@4.[151,1436,179,1838]" captionTargetPageId="4" captionText="FIGURES 112. Habitus of Linoderus smetanai sp. n.: (1) male holotype and (2) female paratype. Linoderus smetanai, male and female genitalia: (3) male sternite 8; (4) sternite 9; (5) male tergite 10; (6) aedeagus in ventral view; (7) aedeagus in lateral view; (8) male lateral tergal sclerite 9; (9) female tergite 10; (10) female sternite 8; (11) gonocoxites of female genital segment; (12) female lateral tergal sclerites 9." figureDoi="http://doi.org/10.5281/zenodo.5635986" httpUri="https://zenodo.org/record/5635986/files/figure.png" pageId="2" pageNumber="441">Figs. 1, 2</figureCitation>
). Head slightly longer than wide (HW/HL = 0.930.94), moderately wider than pronotum (HW/PW = 1.161.18). Eyes distinctly shorter than temples (EL/TL = 0.65; both sexes) seen from above (
<figureCitation id="13747DD1FFF1310A37DAF639BC26FD11" box="[763,873,619,645]" captionStart="FIGURES 112" captionStartId="4.[151,264,1864,1888]" captionTargetBox="[151,1413,179,1837]" captionTargetId="figure-23@4.[151,1436,179,1838]" captionTargetPageId="4" captionText="FIGURES 112. Habitus of Linoderus smetanai sp. n.: (1) male holotype and (2) female paratype. Linoderus smetanai, male and female genitalia: (3) male sternite 8; (4) sternite 9; (5) male tergite 10; (6) aedeagus in ventral view; (7) aedeagus in lateral view; (8) male lateral tergal sclerite 9; (9) female tergite 10; (10) female sternite 8; (11) gonocoxites of female genital segment; (12) female lateral tergal sclerites 9." figureDoi="http://doi.org/10.5281/zenodo.5635986" httpUri="https://zenodo.org/record/5635986/files/figure.png" pageId="2" pageNumber="441">Figs. 1, 2</figureCitation>
). Antennae with 1st antennomere moderately longer than half of head length (A1/HL = 0.66; both sexes), moderately shorter than 2nd and 3rd combined, 3rd about 1.5 × as long as 2nd. Labial palpus with 2nd palpomere about twice as long as 1st, about as long as 3rd. Maxillary palpus with 4th palpomere (apical) 1.8 × as long as 3rd. Neck about 0.4 × as wide as head at widest point. Pronotum distinctly longer than wide (PW/PL = 0.710.72), dorsal rows of punctures each with four punctures, 2nd and 3rd more distant to each other than 2nd to 1st and 3rd to 4th. Elytra at sides about 1.5 × as long as elytra along suture (EtL/Etl= 1.57; both sexes). First metatarsomere not sexually dimorphic in length, distinctly longer than 5th in both sexes (S1/S5 = 1.551.66). Abdominal tergites 35 with posterior basal transverse carina complete and straight, space behind not distinctly punctate. Male sexual characters: Sternite 8 with medially produced and subangulately emarginate projection (
<figureCitation id="13747DD1FFF1310A34BAF7FDBEAFFC5D" box="[411,480,943,969]" captionStart="FIGURES 112" captionStartId="4.[151,264,1864,1888]" captionTargetBox="[151,1413,179,1837]" captionTargetId="figure-23@4.[151,1436,179,1838]" captionTargetPageId="4" captionText="FIGURES 112. Habitus of Linoderus smetanai sp. n.: (1) male holotype and (2) female paratype. Linoderus smetanai, male and female genitalia: (3) male sternite 8; (4) sternite 9; (5) male tergite 10; (6) aedeagus in ventral view; (7) aedeagus in lateral view; (8) male lateral tergal sclerite 9; (9) female tergite 10; (10) female sternite 8; (11) gonocoxites of female genital segment; (12) female lateral tergal sclerites 9." figureDoi="http://doi.org/10.5281/zenodo.5635986" httpUri="https://zenodo.org/record/5635986/files/figure.png" pageId="2" pageNumber="441">Fig. 3</figureCitation>
