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<document id="BDF6AEDAA7A5D603D7DAF8FB27C16F39" ID-DOI="10.11646/zootaxa.5489.1.4" ID-ISSN="1175-5326" ID-Zenodo-Dep="13211341" ID-ZooBank="373DBA20-A1A7-4A2D-856C-67BF13D83C41" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722790258077" checkinUser="plazi" docAuthor="Okajima, Shûji &amp; Masumoto, Masami" docDate="2024" docId="03D2383B851DC324D9B2A50EAD6D70EF" docLanguage="en" docName="zootaxa.5489.1.4.pdf" docOrigin="Zootaxa 5489 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5489.1.4" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hoplandrothrips flavicornis Okajima &amp; Masumoto, 2024, sp. n." docType="treatment" docVersion="2" lastPageNumber="44" masterDocId="FFEB40438508C332D925A153A8767142" masterDocTitle="The genus Hoplandrothrips and its relatives (Thysanoptera: Phlaeothripidae) from Southeast Asia and Taiwan" masterLastPageNumber="91" masterPageNumber="22" pageNumber="43" updateTime="1722791853120" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="BC028123021524EE709284DC0333E591">The genus Hoplandrothrips and its relatives (Thysanoptera: Phlaeothripidae) from Southeast Asia and Taiwan</mods:title>
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<mods:namePart id="265FBCE7516098046E8A9A8E0408C2EB">Okajima, Shûji</mods:namePart>
<mods:nameIdentifier id="46FF0446FE9EDA5DEF637E65A956D2A8" type="ORCID">0000-0001-7249-671X</mods:nameIdentifier>
<mods:affiliation id="D70FEFB2F1D2B56671964B3A91905CA5">Laboratory of Entomology, Tokyo University of Agriculture, 1737 Funako, Atsugi, Kanagawa, 243 - 0034 Japan.</mods:affiliation>
<mods:nameIdentifier id="99DBD31B84707A277EE8280403F59670" type="email">7okajimas2@gmail.com</mods:nameIdentifier>
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<mods:namePart id="D7677573E4989C92B9246C4692977563">Masumoto, Masami</mods:namePart>
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<mods:affiliation id="2B44E6945B69FD6E67597E52C2663548">Laboratory of Entomology, Tokyo University of Agriculture, 1737 Funako, Atsugi, Kanagawa, 243 - 0034 Japan.</mods:affiliation>
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<mods:date id="1A0B3CA66A3399B591EF57AB623B55C2">2024</mods:date>
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<treatment id="03D2383B851DC324D9B2A50EAD6D70EF" LSID="urn:lsid:plazi:treatment:03D2383B851DC324D9B2A50EAD6D70EF" httpUri="http://treatment.plazi.org/id/03D2383B851DC324D9B2A50EAD6D70EF" lastPageId="22" lastPageNumber="44" pageId="21" pageNumber="43">
<subSubSection id="C361DAA6851DC327D9B2A50EA90275D9" pageId="21" pageNumber="43" type="nomenclature">
<paragraph id="8BC4892D851DC327D9B2A50EAA457535" blockId="21.[151,563,1117,1179]" box="[151,563,1117,1144]" pageId="21" pageNumber="43">
<heading id="D08C3E41851DC327D9B2A50EAA457535" bold="true" box="[151,563,1117,1144]" fontSize="11" level="1" pageId="21" pageNumber="43" reason="1">
<emphasis id="B90F553F851DC327D9B2A50EAA457535" bold="true" box="[151,563,1117,1144]" pageId="21" pageNumber="43">
<taxonomicName id="4C7BF2AE851DC327D9B2A50EA99C753A" authorityName="Okajima &amp; Masumoto" authorityYear="2024" box="[151,490,1117,1144]" class="Insecta" family="Phlaeothripidae" genus="Hoplandrothrips" kingdom="Animalia" order="Thysanoptera" pageId="21" pageNumber="43" phylum="Arthropoda" rank="species" species="flavicornis" status="sp. nov.">
