212 lines
34 KiB
XML
212 lines
34 KiB
XML
<document id="284F720C3A11ED794CB23164620EBF49" ID-CLB-Dataset="233753" ID-DOI="10.11646/zootaxa.5254.2.2" ID-GBIF-Dataset="f252562c-4717-47ac-a322-24b63bd4ef49" ID-ISSN="1175-5326" ID-Zenodo-Dep="7727356" ID-ZooBank="413FCD68-A4B1-4EA0-A58F-63C9993AC08F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1678695365183" checkinUser="plazi" docAuthor="Soares, Matheus M. M., Capellari, Renato S. & Ale-Rocha, Rosaly" docDate="2023" docId="03D787AAB66FFFA0FF2AF9B8FD64FDC9" docLanguage="en" docName="zootaxa.5254.2.2.pdf" docOrigin="Zootaxa 5254 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5254.2.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Paraclius latipes Loew 1864" docType="treatment" docVersion="8" lastPageNumber="191" masterDocId="FFEEFFD2B666FFAAFFBDFFE2FFE3FFD0" masterDocTitle="New or little-known Neotropical Dolichopodidae (Diptera) (II): genus Hercostomus Loew excluded from the Neotropics" masterLastPageNumber="208" masterPageNumber="181" pageNumber="190" updateTime="1699476502084" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods id="3811213EB0C52EB21AC2093A1FF2E5C8" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="5F7FF6D7A73C213755B338D531A14768">
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<mods:title id="85CF00AFCBECD7210497954951A0F057">New or little-known Neotropical Dolichopodidae (Diptera) (II): genus Hercostomus Loew excluded from the Neotropics</mods:title>
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<mods:name id="8EE178F2BF42768AC099295B7E10F221" type="personal">
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<mods:roleTerm id="E543B8CE813C7AEB19AEC9043B4201F6">Author</mods:roleTerm>
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<mods:namePart id="5AD82225F695B3729940530B6B333061">Soares, Matheus M. M.</mods:namePart>
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<mods:nameIdentifier id="9B988E580F8805AA529E0C559663844F" type="ORCID">0000-0002-2355-1441</mods:nameIdentifier>
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<mods:affiliation id="52D53FC263E4656095A84C1BB46078A4">Graduate Program in Entomology (PPG-Ent), Instituto Nacional de Pesquisas da Amazônia (INPA). & matheusmmsoares @ gmail. com; https: // orcid. org / 0000 - 0002 - 2355 - 1441</mods:affiliation>
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<mods:nameIdentifier id="06D1C07E594A80002A132A9794399A19" type="email">matheusmmsoares@gmail.com</mods:nameIdentifier>
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<mods:name id="EF0B81901F231FE9D731679D8703721E" type="personal">
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<mods:roleTerm id="3B3C99730EFBB89241EDC3E1553AEDD1">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="8C71333F07E3D64ACCCDE9BF2AE3DB48">Capellari, Renato S.</mods:namePart>
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<mods:nameIdentifier id="F35857D96154564871293665E9D294D4" type="ORCID">0000-0002-8410-9235</mods:nameIdentifier>
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<mods:affiliation id="295CE83227FD2CE4FEBA064DF9623688">Instituto Federal do Triângulo Mineiro, Campus Uberaba, Rua João Batista Ribeiro 4000, Distrito Industrial II, Uberaba, Minas Gerais 38064 - 790, Brazil. & rscapellari @ gmail. com; https: // orcid. org / 0000 - 0002 - 8410 - 9235</mods:affiliation>
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<mods:nameIdentifier id="7921CF7EF2B2F6E0E3ABF157DB1C0659" type="email">rscapellari@gmail.com</mods:nameIdentifier>
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<mods:name id="D7C5C1422F12FC9DE52C0B66C5DF9E37" type="personal">
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<mods:roleTerm id="2DFBC24E3A435D928444DA4AE3175711">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="5B694F99E3C826216B83E26FE37061AB">Ale-Rocha, Rosaly</mods:namePart>
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<mods:nameIdentifier id="6D0662A91A7DDE798F3F41E1933EAD25" type="ORCID">0000-0001-9874-9770</mods:nameIdentifier>
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<mods:affiliation id="2E8BAEC63FBE9DB18843633C259EAD49">Coordenação de Biodiversidade (COBIO, INPA), Av. André Araújo, 2936, Petrópolis, CEP 69067 - 375, Manaus, Amazonas, Brazil. & alerocha @ inpa. gov. br; https: // orcid. org / 0000 - 0001 - 9874 - 9770</mods:affiliation>