). Sternite 9 with basal portion distinctly asymmetrical, 1.8 × as long as distal portion and deeply emarginate apically, with several apical setae at each side of emargination (
<figureCitation id="13747DD1FFF1310A31A5F781BB83FC79" box="[1156,1228,979,1005]" captionStart="FIGURES 112" captionStartId="4.[151,264,1864,1888]" captionTargetBox="[151,1413,179,1837]" captionTargetId="figure-23@4.[151,1436,179,1838]" captionTargetPageId="4" captionText="FIGURES 112. Habitus of Linoderus smetanai sp. n.: (1) male holotype and (2) female paratype. Linoderus smetanai, male and female genitalia: (3) male sternite 8; (4) sternite 9; (5) male tergite 10; (6) aedeagus in ventral view; (7) aedeagus in lateral view; (8) male lateral tergal sclerite 9; (9) female tergite 10; (10) female sternite 8; (11) gonocoxites of female genital segment; (12) female lateral tergal sclerites 9." figureDoi="http://doi.org/10.5281/zenodo.5635986" httpUri="https://zenodo.org/record/5635986/files/figure.png" pageId="2" pageNumber="441">Fig. 4</figureCitation>
). Genital segment with lateral tergal sclerites 9 (styli) dorsoventrally flattened and moderately dilated, with a distinct latero-apical emargination (
<figureCitation id="13747DD1FFF1310A341AF049BE30FBA1" box="[315,383,1051,1077]" captionStart="FIGURES 112" captionStartId="4.[151,264,1864,1888]" captionTargetBox="[151,1413,179,1837]" captionTargetId="figure-23@4.[151,1436,179,1838]" captionTargetPageId="4" captionText="FIGURES 112. Habitus of Linoderus smetanai sp. n.: (1) male holotype and (2) female paratype. Linoderus smetanai, male and female genitalia: (3) male sternite 8; (4) sternite 9; (5) male tergite 10; (6) aedeagus in ventral view; (7) aedeagus in lateral view; (8) male lateral tergal sclerite 9; (9) female tergite 10; (10) female sternite 8; (11) gonocoxites of female genital segment; (12) female lateral tergal sclerites 9." figureDoi="http://doi.org/10.5281/zenodo.5635986" httpUri="https://zenodo.org/record/5635986/files/figure.png" pageId="2" pageNumber="441">Fig. 8</figureCitation>
); tergite 10 subtruncate apically. Aedeagus with median lobe gradually narrowed from about distal half of paramere length and with acute apex; apex of paramere distinctly notched at middle (
<figureCitation id="13747DD1FFF1310A31D5F06DBA2EFBCD" box="[1268,1377,1087,1113]" captionStart="FIGURES 112" captionStartId="4.[151,264,1864,1888]" captionTargetBox="[151,1413,179,1837]" captionTargetId="figure-23@4.[151,1436,179,1838]" captionTargetPageId="4" captionText="FIGURES 112. Habitus of Linoderus smetanai sp. n.: (1) male holotype and (2) female paratype. Linoderus smetanai, male and female genitalia: (3) male sternite 8; (4) sternite 9; (5) male tergite 10; (6) aedeagus in ventral view; (7) aedeagus in lateral view; (8) male lateral tergal sclerite 9; (9) female tergite 10; (10) female sternite 8; (11) gonocoxites of female genital segment; (12) female lateral tergal sclerites 9." figureDoi="http://doi.org/10.5281/zenodo.5635986" httpUri="https://zenodo.org/record/5635986/files/figure.png" pageId="2" pageNumber="441">Figs. 6, 7</figureCitation>
). Female sexual characters: Sternite 8 with hind margin projected medially (