<emphasis id="B90F553F851DC327D9B2A50EA99C753A" bold="true" box="[151,490,1117,1144]" italics="true" pageId="21" pageNumber="43">Hoplandrothrips flavicornis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23CE844851DC327D8D4A50EAA457535" box="[497,563,1117,1143]" pageId="21" pageNumber="43" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BC4892D851DC327D9B2A5D2A90275D9" blockId="21.[151,563,1117,1179]" box="[151,372,1153,1179]" pageId="21" pageNumber="43">(Figs, 28, 170176)</paragraph>
</subSubSection>
<subSubSection id="C361DAA6851DC324D9B2A59AA9F371F3" lastPageId="22" lastPageNumber="44" pageId="21" pageNumber="43" type="description">
<paragraph id="8BC4892D851DC327D9B2A59AABF277BD" blockId="21.[151,1437,1225,2008]" pageId="21" pageNumber="43">
<emphasis id="B90F553F851DC327D9B2A59AA9F875A6" box="[151,398,1225,1252]" italics="true" pageId="21" pageNumber="43">Female (macroptera).</emphasis>
<materialsCitation id="3B138370851DC327D8B1A59AABF677BD" collectionCode="II, III, IV, VI, VII, VIII" collectorName="Maxillary &amp; Pronotum &amp; Metanotal &amp; Fore &amp; Tergite IX &amp; Tube &amp; Terminal" pageId="21" pageNumber="43" specimenCount="2" typeStatus="holotype">
Distended body length:
<quantity id="4C8324C8851DC327DB84A59AAB5275A1" box="[673,804,1225,1251]" metricMagnitude="-3" metricUnit="m" metricValue="2.65" metricValueMax="2.7" metricValueMin="2.6" pageId="21" pageNumber="43" unit="mm" value="2.65" valueMax="2.7" valueMin="2.6">2.62.7mm</quantity>
. Body uniformly brown, with red hypodermal pigments. Antennal segments I
<collectionCode id="ED6A11E8851DC327D8AFA5BDA9EA744A" box="[394,412,1262,1288]" pageId="21" pageNumber="43">II</collectionCode>
dark brown, slightly darker than head, segment
<collectionCode id="ED6A11E8851DC327DA94A5BDABBB744A" box="[945,973,1262,1288]" pageId="21" pageNumber="43">III</collectionCode>
yellow, segments
<collectionCode id="ED6A11E8851DC327DDBCA5BDACC0744A" box="[1177,1206,1262,1288]" pageId="21" pageNumber="43">IV</collectionCode>
<collectionCode id="ED6A11E8851DC327DDE0A5BDAC94744A" box="[1221,1250,1262,1288]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="21" pageNumber="43">VI</collectionCode>
pale brown with basal 1/2 yellowish, segment
<collectionCode id="ED6A11E8851DC327D8C6A441AA7C746E" box="[483,522,1298,1324]" pageId="21" pageNumber="43">VII</collectionCode>
pale brown with basal 1/3 yellowish, segment
<collectionCode id="ED6A11E8851DC327DD39A441AC3A746E" box="[1052,1100,1298,1324]" pageId="21" pageNumber="43">VIII</collectionCode>
brown. Femora brown; tibiae and tarsi yellow. Fore wings almost clear. All prominent setae pale. Head (
<figureCitation id="134095A8851DC327DACEA466AC3A740D" box="[1003,1100,1333,1359]" captionStart="FIGURES 170176" captionStartId="54.[151,264,1943,1967]" captionTargetBox="[282,1306,182,1919]" captionTargetId="figure-17@54.[282,1306,181,1919]" captionTargetPageId="54" captionText="FIGURES 170176. Hoplandrothrips flavicornis sp. n., female. (170) Head and prothorax; (171) antenna; (172) antennal segments IIIIV; (173) metanotum; (174) pelta; (175) sub-basal wing setae; (176) tube." figureDoi="http://doi.org/10.5281/zenodo.13211401" httpUri="https://zenodo.org/record/13211401/files/figure.png" pageId="21" pageNumber="43">Fig. 170</figureCitation>
) elongate, 1.43 times as long as wide, dorsal surface probably smooth, sculptured laterally. Cheeks weakly rounded, constricted behind eyes, serrated, each with a few setae. Postocular setae well-developed, almost as long as eyes, expanded. Eyes about 1/3 as long as head, with at least one somewhat large ommatidium postero-laterally. Antennae (
