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<document id="752116E4E0AF717732FEC116C0FCD8E2" ID-DOI="10.5281/zenodo.6391931" ID-ISSN="1942-1354" ID-Zenodo-Dep="6391931" ID-ZooBank="25B9843D-F0AA-4100-8E8E-DA0030EBE56D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1648504512204" checkinUser="felipe" docAuthor="Skelley, Paul E., Clavijo-Bustos, Julian &amp; Keller, Oliver" docDate="2022" docId="039798726509EE66FF35FC55FA0AF931" docLanguage="en" docName="InsectaMundi.2022.0915.1-14.pdf" docOrigin="Insecta Mundi 2022 (915)" docStyle="DocumentStyle:F9976E9B7F8A094D890004515A7A0592.3:InsectaMundi.2020-.journal_article" docStyleId="F9976E9B7F8A094D890004515A7A0592" docStyleName="InsectaMundi.2020-.journal_article" docStyleVersion="3" docTitle="Termitodius chaki Reyes-Castillo and Martinez" docType="treatment" docVersion="3" lastPageNumber="5" masterDocId="FFAEE00A650FEE60FFF5FFA8FF91FF91" masterDocTitle="Extinct or extant? A new species of Termitodius Wasmann, 1894, (Coleoptera: Scarabaeidae: Aphodiinae: Rhyparini) with a short review of the genus" masterLastPageNumber="14" masterPageNumber="1" pageNumber="5" updateTime="1699308853719" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="D19F9AFB9B604EF53CBFE954A8A63C40">Extinct or extant? A new species of Termitodius Wasmann, 1894, (Coleoptera: Scarabaeidae: Aphodiinae: Rhyparini) with a short review of the genus</mods:title>
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<mods:namePart id="F69D71BF3007CEE08E6134079B017A0E">Skelley, Paul E.</mods:namePart>
<mods:affiliation id="AA566FAD37C163F447E5B6F3032D0A3C">Florida State Collection of Arthropods Florida Department of Agriculture and Consumer Services P. O. Box 147100 Gainesville, FL 32614 - 7100</mods:affiliation>
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<mods:namePart id="49292716D366E3C435D81D76B8BED230">Clavijo-Bustos, Julian</mods:namePart>
<mods:affiliation id="D62C4685B06F594A64E4AE6CF36C158E">Entomology section, Biological Collections Instituto de Investigación de Recursos Biológicos Alexander von Humboldt Villa de Leyva, Boyacá, Colombia</mods:affiliation>
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<mods:namePart id="8CABF412DC5492A8E079DDBDF793F0BE">Keller, Oliver</mods:namePart>
<mods:affiliation id="46C3E6B69D4B0817C6C99EB01D8F78DD">Florida State Collection of Arthropods Florida Department of Agriculture and Consumer Services P. O. Box 147100 Gainesville, FL 32614 - 7100</mods:affiliation>
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<mods:title id="0B6B7E3B252856FF2D1685AF3D9A3563">Insecta Mundi</mods:title>
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<mods:date id="71797FD9E7B8E128970CFF0B9ACA5650">2022</mods:date>
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<paragraph id="8B8129646509EE66FF35FC55FC84FB8B" blockId="6.[192,789,1021,1084]" box="[192,789,1021,1051]" pageId="6" pageNumber="5">
<heading id="D0C99E086509EE66FF35FC55FC84FB8B" box="[192,789,1021,1051]" fontSize="12" level="2" pageId="6" pageNumber="5" reason="2">