<figureCitation id="13747DD1FFF1310A3696F031BB46FBE9" box="[951,1033,1123,1149]" captionStart="FIGURES 112" captionStartId="4.[151,264,1864,1888]" captionTargetBox="[151,1413,179,1837]" captionTargetId="figure-23@4.[151,1436,179,1838]" captionTargetPageId="4" captionText="FIGURES 112. Habitus of Linoderus smetanai sp. n.: (1) male holotype and (2) female paratype. Linoderus smetanai, male and female genitalia: (3) male sternite 8; (4) sternite 9; (5) male tergite 10; (6) aedeagus in ventral view; (7) aedeagus in lateral view; (8) male lateral tergal sclerite 9; (9) female tergite 10; (10) female sternite 8; (11) gonocoxites of female genital segment; (12) female lateral tergal sclerites 9." figureDoi="http://doi.org/10.5281/zenodo.5635986" httpUri="https://zenodo.org/record/5635986/files/figure.png" pageId="2" pageNumber="441">Fig. 10</figureCitation>
). Genital segment with lateral tergal sclerites 9 (styli) dorsoventrally flattened and not distinctly dilated, with a moderate latero-apical emargination (
<figureCitation id="13747DD1FFF1310A304EF0D5BFF9FB51" captionStart="FIGURES 112" captionStartId="4.[151,264,1864,1888]" captionTargetBox="[151,1413,179,1837]" captionTargetId="figure-23@4.[151,1436,179,1838]" captionTargetPageId="4" captionText="FIGURES 112. Habitus of Linoderus smetanai sp. n.: (1) male holotype and (2) female paratype. Linoderus smetanai, male and female genitalia: (3) male sternite 8; (4) sternite 9; (5) male tergite 10; (6) aedeagus in ventral view; (7) aedeagus in lateral view; (8) male lateral tergal sclerite 9; (9) female tergite 10; (10) female sternite 8; (11) gonocoxites of female genital segment; (12) female lateral tergal sclerites 9." figureDoi="http://doi.org/10.5281/zenodo.5635986" httpUri="https://zenodo.org/record/5635986/files/figure.png" pageId="2" pageNumber="441">Fig. 12</figureCitation>
); tergum 10 subangulate to arcuate apically (
<figureCitation id="13747DD1FFF1310A3784F0F9BDA7FB51" box="[677,744,1195,1221]" captionStart="FIGURES 112" captionStartId="4.[151,264,1864,1888]" captionTargetBox="[151,1413,179,1837]" captionTargetId="figure-23@4.[151,1436,179,1838]" captionTargetPageId="4" captionText="FIGURES 112. Habitus of Linoderus smetanai sp. n.: (1) male holotype and (2) female paratype. Linoderus smetanai, male and female genitalia: (3) male sternite 8; (4) sternite 9; (5) male tergite 10; (6) aedeagus in ventral view; (7) aedeagus in lateral view; (8) male lateral tergal sclerite 9; (9) female tergite 10; (10) female sternite 8; (11) gonocoxites of female genital segment; (12) female lateral tergal sclerites 9." figureDoi="http://doi.org/10.5281/zenodo.5635986" httpUri="https://zenodo.org/record/5635986/files/figure.png" pageId="2" pageNumber="441">Fig. 9</figureCitation>
); second gonocoxites each with a long macroseta, with minute stylus (
<figureCitation id="13747DD1FFF1310A35C9F09DBE78FB7D" box="[232,311,1231,1257]" captionStart="FIGURES 112" captionStartId="4.[151,264,1864,1888]" captionTargetBox="[151,1413,179,1837]" captionTargetId="figure-23@4.[151,1436,179,1838]" captionTargetPageId="4" captionText="FIGURES 112. Habitus of Linoderus smetanai sp. n.: (1) male holotype and (2) female paratype. Linoderus smetanai, male and female genitalia: (3) male sternite 8; (4) sternite 9; (5) male tergite 10; (6) aedeagus in ventral view; (7) aedeagus in lateral view; (8) male lateral tergal sclerite 9; (9) female tergite 10; (10) female sternite 8; (11) gonocoxites of female genital segment; (12) female lateral tergal sclerites 9." figureDoi="http://doi.org/10.5281/zenodo.5635986" httpUri="https://zenodo.org/record/5635986/files/figure.png" pageId="2" pageNumber="441">Fig. 11</figureCitation>
) bearing one long apical macroseta.