<figureCitation id="134095A8851DC327DDB9A4F2AD3874F9" box="[1180,1358,1441,1467]" captionStart="FIGURES 170176" captionStartId="54.[151,264,1943,1967]" captionTargetBox="[282,1306,182,1919]" captionTargetId="figure-17@54.[282,1306,181,1919]" captionTargetPageId="54" captionText="FIGURES 170176. Hoplandrothrips flavicornis sp. n., female. (170) Head and prothorax; (171) antenna; (172) antennal segments IIIIV; (173) metanotum; (174) pelta; (175) sub-basal wing setae; (176) tube." figureDoi="http://doi.org/10.5281/zenodo.13211401" httpUri="https://zenodo.org/record/13211401/files/figure.png" pageId="21" pageNumber="43">Figs 171 &amp; 172</figureCitation>
) about 1.9 times as long as head; segment
<collectionCode id="ED6A11E8851DC327DB05A495AA2674A2" box="[544,592,1478,1504]" pageId="21" pageNumber="43">VIII</collectionCode>
long, distinctly constricted at base, pedicellate; segment
<collectionCode id="ED6A11E8851DC327DDEBA495AC9C74A2" box="[1230,1258,1478,1504]" pageId="21" pageNumber="43">III</collectionCode>
about 2.0 times as long as wide, almost as long as segment
<collectionCode id="ED6A11E8851DC327DBA0A4B9AAD47746" box="[645,674,1514,1540]" pageId="21" pageNumber="43">IV</collectionCode>
, with three sense cones.
<collectorName id="268EECFB851DC327DA9FA4BAAC5E7741" box="[954,1064,1513,1539]" pageId="21" pageNumber="43">Maxillary</collectorName>
stylets rather short, not reaching middle of head, close together, 78μm apart at middle.
<collectorName id="268EECFB851DC327DA34A75DAB09776A" box="[785,895,1550,1576]" pageId="21" pageNumber="43">Pronotum</collectorName>
0.67 times as long as head, 1.56 times as wide as long; am setae reduced, about 0.5 times as long as aa, but expanded, remaining four pairs of prominent setae elongate, expanded.
<collectorName id="268EECFB851DC327D85AA706A987772D" box="[383,497,1621,1647]" pageId="21" pageNumber="43">Metanotal</collectorName>
median pair of setae acute, 4345μm apart from anterior margin, 70μm apart from one another.
<collectorName id="268EECFB851DC327D80CA729A92A77D6" box="[297,348,1658,1684]" pageId="21" pageNumber="43">Fore</collectorName>
tarsal tooth distinct.
<collectorName id="268EECFB851DC327DB6EA729AA0877D6" box="[587,638,1658,1684]" pageId="21" pageNumber="43">Fore</collectorName>
wings each with 58 duplicated cilia,
<quantity id="4C8324C8851DC327DD17A72AAC0C77D1" box="[1074,1146,1657,1683]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" metricValueMax="2.032" metricValueMin="1.524" pageId="21" pageNumber="43" unit="in" value="7.0" valueMax="8.0" valueMin="6.0">68 in</quantity>
<typeStatus id="54C0378F851DC327DDA4A72AAC9177D1" box="[1153,1255,1657,1683]" pageId="21" pageNumber="43" type="holotype">holotype</typeStatus>
; three sub-basal setae (
<figureCitation id="134095A8851DC327D9C5A7CEA93777FA" box="[224,321,1693,1720]" captionStart="FIGURES 170176" captionStartId="54.[151,264,1943,1967]" captionTargetBox="[282,1306,182,1919]" captionTargetId="figure-17@54.[282,1306,181,1919]" captionTargetPageId="54" captionText="FIGURES 170176. Hoplandrothrips flavicornis sp. n., female. (170) Head and prothorax; (171) antenna; (172) antennal segments IIIIV; (173) metanotum; (174) pelta; (175) sub-basal wing setae; (176) tube." figureDoi="http://doi.org/10.5281/zenodo.13211401" httpUri="https://zenodo.org/record/13211401/files/figure.png" pageId="21" pageNumber="43">Fig. 175</figureCitation>
) expanded, S2 the longest.