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<mods:nameIdentifier id="9BF6A92E55A19ADAB6381AE8238737EF" type="email">alerocha@inpa.gov.br</mods:nameIdentifier>
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<mods:title id="A779605F9A7CA6869ED69D6E959AAF29">Zootaxa</mods:title>
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<mods:part id="C5FFD1668B05F26E2EB0EA14C07C3FD7">
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<mods:date id="69BC29789428630CB7E41930E4C7E45C">2023</mods:date>
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<mods:number id="25CD4FFFE963F71342F0CA464E07A3E6">2023-03-10</mods:number>
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<mods:number id="57F272C3186CC307DED8E262E10C5564">5254</mods:number>
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<mods:url id="89F658D49BBED0D5711AF7523AC70814">http://dx.doi.org/10.11646/zootaxa.5254.2.2</mods:url>
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<mods:classification id="A64B91C9E6F8A4EC431C26366BC1E3EC">journal article</mods:classification>
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<treatment id="03D787AAB66FFFA0FF2AF9B8FD64FDC9" ID-DOI="http://doi.org/10.5281/zenodo.7730105" ID-GBIF-Taxon="209438281" ID-Zenodo-Dep="7730105" LSID="urn:lsid:plazi:treatment:03D787AAB66FFFA0FF2AF9B8FD64FDC9" httpUri="http://treatment.plazi.org/id/03D787AAB66FFFA0FF2AF9B8FD64FDC9" lastPageId="10" lastPageNumber="191" pageId="9" pageNumber="190">
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<subSubSection id="C3646537B66FFFA3FF2AF9B8FDEEF9A5" box="[151,525,1626,1653]" pageId="9" pageNumber="190" type="nomenclature">
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<paragraph id="8BC136BCB66FFFA3FF2AF9B8FDEEF9A5" blockId="9.[151,525,1626,1653]" box="[151,525,1626,1653]" pageId="9" pageNumber="190">
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<heading id="D08981D0B66FFFA3FF2AF9B8FDEEF9A5" bold="true" box="[151,525,1626,1653]" fontSize="11" level="1" pageId="9" pageNumber="190" reason="1">
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<emphasis id="B90AEAAEB66FFFA3FF2AF9B8FDEEF9A5" bold="true" box="[151,525,1626,1653]" pageId="9" pageNumber="190">
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The
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<taxonomicName id="4C7E4D3FB66FFFA3FF72F9B8FDEEF9A5" ID-CoL="BDMD9" authorityName="Loew" authorityYear="1864" baseAuthorityName="Aldrich" baseAuthorityYear="1901" box="[207,525,1626,1653]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="190" phylum="Arthropoda" rank="species" species="latipes">
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<emphasis id="B90AEAAEB66FFFA3FF72F9B8FEFEF9A5" bold="true" box="[207,285,1626,1653]" italics="true" pageId="9" pageNumber="190">latipes</emphasis>
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-group of
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<emphasis id="B90AEAAEB66FFFA3FE24F9B8FDEEF9A5" bold="true" box="[409,525,1626,1653]" italics="true" pageId="9" pageNumber="190">Paraclius</emphasis>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3646537B66FFFA0FF2AF940FE48FF05" lastPageId="10" lastPageNumber="191" pageId="9" pageNumber="190" type="diagnosis">
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<paragraph id="8BC136BCB66FFFA3FF2AF940FAECF80D" blockId="9.[151,1437,1698,2013]" pageId="9" pageNumber="190">
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<emphasis id="B90AEAAEB66FFFA3FF2AF940FE91F96C" bold="true" box="[151,370,1698,1724]" pageId="9" pageNumber="190">Diagnosis (males).</emphasis>
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The group is here proposed to accommodate the species with the following combination of characters: fore tarsomeres from apical 1/2–2/3 of It
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<subScript id="17FA34F9B66FFFA3FD57F93BFD11F937" attach="left" box="[746,754,1753,1767]" fontSize="6" pageId="9" pageNumber="190">1</subScript>
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with conspicuous pile of short whitish setae (