</paragraph>
<paragraph id="8BF06154FFF1310A35E6F0A0BC4BFAA5" blockId="2.[151,1437,546,1797]" pageId="2" pageNumber="441">
<emphasis id="B93BBD46FFF1310A35E6F0A0BD48FA98" bold="true" box="[199,519,1266,1292]" pageId="2" pageNumber="441">Geographical distribution.</emphasis>
<taxonomicName id="4C4F1AD7FFF1310A3731F0A1BDBEFA98" authorityName="Chani-Posse" authorityYear="2021" box="[528,753,1267,1293]" class="Insecta" family="Staphylinidae" genus="Linoderus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="441" phylum="Arthropoda" rank="species" species="smetanai">
<emphasis id="B93BBD46FFF1310A3731F0A1BDBEFA98" box="[528,753,1267,1293]" italics="true" pageId="2" pageNumber="441">Linoderus smetanai</emphasis>
</taxonomicName>
has been recorded from four different localities in Western
<collectingCountry id="F35821C4FFF1310A35B6F145BE5EFAA5" box="[151,273,1303,1329]" name="Costa Rica" pageId="2" pageNumber="441">Costa Rica</collectingCountry>
within the
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range of elevation.
</paragraph>
</subSubSection>
<subSubSection id="C35532DFFFF1310A35E6F169BC61FAED" pageId="2" pageNumber="441" type="etymology">
<paragraph id="8BF06154FFF1310A35E6F169BC61FAED" blockId="2.[151,1437,546,1797]" pageId="2" pageNumber="441">
<emphasis id="B93BBD46FFF1310A35E6F169BE01FAC1" bold="true" box="[199,334,1339,1365]" pageId="2" pageNumber="441">Etymology.</emphasis>
The specific name is dedicated to Dr. Aleš Smetana in recognition of his pioneering work and great legacy in the modern study of the subfamily
<taxonomicName id="4C4F1AD7FFF1310A37ADF10DBC65FAED" authorityName="Latreille" authorityYear="1802" box="[652,810,1375,1401]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="441" phylum="Arthropoda" rank="subFamily" subFamily="Staphylininae">Staphylininae</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C35532DFFFF1310A35E6F1D1BFA6FA55" pageId="2" pageNumber="441" type="discussion">
<paragraph id="8BF06154FFF1310A35E6F1D1BFA6FA55" blockId="2.[151,1437,546,1797]" pageId="2" pageNumber="441">
<emphasis id="B93BBD46FFF1310A35E6F1D1BE74FA09" bold="true" box="[199,315,1411,1437]" pageId="2" pageNumber="441">Remarks.</emphasis>
The new species fully fits into the generic diagnosis provided by
<bibRefCitation id="EFDE1CA5FFF1310A3169F1D1BFAAFA55" author="Chani-Posse, M. &amp; Solodovnikov, A." pageId="2" pageNumber="441" pagination="83 - 97" refId="ref4307" refString="Chani-Posse, M. &amp; Solodovnikov, A. (2017) Tackling generic limits for the Neotropical Philonthina with a phylogenetic approach: revision of the genera Linoderus Sharp and Odontolinus Sharp (Coleoptera: Staphylinidae). Arthropod Systematics &amp; Phylogeny, 75 (1), 83 - 97." type="journal article" year="2017">Chani-Posse &amp; Solodovnikov (2017)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C35532DFFFF1310A35E6F199BE90F891" pageId="2" pageNumber="441" type="materials_examined">
<paragraph id="8BF06154FFF1310A35E6F199BE90F891" blockId="2.[151,1437,546,1797]" pageId="2" pageNumber="441">
<emphasis id="B93BBD46FFF1310A35E6F199BE20FA70" bold="true" box="[199,367,1482,1509]" pageId="2" pageNumber="441">Type material</emphasis>
.