<collectorName id="268EECFB851DC327DB58A7CEAA8477FA" box="[637,754,1693,1720]" pageId="21" pageNumber="43">Tergite IX</collectorName>
posteromarginal setae longer than tube, sharply pointed, S2 longer than S1, intermediate setae shorter than 1/2 of S1.
<collectorName id="268EECFB851DC327DA34A792AB3C7799" box="[785,842,1729,1755]" pageId="21" pageNumber="43">Tube</collectorName>
(
<figureCitation id="134095A8851DC327DA7DA792ABC07799" box="[856,950,1729,1755]" captionStart="FIGURES 170176" captionStartId="54.[151,264,1943,1967]" captionTargetBox="[282,1306,182,1919]" captionTargetId="figure-17@54.[282,1306,181,1919]" captionTargetPageId="54" captionText="FIGURES 170176. Hoplandrothrips flavicornis sp. n., female. (170) Head and prothorax; (171) antenna; (172) antennal segments IIIIV; (173) metanotum; (174) pelta; (175) sub-basal wing setae; (176) tube." figureDoi="http://doi.org/10.5281/zenodo.13211401" httpUri="https://zenodo.org/record/13211401/files/figure.png" pageId="21" pageNumber="43">Fig. 176</figureCitation>
) about 0.6 times as long as head, 1.92 times as long as wide in
<typeStatus id="54C0378F851DC327D842A7B6A9BB77BD" box="[359,461,1765,1791]" pageId="21" pageNumber="43" type="holotype">holotype</typeStatus>
.
<collectorName id="268EECFB851DC327D8F3A7B6AA4D77BD" box="[470,571,1765,1791]" pageId="21" pageNumber="43">Terminal</collectorName>
setae much longer than tube
</materialsCitation>
.
</paragraph>
<paragraph id="8BC4892D851DC327D9E2A65AACD97695" blockId="21.[151,1437,1225,2008]" pageId="21" pageNumber="43">
<emphasis id="B90F553F851DC327D9E2A65AA9007661" bold="true" box="[199,374,1801,1827]" pageId="21" pageNumber="43">Measurements</emphasis>
(
<typeStatus id="54C0378F851DC327D8A0A65AA99F7661" box="[389,489,1801,1827]" pageId="21" pageNumber="43" type="holotype">holotype</typeStatus>
female in μm). Body length about 2700 (distended). Head length 265, from anterior margin of eyes 240, width across eyes 175, maximum width across cheeks 185; eyes length 88, width 5558; postocular setae 8890. Antenna total length about 500, segments IVIII length (width) as follows: 50 (48), 53 (35), 73 (37), 72 (37), 65 (31), 62 (27), 55 (22), 43 (12). Pronotum length 179, width 280. Setae on prothorax: am 28, aa 60, ml 70, pa 70, epim 6567, cox 50. Fore wing length 940. Sub-basal wing setae:
<geoCoordinate id="EE4FEFEA851DC327DD1EA6CAAC0B76F6" box="[1083,1149,1945,1972]" degrees="1" direction="south" minutes="50" orientation="latitude" pageId="21" pageNumber="43" precision="925" value="-1.8333333">S1 50</geoCoordinate>
,
<geoCoordinate id="EE4FEFEA851DC327DDACA6CAACBF76F6" box="[1161,1225,1945,1972]" degrees="2" direction="south" minutes="58" orientation="latitude" pageId="21" pageNumber="43" precision="925" value="-2.9666667">S2 58</geoCoordinate>
60,
<geoCoordinate id="EE4FEFEA851DC327DC25A6CAAD3276F6" box="[1280,1348,1945,1972]" degrees="3" direction="south" minutes="50" orientation="latitude" pageId="21" pageNumber="43" precision="925" value="-3.8333333">S3 50</geoCoordinate>
. Tergite IX setae: S1 160170, S2 180185. Tube length 160, maximum width 83; terminal setae 230.
</paragraph>
<paragraph id="8BC4892D851EC324D9E2A1C4A9F371F3" blockId="22.[151,1437,151,429]" box="[199,389,151,177]" pageId="22" pageNumber="44">
<emphasis id="B90F553F851EC324D9E2A1C4A97771F3" box="[199,257,151,177]" italics="true" pageId="22" pageNumber="44">Male</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C361DAA6851EC324D9E2A1E8AC9171BB" pageId="22" pageNumber="44" type="materials_examined">
<paragraph id="8BC4892D851EC324D9E2A1E8AC9171BB" blockId="22.[151,1437,151,429]" pageId="22" pageNumber="44">
<emphasis id="B90F553F851EC324D9E2A1E8A93B7196" bold="true" box="[199,333,186,213]" pageId="22" pageNumber="44">Type series</emphasis>
.
<materialsCitation id="3B138370851EC324D878A1E8ADEE7197" box="[349,1432,186,213]" collectingDate="1976-07-26" country="Malaysia" location="Peninsular" municipality="Tapah" pageId="22" pageNumber="44" specimenCount="1" typeStatus="holotype">
<typeStatus id="54C0378F851EC324D878A1E8A9BC7197" box="[349,458,187,213]" pageId="22" pageNumber="44" type="holotype">Holotype</typeStatus>
: macropterous female,
<emphasis id="B90F553F851EC324DBF7A1E9ABBE7196" bold="true" box="[722,968,186,212]" pageId="22" pageNumber="44">
<location id="8EA4DFF6851EC324DBF7A1E9AB257196" LSID="urn:lsid:plazi:treatment:03D2383B851DC324D9B2A50EAD6D70EF:8EA4DFF6851EC324DBF7A1E9AB257196" box="[722,851,186,212]" country="Malaysia" municipality="Tapah" name="Peninsular" pageId="22" pageNumber="44">Peninsular</location>
<collectingCountry id="F36CC9BD851EC324DA7EA1E9ABBE7196" box="[859,968,186,212]" name="Malaysia" pageId="22" pageNumber="44">Malaysia</collectingCountry>
</emphasis>
,
<collectingMunicipality id="6BA01357851EC324DAF2A1E8AC687197" box="[983,1054,187,213]" pageId="22" pageNumber="44">Tapah</collectingMunicipality>
, on dead leaves,
<date id="FFC5AFED851EC324DDC7A1E8AD127197" box="[1250,1380,187,213]" pageId="22" pageNumber="44" value="1976-07-26">
<collectingDate id="EF815605851EC324DDC7A1E8AD127197" box="[1250,1380,187,213]" pageId="22" pageNumber="44" value="1976-07-26">26.vii.1976</collectingDate>
</date>
, SO
</materialsCitation>
.
<materialsCitation id="3B138370851EC324D9B2A18CAC9571BB" box="[151,1251,222,249]" collectingDate="1976-07-27" country="Malaysia" location="Peninsular" municipality="Tapah" pageId="22" pageNumber="44" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<typeStatus id="54C0378F851EC324D9B2A18CA88B71BB" box="[151,253,223,249]" pageId="22" pageNumber="44" type="paratype">Paratype</typeStatus>
:
<emphasis id="B90F553F851EC324D82DA18DA98B71BA" bold="true" box="[264,509,222,248]" pageId="22" pageNumber="44">
<location id="8EA4DFF6851EC324D82DA18DA9FF71BA" LSID="urn:lsid:plazi:treatment:03D2383B851DC324D9B2A50EAD6D70EF:8EA4DFF6851EC324D82DA18DA9FF71BA" box="[264,393,222,248]" country="Malaysia" municipality="Tapah" name="Peninsular" pageId="22" pageNumber="44">Peninsular</location>
<collectingCountry id="F36CC9BD851EC324D8B5A18DA98B71BA" box="[400,509,222,248]" name="Malaysia" pageId="22" pageNumber="44">Malaysia</collectingCountry>
</emphasis>
, NE
<quantity id="4C8324C8851EC324DB12A18CAA0171BB" box="[567,631,223,249]" metricMagnitude="4" metricUnit="m" metricValue="1.3" pageId="22" pageNumber="44" unit="km" value="13.0">13km</quantity>
from
<collectingMunicipality id="6BA01357851EC324DB9FA18CAB7771BB" box="[698,769,223,249]" pageId="22" pageNumber="44">Tapah</collectingMunicipality>
,
<specimenCount id="9D7D42A4851EC324DA29A18CAB0671BB" box="[780,880,223,249]" count="1" pageId="22" pageNumber="44" type="female">1 female</specimenCount>
, on dead leaves,
<date id="FFC5AFED851EC324DD0BA18CACC671BB" box="[1070,1200,223,249]" pageId="22" pageNumber="44" value="1976-07-27">
<collectingDate id="EF815605851EC324DD0BA18CACC671BB" box="[1070,1200,223,249]" pageId="22" pageNumber="44" value="1976-07-27">27.vii.1976</collectingDate>
</date>
, SO
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C361DAA6851EC324D9E2A050AD6D70EF" pageId="22" pageNumber="44" type="discussion">
<paragraph id="8BC4892D851EC324D9E2A050AD6D70EF" blockId="22.[151,1437,151,429]" pageId="22" pageNumber="44">
<emphasis id="B90F553F851EC324D9E2A050A942705F" bold="true" box="[199,308,259,285]" pageId="22" pageNumber="44">Remarks</emphasis>
.
<taxonomicName id="4C7BF2AE851EC324D864A050A9AC705F" authorityName="Okajima &amp; Masumoto" authorityYear="2024" box="[321,474,259,285]" class="Insecta" family="Phlaeothripidae" genus="Hoplandrothrips" kingdom="Animalia" order="Thysanoptera" pageId="22" pageNumber="44" phylum="Arthropoda" rank="species" species="flavicornis">
<emphasis id="B90F553F851EC324D864A050A9AC705F" box="[321,474,259,285]" italics="true" pageId="22" pageNumber="44">H. flavicornis</emphasis>
</taxonomicName>
is somewhat similar to
<taxonomicName id="4C7BF2AE851EC324DBFAA050ABFC705F" authorityName="Okajima &amp; Masumoto" authorityYear="2024" box="[735,906,259,285]" class="Insecta" family="Phlaeothripidae" genus="Hoplandrothrips" kingdom="Animalia" order="Thysanoptera" pageId="22" pageNumber="44" phylum="Arthropoda" rank="species" species="graminicola" status="sp. n.">
<emphasis id="B90F553F851EC324DBFAA050ABFC705F" box="[735,906,259,285]" italics="true" pageId="22" pageNumber="44">H. graminicola</emphasis>
</taxonomicName>
<emphasis id="B90F553F851EC324DAAAA050ABB8705F" bold="true" box="[911,974,259,285]" pageId="22" pageNumber="44">
<taxonomicNameLabel id="A23CE844851EC324DAAAA050ABB8705F" box="[911,974,259,285]" pageId="22" pageNumber="44" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
described below from
<collectingCountry id="F36CC9BD851EC324DDEEA050AD59705F" box="[1227,1327,259,285]" name="Thailand" pageId="22" pageNumber="44">Thailand</collectingCountry>
in having the posteromarginal setae on abdominal tergite IX long and pointed and the head elongated with rather short maxillary stylets. However, it can easily be distinguished from the latter by the paler antennal segment III, pedicellate antennal segment VIII, and the sub-basal wing setae S3 expanded. Unfortunately, the prosternal sclerites of this species could not be observed, because both the
<typeStatus id="54C0378F851EC324DB3DA0C0AA0C70EF" box="[536,634,403,429]" pageId="22" pageNumber="44" type="holotype">holotype</typeStatus>
and
<typeStatus id="54C0378F851EC324DB94A0C0AB6770EF" box="[689,785,403,429]" pageId="22" pageNumber="44" type="paratype">paratype</typeStatus>
females were not macerated in KOH solution.
</paragraph>
</subSubSection>
</treatment>
</document>