<taxonomicName id="4C3E52E76509EE66FF35FC55FC84FB8B" authority="Reyes-Castillo and Martinez" authorityName="Reyes-Castillo and Martinez" box="[192,789,1021,1051]" class="Insecta" family="Aphodiidae" genus="Termitodius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="chaki">
<emphasis id="B94AF5766509EE66FF35FC55FC84FB8B" bold="true" box="[192,789,1021,1051]" pageId="6" pageNumber="5">
<emphasis id="B94AF5766509EE66FF35FC55FE32FB8B" bold="true" box="[192,419,1021,1050]" italics="true" pageId="6" pageNumber="5">Termitodius chaki</emphasis>
Reyes-Castillo and Martínez
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3247AEF6509EE66FF35FB89FED3FBAD" box="[192,322,1057,1084]" pageId="6" pageNumber="5" type="description">
<paragraph id="8B8129646509EE66FF35FB89FED3FBAD" blockId="6.[192,789,1021,1084]" box="[192,322,1057,1084]" pageId="6" pageNumber="5">
<figureCitation id="130535E16509EE66FF35FB89FED3FBAD" box="[192,322,1057,1084]" captionStart="Figures 110" captionStartId="5.[192,279,1866,1892]" captionTargetBox="[192,1444,995,1836]" captionTargetId="figure-673@5.[192,1444,995,1836]" captionTargetPageId="5" captionText="Figures 110. Termitodius species dorsal habitus and lateral pronotum. 12) T. coronatus, cotype. 34) T. araujoi. 56) T. chaki, paratype. 78) T. woodruffi, copal paratype. 910) T. woodruffi, holotype." figureDoi="http://doi.org/10.5281/zenodo.6391935" httpUri="https://zenodo.org/record/6391935/files/figure.png" pageId="6" pageNumber="5">Figures 56</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3247AEF6509EE66FF15FBE1FBC8FB11" pageId="6" pageNumber="5" type="reference_group">
<paragraph id="8B8129646509EE66FF15FBE1FBC8FB11" blockId="6.[224,1440,1097,1152]" pageId="6" pageNumber="5">
<treatmentCitationGroup id="AB2E0E4A6509EE66FF15FBE1FBC8FB11" pageId="6" pageNumber="5">
<taxonomicName id="4C3E52E76509EE66FF15FBE1FC3EFBF0" authority="Reyes-Castillo and Martinez 1979: 125 - 128" authorityName="Reyes-Castillo and Martinez" authorityPageNumber="125 - 128" authorityYear="1979" box="[224,943,1097,1122]" class="Insecta" family="Aphodiidae" genus="Termitodius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="chaki">
<treatmentCitation id="0A9F0F756509EE66FF15FBE1FCDBFBF0" author="Reyes-Castillo P &amp; Martinez A." box="[224,842,1097,1122]" page="128" pageId="6" pageNumber="5" year="1979">
<emphasis id="B94AF5766509EE66FF15FBE1FE00FBF0" box="[224,401,1097,1121]" italics="true" pageId="6" pageNumber="5">Termitodius chaki</emphasis>
<bibRefCitation id="EFAF54956509EE66FE63FBE1FCDBFBF0" author="Reyes-Castillo P &amp; Martinez A." box="[406,842,1097,1122]" pageId="6" pageNumber="5" pagination="115 - 133" refId="ref8839" refString="Reyes-Castillo P, Martinez A. 1979. Nuevos Rhyparini Neotropicales, con notas sobre su biologia (Coleoptera, Scarabeidae, Aphodiinae). Folia Entomologica Mexicana 41: 115 - 133." type="journal article" year="1979">Reyes-Castillo and Martínez 1979: 125128</bibRefCitation>
</treatmentCitation>
(
<collectingCountry id="F32969F46509EE66FCA3FBE1FC36FBF0" box="[854,935,1097,1121]" name="Mexico" pageId="6" pageNumber="5">Mexico</collectingCountry>
)
</taxonomicName>
.