<typeStatus id="54F4DFF6FFF1310A345EF199BEA8FA71" box="[383,487,1483,1509]" pageId="2" pageNumber="441" type="holotype">Holotype</typeStatus>
<specimenCount id="9D49AADDFFF1310A34CEF199BD4BFA71" box="[495,516,1483,1509]" pageId="2" pageNumber="441" type="male"></specimenCount>
(
<figureCitation id="13747DD1FFF1310A3734F199BD17FA71" box="[533,600,1483,1509]" captionStart="FIGURES 112" captionStartId="4.[151,264,1864,1888]" captionTargetBox="[151,1413,179,1837]" captionTargetId="figure-23@4.[151,1436,179,1838]" captionTargetPageId="4" captionText="FIGURES 112. Habitus of Linoderus smetanai sp. n.: (1) male holotype and (2) female paratype. Linoderus smetanai, male and female genitalia: (3) male sternite 8; (4) sternite 9; (5) male tergite 10; (6) aedeagus in ventral view; (7) aedeagus in lateral view; (8) male lateral tergal sclerite 9; (9) female tergite 10; (10) female sternite 8; (11) gonocoxites of female genital segment; (12) female lateral tergal sclerites 9." figureDoi="http://doi.org/10.5281/zenodo.5635986" httpUri="https://zenodo.org/record/5635986/files/figure.png" pageId="2" pageNumber="441">Fig. 1</figureCitation>
), with labels: [white label]´CR. Pnt.
<quantity id="4CB7CCB1FFF1310A3128F198BB18FA71" box="[1033,1111,1482,1509]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="441" unit="m" value="1500.0">1500m</quantity>
/ Reserva de / Monteverde /
<date id="FFF14794FFF1310A35B6F1BDBE66F99D" box="[151,297,1519,1545]" pageId="2" pageNumber="441" value="1979-06-01" valueMax="1979-06-04" valueMin="1979-06-01">1-4.VI. 1979</date>
/ JM&amp;BA Campbell´, [red label]´
<materialsCitation id="3B276B09FFF1310A37E1F1BDBD04F9C5" ID-GBIF-Occurrence="3396021303" collectingDate="1979-06-27" collectingDateMax="2001-06-17" collectingDateMin="1979-06-27" collectionCode="CNC, SEMC, FMNH" collectorName="JM&amp;BA Campbell" country="Costa Rica" elevation="1480" latitude="10.3450165" location="Monteverde" longLatPrecision="1" longitude="-84.79832" municipality="Reserva" pageId="2" pageNumber="441" specimenCount="2" specimenCount-female="1" stateProvince="Cartago" typeStatus="holotype">
<typeStatus id="54F4DFF6FFF1310A37E1F1BDBC64F99D" box="[704,811,1519,1545]" pageId="2" pageNumber="441" type="holotype">Holotype</typeStatus>
<taxonomicName id="4C4F1AD7FFF1310A3615F1BDBBA3F99D" authority="Chani-Posse 2021" authorityName="Chani-Posse" authorityYear="2021" box="[820,1260,1519,1545]" class="Insecta" family="Staphylinidae" genus="Linoderus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="441" phylum="Arthropoda" rank="species" species="smetanai">
<emphasis id="B93BBD46FFF1310A3615F1BDBB5BF99C" box="[820,1044,1519,1545]" italics="true" pageId="2" pageNumber="441">Linoderus smetanai</emphasis>
Chani-Posse 2021
</taxonomicName>
´(
<collectionCode id="ED5EF991FFF1310A3024F1BDBA0BF99D" box="[1285,1348,1519,1545]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="2" pageNumber="441">CNC</collectionCode>
). Six
<specimenCount id="9D49AADDFFF1310A30A6F1BDBAD3F99D" box="[1415,1436,1519,1545]" pageId="2" pageNumber="441" type="female"></specimenCount>
<typeStatus id="54F4DFF6FFF1310A35B6F241BE4DF9B9" box="[151,258,1555,1581]" pageId="2" pageNumber="441" type="paratype">paratypes</typeStatus>