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<figureCitation id="13452A39B66FFFA3FABDF924FAB0F930" box="[1280,1363,1734,1760]" captionStart="FIGURE 8" captionStartId="14.[151,250,1943,1967]" captionTargetBox="[164,1424,181,1920]" captionTargetId="figure-22@14.[164,1424,181,1920]" captionTargetPageId="14" captionText="FIGURE 8. Paraclius latipes (Aldrich), identified male specimen from Costa Rica. A. Habitus, lateral view; B. Abdomen and hypopygium, lateral view; C. Head, anterior view; D. Left antenna, inner view; E–G. Tarsus I in dorsal, anterior and ventral views, respectively (arrow pointing to ventral whitish pile on tarsus I); H. Left wing." figureDoi="http://doi.org/10.5281/zenodo.7727382" httpUri="https://zenodo.org/record/7727382/files/figure.png" pageId="9" pageNumber="190">Fig. 8F</figureCitation>
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,
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<figureCitation id="13452A39B66FFFA3FADCF924FA6CF930" box="[1377,1423,1734,1760]" captionStart="FIGURE 11" captionStartId="18.[151,250,1943,1967]" captionTargetBox="[178,1409,181,1919]" captionTargetId="figure-22@18.[178,1409,181,1919]" captionTargetPageId="18" captionText="FIGURE 11. Paraclius norrbomi sp. nov., male holotype (CNC) (before dissection). A. Habitus, lateral view; B. Head, anterior view; C. Head, lateral view; D. Abdomen and hypopygium, lateral view; E, F. Tarsus I, ventral and anterior views, respectively; G. Left wing." figureDoi="http://doi.org/10.5281/zenodo.7727394" httpUri="https://zenodo.org/record/7727394/files/figure.png" pageId="9" pageNumber="190">11F</figureCitation>
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); arista-like stylus dorsal, arising at basal 1/2 of postpedicel (
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<figureCitation id="13452A39B66FFFA3FC9AF908FC60F8D4" box="[807,899,1770,1796]" captionStart="FIGURE 8" captionStartId="14.[151,250,1943,1967]" captionTargetBox="[164,1424,181,1920]" captionTargetId="figure-22@14.[164,1424,181,1920]" captionTargetPageId="14" captionText="FIGURE 8. Paraclius latipes (Aldrich), identified male specimen from Costa Rica. A. Habitus, lateral view; B. Abdomen and hypopygium, lateral view; C. Head, anterior view; D. Left antenna, inner view; E–G. Tarsus I in dorsal, anterior and ventral views, respectively (arrow pointing to ventral whitish pile on tarsus I); H. Left wing." figureDoi="http://doi.org/10.5281/zenodo.7727382" httpUri="https://zenodo.org/record/7727382/files/figure.png" pageId="9" pageNumber="190">Figs 8D</figureCitation>
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,
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<figureCitation id="13452A39B66FFFA3FC33F908FC23F8D4" box="[910,960,1770,1796]" captionStart="FIGURE 10" captionStartId="16.[151,250,1744,1768]" captionTargetBox="[151,1435,555,1719]" captionTargetId="figure-112@16.[151,1435,555,1719]" captionTargetPageId="16" captionText="FIGURE 10. Paraclius panamensis (Van Duzee) comb. nov., male holotype (AMNH). A. Habitus, lateral view; B. Head and thorax, lateral view; C. Labels; D. Head, anterior view; E. Abdomen and hypopygium, lateral view. Images originally lacked scale bars. Photographs by Courtney Richenbacher, AMNH." figureDoi="http://doi.org/10.5281/zenodo.7727390" httpUri="https://zenodo.org/record/7727390/files/figure.png" pageId="9" pageNumber="190">10B</figureCitation>
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,
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<figureCitation id="13452A39B66FFFA3FC77F908FC19F8D4" box="[970,1018,1770,1796]" captionStart="FIGURE 11" captionStartId="18.[151,250,1943,1967]" captionTargetBox="[178,1409,181,1919]" captionTargetId="figure-22@18.[178,1409,181,1919]" captionTargetPageId="18" captionText="FIGURE 11. Paraclius norrbomi sp. nov., male holotype (CNC) (before dissection). A. Habitus, lateral view; B. Head, anterior view; C. Head, lateral view; D. Abdomen and hypopygium, lateral view; E, F. Tarsus I, ventral and anterior views, respectively; G. Left wing." figureDoi="http://doi.org/10.5281/zenodo.7727394" httpUri="https://zenodo.org/record/7727394/files/figure.png" pageId="9" pageNumber="190">11C</figureCitation>
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); femur III with 1 strong anterodorsal and 1 long anteroventral preapical setae; cell dm ending at middle of wing; vein M
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<subScript id="17FA34F9B66FFFA3FB80F8C3FBA6F8FF" attach="left" box="[1085,1093,1825,1839]" fontSize="6" pageId="9" pageNumber="190">1</subScript>
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gradually angled towards R
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<subScript id="17FA34F9B66FFFA3FA3EF8C2FA7EF8FE" attach="left" box="[1411,1437,1824,1838]" fontSize="6" pageId="9" pageNumber="190">4+5</subScript>