<treatmentCitation id="0A9F0F756509EE66FC42FBE1FB17FBF3" author="Chalumeau FE" box="[951,1158,1097,1122]" page="13" pageId="6" pageNumber="5" year="1981">
<bibRefCitation id="EFAF54956509EE66FC42FBE1FB17FBF3" author="Chalumeau FE" box="[951,1158,1097,1122]" pageId="6" pageNumber="5" pagination="13 - 16" refId="ref7528" refString="Chalumeau FE. 1981. Capture de Termitodius peregrinus Hinton en Guyane et notes diverses (Coleoptera: Scarabaeidae: Aphodiinae: Rhypariini). Bulletin de la Societe Entomologique de Mulhouse 1981: 13 - 16." type="journal article" year="1981">Chalumeau 1981: 13</bibRefCitation>
</treatmentCitation>
, 16;
<treatmentCitation id="0A9F0F756509EE66FB47FBE1FAFCFBF3" author="Dellacasa M." box="[1202,1389,1097,1122]" page="34" pageId="6" pageNumber="5" year="1988">
<bibRefCitation id="EFAF54956509EE66FB47FBE1FAFCFBF3" author="Dellacasa M." box="[1202,1389,1097,1122]" pageId="6" pageNumber="5" pagination="1 - 455" refId="ref7654" refString="Dellacasa M. 1988 [1987]. Contribution to a world-wide catalogue of Aegialiidae, Aphodiidae, Aulonocnemidae, Termitotrogidae (Coleoptera Scarabaeoidea). Memorie della Societa Entomologica Italiana 66: 1 - 455." type="journal article" year="1988">Dellacasa 1988: 34</bibRefCitation>
</treatmentCitation>
, 267, 426;
<treatmentCitation id="0A9F0F756509EE66FEDAFBC0FE66FB11" author="Dellacasa M." box="[303,503,1128,1152]" page="301" pageId="6" pageNumber="5" year="1989">
<bibRefCitation id="EFAF54956509EE66FEDAFBC0FE66FB11" author="Dellacasa M." box="[303,503,1128,1152]" pageId="6" pageNumber="5" pagination="291 - 316" refId="ref7691" refString="Dellacasa M. 1989 [1988]. Contribution to a world-wide catalogue of Aegialiidae, Aphodiidae, Aulonocnemidae, Termitotrogidae (Coleoptera Scarabaeoidea). Addenda et corrigenda (First note). Memorie della Societa Entomologica Italiana 67 (2): 291 - 316." type="journal article" year="1989">Dellacasa 1989: 301</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0A9F0F756509EE66FDF4FBC0FD7CFB11" author="Galante E &amp; Stebnicka Z &amp; Verdu JR" box="[513,749,1128,1152]" page="309" pageId="6" pageNumber="5" year="2003">
<bibRefCitation id="EFAF54956509EE66FDF4FBC0FD7CFB11" author="Galante E &amp; Stebnicka Z &amp; Verdu JR" box="[513,749,1128,1152]" pageId="6" pageNumber="5" pagination="283 - 312" refId="ref7927" refString="Galante E, Stebnicka Z, Verdu JR. 2003. The Aphodiinae and Rhyparinae (Coleoptera: Scarabaeidae) in southern states of Mexico (Chiapas, Oaxaca, Puebla and Veracruz). Acta Zoologica Cracoviensia 46 (3): 283 - 312." type="journal article" year="2003">Galante et al. 2003: 309</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0A9F0F756509EE66FD02FBC0FC24FB11" author="Howden HF" box="[759,949,1128,1152]" page="393" pageId="6" pageNumber="5" year="2003">
<bibRefCitation id="EFAF54956509EE66FD02FBC0FC24FB11" author="Howden HF" box="[759,949,1128,1152]" pageId="6" pageNumber="5" pagination="391 - 402" refId="ref8083" refString="Howden HF. 2003. Two new genera of New World Rhyparini (Coleoptera: Scarabaeidae: Aphodiinae). Sociobiology 42 (2): 391 - 402." type="journal article" year="2003">Howden 2003: 393</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0A9F0F756509EE66FC4AFBC0FBC7FB11" author="Skelley PE" box="[959,1110,1128,1152]" page="8" pageId="6" pageNumber="5" year="2007">
