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<emphasis id="B96334101169FFE9E3DB66D7FE1128C0" bold="true" box="[151,411,1491,1518]" italics="true" pageId="6" pageNumber="99">Geothelphusa takuan</emphasis>
<bibRefCitation id="EF8695F31169FFE9E2EF66D7FD1F28C0" author="Shy, J. - Y. &amp; Ng, P. K. L. &amp; Yu, H. - P." box="[419,661,1491,1518]" pageId="6" pageNumber="99" pagination="781 - 846" refId="ref8705" refString="Shy, J. - Y., Ng, P. K. L. &amp; Yu, H. - P. (1994) Crabs of the genus Geothelphusa Stimpson, 1858 (Crustacea: Decapoda: Brachyura: Potamidae) from Taiwan, with descriptions of 25 new species. Raffles Bulletin of Zoology, 42, 781 - 846." type="journal article" year="1994">Shy, Ng &amp; Yu, 1994</bibRefCitation>
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(
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)
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<taxonomicName id="4C1793811169FFE9E3DB6538FDF22B7A" ID-CoL="3FT4R" authority="Shy, Ng &amp; Yu, 1994: 823" authorityName="Shy, Ng &amp; Yu" authorityPageNumber="823" authorityYear="1994" box="[151,632,1596,1621]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="99" phylum="Arthropoda" rank="species" species="takuan">
<emphasis id="B96334101169FFE9E3DB6538FEE52B7A" box="[151,367,1596,1620]" italics="true" pageId="6" pageNumber="99">Geothelphusa takuan</emphasis>
<bibRefCitation id="EF8695F31169FFE9E23A6538FDF22B7A" author="Shy, J. - Y. &amp; Ng, P. K. L. &amp; Yu, H. - P." box="[374,632,1596,1621]" pageId="6" pageNumber="99" pagination="781 - 846" refId="ref8705" refString="Shy, J. - Y., Ng, P. K. L. &amp; Yu, H. - P. (1994) Crabs of the genus Geothelphusa Stimpson, 1858 (Crustacea: Decapoda: Brachyura: Potamidae) from Taiwan, with descriptions of 25 new species. Raffles Bulletin of Zoology, 42, 781 - 846." type="journal article" year="1994">Shy, Ng &amp; Yu, 1994: 823</bibRefCitation>
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, fig. 20af [
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locality: Daguan, Fusing,
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<bibRefCitation id="EF8695F31169FFE9E7D66538FAA42B7A" author="Dai, A. - Y." box="[1178,1326,1596,1620]" pageId="6" pageNumber="99" refId="ref6531" refString="Dai, A. - Y. (1999) Fauna Sinica. Arthropoda: Crustacea: Malacostraca: Decapoda: Parathelphusidae, Potamidae. Science Press, Beijing, 501 pp., 30 pls. [in Chinese]" type="book" year="1999">Dai 1999: 387</bibRefCitation>
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;
<treatmentCitation id="0AB6CE131169FFE9E6756538FEA92B5D" author="Shy, J. - Y. &amp; Yu, H. - P." page="75" pageId="6" pageNumber="99" year="1999">
<bibRefCitation id="EF8695F31169FFE9E6756538FEA92B5D" author="Shy, J. - Y. &amp; Yu, H. - P." pageId="6" pageNumber="99" refId="ref8898" refString="Shy, J. - Y. &amp; Yu, H. - P. (1999) Freshwater Crabs of Taiwan. National Museum of Marine Biology / Aquarium, Pingtung, ii + 114 pp. [in Chinese]" type="book" year="1999">Shy &amp; Yu 1999: 75</bibRefCitation>
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, 2 un-numbered figs.;
<treatmentCitation id="0AB6CE131169FFE9E146655FFD4D2B5D" author="Ng, P. K. L. &amp; Wang, C. - H. &amp; Ho, P. - H. &amp; Shih, H. - T." box="[522,711,1627,1651]" page="51" pageId="6" pageNumber="99" year="2001">
<bibRefCitation id="EF8695F31169FFE9E146655FFD4D2B5D" author="Ng, P. K. L. &amp; Wang, C. - H. &amp; Ho, P. - H. &amp; Shih, H. - T." box="[522,711,1627,1651]" pageId="6" pageNumber="99" pagination="1 - 86" refId="ref8049" refString="Ng, P. K. L., Wang, C. - H., Ho, P. - H. &amp; Shih, H. - T. (2001) An annotated checklist of brachyuran crabs from Taiwan (Crustacea: Decapoda). National Taiwan Museum Special Publication Series, 11, 1 - 86." type="journal article" year="2001">
Ng
<emphasis id="B96334101169FFE9E17C6558FDD42B5D" box="[560,606,1627,1651]" italics="true" pageId="6" pageNumber="99">et al</emphasis>
. 2001: 51
</bibRefCitation>
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;
<treatmentCitation id="0AB6CE131169FFE9E19E655FFC372B5D" author="Chen, W. - J. &amp; Cheng, J. - H. &amp; Chen, T. - I. &amp; Hsu, M. J." box="[722,957,1627,1651]" page="881" pageId="6" pageNumber="99" year="2007">
<bibRefCitation id="EF8695F31169FFE9E19E655FFC372B5D" author="Chen, W. - J. &amp; Cheng, J. - H. &amp; Chen, T. - I. &amp; Hsu, M. J." box="[722,957,1627,1651]" pageId="6" pageNumber="99" pagination="861 - 889" refId="ref5996" refString="Chen, W. - J., Cheng, J. - H., Chen, T. - I. &amp; Hsu, M. J. (2007 a) A comparison of the micromorphology of the G 1 of freshwater crabs of the genus Geothelphusa (Brachyura, Potamidae) from Taiwan. Crustaceana, 80, 861 - 889. https: // doi. org / 10.1163 / 156854007781363105" type="journal article" year="2007">
Chen
<emphasis id="B96334101169FFE9E0436558FCB62B5D" box="[783,828,1627,1651]" italics="true" pageId="6" pageNumber="99">et al</emphasis>
. 2007a: 881
</bibRefCitation>
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, fig. 8A1A3;
<treatmentCitation id="0AB6CE131169FFE9E71B655FFAC92B5D" author="Chen, W. - J. &amp; Hsu, M. J. &amp; Cheng, J. - H." box="[1111,1347,1627,1651]" page="906" pageId="6" pageNumber="99" year="2007">