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(
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<figureCitation id="13452A39B66FFFA3FF22F8D0FF1FF89C" box="[159,252,1842,1868]" captionStart="FIGURE 6" captionStartId="11.[151,250,1428,1452]" captionTargetBox="[151,1435,181,1404]" captionTargetId="figure-18@11.[151,1435,181,1404]" captionTargetPageId="11" captionText="FIGURE 6. Paraclius aberrans Robinson, male holotype (NMNH). A–C. Habitus in lateral, anterior and dorsal views, respectively; D. Labels. Images B and C originally lacked scale bars. Photographs by Alyssa Seemann, NMNH." figureDoi="http://doi.org/10.5281/zenodo.7727376" httpUri="https://zenodo.org/record/7727376/files/figure.png" pageId="9" pageNumber="190">Figs 6A</figureCitation>
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,
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<figureCitation id="13452A39B66FFFA3FEBBF8D1FECBF89D" box="[262,296,1843,1869]" captionStart="FIGURE 7" captionStartId="13.[151,250,1867,1891]" captionTargetBox="[151,1435,849,1844]" captionTargetId="figure-357@13.[151,1435,849,1844]" captionTargetPageId="13" captionText="FIGURE 7. Paraclius latipes (Aldrich), male syntype (NMNH). A. Habitus, lateral view; B. Head, anterior view; C. Labels; D. Tarsus I, posterior view; E. Left wing. Images B, D and E originally lacked scale bars. Photographs by Alyssa Seemann, NMNH." figureDoi="http://doi.org/10.5281/zenodo.7727378" httpUri="https://zenodo.org/record/7727378/files/figure.png" pageId="9" pageNumber="190">7E</figureCitation>
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,
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<figureCitation id="13452A39B66FFFA3FE8EF8D0FEB4F89C" box="[307,343,1842,1868]" captionStart="FIGURE 8" captionStartId="14.[151,250,1943,1967]" captionTargetBox="[164,1424,181,1920]" captionTargetId="figure-22@14.[164,1424,181,1920]" captionTargetPageId="14" captionText="FIGURE 8. Paraclius latipes (Aldrich), identified male specimen from Costa Rica. A. Habitus, lateral view; B. Abdomen and hypopygium, lateral view; C. Head, anterior view; D. Left antenna, inner view; E–G. Tarsus I in dorsal, anterior and ventral views, respectively (arrow pointing to ventral whitish pile on tarsus I); H. Left wing." figureDoi="http://doi.org/10.5281/zenodo.7727382" httpUri="https://zenodo.org/record/7727382/files/figure.png" pageId="9" pageNumber="190">8H</figureCitation>
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,
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<figureCitation id="13452A39B66FFFA3FEDFF8D0FE77F89C" box="[354,404,1842,1868]" captionStart="FIGURE 11" captionStartId="18.[151,250,1943,1967]" captionTargetBox="[178,1409,181,1919]" captionTargetId="figure-22@18.[178,1409,181,1919]" captionTargetPageId="18" captionText="FIGURE 11. Paraclius norrbomi sp. nov., male holotype (CNC) (before dissection). A. Habitus, lateral view; B. Head, anterior view; C. Head, lateral view; D. Abdomen and hypopygium, lateral view; E, F. Tarsus I, ventral and anterior views, respectively; G. Left wing." figureDoi="http://doi.org/10.5281/zenodo.7727394" httpUri="https://zenodo.org/record/7727394/files/figure.png" pageId="9" pageNumber="190">11G</figureCitation>
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); hypopygium: cercus remarkably short and branched, dorsal branch shorter than ventral one (
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<figureCitation id="13452A39B66FFFA3FF22F8B4FF1BF8A0" box="[159,248,1878,1904]" captionStart="FIGURE 9" captionStartId="15.[151,250,1787,1811]" captionTargetBox="[151,1435,381,1764]" captionTargetId="figure-91@15.[151,1435,381,1764]" captionTargetPageId="15" captionText="FIGURE 9. Paraclius latipes (Aldrich), hypopygium, identified male specimen from Costa Rica. A, B. Hypopygium, not dissected and dissected, respectively, left lateral view; C. Internal appendages, right lateral view; D. Hypandrium, right lateral view; E. Right cercus, outer view; F. Right ventral and dorsal lobes of surstylus, inner view; G. Postgonite, ventral view. Abbreviations: apv lb = apicoventral epandrial lobe; bs sp = basal sclerite of sperm pump; cerc = cercus; d sur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; sp = sperm pump; v sur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.7727388" httpUri="https://zenodo.org/record/7727388/files/figure.png" pageId="9" pageNumber="190">Figs 9E</figureCitation>