with labels: [white label]´CR. Pnt. 5000 /
<collectingMunicipality id="6B94FB2EFFF1310A37D2F241BC02F9B9" box="[755,845,1555,1581]" pageId="2" pageNumber="441">Reserva</collectingMunicipality>
de /
<location id="8E90378FFFF1310A36A6F241BB41F9B9" LSID="urn:lsid:plazi:treatment:03E6D042FFF2310A35B6F361BE90F891:8E90378FFFF1310A36A6F241BB41F9B9" box="[903,1038,1555,1581]" country="Costa Rica" latitude="10.3450165" longLatPrecision="1" longitude="-84.79832" municipality="Reserva" name="Monteverde" pageId="2" pageNumber="441" stateProvince="Cartago">Monteverde</location>
/
<date id="FFF14794FFF1310A3104F241BB9DF9B9" box="[1061,1234,1555,1581]" pageId="2" pageNumber="441" value="1979-06-27" valueMax="1979-06-29" valueMin="1979-06-27">
<collectingDate id="EFB5BE7CFFF1310A3104F241BB9DF9B9" box="[1061,1234,1555,1581]" pageId="2" pageNumber="441" value="1979-06-27" valueMax="1979-06-29" valueMin="1979-06-27">27-29.VI. 1979</collectingDate>
</date>
/ JM&amp;BA Campbell´, 1 (
<collectionCode id="ED5EF991FFF1310A35D9F265BE7AF9C5" box="[248,309,1591,1617]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="2" pageNumber="441">CNC</collectionCode>
) (
<figureCitation id="13747DD1FFF1310A346FF265BEDBF9C5" box="[334,404,1591,1617]" captionStart="FIGURES 112" captionStartId="4.[151,264,1864,1888]" captionTargetBox="[151,1413,179,1837]" captionTargetId="figure-23@4.[151,1436,179,1838]" captionTargetPageId="4" captionText="FIGURES 112. Habitus of Linoderus smetanai sp. n.: (1) male holotype and (2) female paratype. Linoderus smetanai, male and female genitalia: (3) male sternite 8; (4) sternite 9; (5) male tergite 10; (6) aedeagus in ventral view; (7) aedeagus in lateral view; (8) male lateral tergal sclerite 9; (9) female tergite 10; (10) female sternite 8; (11) gonocoxites of female genital segment; (12) female lateral tergal sclerites 9." figureDoi="http://doi.org/10.5281/zenodo.5635986" httpUri="https://zenodo.org/record/5635986/files/figure.png" pageId="2" pageNumber="441">Fig. 2</figureCitation>
); [white label]´
</materialsCitation>
<materialsCitation id="3B276B09FFF1310A376BF265BFD1F90D" ID-GBIF-Occurrence="3396021302" collectingDate="1979-06-27" collectingDateMax="2001-06-17" collectingDateMin="1979-06-27" collectionCode="CNC, SEMC, FMNH" collectorName="J. Ashe &amp; R. Brooks &amp; Z. Falin" country="Costa Rica" elevation="1480" latitude="10.3450165" location="Monteverde" longLatPrecision="1" longitude="-84.79832" municipality="Reserva" pageId="2" pageNumber="441" specimenCount="2" specimenCount-female="1" stateProvince="Cartago" typeStatus="holotype">
<collectingCountry id="F35821C4FFF1310A376BF265BD82F9C5" box="[586,717,1591,1617]" name="Costa Rica" pageId="2" pageNumber="441">Costa Rica</collectingCountry>
:
<collectingRegion id="498BAFB6FFF1310A37F8F265BC7DF9C5" box="[729,818,1591,1617]" country="Costa Rica" name="Cartago" pageId="2" pageNumber="441">Cartago</collectingRegion>
<collectorName id="26BA0482FFF1310A361BF265BC8AF9C5" box="[826,965,1591,1617]" pageId="2" pageNumber="441">P.N. Tapanti</collectorName>
/