<bibRefCitation id="EF8695F31169FFE9E71B655FFAC92B5D" author="Chen, W. - J. &amp; Hsu, M. J. &amp; Cheng, J. - H." box="[1111,1347,1627,1651]" pageId="6" pageNumber="99" pagination="897 - 907" refId="ref6072" refString="Chen, W. - J., Hsu, M. J. &amp; Cheng, J. - H. (2007 b) A new species of alpine freshwater crab (Brachyura: Potamidae) from Siangyang National Forest Recreation area, Taiwan. Crustaceana, 80, 897 - 907. https: // doi. org / 10.1163 / 156854007781681292" type="journal article" year="2007">
Chen
<emphasis id="B96334101169FFE9E7DF6558FB4B2B5D" box="[1171,1217,1627,1651]" italics="true" pageId="6" pageNumber="99">et al</emphasis>
. 2007b: 906
</bibRefCitation>
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, fig. 4B;
<treatmentCitation id="0AB6CE131169FFE9E38B657EFE292BBC" author="Shy, J. - Y. &amp; Lee, P. - W." box="[199,419,1658,1682]" page="184" pageId="6" pageNumber="99" year="2009">
<bibRefCitation id="EF8695F31169FFE9E38B657EFE292BBC" author="Shy, J. - Y. &amp; Lee, P. - W." box="[199,419,1658,1682]" pageId="6" pageNumber="99" refId="ref8667" refString="Shy, J. - Y. &amp; Lee, P. - W. (2009) Guide to the Taiwanese Freshwater Crabs. Morning Star Publishing, Taipei, 221 pp. [in Chinese]" type="book" year="2009">Shy &amp; Lee 2009: 184</bibRefCitation>
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, 8 un-numbered figs.; Shih
<emphasis id="B96334101169FFE9E1F1657EFD632BBC" box="[701,745,1658,1682]" italics="true" pageId="6" pageNumber="99">et al</emphasis>
. 2011: 461;
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<bibRefCitation id="EF8695F31169FFE9E024657EFBA92BBC" author="Ng, P. K. L. &amp; Shih, H. - T. &amp; Ho, P. - H. &amp; Wang, C. - H." box="[872,1059,1658,1682]" pageId="6" pageNumber="99" pagination="1 - 208" refId="ref7877" refString="Ng, P. K. L., Shih, H. - T., Ho, P. - H. &amp; Wang, C. - H. (2017) An updated annotated checklist of brachyuran crabs from Taiwan (Crustacea: Decapoda). Journal of the National Taiwan Museum, 70 (3 / 4), 1 - 208. https: // doi. org / 10.6532 / JNTM. 201712 _ 70 (3; 4). 01" type="journal article" year="2017">
Ng
<emphasis id="B96334101169FFE9E0C2657EFC302BBC" box="[910,954,1658,1682]" italics="true" pageId="6" pageNumber="99">et al</emphasis>
. 2017: 81
</bibRefCitation>
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;
<treatmentCitation id="0AB6CE131169FFE9E761657EFB762BBF" author="Shy, J. - Y. &amp; Shih, H. - T. &amp; Ng, P. K. L." box="[1069,1276,1657,1682]" page="154" pageId="6" pageNumber="99" year="2020">
<bibRefCitation id="EF8695F31169FFE9E761657EFB762BBF" author="Shy, J. - Y. &amp; Shih, H. - T. &amp; Ng, P. K. L." box="[1069,1276,1657,1682]" pageId="6" pageNumber="99" refId="ref8837" refString="Shy, J. - Y., Shih, H. - T. &amp; Ng, P. K. L. (2020) Crustacean Fauna of Taiwan: Brachyuran Crabs. Vol. III. Freshwater Crabs - Potamidae, Gecarcinucidae. National Penghu University of Science and Technology, Penghu, 26 + 232 pp." type="book" year="2020">
Shy
<emphasis id="B96334101169FFE9E717657EFB022BBC" box="[1115,1160,1658,1682]" italics="true" pageId="6" pageNumber="99">et al</emphasis>
. 2020: 154
</bibRefCitation>
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, figs. 119121.
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</paragraph>
<paragraph id="8BA8E8021169FFE9E3DB659CFACC2BE1" blockId="6.[151,1436,1596,1743]" pageId="6" pageNumber="99">
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<treatmentCitation id="0AB6CE131169FFE9E3DB659CFDDD2B9E" author="Tan, S. H. &amp; Liu, H. - C." box="[151,599,1688,1712]" page="288" pageId="6" pageNumber="99" year="1998">
<taxonomicName id="4C1793811169FFE9E3DB659CFDDD2B9E" ID-CoL="3FT3P" authority="Tan &amp; Liu, 1998: 288" authorityName="Tan &amp; Liu" authorityPageNumber="288" authorityYear="1998" box="[151,599,1688,1712]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="99" phylum="Arthropoda" rank="species" species="hirsuta">
<emphasis id="B96334101169FFE9E3DB659CFEF92B9E" box="[151,371,1688,1712]" italics="true" pageId="6" pageNumber="99">Geothelphusa hirsuta</emphasis>
<bibRefCitation id="EF8695F31169FFE9E234659DFDDD2B9E" author="Tan, S. H. &amp; Liu, H. - C." box="[376,599,1688,1712]" pageId="6" pageNumber="99" pagination="286 - 290" refId="ref9073" refString="Tan, S. H. &amp; Liu, H. - C. (1998) Two new species of Geothelphusa (Decapoda: Brachyura: Potamidae) from Taiwan. Zoological Studies, 37, 286 - 290." type="journal article" year="1998">Tan &amp; Liu, 1998: 288</bibRefCitation>
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, fig. 2AE [
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locality: Tai-an,
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,
<collectingCountry id="F300A8921169FFE9E702659CFB142B9E" box="[1102,1182,1688,1712]" name="Taiwan" pageId="6" pageNumber="99">Taiwan</collectingCountry>
];
<treatmentCitation id="0AB6CE131169FFE9E7E2659DFAE02B9E" author="Ng, P. K. L. &amp; Wang, C. - H. &amp; Ho, P. - H. &amp; Shih, H. - T." box="[1198,1386,1688,1713]" page="50" pageId="6" pageNumber="99" year="2001">
<bibRefCitation id="EF8695F31169FFE9E7E2659DFAE02B9E" author="Ng, P. K. L. &amp; Wang, C. - H. &amp; Ho, P. - H. &amp; Shih, H. - T." box="[1198,1386,1688,1713]" pageId="6" pageNumber="99" pagination="1 - 86" refId="ref8049" refString="Ng, P. K. L., Wang, C. - H., Ho, P. - H. &amp; Shih, H. - T. (2001) An annotated checklist of brachyuran crabs from Taiwan (Crustacea: Decapoda). National Taiwan Museum Special Publication Series, 11, 1 - 86." type="journal article" year="2001">
Ng