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,
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<figureCitation id="13452A39B66FFFA3FEBFF8B4FED4F8A0" box="[258,311,1878,1904]" captionStart="FIGURE 12" captionStartId="19.[151,250,1824,1848]" captionTargetBox="[243,1344,813,1801]" captionTargetId="figure-330@19.[243,1344,813,1801]" captionTargetPageId="19" captionText="FIGURE 12. Paraclius norrbomi sp. nov., male holotype(CNC).A, B. Hypopygium, not macerated and macerated, respectively, left lateral view; C. Internal appendages, right lateral view; D. left cercus, outer view; E. Hypandrium, right lateral view; F. Right ventral lobe of surstylus, inner view; G. Right dorsal lobe of surstylus, inner view. Abbreviations: apv lb = apicoventral epandrial lobe; bs sp = basal sclerite of sperm pump; cerc = cercus; d sur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; prct = proctiger; sp = sperm pump; v sur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.7727398" httpUri="https://zenodo.org/record/7727398/files/figure.png" pageId="9" pageNumber="190">12D</figureCitation>
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). In addition to the preceding features, it remains to be investigated if the phallus presents a narrowing which clearly delimits two regions (with distal segment serrate) (
|
||
<figureCitation id="13452A39B66FFFA3FCC3F898FC39F844" box="[894,986,1914,1940]" captionStart="FIGURE 9" captionStartId="15.[151,250,1787,1811]" captionTargetBox="[151,1435,381,1764]" captionTargetId="figure-91@15.[151,1435,381,1764]" captionTargetPageId="15" captionText="FIGURE 9. Paraclius latipes (Aldrich), hypopygium, identified male specimen from Costa Rica. A, B. Hypopygium, not dissected and dissected, respectively, left lateral view; C. Internal appendages, right lateral view; D. Hypandrium, right lateral view; E. Right cercus, outer view; F. Right ventral and dorsal lobes of surstylus, inner view; G. Postgonite, ventral view. Abbreviations: apv lb = apicoventral epandrial lobe; bs sp = basal sclerite of sperm pump; cerc = cercus; d sur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; sp = sperm pump; v sur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.7727388" httpUri="https://zenodo.org/record/7727388/files/figure.png" pageId="9" pageNumber="190">Figs 9C</figureCitation>
|
||
,
|
||
<figureCitation id="13452A39B66FFFA3FC55F898FBF8F844" box="[1000,1051,1914,1940]" captionStart="FIGURE 12" captionStartId="19.[151,250,1824,1848]" captionTargetBox="[243,1344,813,1801]" captionTargetId="figure-330@19.[243,1344,813,1801]" captionTargetPageId="19" captionText="FIGURE 12. Paraclius norrbomi sp. nov., male holotype(CNC).A, B. Hypopygium, not macerated and macerated, respectively, left lateral view; C. Internal appendages, right lateral view; D. left cercus, outer view; E. Hypandrium, right lateral view; F. Right ventral lobe of surstylus, inner view; G. Right dorsal lobe of surstylus, inner view. Abbreviations: apv lb = apicoventral epandrial lobe; bs sp = basal sclerite of sperm pump; cerc = cercus; d sur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; prct = proctiger; sp = sperm pump; v sur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.7727398" httpUri="https://zenodo.org/record/7727398/files/figure.png" pageId="9" pageNumber="190">12D</figureCitation>
|
||
), hypandrium asymmetrical (with one wide ventral projection near/or at apex) (
|
||
<figureCitation id="13452A39B66FFFA3FD37F87CFD05F868" box="[650,742,1950,1976]" captionStart="FIGURE 9" captionStartId="15.[151,250,1787,1811]" captionTargetBox="[151,1435,381,1764]" captionTargetId="figure-91@15.[151,1435,381,1764]" captionTargetPageId="15" captionText="FIGURE 9. Paraclius latipes (Aldrich), hypopygium, identified male specimen from Costa Rica. A, B. Hypopygium, not dissected and dissected, respectively, left lateral view; C. Internal appendages, right lateral view; D. Hypandrium, right lateral view; E. Right cercus, outer view; F. Right ventral and dorsal lobes of surstylus, inner view; G. Postgonite, ventral view. Abbreviations: apv lb = apicoventral epandrial lobe; bs sp = basal sclerite of sperm pump; cerc = cercus; d sur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; sp = sperm pump; v sur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.7727388" httpUri="https://zenodo.org/record/7727388/files/figure.png" pageId="9" pageNumber="190">Figs 9D</figureCitation>