<date id="FFF14794FFF1310A36FCF265BBD8F9C5" box="[989,1175,1591,1617]" pageId="2" pageNumber="441" value="2000-07-17" valueMax="2000-07-20" valueMin="2000-07-17">
<collectingDate id="EFB5BE7CFFF1310A36FCF265BBD8F9C5" box="[989,1175,1591,1617]" pageId="2" pageNumber="441" value="2000-07-17" valueMax="2000-07-20" valueMin="2000-07-17">17-20 JUL 2000</collectingDate>
</date>
<collectorName id="26BA0482FFF1310A31BEF265BBBBF9C5" box="[1183,1268,1591,1617]" pageId="2" pageNumber="441">J. Ashe</collectorName>
,
<collectorName id="26BA0482FFF1310A3020F265BA37F9C5" box="[1281,1400,1591,1617]" pageId="2" pageNumber="441">R. Brooks</collectorName>
,
<collectorName id="26BA0482FFF1310A30A5F265BF9EF9E1" pageId="2" pageNumber="441">Z. Falin</collectorName>
/
<quantity id="4CB7CCB1FFF1310A35CBF209BE77F9E1" box="[234,312,1627,1653]" metricMagnitude="3" metricUnit="m" metricValue="1.48" pageId="2" pageNumber="441" unit="m" value="1480.0">
<elevation id="00628667FFF1310A35CBF209BE77F9E1" box="[234,312,1627,1653]" metricMagnitude="3" metricUnit="m" metricValue="1.48" pageId="2" pageNumber="441" unit="m" value="1480.0">1480m</elevation>
</quantity>
<geoCoordinate id="EE7B0793FFF1310A3460F209BE82F9E1" box="[321,461,1627,1653]" degrees="9" direction="north" minutes="43" orientation="latitude" pageId="2" pageNumber="441" precision="15" seconds="12" value="9.719999">9° 43´12´´N</geoCoordinate>
,
<geoCoordinate id="EE7B0793FFF1310A34FAF209BD36F9E1" box="[475,633,1627,1653]" degrees="83" direction="west" minutes="46" orientation="longitude" pageId="2" pageNumber="441" precision="15" seconds="36" value="-83.77667">83°46´36´´W</geoCoordinate>
/ ex.
<collectingMethod id="520E1943FFF1310A379CF209BCD0F9E1" box="[701,927,1627,1653]" pageId="2" pageNumber="441">Flight intercept trap</collectingMethod>
CR1ABFOO 194´, 2 (
<collectionCode id="ED5EF991FFF1310A3184F209BBB8F9E1" box="[1189,1271,1627,1653]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="2" pageNumber="441" type="Museum">SEMC</collectionCode>
); [white label]´
</materialsCitation>
<materialsCitation id="3B276B09FFF1310A35BFF22DBBE0F975" ID-GBIF-Occurrence="3396021301" collectingDate="1979-06-27" collectingDateMax="2001-06-17" collectingDateMin="1979-06-27" collectionCode="CNC, SEMC, FMNH" country="Costa Rica" elevation="1480" latitude="10.3450165" location="Monteverde" longLatPrecision="1" longitude="-84.79832" municipality="Reserva" pageId="2" pageNumber="441" specimenCount="2" specimenCount-female="1" stateProvince="Cartago" typeStatus="holotype">
<collectingCountry id="F35821C4FFF1310A35BFF22DBE6EF90D" box="[158,289,1663,1689]" name="Costa Rica" pageId="2" pageNumber="441">Costa Rica</collectingCountry>
:
<collectingRegion id="498BAFB6FFF1310A340CF22DBEBEF90D" box="[301,497,1663,1689]" country="Costa Rica" name="Guanacaste" pageId="2" pageNumber="441">Guanacaste Prov.</collectingRegion>
<locationDeviation id="9A427876FFF1310A34D8F22DBD8AF90D" box="[505,709,1663,1689]" location="Sta Elena" pageId="2" pageNumber="441">
<quantity id="4CB7CCB1FFF1310A34D8F22DBD7BF90D" box="[505,564,1663,1689]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="441" unit="km" value="8.0">8 km</quantity>
N Sta Elena
</locationDeviation>
/ Santa Elena Forest Res.
<date id="FFF14794FFF1310A36D7F22DBBD1F90D" box="[1014,1182,1663,1689]" pageId="2" pageNumber="441" value="2001-06-11" valueMax="2001-06-17" valueMin="2001-06-11">
<collectingDate id="EFB5BE7CFFF1310A36D7F22DBBD1F90D" box="[1014,1182,1663,1689]" pageId="2" pageNumber="441" value="2001-06-11" valueMax="2001-06-17" valueMin="2001-06-11">11-17-VI-2001</collectingDate>
</date>
/
<collectorName id="26BA0482FFF1310A3196F22DBA0AF90D" box="[1207,1349,1663,1689]" pageId="2" pageNumber="441">S. &amp; J. Peck</collectorName>
<quantity id="4CB7CCB1FFF1310A306FF22DBAD2F90D" box="[1358,1437,1663,1689]" metricMagnitude="3" metricUnit="m" metricValue="1.64" pageId="2" pageNumber="441" unit="m" value="1640.0">
<elevation id="00628667FFF1310A306FF22DBAD2F90D" box="[1358,1437,1663,1689]" metricMagnitude="3" metricUnit="m" metricValue="1.64" pageId="2" pageNumber="441" unit="m" value="1640.0">1640m</elevation>
</quantity>
<geoCoordinate id="EE7B0793FFF1310A35B6F2F1BE7CF929" box="[151,307,1699,1725]" degrees="10" direction="north" minutes="20.701" orientation="latitude" pageId="2" pageNumber="441" precision="1" value="10.3450165">10° 20.701´N</geoCoordinate>
,
<geoCoordinate id="EE7B0793FFF1310A3460F2F1BEAFF929" box="[321,480,1699,1725]" degrees="84" direction="west" minutes="47.899" orientation="longitude" pageId="2" pageNumber="441" precision="1" value="-84.79832">84° 47.899´W</geoCoordinate>
/ ex. FIT CR1P01 004´, 1 (
<collectionCode id="ED5EF991FFF1310A363DF2F1BC21F929" box="[796,878,1699,1725]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="2" pageNumber="441" type="Museum">SEMC</collectionCode>
); [white label]´
<collectorName id="26BA0482FFF1310A3103F2F1BBE1F929" box="[1058,1198,1699,1725]" pageId="2" pageNumber="441">Vara Blanca</collectorName>
VIII.38 /
<collectorName id="26BA0482FFF1310A3000F2F1BE4AF975" pageId="2" pageNumber="441">Field Mus. Nat. Hist.</collectorName>
1966
<collectorName id="26BA0482FFF1310A346BF295BD7CF975" box="[330,563,1735,1761]" pageId="2" pageNumber="441">A. Bierig Collection</collectorName>
,
<collectorName id="26BA0482FFF1310A371CF295BDDEF975" box="[573,657,1735,1761]" pageId="2" pageNumber="441">Acc. Z.</collectorName>
13812´, 1 (
<collectionCode id="ED5EF991FFF1310A3635F295BC24F975" box="[788,875,1735,1761]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="2" pageNumber="441" type="Museum">FMNH</collectionCode>
); all of them with [red label]
</materialsCitation>
´
<typeStatus id="54F4DFF6FFF1310A319FF295BA6CF975" box="[1214,1315,1735,1761]" pageId="2" pageNumber="441" type="paratype">Paratype</typeStatus>
<taxonomicName id="4C4F1AD7FFF1310A300AF295BE9CF891" authority="Chani-Posse 2021" authorityName="Chani-Posse" authorityYear="2021" class="Insecta" family="Staphylinidae" genus="Linoderus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="441" phylum="Arthropoda" rank="species" species="smetanai">
<emphasis id="B93BBD46FFF1310A300AF295BFB3F890" italics="true" pageId="2" pageNumber="441">Linoderus smetanai</emphasis>
Chani-Posse 2021
</taxonomicName>
´.
</paragraph>
</subSubSection>
</treatment>
</document>