<emphasis id="B96334101169FFE9E798659DFA8B2B9E" box="[1236,1281,1688,1712]" italics="true" pageId="6" pageNumber="99">et al</emphasis>
. 2001: 50
</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AB6CE131169FFE9E639659CFEFC2BE1" author="Shy, J. - Y. &amp; Lee, P. - W." page="137" pageId="6" pageNumber="99" year="2009">
<bibRefCitation id="EF8695F31169FFE9E639659CFEFC2BE1" author="Shy, J. - Y. &amp; Lee, P. - W." pageId="6" pageNumber="99" refId="ref8667" refString="Shy, J. - Y. &amp; Lee, P. - W. (2009) Guide to the Taiwanese Freshwater Crabs. Morning Star Publishing, Taipei, 221 pp. [in Chinese]" type="book" year="2009">Shy &amp; Lee 2009: 137</bibRefCitation>
</treatmentCitation>
, 7 un-numbered figs.; Shih
<emphasis id="B96334101169FFE9E1C365BCFD362BE1" box="[655,700,1719,1743]" italics="true" pageId="6" pageNumber="99">et al</emphasis>
. 2011: 461;
<treatmentCitation id="0AB6CE131169FFE9E07765B3FC7F2BE1" author="Ng, P. K. L. &amp; Shih, H. - T. &amp; Ho, P. - H. &amp; Wang, C. - H." box="[827,1013,1719,1743]" page="79" pageId="6" pageNumber="99" year="2017">
<bibRefCitation id="EF8695F31169FFE9E07765B3FC7F2BE1" author="Ng, P. K. L. &amp; Shih, H. - T. &amp; Ho, P. - H. &amp; Wang, C. - H." box="[827,1013,1719,1743]" pageId="6" pageNumber="99" pagination="1 - 208" refId="ref7877" refString="Ng, P. K. L., Shih, H. - T., Ho, P. - H. &amp; Wang, C. - H. (2017) An updated annotated checklist of brachyuran crabs from Taiwan (Crustacea: Decapoda). Journal of the National Taiwan Museum, 70 (3 / 4), 1 - 208. https: // doi. org / 10.6532 / JNTM. 201712 _ 70 (3; 4). 01" type="journal article" year="2017">
Ng
<emphasis id="B96334101169FFE9E02C65BCFC062BE1" box="[864,908,1719,1743]" italics="true" pageId="6" pageNumber="99">et al</emphasis>
. 2017: 79
</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AB6CE131169FFE9E74C65B3FB482BE1" author="Shy, J. - Y. &amp; Shih, H. - T. &amp; Ng, P. K. L." box="[1024,1218,1719,1743]" page="82" pageId="6" pageNumber="99" year="2020">
<bibRefCitation id="EF8695F31169FFE9E74C65B3FB482BE1" author="Shy, J. - Y. &amp; Shih, H. - T. &amp; Ng, P. K. L." box="[1024,1218,1719,1743]" pageId="6" pageNumber="99" refId="ref8837" refString="Shy, J. - Y., Shih, H. - T. &amp; Ng, P. K. L. (2020) Crustacean Fauna of Taiwan: Brachyuran Crabs. Vol. III. Freshwater Crabs - Potamidae, Gecarcinucidae. National Penghu University of Science and Technology, Penghu, 26 + 232 pp." type="book" year="2020">
Shy
<emphasis id="B96334101169FFE9E76165BCFBD02BE1" box="[1069,1114,1719,1743]" italics="true" pageId="6" pageNumber="99">et al</emphasis>
. 2020: 82
</bibRefCitation>
</treatmentCitation>
, figs. 6668.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C30DBB891169FFE8E3DB65F9FDFE28C0" lastPageId="7" lastPageNumber="100" pageId="6" pageNumber="99" type="materials_examined">
<paragraph id="8BA8E8021169FFE8E3DB65F9FA0B2C33" blockId="6.[151,1437,1789,2032]" lastBlockId="7.[151,1436,151,285]" lastPageId="7" lastPageNumber="100" pageId="6" pageNumber="99">
<emphasis id="B96334101169FFE9E3DB65F9FEF22A39" bold="true" box="[151,376,1789,1815]" pageId="6" pageNumber="99">Material examined</emphasis>
.
<emphasis id="B96334101169FFE9E2C965F9FE402A39" bold="true" box="[389,458,1789,1815]" pageId="6" pageNumber="99">Types</emphasis>
:
<materialsCitation id="3B7FE25F1169FFE9E29465F9FB1A2A15" ID-GBIF-Occurrence="3395042311" accessionNumber="F10205" collectingDate="1992-11-01" collectionCode="NTOU" collectorName="J.-Y. Shy &amp; W.-L. Tsay" country="Taiwan" county="Taoyuan Co." location="Taoyuan Co." pageId="6" pageNumber="99" specimenCount="1" typeStatus="holotype">
<typeStatus id="54AC56A01169FFE9E29465F9FDB02A39" box="[472,570,1789,1815]" pageId="6" pageNumber="99" type="holotype">holotype</typeStatus>
male (
<collectionCode id="ED0670C71169FFE9E1CF65F9FD582A39" box="[643,722,1789,1815]" country="Taiwan" httpUri="http://grbio.org/cool/be15-j6k8" name="Institute of Marine Biology, National Taiwan Ocean University" pageId="6" pageNumber="99">NTOU</collectionCode>
F10205),
<collectorName id="26E28DD41169FFE9E00865FAFC112A36" box="[836,923,1790,1816]" pageId="6" pageNumber="99">Daguan</collectorName>
(=
<collectorName id="26E28DD41169FFE9E0F365F9FB9C2A36" box="[959,1046,1789,1816]" pageId="6" pageNumber="99">Takuan</collectorName>
),
<collectorName id="26E28DD41169FFE9E76B65F9FBF92A39" box="[1063,1139,1789,1815]" pageId="6" pageNumber="99">Fusing</collectorName>
(=
<collectorName id="26E28DD41169FFE9E7D465F9FB7D2A36" box="[1176,1271,1789,1816]" pageId="6" pageNumber="99">Fuhsing</collectorName>
),
<collectingCounty id="62C9908E1169FFE9E64465FAFA122A39" box="[1288,1432,1789,1816]" pageId="6" pageNumber="99">Taoyuan Co.</collectingCounty>
,
<collectingCountry id="F300A8921169FFE9E3DB6425FF662A15" box="[151,236,1825,1851]" name="Taiwan" pageId="6" pageNumber="99">Taiwan</collectingCountry>
, coll.
<collectorName id="26E28DD41169FFE9E27C6426FE1C2A15" box="[304,406,1825,1852]" pageId="6" pageNumber="99">J.-Y. Shy</collectorName>
&amp;
<collectorName id="26E28DD41169FFE9E2F56426FDBC2A12" box="[441,566,1826,1852]" pageId="6" pageNumber="99">W.-L. Tsay</collectorName>
,
<date id="FFA9CEC21169FFE9E10D6425FD472A15" box="[577,717,1825,1852]" pageId="6" pageNumber="99" value="1992-11-01">
<collectingDate id="EFED372A1169FFE9E10D6425FD472A15" box="[577,717,1825,1852]" pageId="6" pageNumber="99" value="1992-11-01">1 Nov. 1992</collectingDate>
</date>
.
<taxonomicName id="4C1793811169FFE9E1956425FB1A2A15" authority="Tan &amp; Liu, 1998" authorityName="Tan &amp; Liu" authorityYear="1998" box="[729,1168,1825,1852]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="99" phylum="Arthropoda" rank="species" species="hirsuta">
<emphasis id="B96334101169FFE9E1956425FC402A15" box="[729,970,1825,1851]" italics="true" pageId="6" pageNumber="99">Geothelphusa hirsuta</emphasis>
<bibRefCitation id="EF8695F31169FFE9E09D6426FB1A2A15" author="Tan, S. H. &amp; Liu, H. - C." box="[977,1168,1825,1852]" pageId="6" pageNumber="99" pagination="286 - 290" refId="ref9073" refString="Tan, S. H. &amp; Liu, H. - C. (1998) Two new species of Geothelphusa (Decapoda: Brachyura: Potamidae) from Taiwan. Zoological Studies, 37, 286 - 290." type="journal article" year="1998">Tan &amp; Liu, 1998</bibRefCitation>
</taxonomicName>
</materialsCitation>
:
<materialsCitation id="3B7FE25F1169FFE9E7D76425FD182AAD" ID-GBIF-Occurrence="3395042306" collectingDate="1993-11-13" collectionCode="TMCD" collectorName="H. - C. Liu" country="Taiwan" county="Others" elevation="2060" latitude="24.708029" location="Lalashan" longLatPrecision="1" longitude="121.43589" municipality="Daguanshan" pageId="6" pageNumber="99" specimenCode="TMCD 3285, TMCD 3292" specimenCount="7" specimenCount-female="3" specimenCount-male="4" stateProvince="Taoyuan" typeStatus="holotype">
<typeStatus id="54AC56A01169FFE9E7D76425FB772A15" box="[1179,1277,1825,1851]" pageId="6" pageNumber="99" type="holotype">holotype</typeStatus>
male (
<specimenCode id="DBB140791169FFE9E6056425FF592A4E" collectionCode="TMCD" pageId="6" pageNumber="99">TMCD 3285</specimenCode>
),
<specimenCount id="9D11238B1169FFE9E3AA6441FE1D2A71" box="[230,407,1861,1888]" pageId="6" pageNumber="99" type="male" typeStatus="paratype">1 paratype male</specimenCount>
(
<specimenCode id="DBB140791169FFE9E2E96441FDB02A71" box="[421,570,1861,1887]" collectionCode="TMCD" pageId="6" pageNumber="99">TMCD 3292</specimenCode>
),
<collectorName id="26E28DD41169FFE9E1006441FD212A71" box="[588,683,1861,1887]" pageId="6" pageNumber="99">Guanwu</collectorName>
(=
<collectorName id="26E28DD41169FFE9E19D6442FCBE2A4E" box="[721,820,1862,1888]" pageId="6" pageNumber="99">Kuanwu</collectorName>
),
<collectorName id="26E28DD41169FFE9E0096441FC052A71" box="[837,911,1861,1887]" pageId="6" pageNumber="99">Tai-an</collectorName>
,
<collectorName id="26E28DD41169FFE9E0D66441FC6F2A71" box="[922,997,1861,1887]" pageId="6" pageNumber="99">Maioli</collectorName>
(
<geoCoordinate id="EE238EC51169FFE9E0BF6441FB182A4E" box="[1011,1170,1861,1888]" degrees="24" direction="north" minutes="30" orientation="latitude" pageId="6" pageNumber="99" precision="1" seconds="17.5" value="24.504862">24°3017.5”N</geoCoordinate>
<geoCoordinate id="EE238EC51169FFE9E7D46441FACE2A71" box="[1176,1348,1861,1887]" degrees="121" direction="east" minutes="07" orientation="longitude" pageId="6" pageNumber="99" precision="1" seconds="09.9" value="121.119415">121°0709.9”E</geoCoordinate>
), elevation of
<quantity id="4CEF45E71169FFE9E3AA646DFEB62AAA" box="[230,316,1897,1924]" metricMagnitude="3" metricUnit="m" metricValue="2.06" pageId="6" pageNumber="99" unit="m" value="2060.0">
<elevation id="003A0F311169FFE9E3AA646DFEB62AAA" box="[230,316,1897,1924]" metricMagnitude="3" metricUnit="m" metricValue="2.06" pageId="6" pageNumber="99" unit="m" value="2060.0">2060 m</elevation>
</quantity>
, coll.
<collectorName id="26E28DD41169FFE9E231646DFE612AAD" box="[381,491,1897,1923]" pageId="6" pageNumber="99">H.-C. Liu</collectorName>
,
<date id="FFA9CEC21169FFE9E2B9646DFD072AAD" box="[501,653,1897,1924]" pageId="6" pageNumber="99" value="1993-11-13">
<collectingDate id="EFED372A1169FFE9E2B9646DFD072AAD" box="[501,653,1897,1924]" pageId="6" pageNumber="99" value="1993-11-13">13 Nov. 1993</collectingDate>
</date>
.
</materialsCitation>
<materialsCitation id="3B7FE25F1169FFE9E1DB646DFABE2AE2" ID-GBIF-Occurrence="3395042304" collectingDate="1993-11-13" collectionCode="TMCD" collectorName="H.- T. Shih" country="Taiwan" county="Others" elevation="2060" latitude="24.708029" location="Lalashan" longLatPrecision="1" longitude="121.43589" municipality="Daguanshan" pageId="6" pageNumber="99" specimenCode="TMCD 3285, TMCD 3292" specimenCount="7" specimenCount-female="3" specimenCount-male="4" stateProvince="Taoyuan" typeStatus="holotype">
<emphasis id="B96334101169FFE9E1DB646DFD632AAD" bold="true" box="[663,745,1897,1923]" pageId="6" pageNumber="99">
<collectingCounty id="62C9908E1169FFE9E1DB646DFD632AAD" box="[663,745,1897,1923]" pageId="6" pageNumber="99">Others</collectingCounty>
</emphasis>
:
<specimenCount id="9D11238B1169FFE9E1BB646DFCBF2AAA" box="[759,821,1897,1924]" pageId="6" pageNumber="99" type="male">3 ♂♂</specimenCount>
(12.0
<quantity id="4CEF45E71169FFE9E0CF646DFC602AAA" box="[899,1002,1897,1924]" metricMagnitude="-2" metricUnit="m" metricValue="1.6199999999999999" pageId="6" pageNumber="99" unit="mm" value="16.2">16.2 mm</quantity>
),
<specimenCount id="9D11238B1169FFE9E0B0646DFBB02AAA" box="[1020,1082,1897,1924]" pageId="6" pageNumber="99" type="female">3 ♀♀</specimenCount>
(
<quantity id="4CEF45E71169FFE9E704646DFB662AAA" box="[1096,1260,1897,1924]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" metricValueMax="1.59" metricValueMin="1.21" pageId="6" pageNumber="99" unit="mm" value="14.0" valueMax="15.9" valueMin="12.1">12.115.9 mm</quantity>
) (NCHUZOOL 13285),
<collectingMunicipality id="6BCC72781169FFE9E3B86489FEF42A89" box="[244,382,1933,1959]" pageId="6" pageNumber="99">Daguanshan</collectingMunicipality>
(=
<location id="8EC8BED91169FFE9E2E86489FD872A86" LSID="urn:lsid:plazi:treatment:03BE59141169FFE8E3DB66D7FE7A2ACB:8EC8BED91169FFE9E2E86489FD872A86" box="[420,525,1933,1960]" country="Taiwan" county="Others" latitude="24.708029" longLatPrecision="1" longitude="121.43589" municipality="Daguanshan" name="Lalashan" pageId="6" pageNumber="99" stateProvince="Taoyuan">Lalashan</location>
),
<location id="8EC8BED91169FFE9E16C6489FDE52A89" LSID="urn:lsid:plazi:treatment:03BE59141169FFE8E3DB66D7FE7A2ACB:8EC8BED91169FFE9E16C6489FDE52A89" box="[544,623,1933,1959]" country="Taiwan" county="Others" latitude="24.708029" longLatPrecision="1" longitude="121.43589" municipality="Daguanshan" name="Fusing" pageId="6" pageNumber="99" stateProvince="Taoyuan">Fusing</location>
,
<collectingRegion id="49D326E01169FFE9E135648AFD532A86" box="[633,729,1934,1960]" country="Taiwan" name="Taoyuan" pageId="6" pageNumber="99">Taoyuan</collectingRegion>
(
<geoCoordinate id="EE238EC51169FFE9E1A46489FC032A89" box="[744,905,1933,1959]" degrees="24" direction="north" minutes="42" orientation="latitude" pageId="6" pageNumber="99" precision="1" seconds="28.9" value="24.708029">24°4228.9”N</geoCoordinate>
,
<geoCoordinate id="EE238EC51169FFE9E0D86489FBCB2A86" box="[916,1089,1933,1960]" degrees="121" direction="east" minutes="26" orientation="longitude" pageId="6" pageNumber="99" precision="1" seconds="09.2" value="121.43589">121°2609.2”E</geoCoordinate>
), elevation of
<quantity id="4CEF45E71169FFE9E7936489FABF2A86" box="[1247,1333,1933,1960]" metricMagnitude="3" metricUnit="m" metricValue="1.65" pageId="6" pageNumber="99" unit="m" value="1650.0">
<elevation id="003A0F311169FFE9E7936489FABF2A86" box="[1247,1333,1933,1960]" metricMagnitude="3" metricUnit="m" metricValue="1.65" pageId="6" pageNumber="99" unit="m" value="1650.0">1650 m</elevation>
</quantity>
, coll.
<collectorName id="26E28DD41169FFE9E634648AFF602AE5" pageId="6" pageNumber="99">H.- T. Shih</collectorName>
,
<date id="FFA9CEC21169FFE9E3BA64B5FE092AE5" box="[246,387,1969,1996]" pageId="6" pageNumber="99" value="2000-12-04">
<collectingDate id="EFED372A1169FFE9E3BA64B5FE092AE5" box="[246,387,1969,1996]" pageId="6" pageNumber="99" value="2000-12-04">4 Dec. 2000</collectingDate>
</date>
;
<specimenCount id="9D11238B1169FFE9E2C264B5FE472AE2" box="[398,461,1969,1996]" pageId="6" pageNumber="99" type="male">3 ♂♂</specimenCount>
(12.4 × 10.417.4 × 13.0 mm),
<specimenCount id="9D11238B1169FFE9E07A64B5FCEA2AE2" box="[822,864,1969,1996]" pageId="6" pageNumber="99" type="female">1 ♀</specimenCount>
(17.3 ×
<quantity id="4CEF45E71169FFE9E08C64B5FBAC2AE2" box="[960,1062,1969,1996]" metricMagnitude="-2" metricUnit="m" metricValue="1.28" pageId="6" pageNumber="99" unit="mm" value="12.8">12.8 mm</quantity>
) (NCHUZOOL 16435),
</materialsCitation>
<materialsCitation id="3B7FE25F1169FFE9E67064B6FC1D2ADE" ID-GBIF-Occurrence="3395042302" collectingDate="2000-12-04" collectorName="H. - T. Shih" country="Taiwan" elevation="1650" latitude="24.708029" location="Fusing" longLatPrecision="1" longitude="121.43589" municipality="Lalashan" pageId="6" pageNumber="99" specimenCount="4" specimenCount-female="1" specimenCount-male="3" stateProvince="Taoyuan" typeStatus="holotype">
Daguanshan (=
<collectingMunicipality id="6BCC72781169FFE9E3BD64D1FED02ADE" box="[241,346,2005,2032]" pageId="6" pageNumber="99">Lalashan</collectingMunicipality>
),
<location id="8EC8BED91169FFE9E22164D1FE362AC1" LSID="urn:lsid:plazi:treatment:03BE59141169FFE8E3DB66D7FE7A2ACB:8EC8BED91169FFE9E22164D1FE362AC1" box="[365,444,2005,2031]" country="Taiwan" latitude="24.708029" longLatPrecision="1" longitude="121.43589" municipality="Lalashan" name="Fusing" pageId="6" pageNumber="99" stateProvince="Taoyuan">Fusing</location>
,
<collectingRegion id="49D326E01169FFE9E28A64D2FDAC2ADE" box="[454,550,2006,2032]" country="Taiwan" name="Taoyuan" pageId="6" pageNumber="99">Taoyuan</collectingRegion>
(
<geoCoordinate id="EE238EC51169FFE9E17964D1FD5D2AC1" box="[565,727,2005,2031]" degrees="24" direction="north" minutes="42" orientation="latitude" pageId="6" pageNumber="99" precision="1" seconds="28.9" value="24.708029">24°4228.9”N</geoCoordinate>
,
<geoCoordinate id="EE238EC51169FFE9E1AE64D1FC042ADE" box="[738,910,2005,2032]" degrees="121" direction="east" minutes="26" orientation="longitude" pageId="6" pageNumber="99" precision="1" seconds="09.2" value="121.43589">121°2609.2”E</geoCoordinate>
)
</materialsCitation>
,
<materialsCitation id="3B7FE25F1169FFE8E0EE64D1FD372DFB" ID-GBIF-Occurrence="3395042303" collectingDate="2010-09-07" collectorName="Y. - H. Wang" country="Taiwan" elevation="1650" lastPageId="7" lastPageNumber="100" latitude="24.578222" location="Jianshih" longLatPrecision="1" longitude="121.346504" municipality="Smangus" pageId="6" pageNumber="99" specimenCount="5" specimenCount-female="1" specimenCount-male="4" stateProvince="Hsinchu" typeStatus="holotype">
elevation of
<quantity id="4CEF45E71169FFE9E76264D1FB0E2ADE" box="[1070,1156,2005,2032]" metricMagnitude="3" metricUnit="m" metricValue="1.65" pageId="6" pageNumber="99" unit="m" value="1650.0">
<elevation id="003A0F311169FFE9E76264D1FB0E2ADE" box="[1070,1156,2005,2032]" metricMagnitude="3" metricUnit="m" metricValue="1.65" pageId="6" pageNumber="99" unit="m" value="1650.0">1650 m</elevation>
</quantity>
, coll.
<collectorName id="26E28DD41169FFE9E78A64D2FAC02ADE" box="[1222,1354,2006,2032]" pageId="6" pageNumber="99">Y.-H. Wang</collectorName>
<emphasis id="B96334101169FFE9E61C64D2FA0B2AC1" box="[1360,1409,2005,2031]" italics="true" pageId="6" pageNumber="99">et al</emphasis>
.
<date id="FFA9CEC21169FFE8E6C264D2FE812D9F" lastPageId="7" lastPageNumber="100" pageId="6" pageNumber="99" value="2010-09-07">
<collectingDate id="EFED372A1169FFE8E6C264D2FE812D9F" lastPageId="7" lastPageNumber="100" pageId="6" pageNumber="99" value="2010-09-07">7 Sep. 2010</collectingDate>
</date>
;
<specimenCount id="9D11238B1168FFE8E25A6393FEDF2D9F" box="[278,341,151,177]" pageId="7" pageNumber="100" type="male">4 ♂♂</specimenCount>
(13.2 × 10.8 16.1 ×
<quantity id="4CEF45E71168FFE8E1046393FD252D9F" box="[584,687,151,177]" metricMagnitude="-2" metricUnit="m" metricValue="1.21" pageId="7" pageNumber="100" unit="mm" value="12.1">12.1 mm</quantity>
),
<specimenCount id="9D11238B1168FFE8E18E6393FD662D9F" box="[706,748,151,177]" pageId="7" pageNumber="100" type="female">1 ♀</specimenCount>
(16.2 ×
<quantity id="4CEF45E71168FFE8E0076393FC3A2D9F" box="[843,944,151,177]" metricMagnitude="-2" metricUnit="m" metricValue="1.34" pageId="7" pageNumber="100" unit="mm" value="13.4">13.4 mm</quantity>
) (NCHUZOOL 17041),
<collectingMunicipality id="6BCC72781168FFE8E7896393FAA42D9F" box="[1221,1326,151,177]" pageId="7" pageNumber="100">Smangus</collectingMunicipality>
,
<location id="8EC8BED91168FFE8E6756393FA122D9F" LSID="urn:lsid:plazi:treatment:03BE59141169FFE8E3DB66D7FE7A2ACB:8EC8BED91168FFE8E6756393FA122D9F" box="[1337,1432,151,177]" country="Taiwan" latitude="24.578222" longLatPrecision="1" longitude="121.346504" municipality="Smangus" name="Jianshih" pageId="7" pageNumber="100" stateProvince="Hsinchu">Jianshih</location>
,
<collectingRegion id="49D326E01168FFE8E3DB63BFFEDA2DFB" box="[151,336,187,213]" country="Taiwan" name="Hsinchu" pageId="7" pageNumber="100">Hsinchu County</collectingRegion>
(
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,
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elevation of
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, coll.
<collectorName id="26E28DD41168FFE8E74D63BFFBD92DFB" box="[1025,1107,187,213]" pageId="7" pageNumber="100">J.-J. Li</collectorName>
,
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<collectingDate id="EFED372A1168FFE8E72E63BFFB642DFB" box="[1122,1262,187,213]" pageId="7" pageNumber="100" value="2021-04-03">3 Apr. 2021</collectingDate>
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;
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(14.8 × 11.618.4 ×
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),
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(16.5 × 13.320.2 ×
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) (NCHUZOOL 13432),
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,
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,
<collectingRegion id="49D326E01168FFE8E67363DBFF612C33" country="Taiwan" name="Hsinchu" pageId="7" pageNumber="100">Hsinchu County</collectingRegion>
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,
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coll.
<date id="FFA9CEC21168FFE8E2626207FE332C33" box="[302,441,259,285]" pageId="7" pageNumber="100" value="2001-12-01">
<collectingDate id="EFED372A1168FFE8E2626207FE332C33" box="[302,441,259,285]" pageId="7" pageNumber="100" value="2001-12-01">1 Dec. 2001</collectingDate>
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;
<specimenCount id="9D11238B1168FFE8E2896207FE652C33" box="[453,495,259,285]" pageId="7" pageNumber="100" type="male">1 ♂</specimenCount>
(18.7 ×
<quantity id="4CEF45E71168FFE8E1026207FD392C33" box="[590,691,259,285]" metricMagnitude="-2" metricUnit="m" metricValue="1.43" pageId="7" pageNumber="100" unit="mm" value="14.3">14.3 mm</quantity>
) (NCHUZOOL 13431),
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Road,
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,
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.
</paragraph>
<caption id="DF68B88A1168FFE8E3DB67F9FE4F28AF" ID-DOI="http://doi.org/10.5281/zenodo.5606785" ID-Zenodo-Dep="5606785" httpUri="https://zenodo.org/record/5606785/files/figure.png" pageId="7" pageNumber="100" startId="7.[151,250,1277,1301]" targetBox="[172,1347,329,1244]" targetPageId="7">
<paragraph id="8BA8E8021168FFE8E3DB67F9FE4F28AF" blockId="7.[151,1436,1277,1409]" pageId="7" pageNumber="100">
<emphasis id="B96334101168FFE8E3DB67F9FE98283B" bold="true" box="[151,274,1277,1301]" pageId="7" pageNumber="100">FIGURE 5.</emphasis>
A Bayesian Inference (BI) tree of
<taxonomicName id="4C1793811168FFE8E12067F9FCC9283B" authority="Shy &amp; Shih &amp; Mao, 2021" authorityName="Shy &amp; Shih &amp; Mao" authorityYear="2021" box="[620,835,1277,1301]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="100" phylum="Arthropoda" rank="species" species="boreas" status="sp. nov.">
<emphasis id="B96334101168FFE8E12067F9FCC9283B" box="[620,835,1277,1301]" italics="true" pageId="7" pageNumber="100">Geothelphusa boreas</emphasis>
</taxonomicName>
<emphasis id="B96334101168FFE8E00567F9FC13283B" bold="true" box="[841,921,1277,1301]" pageId="7" pageNumber="100">
<taxonomicNameLabel id="A250896B1168FFE8E00567F9FC13283B" box="[841,921,1277,1301]" pageId="7" pageNumber="100" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and related montane species from northern Taiwan based on cytochrome oxidase I (COI) marker. Probability values at the nodes represent the support values for BI and maximum evolution (ME). For haplotype abbreviations, see Table 1. Corresponding numbers of localities in Fig. 1 are in parenthesis behind haplotype abbreviations.
</paragraph>
</caption>
<paragraph id="8BA8E8021168FFE8E38B66ABFDFE28C0" blockId="7.[151,1437,1455,2022]" pageId="7" pageNumber="100">
<emphasis id="B96334101168FFE8E38B66ABFE0328E4" bold="true" box="[199,393,1455,1482]" pageId="7" pageNumber="100">Ecological notes</emphasis>
: In Daguanshan,
<taxonomicName id="4C1793811168FFE8E11866ABFCCE28E7" authorityName="Shy, Ng &amp; Yu" authorityYear="1994" box="[596,836,1455,1481]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="100" phylum="Arthropoda" rank="species" species="takuan">
<emphasis id="B96334101168FFE8E11866ABFCCE28E7" box="[596,836,1455,1481]" italics="true" pageId="7" pageNumber="100">Geothelphusa takuan</emphasis>
</taxonomicName>
is sympatric with
<taxonomicName id="4C1793811168FFE8E75766B4FBE128E7" authorityName="Shy, Ng &amp; Yu" authorityYear="1994" box="[1051,1131,1455,1482]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="100" phylum="Arthropoda" rank="species" species="ilan">
<emphasis id="B96334101168FFE8E75766B4FBE128E7" box="[1051,1131,1455,1482]" italics="true" pageId="7" pageNumber="100">G. ilan</emphasis>
</taxonomicName>
. In Smangus,
<taxonomicName id="4C1793811168FFE8E65D66B4FA0B28E7" authorityName="Shy, Ng &amp; Yu" authorityYear="1994" box="[1297,1409,1455,1482]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="100" phylum="Arthropoda" rank="species" species="takuan">
<emphasis id="B96334101168FFE8E65D66B4FA0B28E7" box="[1297,1409,1455,1482]" italics="true" pageId="7" pageNumber="100">G. takuan</emphasis>
</taxonomicName>
is sympatric with
<taxonomicName id="4C1793811168FFE8E20966D0FE1E28C3" authorityName="Shy, Ng &amp; Yu" authorityYear="1994" box="[325,404,1491,1518]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="100" phylum="Arthropoda" rank="species" species="ilan">
<emphasis id="B96334101168FFE8E20966D0FE1E28C3" box="[325,404,1491,1518]" italics="true" pageId="7" pageNumber="100">G. ilan</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1793811168FFE8E2ED66D0FE7128C3" authorityName="Shy, Shih &amp; Mao" authorityYear="2014" box="[417,507,1491,1518]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="100" phylum="Arthropoda" rank="species" species="cilan">
<emphasis id="B96334101168FFE8E2ED66D0FE7128C3" box="[417,507,1491,1518]" italics="true" pageId="7" pageNumber="100">G. cilan</emphasis>
</taxonomicName>
and
<emphasis id="B96334101168FFE8E17E66D0FDCC28C0" box="[562,582,1492,1518]" italics="true" pageId="7" pageNumber="100">G</emphasis>
. sp.