|
||
,
|
||
<figureCitation id="13452A39B66FFFA3FD4CF87CFCC3F868" box="[753,800,1950,1976]" captionStart="FIGURE 12" captionStartId="19.[151,250,1824,1848]" captionTargetBox="[243,1344,813,1801]" captionTargetId="figure-330@19.[243,1344,813,1801]" captionTargetPageId="19" captionText="FIGURE 12. Paraclius norrbomi sp. nov., male holotype(CNC).A, B. Hypopygium, not macerated and macerated, respectively, left lateral view; C. Internal appendages, right lateral view; D. left cercus, outer view; E. Hypandrium, right lateral view; F. Right ventral lobe of surstylus, inner view; G. Right dorsal lobe of surstylus, inner view. Abbreviations: apv lb = apicoventral epandrial lobe; bs sp = basal sclerite of sperm pump; cerc = cercus; d sur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; prct = proctiger; sp = sperm pump; v sur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.7727398" httpUri="https://zenodo.org/record/7727398/files/figure.png" pageId="9" pageNumber="190">12E</figureCitation>
|
||
) and the postgonite bifurcated at apex (
|
||
<figureCitation id="13452A39B66FFFA3FB68F87CFAD2F868" box="[1237,1329,1950,1976]" captionStart="FIGURE 9" captionStartId="15.[151,250,1787,1811]" captionTargetBox="[151,1435,381,1764]" captionTargetId="figure-91@15.[151,1435,381,1764]" captionTargetPageId="15" captionText="FIGURE 9. Paraclius latipes (Aldrich), hypopygium, identified male specimen from Costa Rica. A, B. Hypopygium, not dissected and dissected, respectively, left lateral view; C. Internal appendages, right lateral view; D. Hypandrium, right lateral view; E. Right cercus, outer view; F. Right ventral and dorsal lobes of surstylus, inner view; G. Postgonite, ventral view. Abbreviations: apv lb = apicoventral epandrial lobe; bs sp = basal sclerite of sperm pump; cerc = cercus; d sur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; sp = sperm pump; v sur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.7727388" httpUri="https://zenodo.org/record/7727388/files/figure.png" pageId="9" pageNumber="190">Figs 9G</figureCitation>
|
||
,
|
||
<figureCitation id="13452A39B66FFFA3FA81F87CFA8FF868" box="[1340,1388,1950,1976]" captionStart="FIGURE 12" captionStartId="19.[151,250,1824,1848]" captionTargetBox="[243,1344,813,1801]" captionTargetId="figure-330@19.[243,1344,813,1801]" captionTargetPageId="19" captionText="FIGURE 12. Paraclius norrbomi sp. nov., male holotype(CNC).A, B. Hypopygium, not macerated and macerated, respectively, left lateral view; C. Internal appendages, right lateral view; D. left cercus, outer view; E. Hypandrium, right lateral view; F. Right ventral lobe of surstylus, inner view; G. Right dorsal lobe of surstylus, inner view. Abbreviations: apv lb = apicoventral epandrial lobe; bs sp = basal sclerite of sperm pump; cerc = cercus; d sur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; prct = proctiger; sp = sperm pump; v sur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.7727398" httpUri="https://zenodo.org/record/7727398/files/figure.png" pageId="9" pageNumber="190">12C</figureCitation>
|
||
) are exclusive features of
|
||
<taxonomicName id="4C7E4D3FB66FFFA3FE3BF821FE0FF80C" baseAuthorityName="Aldrich" baseAuthorityYear="1901" box="[390,492,1986,2012]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="190" phylum="Arthropoda" rank="species" species="latipes">
|
||
<emphasis id="B90AEAAEB66FFFA3FE3BF821FE0FF80C" box="[390,492,1986,2012]" italics="true" pageId="9" pageNumber="190">P. latipes</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7E4D3FB66FFFA3FD9FF821FD4BF80C" authority="Soares & Capellari & Ale-Rocha, 2023" authorityName="Soares & Capellari & Ale-Rocha" authorityYear="2023" box="[546,680,1986,2012]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="190" phylum="Arthropoda" rank="species" species="norrbomi" status="sp. nov.">
|
||
<emphasis id="B90AEAAEB66FFFA3FD9FF821FD4BF80C" box="[546,680,1986,2012]" italics="true" pageId="9" pageNumber="190">P. norrbomi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B90AEAAEB66FFFA3FD12F820FCEBF80C" bold="true" box="[687,776,1986,2012]" pageId="9" pageNumber="190">
|
||
<taxonomicNameLabel id="A23957D5B66FFFA3FD12F820FCEBF80C" box="[687,776,1986,2012]" pageId="9" pageNumber="190" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
or part of the ground-plan of the entire group.