<bibRefCitation id="EFAF54956509EE66FC4AFBC0FBC7FB11" author="Skelley PE" box="[959,1110,1128,1152]" pageId="6" pageNumber="5" pagination="1 - 9" refId="ref9150" refString="Skelley PE. 2007. Generic limits of the Rhyparini with respect to the genus Termitodius Wasmann, 1894 (Coleoptera: Scarabaeidae: Aphodiinae). Insecta Mundi 0009: 1 - 9." type="book chapter" year="2007">Skelley 2007: 8</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3247AEF6509EE66FF35FB27FD24FA9E" pageId="6" pageNumber="5" type="diagnosis">
<paragraph id="8B8129646509EE66FF35FB27FD24FA9E" blockId="6.[191,1441,1167,1295]" pageId="6" pageNumber="5">
<emphasis id="B94AF5766509EE66FF35FB27FEADFB38" bold="true" box="[192,316,1167,1193]" pageId="6" pageNumber="5">Diagnosis.</emphasis>
A species of
<taxonomicName id="4C3E52E76509EE66FE2DFB38FDCDFB3B" authorityName="Wasmann" authorityYear="1894" box="[472,604,1168,1194]" class="Insecta" family="Aphodiidae" genus="Termitodius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B94AF5766509EE66FE2DFB38FDCDFB3B" box="[472,604,1168,1194]" italics="true" pageId="6" pageNumber="5">Termitodius</emphasis>
</taxonomicName>
with the pronotal paramedian costa having the anterior lobe elongate in dorsal view and anteriorly sloping in lateral view (
<figureCitation id="130535E16509EE66FD01FB19FCC5FB5D" box="[756,852,1201,1228]" captionStart="Figures 110" captionStartId="5.[192,279,1866,1892]" captionTargetBox="[192,1444,995,1836]" captionTargetId="figure-673@5.[192,1444,995,1836]" captionTargetPageId="5" captionText="Figures 110. Termitodius species dorsal habitus and lateral pronotum. 12) T. coronatus, cotype. 34) T. araujoi. 56) T. chaki, paratype. 78) T. woodruffi, copal paratype. 910) T. woodruffi, holotype." figureDoi="http://doi.org/10.5281/zenodo.6391935" httpUri="https://zenodo.org/record/6391935/files/figure.png" pageId="6" pageNumber="5">Fig. 56</figureCitation>
), posterior part of discolateral pronotal costa absent medially, elytra intercostal areas entirely with transverse wrinkles, the nearly globe-like caudal bulbs on the elytra, and in distribution (
<collectingCountry id="F32969F46509EE66FE6AFB5CFE65FA9F" box="[415,500,1268,1294]" name="Mexico" pageId="6" pageNumber="5">Mexico</collectingCountry>
and
<collectingCountry id="F32969F46509EE66FDDEFB5CFD38FA9E" box="[555,681,1268,1295]" name="Guatemala" pageId="6" pageNumber="5">Guatemala</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="C3247AEF6509EE66FF35FAB6FB54F99B" pageId="6" pageNumber="5" type="materials_examined">
<paragraph id="8B8129646509EE66FF35FAB6FA31FAED" blockId="6.[192,1441,1310,1404]" pageId="6" pageNumber="5">
<emphasis id="B94AF5766509EE66FF35FAB6FE93FAA9" bold="true" box="[192,258,1310,1336]" pageId="6" pageNumber="5">Type.</emphasis>
<materialsCitation id="3B5623396509EE66FEF3FAB6FA31FAED" collectingDate="1977-01-25" collectionCode="IEXA" collectorName="P. Reyes" country="Mexico" elevation="350" location="Selva Lacandona" municipality="Lacanja-Chansayab" pageId="6" pageNumber="5" specimenCount="350" stateProvince="Chiapas" typeStatus="holotype">
<typeStatus id="548597C66509EE66FEF3FAB6FEFFFAA9" box="[262,366,1310,1336]" pageId="6" pageNumber="5" type="holotype">Holotype</typeStatus>
male and
<typeStatus id="548597C66509EE66FE15FAB6FDA8FAA9" box="[480,569,1310,1336]" pageId="6" pageNumber="5" type="allotype">allotype</typeStatus>
female: “
<collectingCountry id="F32969F46509EE66FD6AFAB6FD67FAA9" box="[671,758,1310,1336]" name="Mexico" pageId="6" pageNumber="5">México</collectingCountry>
:
<collectingMunicipality id="6BE5B31E6509EE66FCF5FAB6FC4DFAA9" box="[768,988,1310,1336]" pageId="6" pageNumber="5">Lacanjá-Chansayab</collectingMunicipality>
,
<location id="8EE17FBF6509EE66FC10FAB6FB32FAA9" LSID="urn:lsid:plazi:treatment:039798726509EE66FF35FC55FA0AF931:8EE17FBF6509EE66FC10FAB6FB32FAA9" box="[997,1187,1310,1336]" country="Mexico" municipality="Lacanja-Chansayab" name="Selva Lacandona" pageId="6" pageNumber="5" stateProvince="Chiapas">Selva Lacandona</location>
,
<collectingRegion id="49FAE7866509EE66FB58FAB6FA9BFAA9" box="[1197,1290,1310,1336]" country="Mexico" name="Chiapas" pageId="6" pageNumber="5">Chiapas</collectingRegion>
,
<date id="FF800FA46509EE66FAE6FAB6FA12FAA8" box="[1299,1411,1310,1337]" pageId="6" pageNumber="5" value="1977-01-25">
<collectingDate id="EFC4F64C6509EE66FAE6FAB6FA12FAA8" box="[1299,1411,1310,1337]" pageId="6" pageNumber="5" value="1977-01-25">25-I-1977</collectingDate>
</date>
,
<collectorName id="26CB4CB26509EE66FA7BFAB7FE92FACB" pageId="6" pageNumber="5">P. Reyes</collectorName>
, col.,
<emphasis id="B94AF5766509EE66FECAFAE8FEC7FAC8" box="[319,342,1344,1369]" italics="true" pageId="6" pageNumber="5">in</emphasis>
nido de
<taxonomicName id="4C3E52E76509EE66FE41FAE8FDACFACB" authorityName="Wasmann" authorityYear="1896" box="[436,573,1344,1370]" class="Insecta" family="Rhinotermitidae" genus="Coptotermes" kingdom="Animalia" order="Blattodea" pageId="6" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B94AF5766509EE66FE41FAE8FDACFACB" box="[436,573,1344,1370]" italics="true" pageId="6" pageNumber="5">Coptotermes</emphasis>
</taxonomicName>
af.
<taxonomicName id="4C3E52E76509EE66FD96FAE8FC9AFACA" authority="(Lin.)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[611,779,1344,1371]" class="Insecta" family="Rhinotermitidae" genus="Coptotermes" kingdom="Animalia" order="Blattodea" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="testaceus">
<emphasis id="B94AF5766509EE66FD96FAE8FD52FAC8" box="[611,707,1344,1369]" italics="true" pageId="6" pageNumber="5">testaceus</emphasis>
(Lin.)
</taxonomicName>
, selva alta perennefolia, alt.
<specimenCount id="9D38E2ED6509EE66FBB5FA97FB15FACB" box="[1088,1156,1343,1370]" pageId="6" pageNumber="5" type="generic">
<quantity id="4CC684816509EE66FBB5FA97FB11FACB" box="[1088,1152,1343,1370]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="6" pageNumber="5" unit="m" value="350.0">
<elevation id="0013CE576509EE66FBB5FA97FB11FACB" box="[1088,1152,1343,1370]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="6" pageNumber="5" unit="m" value="350.0">350m</elevation>
</quantity>
.
</specimenCount>
<bibRefCitation id="EFAF54956509EE66FB67FAE8FED6FAEA" author="Reyes-Castillo P &amp; Martinez A." pageId="6" pageNumber="5" pagination="115 - 133" refId="ref8839" refString="Reyes-Castillo P, Martinez A. 1979. Nuevos Rhyparini Neotropicales, con notas sobre su biologia (Coleoptera, Scarabeidae, Aphodiinae). Folia Entomologica Mexicana 41: 115 - 133." type="journal article" year="1979">Reyes-Castillo and Martínez (1979)</bibRefCitation>
state the
<typeStatus id="548597C66509EE66FE45FAC9FD83FAEA" box="[432,530,1377,1403]" pageId="6" pageNumber="5" type="holotype">holotype</typeStatus>
is deposited in the collection of
<collectorName id="26CB4CB26509EE66FC8FFACAFC47FAED" box="[890,982,1378,1404]" pageId="6" pageNumber="5">P. Reyes</collectorName>
, now deposited in
<collectionCode id="ED2FB1A16509EE66FB53FAC9FB77FAEA" box="[1190,1254,1377,1403]" pageId="6" pageNumber="5">IEXA</collectionCode>
(not examined).