</paragraph>
</subSubSection>
<subSubSection id="C30DBB891168FFE8E38B66FCFE7A2ACB" pageId="7" pageNumber="100" type="discussion">
<paragraph id="8BA8E8021168FFE8E38B66FCFE7A2ACB" blockId="7.[151,1437,1455,2022]" pageId="7" pageNumber="100">
<emphasis id="B96334101168FFE8E38B66FCFEBE2B3C" bold="true" box="[199,308,1528,1554]" pageId="7" pageNumber="100">Remarks</emphasis>
: Based on the figure in
<bibRefCitation id="EF8695F31168FFE8E10866FCFC8C2B3C" author="Tan, S. H. &amp; Liu, H. - C." box="[580,774,1527,1554]" pageId="7" pageNumber="100" pagination="286 - 290" refId="ref9073" refString="Tan, S. H. &amp; Liu, H. - C. (1998) Two new species of Geothelphusa (Decapoda: Brachyura: Potamidae) from Taiwan. Zoological Studies, 37, 286 - 290." type="journal article" year="1998">Tan &amp; Liu (1998</bibRefCitation>
: fig. 2), the G1 of
<taxonomicName id="4C1793811168FFE8E09B66FCFBC22B3F" authorityName="Tan &amp; Liu" authorityYear="1998" box="[983,1096,1527,1554]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="100" phylum="Arthropoda" rank="species" species="hirsuta">
<emphasis id="B96334101168FFE8E09B66FCFBC22B3F" box="[983,1096,1527,1554]" italics="true" pageId="7" pageNumber="100">G. hirsuta</emphasis>
</taxonomicName>
with the subterminal segment curves outwards, which is different from the inward curved subterminal segment of
<taxonomicName id="4C1793811168FFE8E7166518FA122B18" authority="(Shy et al. 1994)" baseAuthorityName="Shy" baseAuthorityYear="1994" box="[1114,1432,1563,1590]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="100" phylum="Arthropoda" rank="species" species="takuan">
<emphasis id="B96334101168FFE8E7166518FB402B1B" box="[1114,1226,1563,1590]" italics="true" pageId="7" pageNumber="100">G. takuan</emphasis>
(
<bibRefCitation id="EF8695F31168FFE8E797651FFA1A2B18" author="Shy, J. - Y. &amp; Ng, P. K. L. &amp; Yu, H. - P." box="[1243,1424,1563,1590]" pageId="7" pageNumber="100" pagination="781 - 846" refId="ref8705" refString="Shy, J. - Y., Ng, P. K. L. &amp; Yu, H. - P. (1994) Crabs of the genus Geothelphusa Stimpson, 1858 (Crustacea: Decapoda: Brachyura: Potamidae) from Taiwan, with descriptions of 25 new species. Raffles Bulletin of Zoology, 42, 781 - 846." type="journal article" year="1994">
Shy
<emphasis id="B96334101168FFE8E65D6518FAC12B1B" box="[1297,1355,1563,1589]" italics="true" pageId="7" pageNumber="100">et al.</emphasis>
1994
</bibRefCitation>
)
</taxonomicName>
. However, after reexamining on the
<typeStatus id="54AC56A01168FFE8E16E653BFD0E2B77" box="[546,644,1599,1625]" pageId="7" pageNumber="100" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C1793811168FFE8E1E56544FC932B77" authorityName="Tan &amp; Liu" authorityYear="1998" box="[681,793,1599,1626]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="100" phylum="Arthropoda" rank="species" species="hirsuta">
<emphasis id="B96334101168FFE8E1E56544FC932B77" box="[681,793,1599,1626]" italics="true" pageId="7" pageNumber="100">G. hirsuta</emphasis>
</taxonomicName>
(TMCD 3285), we found both the right and left G1s (and G2s) were dissected and stored in small tubes separately. By comparing the right G1 detached from the adult male
<typeStatus id="54AC56A01168FFE8E3DB658CFF7D2B8C" box="[151,247,1672,1698]" pageId="7" pageNumber="100" type="paratype">paratype</typeStatus>
(TMCD 3292), the labels of left and right on the tubes of
<typeStatus id="54AC56A01168FFE8E0C56583FC612B8F" box="[905,1003,1671,1697]" pageId="7" pageNumber="100" type="holotype">holotype</typeStatus>
are confirmed to be actually reversed. That is, the figure and description for the G1 and G
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<bibRefCitation id="EF8695F31168FFE8E05C65A8FC5D2BE8" author="Tan, S. H. &amp; Liu, H. - C." box="[784,983,1707,1734]" pageId="7" pageNumber="100" pagination="286 - 290" refId="ref9073" refString="Tan, S. H. &amp; Liu, H. - C. (1998) Two new species of Geothelphusa (Decapoda: Brachyura: Potamidae) from Taiwan. Zoological Studies, 37, 286 - 290." type="journal article" year="1998">Tan &amp; Liu (1998)</bibRefCitation>
are actually left ones instead. Our reexamination of the right G1s of the
<typeStatus id="54AC56A01168FFE8E15965CBFDFD2BC7" box="[533,631,1743,1769]" pageId="7" pageNumber="100" type="holotype">holotype</typeStatus>
(
<figureCitation id="132CF4871168FFE8E1C465CBFC882BC4" box="[648,770,1743,1770]" captionStart="FIGURE 4" captionStartId="6.[151,250,1349,1373]" captionTargetBox="[279,1306,521,1284]" captionTargetId="figure-193@6.[166,1421,453,1329]" captionTargetPageId="6" captionText="FIGURE 4. Male right G1s of Geothelphusa takuan Shy, Yu &amp; Ng, 1994 (holotype, NTOU F10205; A, B) and G. hirsuta Tan &amp; Liu, 1998 (holotype, TMCD 3285; C, D). A, C, ventral view; B, D, dorsal view. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5606781" httpUri="https://zenodo.org/record/5606781/files/figure.png" pageId="7" pageNumber="100">Fig. 4C, D</figureCitation>
) and
<typeStatus id="54AC56A01168FFE8E00F65D4FC292BC4" box="[835,931,1744,1770]" pageId="7" pageNumber="100" type="paratype">paratype</typeStatus>
of
<taxonomicName id="4C1793811168FFE8E08765D4FBB42BC7" authorityName="Tan &amp; Liu" authorityYear="1998" box="[971,1086,1743,1770]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="100" phylum="Arthropoda" rank="species" species="hirsuta">
<emphasis id="B96334101168FFE8E08765D4FBB42BC7" box="[971,1086,1743,1770]" italics="true" pageId="7" pageNumber="100">G. hirsuta</emphasis>
</taxonomicName>
, found no difference from that of the
<typeStatus id="54AC56A01168FFE8E3AD65F7FEC92A23" box="[225,323,1779,1805]" pageId="7" pageNumber="100" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C1793811168FFE8E22065F0FE512A23" box="[364,475,1779,1806]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="100" phylum="Arthropoda" rank="species" species="takuan">
<emphasis id="B96334101168FFE8E22065F0FE512A23" box="[364,475,1779,1806]" italics="true" pageId="7" pageNumber="100">G. takuan</emphasis>
</taxonomicName>
(NTOU F10205) (
<figureCitation id="132CF4871168FFE8E1FE65F7FCA52A20" box="[690,815,1779,1806]" captionStart="FIGURE 4" captionStartId="6.[151,250,1349,1373]" captionTargetBox="[279,1306,521,1284]" captionTargetId="figure-193@6.[166,1421,453,1329]" captionTargetPageId="6" captionText="FIGURE 4. Male right G1s of Geothelphusa takuan Shy, Yu &amp; Ng, 1994 (holotype, NTOU F10205; A, B) and G. hirsuta Tan &amp; Liu, 1998 (holotype, TMCD 3285; C, D). A, C, ventral view; B, D, dorsal view. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5606781" httpUri="https://zenodo.org/record/5606781/files/figure.png" pageId="7" pageNumber="100">Fig. 4A, B</figureCitation>
). We also find that they differ only in the anterolateral margins, with
<taxonomicName id="4C1793811168FFE8E274641CFE232A1F" authorityName="Tan &amp; Liu" authorityYear="1998" box="[312,425,1815,1842]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="100" phylum="Arthropoda" rank="species" species="hirsuta">
<emphasis id="B96334101168FFE8E274641CFE232A1F" box="[312,425,1815,1842]" italics="true" pageId="7" pageNumber="100">G. hirsuta</emphasis>
</taxonomicName>
having lined with relatively clear granules (
<bibRefCitation id="EF8695F31168FFE8E0DB6413FBCD2A1C" author="Shy, J. - Y. &amp; Shih, H. - T. &amp; Ng, P. K. L." box="[919,1095,1815,1842]" pageId="7" pageNumber="100" refId="ref8837" refString="Shy, J. - Y., Shih, H. - T. &amp; Ng, P. K. L. (2020) Crustacean Fauna of Taiwan: Brachyuran Crabs. Vol. III. Freshwater Crabs - Potamidae, Gecarcinucidae. National Penghu University of Science and Technology, Penghu, 26 + 232 pp." type="book" year="2020">
Shy
<emphasis id="B96334101168FFE8E080641CFB8E2A1F" box="[972,1028,1815,1841]" italics="true" pageId="7" pageNumber="100">et al.</emphasis>
2020
</bibRefCitation>
: fig. 66) and
<taxonomicName id="4C1793811168FFE8E796641CFAC32A1F" box="[1242,1353,1815,1842]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="100" phylum="Arthropoda" rank="species" species="takuan">
<emphasis id="B96334101168FFE8E796641CFAC32A1F" box="[1242,1353,1815,1842]" italics="true" pageId="7" pageNumber="100">G. takuan</emphasis>
</taxonomicName>
having lined with lower granules (
<bibRefCitation id="EF8695F31168FFE8E288643FFDFE2A78" author="Shy, J. - Y. &amp; Shih, H. - T. &amp; Ng, P. K. L." box="[452,628,1851,1878]" pageId="7" pageNumber="100" refId="ref8837" refString="Shy, J. - Y., Shih, H. - T. &amp; Ng, P. K. L. (2020) Crustacean Fauna of Taiwan: Brachyuran Crabs. Vol. III. Freshwater Crabs - Potamidae, Gecarcinucidae. National Penghu University of Science and Technology, Penghu, 26 + 232 pp." type="book" year="2020">
Shy
<emphasis id="B96334101168FFE8E2B46438FDBB2A7B" box="[504,561,1851,1877]" italics="true" pageId="7" pageNumber="100">et al.</emphasis>
2020
</bibRefCitation>
: fig. 119), too small a difference to warrant their recognition as separate taxa. With regard to the genetic similarity for the two species, see “DNA analyses and discussion”. Geographically, although the distance between the
<specimenCount id="9D11238B1168FFE8E16C6480FD082AB0" box="[544,642,1924,1950]" pageId="7" pageNumber="100" type="generic">two type</specimenCount>
localities of
<taxonomicName id="4C1793811168FFE8E0546480FC0D2AB3" box="[792,903,1923,1950]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="100" phylum="Arthropoda" rank="species" species="takuan">
<emphasis id="B96334101168FFE8E0546480FC0D2AB3" box="[792,903,1923,1950]" italics="true" pageId="7" pageNumber="100">G. takuan</emphasis>
</taxonomicName>
(
<figureCitation id="132CF4871168FFE8E0D46487FC542AB0" box="[920,990,1923,1950]" captionStart="FIGURE 1" captionStartId="1.[151,250,1662,1686]" captionTargetBox="[200,1368,750,1619]" captionTargetPageId="1" captionText="FIGURE 1. Collection sites (empty circles) for the Geothelphusa species from northern Taiwan. For locality names, see Table 1. R. = river." figureDoi="http://doi.org/10.5281/zenodo.5606773" httpUri="https://zenodo.org/record/5606773/files/figure.png" pageId="7" pageNumber="100">Fig. 1</figureCitation>
: no. 5) and
<taxonomicName id="4C1793811168FFE8E72B6480FB502AB3" authorityName="Tan &amp; Liu" authorityYear="1998" box="[1127,1242,1923,1950]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="100" phylum="Arthropoda" rank="species" species="hirsuta">
<emphasis id="B96334101168FFE8E72B6480FB502AB3" box="[1127,1242,1923,1950]" italics="true" pageId="7" pageNumber="100">G. hirsuta</emphasis>
</taxonomicName>
(
<figureCitation id="132CF4871168FFE8E7A76487FABB2AB0" box="[1259,1329,1923,1950]" captionStart="FIGURE 1" captionStartId="1.[151,250,1662,1686]" captionTargetBox="[200,1368,750,1619]" captionTargetPageId="1" captionText="FIGURE 1. Collection sites (empty circles) for the Geothelphusa species from northern Taiwan. For locality names, see Table 1. R. = river." figureDoi="http://doi.org/10.5281/zenodo.5606773" httpUri="https://zenodo.org/record/5606773/files/figure.png" pageId="7" pageNumber="100">Fig. 1</figureCitation>
: no. 8) is about
<quantity id="4CEF45E71168FFE8E39064ACFEAF2AEF" box="[220,293,1959,1986]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="7" pageNumber="100" unit="km" value="40.0">40 km</quantity>
, they belongs to the same mountain range, the Hsuehshan Range (
<figureCitation id="132CF4871168FFE8E74D64A3FBCB2AEC" box="[1025,1089,1959,1986]" captionStart="FIGURE 1" captionStartId="1.[151,250,1662,1686]" captionTargetBox="[200,1368,750,1619]" captionTargetPageId="1" captionText="FIGURE 1. Collection sites (empty circles) for the Geothelphusa species from northern Taiwan. For locality names, see Table 1. R. = river." figureDoi="http://doi.org/10.5281/zenodo.5606773" httpUri="https://zenodo.org/record/5606773/files/figure.png" pageId="7" pageNumber="100">Fig. 1</figureCitation>
), and a continuous distribution of a single species is expected.
</paragraph>
</subSubSection>
</treatment>
</document>