|
||
</paragraph>
|
||
<paragraph id="8BC136BCB66CFFA0FF7AFF75FE48FF05" blockId="10.[151,1437,150,537]" pageId="10" pageNumber="191">
|
||
<emphasis id="B90AEAAEB66CFFA0FF7AFF75FE71FF60" bold="true" box="[199,402,150,177]" pageId="10" pageNumber="191">Included species.</emphasis>
|
||
<taxonomicName id="4C7E4D3FB66CFFA0FE25FF75FD04FF61" authority="Robinson" authorityName="Robinson" authorityYear="1964" box="[408,743,151,177]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="191" phylum="Arthropoda" rank="species" species="aberrans">
|
||
<emphasis id="B90AEAAEB66CFFA0FE25FF75FD92FF61" box="[408,625,151,177]" italics="true" pageId="10" pageNumber="191">Paraclius aberrans</emphasis>
|
||
Robinson
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C7E4D3FB66CFFA0FD4FFF75FC2DFF61" authority="(Aldrich)" baseAuthorityName="Aldrich" baseAuthorityYear="1901" box="[754,974,151,177]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="191" phylum="Arthropoda" rank="species" species="latipes">
|
||
<emphasis id="B90AEAAEB66CFFA0FD4FFF75FCE0FF60" box="[754,771,151,176]" italics="true" pageId="10" pageNumber="191">P</emphasis>
|
||
.
|
||
<emphasis id="B90AEAAEB66CFFA0FCACFF75FCBFFF61" box="[785,860,151,177]" italics="true" pageId="10" pageNumber="191">latipes</emphasis>
|
||
(Aldrich)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C7E4D3FB66CFFA0FC64FF75FB81FF61" authority="Soares & Capellari & Ale-Rocha, 2023" authorityName="Soares & Capellari & Ale-Rocha" authorityYear="2023" box="[985,1122,151,177]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="191" phylum="Arthropoda" rank="species" species="norrbomi" status="sp. nov.">
|
||
<emphasis id="B90AEAAEB66CFFA0FC64FF75FC09FF60" box="[985,1002,151,176]" italics="true" pageId="10" pageNumber="191">P</emphasis>
|
||
.
|
||
<emphasis id="B90AEAAEB66CFFA0FC45FF75FB81FF61" box="[1016,1122,151,177]" italics="true" pageId="10" pageNumber="191">norrbomi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B90AEAAEB66CFFA0FBD4FF75FB21FF61" bold="true" box="[1129,1218,151,177]" pageId="10" pageNumber="191">
|
||
<taxonomicNameLabel id="A23957D5B66CFFA0FBD4FF75FB21FF61" box="[1129,1218,151,177]" pageId="10" pageNumber="191" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C7E4D3FB66CFFA0FB44FF75FEC7FF05" authority="(Van Duzee)" authorityName="Soares & Capellari & Ale-Rocha" authorityYear="2023" baseAuthorityName="Van Duzee" baseAuthorityYear="1931" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="191" phylum="Arthropoda" rank="species" species="panamensis" status="comb. nov.">
|
||
<emphasis id="B90AEAAEB66CFFA0FB44FF75FAE9FF60" box="[1273,1290,151,176]" italics="true" pageId="10" pageNumber="191">P</emphasis>
|
||
.