</materialsCitation>
</paragraph>
<paragraph id="8B8129646509EE66FF3DFA23FB54F99B" blockId="6.[192,1440,1419,1548]" pageId="6" pageNumber="5">
<emphasis id="B94AF5766509EE66FF3DFA23FE29FA34" bold="true" box="[200,440,1419,1445]" pageId="6" pageNumber="5">Materials examined.</emphasis>
<materialsCitation id="3B5623396509EE66FE35FA23FDE3FA79" collectingDate="1977-01-28" collectionCode="CMNC, CEMT, FSCA" collectorName="Lacania-Chansayab &amp; Col." country="Mexico" county="Hormiguero" elevation="350" location="Martinez" municipality="Howden" pageId="6" pageNumber="5" specimenCount="8" stateProvince="Chiapas" typeStatus="paratype">
<emphasis id="B94AF5766509EE66FE35FA23FDA1FA34" bold="true" box="[448,560,1419,1445]" pageId="6" pageNumber="5">
<collectingCountry id="F32969F46509EE66FE35FA23FDA1FA34" box="[448,560,1419,1445]" name="Mexico" pageId="6" pageNumber="5">MEXICO</collectingCountry>
</emphasis>
:
<emphasis id="B94AF5766509EE66FDCBFA23FD35FA34" bold="true" box="[574,676,1419,1445]" pageId="6" pageNumber="5">
<collectingRegion id="49FAE7866509EE66FDCBFA23FD0EFA34" box="[574,671,1419,1445]" country="Mexico" name="Chiapas" pageId="6" pageNumber="5">Chiapas</collectingRegion>
:
</emphasis>
<collectorName id="26CB4CB26509EE66FD59FA23FC18FA34" box="[684,905,1419,1445]" pageId="6" pageNumber="5">Lacania-Chansayab</collectorName>
,
<date id="FF800FA46509EE66FC60FA23FB94FA34" box="[917,1029,1419,1445]" pageId="6" pageNumber="5" value="1977-01-28">
<collectingDate id="EFC4F64C6509EE66FC60FA23FB94FA34" box="[917,1029,1419,1445]" pageId="6" pageNumber="5" value="1977-01-28">28-I-1977</collectingDate>
</date>
, P. Reyes-C.
<collectorName id="26CB4CB26509EE66FB6CFA23FB59FA34" box="[1177,1224,1419,1445]" pageId="6" pageNumber="5">Col.</collectorName>
selva alta perennifolia, alt.
<quantity id="4CC684816509EE66FEDCFA04FEE3FA56" box="[297,370,1452,1479]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="6" pageNumber="5" unit="m" value="350.0">
<elevation id="0013CE576509EE66FEDCFA04FEE3FA56" box="[297,370,1452,1479]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="6" pageNumber="5" unit="m" value="350.0">350 m</elevation>
</quantity>
, en
<collectingCounty id="62E051E86509EE66FE54FA04FDBCFA57" box="[417,557,1452,1478]" pageId="6" pageNumber="5">Hormiguero</collectingCounty>
de
<taxonomicName id="4C3E52E76509EE66FDA3FA05FD79FA56" authorityName="Wasmann" authorityYear="1896" box="[598,744,1453,1479]" class="Insecta" family="Rhinotermitidae" genus="Coptotermes" kingdom="Animalia" order="Blattodea" pageId="6" pageNumber="5" phylum="Arthropoda" rank="genus">Coptotermes</taxonomicName>
(
<specimenCount id="9D38E2ED6509EE66FD0DFA04FCEBFA56" box="[760,890,1452,1479]" pageId="6" pageNumber="5" type="generic" typeStatus="paratypes">6 paratypes</specimenCount>
<collectionCode id="ED2FB1A16509EE66FC74FA05FC45FA56" box="[897,980,1453,1479]" pageId="6" pageNumber="5">CMNC</collectionCode>
from
<collectingMunicipality id="6BE5B31E6509EE66FBEEFA04FBEAFA57" box="[1051,1147,1452,1478]" pageId="6" pageNumber="5">Howden</collectingMunicipality>
and
<location id="8EE17FBF6509EE66FB41FA05FA8BFA56" LSID="urn:lsid:plazi:treatment:039798726509EE66FF35FC55FA0AF931:8EE17FBF6509EE66FB41FA05FA8BFA56" box="[1204,1306,1453,1479]" country="Mexico" county="Hormiguero" municipality="Howden" name="Martinez" pageId="6" pageNumber="5" stateProvince="Chiapas">Martínez</location>
collections;
<specimenCount id="9D38E2ED6509EE66FF34FA66FEABFA78" box="[193,314,1486,1513]" pageId="6" pageNumber="5" type="generic" typeStatus="paratype">1 paratype</specimenCount>
<collectionCode id="ED2FB1A16509EE66FEB1FA66FE03FA79" box="[324,402,1486,1512]" pageId="6" pageNumber="5">CEMT</collectionCode>
;
<specimenCount id="9D38E2ED6509EE66FE54FA66FD88FA78" box="[417,537,1486,1513]" pageId="6" pageNumber="5" type="generic" typeStatus="paratype">1 paratype</specimenCount>
<collectionCode id="ED2FB1A16509EE66FDD1FA66FDFAFA79" box="[548,619,1486,1512]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="6" pageNumber="5" type="Museum">FSCA</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B5623396509EE66FD8AFA66FA0DFA78" box="[639,1436,1485,1513]" collectorName="Patulul" country="Guatemala" location="Los Tarrales Private Reserve" pageId="6" pageNumber="5" specimenCount="1" stateProvince="Suchitepequez" typeStatus="paratype">
<emphasis id="B94AF5766509EE66FD8AFA66FC75FA79" bold="true" box="[639,996,1485,1512]" pageId="6" pageNumber="5">
<collectingCountry id="F32969F46509EE66FD8AFA66FCBCFA79" box="[639,813,1486,1512]" name="Guatemala" pageId="6" pageNumber="5">GUATEMALA</collectingCountry>
:
<collectingRegion id="49FAE7866509EE66FCCFFA65FC75FA79" box="[826,996,1485,1512]" country="Guatemala" name="Suchitepequez" pageId="6" pageNumber="5">Suchitepéquez</collectingRegion>
</emphasis>
:
<collectorName id="26CB4CB26509EE66FC00FA66FBD7FA79" box="[1013,1094,1486,1512]" pageId="6" pageNumber="5">Patulul</collectorName>
,
<location id="8EE17FBF6509EE66FBA1FA67FA0DFA78" LSID="urn:lsid:plazi:treatment:039798726509EE66FF35FC55FA0AF931:8EE17FBF6509EE66FBA1FA67FA0DFA78" box="[1108,1436,1486,1513]" country="Guatemala" name="Los Tarrales Private Reserve" pageId="6" pageNumber="5" stateProvince="Suchitepequez">Los Tarrales Private Reserve</location>
</materialsCitation>
;
<quantity id="4CC684816509EE66FF35FA58FE9EF99B" box="[192,271,1520,1546]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="5" unit="m" value="1000.0">1000m</quantity>
; 14°31.942, 91°08.799; w/termites under bark;
<date id="FF800FA46509EE66FCC6FA58FC3AF99B" box="[819,939,1520,1546]" pageId="6" pageNumber="5" value="2009-07-08">8.VII.2009</date>
;
<materialsCitation id="3B5623396509EE66FC43FA58FB50F99B" box="[950,1217,1520,1546]" collectionCode="UNSM" collectorName="Paulsen" country="Guatemala" pageId="6" pageNumber="5" specimenCount="1" stateProvince="Suchitepequez" typeStatus="paratype">
M.J.
<collectorName id="26CB4CB26509EE66FC1FFA58FBD0F99B" box="[1002,1089,1520,1546]" pageId="6" pageNumber="5">Paulsen</collectorName>
(1
<collectionCode id="ED2FB1A16509EE66FB90FA58FB28F99B" box="[1125,1209,1520,1546]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34868" lsid="urn:lsid:biocol.org:col:34868" name="University of Nebraska State Museum" pageId="6" pageNumber="5">UNSM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3247AEF6509EE66FF35F9B1FA89F9C4" pageId="6" pageNumber="5" type="distribution">
<paragraph id="8B8129646509EE66FF35F9B1FA89F9C4" blockId="6.[191,1439,1561,1621]" pageId="6" pageNumber="5">
<emphasis id="B94AF5766509EE66FF35F9B1FECBF9A2" bold="true" box="[192,346,1561,1587]" pageId="6" pageNumber="5">Distribution.</emphasis>
<taxonomicName id="4C3E52E76509EE66FE95F9B1FDB2F9A2" box="[352,547,1561,1587]" class="Insecta" family="Aphodiidae" genus="Termitodius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="chaki">
<emphasis id="B94AF5766509EE66FE95F9B1FDB2F9A2" box="[352,547,1561,1587]" italics="true" pageId="6" pageNumber="5">Termitodius chaki</emphasis>
</taxonomicName>
is distributed in the Mexican transition zone at the
<collectingRegion id="49FAE7866509EE66FBA8F9B1FB26F9A2" box="[1117,1207,1561,1587]" country="Mexico" name="Chiapas" pageId="6" pageNumber="5">Chiapas</collectingRegion>
Highlands province, and in the Veracruzan province of the Mesoamerican dominion of the Brazilian subregion (
<figureCitation id="130535E16509EE66FB4EF993FA9AF9C4" box="[1211,1291,1595,1621]" captionStart="Figure 33" captionStartId="13.[265,341,1323,1349]" captionTargetBox="[243,1386,218,1289]" captionTargetId="figure-206@13.[239,1392,213,1293]" captionTargetPageId="13" captionText="Figure 33. Distribution of Termitodius species." figureDoi="http://doi.org/10.5281/zenodo.6391947" httpUri="https://zenodo.org/record/6391947/files/figure.png" pageId="6" pageNumber="5">Fig. 33</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3247AEF6509EE66FF35F9CCFA0AF931" pageId="6" pageNumber="5" type="discussion">
<paragraph id="8B8129646509EE66FF35F9CCFA0AF931" blockId="6.[192,1440,1636,1697]" pageId="6" pageNumber="5">
<emphasis id="B94AF5766509EE66FF35F9CCFED9F9EF" bold="true" box="[192,328,1636,1662]" pageId="6" pageNumber="5">Comments.</emphasis>
<taxonomicName id="4C3E52E76509EE66FEAFF9CCFD8FF9EF" box="[346,542,1636,1662]" class="Insecta" family="Aphodiidae" genus="Termitodius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="chaki">
<emphasis id="B94AF5766509EE66FEAFF9CCFD8FF9EF" box="[346,542,1636,1662]" italics="true" pageId="6" pageNumber="5">Termitodius chaki</emphasis>
</taxonomicName>
R&amp;M” illustrated in
<bibRefCitation id="EFAF54956509EE66FCFFF9CCFC7DF9EE" author="Galante E &amp; Stebnicka Z &amp; Verdu JR" box="[778,1004,1636,1663]" pageId="6" pageNumber="5" pagination="283 - 312" refId="ref7927" refString="Galante E, Stebnicka Z, Verdu JR. 2003. The Aphodiinae and Rhyparinae (Coleoptera: Scarabaeidae) in southern states of Mexico (Chiapas, Oaxaca, Puebla and Veracruz). Acta Zoologica Cracoviensia 46 (3): 283 - 312." type="journal article" year="2003">Galante et al. (2003)</bibRefCitation>
is not a
<taxonomicName id="4C3E52E76509EE66FBBFF9CCFB5CF9EF" authorityName="Wasmann" authorityYear="1894" box="[1098,1229,1636,1662]" class="Insecta" family="Aphodiidae" genus="Termitodius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B94AF5766509EE66FBBFF9CCFB5CF9EF" box="[1098,1229,1636,1662]" italics="true" pageId="6" pageNumber="5">Termitodius</emphasis>
</taxonomicName>
, it is a new species of
<taxonomicName id="4C3E52E76509EE66FF28F92EFDD0F931" authority="Howden, 2003" authorityName="Howden" authorityYear="2003" box="[221,577,1670,1696]" class="Insecta" family="Aphodiidae" genus="Nanotermitodius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B94AF5766509EE66FF28F92EFE06F931" box="[221,407,1670,1696]" italics="true" pageId="6" pageNumber="5">Nanotermitodius</emphasis>
<bibRefCitation id="EFAF54956509EE66FE68F92EFDD0F931" author="Howden HF" box="[413,577,1670,1696]" pageId="6" pageNumber="5" pagination="391 - 402" refId="ref8083" refString="Howden HF. 2003. Two new genera of New World Rhyparini (Coleoptera: Scarabaeidae: Aphodiinae). Sociobiology 42 (2): 391 - 402." type="journal article" year="2003">Howden, 2003</bibRefCitation>
</taxonomicName>
, that is currently being described. This Oaxacan record needs to be removed.
</paragraph>
</subSubSection>
</treatment>
</document>