|
||
<emphasis id="B90AEAAEB66CFFA0FAA5FF75FA7FFF60" box="[1304,1436,151,176]" italics="true" pageId="10" pageNumber="191">panamensis</emphasis>
|
||
(Van Duzee)
|
||
</taxonomicName>
|
||
<emphasis id="B90AEAAEB66CFFA0FE96FF59FE48FF05" bold="true" box="[299,427,187,213]" pageId="10" pageNumber="191">
|
||
<taxonomicNameLabel id="A23957D5B66CFFA0FE96FF59FE48FF05" box="[299,427,187,213]" pageId="10" pageNumber="191" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3646537B66CFFA0FF7AFF3DFD64FDC9" pageId="10" pageNumber="191" type="discussion">
|
||
<paragraph id="8BC136BCB66CFFA0FF7AFF3DFD64FDC9" blockId="10.[151,1437,150,537]" pageId="10" pageNumber="191">
|
||
<emphasis id="B90AEAAEB66CFFA0FF7AFF3DFED8FF29" bold="true" box="[199,315,223,249]" pageId="10" pageNumber="191">Remarks.</emphasis>
|
||
After gathering information from descriptions, photos of
|
||
<typeStatus id="54C5881EB66CFFA0FC74FF3DFBE7FF29" box="[969,1028,223,249]" pageId="10" pageNumber="191">types</typeStatus>
|
||
and identified material of nearly all New World described species of
|
||
<taxonomicName id="4C7E4D3FB66CFFA0FDB8FEE1FD91FECD" authorityName="Loew" authorityYear="1864" box="[517,626,259,285]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="191" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90AEAAEB66CFFA0FDB8FEE1FD91FECD" box="[517,626,259,285]" italics="true" pageId="10" pageNumber="191">Paraclius</emphasis>
|
||
</taxonomicName>
|
||
, it became clear that
|
||
<taxonomicName id="4C7E4D3FB66CFFA0FCE3FEE1FB9FFECC" authorityName="Soares & Capellari & Ale-Rocha" authorityYear="2023" baseAuthorityName="Soares & Capellari & Ale-Rocha" baseAuthorityYear="2023" box="[862,1148,259,284]" class="Insecta" family="Dolichopodidae" genus="Hercostomus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="191" phylum="Arthropoda" rank="species" species="panamensis">
|
||
<emphasis id="B90AEAAEB66CFFA0FCE3FEE1FB9FFECC" box="[862,1148,259,284]" italics="true" pageId="10" pageNumber="191">Hercostomus panamensis</emphasis>
|
||
</taxonomicName>
|
||
was part of a small clade, sharing distinctive characters with
|
||
<taxonomicName id="4C7E4D3FB66CFFA0FDA3FEC5FCF3FE91" authority="Robinson" authorityName="Robinson" authorityYear="1964" box="[542,784,295,321]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="191" phylum="Arthropoda" rank="species" species="aberrans">
|
||
<emphasis id="B90AEAAEB66CFFA0FDA3FEC5FD7DFE91" box="[542,670,295,321]" italics="true" pageId="10" pageNumber="191">P. aberrans</emphasis>
|
||
Robinson
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7E4D3FB66CFFA0FCFAFEC5FBC3FE91" authority="(Aldrich)" baseAuthorityName="Aldrich" baseAuthorityYear="1901" box="[839,1056,295,321]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="191" phylum="Arthropoda" rank="species" species="latipes">
|
||
<emphasis id="B90AEAAEB66CFFA0FCFAFEC5FC4DFE91" box="[839,942,295,321]" italics="true" pageId="10" pageNumber="191">P. latipes</emphasis>
|
||
(Aldrich)
|
||
</taxonomicName>
|
||
, notably the fore tarsus with short pile of setae (that also occurs elsewhere in other dolichopodine genera, like
|
||
<taxonomicName id="4C7E4D3FB66CFFA0FBBFFEA9FB41FEB5" box="[1026,1186,331,357]" class="Insecta" family="Dolichopodidae" genus="Pelastoneurus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="191" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90AEAAEB66CFFA0FBBFFEA9FB41FEB5" box="[1026,1186,331,357]" italics="true" pageId="10" pageNumber="191">Pelastoneurus</emphasis>
|
||
</taxonomicName>
|
||
) and the shape of the male cercus unlike any other species known to us. Additional characters of the hypandrium, phallus and postgonites are also peculiar in the species for which material was available for dissection and likely synapomorphic for the entire group. Although it is not the main focus of the present paper, we decided to describe a new species of this group from
|
||
<collectingCountry id="F369762CB66CFFA0FE9EFE39FE43FE25" box="[291,416,475,501]" name="Costa Rica" pageId="10" pageNumber="191">Costa Rica</collectingCountry>
|
||
and to redescribe
|
||
<taxonomicName id="4C7E4D3FB66CFFA0FDC6FE39FD07FE25" baseAuthorityName="Aldrich" baseAuthorityYear="1901" box="[635,740,475,501]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="191" phylum="Arthropoda" rank="species" species="latipes">
|
||
<emphasis id="B90AEAAEB66CFFA0FDC6FE39FD07FE25" box="[635,740,475,501]" italics="true" pageId="10" pageNumber="191">P. latipes</emphasis>
|
||
</taxonomicName>
|
||
based on additional material in order to better illustrate the distinctive morphology of the
|
||
<taxonomicName id="4C7E4D3FB66CFFA0FE57FE1DFDD6FDC9" baseAuthorityName="Aldrich" baseAuthorityYear="1901" box="[490,565,511,537]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="191" phylum="Arthropoda" rank="species" species="latipes">
|
||
<emphasis id="B90AEAAEB66CFFA0FE57FE1DFDD6FDC9" box="[490,565,511,537]" italics="true" pageId="10" pageNumber="191">latipes</emphasis>
|
||
</taxonomicName>
|
||
-group.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |